taxonID	type	description	language	source
4A5A3E7BFFDFFF8F55A51150FDD7FEC9.taxon	description	(Figs 1 – 5)	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFDFFF8F55A51150FDD7FEC9.taxon	materials_examined	Type locality. Uganda, Nakiwogo Entebbe. Type material examined. HOLOTYPE: J, ‘ Uganda, Nakiwogo Entebbe, xi. 1961 - vi. 1962, A. S. Hoddeta // HOLO- TYPUS, Pseudohesperus apsilus sp. nov. Hromádka, det., 2009 [red oblong label, printed] ’ (NMPC). PARATYPES: GABON: 1 spec., Konta de Cristal, // Philonthus megalops R. I. Sc. N. B. Coll. et det. A. Fauvel, [white oblong label, handwritten] // Philonthus megalops Tottenham TYPE [ochre oblong label, handwritten] (IRSB); 2 spec., Loango, // Philonthus megalops, R. I. Sc. N. B. Coll. et det. A. Fauvel // Philonthus megalops Tottenham PARATYPE [ochre oblong label, handwritten] (IRSB). IVORY COAST: 4 spec., Danané, Droplou, 10. x. 1980, tamisage ss. fronc mort, Mahnert – Perret (MNHG).	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFDFFF8F55A51150FDD7FEC9.taxon	description	Description. Body length 7.3 mm, length of fore body (to end of elytra) 3.7 mm. Colouration. Head, pronotum and elytra black, scutellum dark carmine, in the middle slightly bluish iridescent, abdomen conspicuously golden, violet and green iridescent, labial palpi brown, palpomeres 1 and 2 of maxillary palpi brown, palpomere 3 yellow-brown, mandibles brown, slightly paler distally, antennae black, femora brown-yellow, tibiae and tarsi dark brown. Head transverse, wider than long (ratio 28: 20), slightly narrowed posteriad, each posterior angle rounded with one small tooth as in P. morio Boheman, 1848, eyes distinctly longer than temples (ratio 11: 6). Distance between medial interocular punctures almost four times as long as distance between medial and lateral puncture, medial punctures distinctly shifted anteriad, each side along impunctate midline with scattered punctures, temporal area with several variably large punctures. Surface without microsculpture. Antennae slender and very long, exceeding posterior margin of pronotum by last four antennomeres when reclined, all antennomeres longer than wide. Relative lengths of antennomeres 1 – 11: 1 = 11 units, 2 = 8 units, 3 = 7 units, 4 – 6 = 6 units, 7 – 10 = 5 units and 11 = 5.5 units. Pronotum slightly convex, as wide as long, parallel-sided, anterior angles conspicuously deflexed, vaguely obtusely rounded, posterior angles markedly rounded. Each dorsal row irregularly punctate, exact number of larger punctures in the dorsal rows difficult to state owing to their irregular nature, each side along narrow impunctate midline with small scattered punctures. Surface without microsculpture. Scutellum densely and finely punctate, punctures as large as eye-facets, separated by one puncture diameter in transverse direction. Elytra combined wider than long (ratio 43: 40), very slightly narrowed posteriad, punctation fine and dense, diameter of punctures smaller than eye-facets, transverse interstices between punctures usually smaller than their diameter. Surface between microsculpture, shiny; setation greyish. Legs. Metatibia longer than metatarsus (ratio 25: 23), relative lengths of metatarsomeres 1 – 5: 1 = 6 units, 2 = 4 units, 3 = 3 units, 4 = 2 units and 5 = 6 units. Abdomen wide, very gradually narrowing towards apex. First three visible tergites with two basal lines, elevated area between lines with scattered punctures. Punctation of basal portion of all tergites coarser than that on elytra, becoming sparser towards posterior margin of each tergite. Surface between punctures without microsculpture; setation similar to that on elytra. Male. Protarsomeres 1 – 4 simple, hardly dilated, each with few modified pale setae laterally. Sternite VIII (Fig. 4), sternite IX (Fig. 5), aedeagus (Figs. 1 – 3). Female. Unknown.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFDFFF8F55A51150FDD7FEC9.taxon	diagnosis	Differential diagnosis. Pseudohesperus apsilus sp. nov. may be separated from P. conradti by longer eyes and antennae, darker antennomere 1 and a different shape of the aedeagus and from P. proselytus by a wider head, longer antennae and denser punctation of the abdomen. Note. I have found this new species in the collection of IRSB, labelled by A. Fauvel and C. E. Tottenham as Philonthus megalops, but neither of them formally described the taxon as new.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFDFFF8F55A51150FDD7FEC9.taxon	etymology	Etymology. The name of this species, a noun in apposition, is the Latin generic name of the African forktail snapper Apsilus fuscus Valenciennes, 1830.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFDFFF8F55A51150FDD7FEC9.taxon	distribution	Distribution. Uganda, Gabon and Ivory Coast.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD8FF8D55741524FE4CFDCE.taxon	description	(Figs. 18 – 22)	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD8FF8D55741524FE4CFDCE.taxon	materials_examined	Type locality. Cameroon, Bafut forest, Nʼguemba. Type material. HOLOTYPE: J, ‘ FʼBAFUT, Nʼguemba, 16. VIII. 1965, Muséum Paris, CAMEROUN, B. de Miré, // Hesperus bafutensis n. sp. L. Levasseur det. 1966 [ochre oblong label, handwritten] ’ (MNHN). Additional material examined. NIGER: 1 J, Abargasit, 3. ii. 1978, R. Macek leg. (NMPC).	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD8FF8D55741524FE4CFDCE.taxon	description	Description. Body length 9.2 mm, length of fore body (to end of elytra) 3.9 mm. Colouration. Head and pronotum black, scutellum and elytra brown-black, suture and posterior margin narrowly brown-yellow, abdomen black-brown, posterior margin of all tergites narrowly red-brown, all tergites with bluish purple iridescence. Maxillary palpi, labial palpi and mandibles brown-yellow, antennae black, base of antennomere 2 yellow-brown, femora brown, tibiae darker, tarsi brown, slightly paler distally. Head rectangular, wider than long (ratio 30: 22), parallel-sided, each posterior angle obtusely rounded, bearing one long black bristle. Eyes flat, as long as temples. Four coarse punctures between eyes, distance between medial interocular punctures three times as long as distance between medial and lateral puncture, medial punctures slightly shifted anteriad. Several variably large punctures placed posterolaterally of the line connecting middle of inner margin of eyes and middle of posterior margin of head. Surface with patches of very fine irregular microsculpture. Antennae slender and very long, exceeding posterior margin of pronotum by antennomeres 10 and 11 when reclined, all antennomeres longer than wide. Relative lengths of antennomeres 1 – 11: 1 = 8 units, 2 = 5 units, 3 = 6 units, 4 – 7 = 5 units, 8 and 9 = 4.5 units, 10 = 4 units and 11 = 5 units. Pronotum highly convex, slightly wider than long (ratio 34: 32), parallel-sided, anterior angles conspicuously deflexed, vaguely obtusely rounded, posterior angles markedly rounded. Each dorsal row with four coarse punctures and many smaller punctures. Each side along impunctate midline with numerous punctures. Surface with microsculpture similar to that on head. Scutellum very finely punctate, punctures somewhat smaller than eye-facets, distance between punctures mostly larger than their diameter. Setation dark. Elytra combined wider than long (ratio 45: 40), widened posteriad. Punctation very fine and dense, punctures as large as eye-facets, separated by 1 – 1.5 diameter of punctures. Surface without microsculpture; setation greyish. Legs. Metatarsus shorter than metatibia (ratio 25: 27), lengths of metatarsomeres 1 – 5: 1 = 7 units, 2 = 4 units, 3 = 3 units, 4 = 2.5 units and 5 = 7 units. Abdomen wide, parallel-sided, very hardly narrowed posteriad. First three visible tergites with two basal lines, elevated area between lines coarsely and densely punctate. Punctation of tergites sparser and coarser than that on elytra, becoming sparser towards apex of each tergite; setation similar to that on elytra. Male. Protarsomeres 1 – 4 simple, each with modified pale setae laterally. Sternite VIII (Fig. 21), sternite IX (Fig. 22), aedeagus (Figs. 18 – 20). Female. Unknown.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD8FF8D55741524FE4CFDCE.taxon	diagnosis	Differential diagnosis. Pseudohesperus bafutensis may be distinguished from the most similar species P. conradti by its wider head, shorter antennae and denser punctation of the scutellum, from P. natalensis by a different colouration of the pronotum and abdomen, narrower elytra and darker legs, from P. varanus sp. nov. by a finer punctation of the elytra and sparser punctation of the abdomen, and from all three species by a different shape of the aedeagus. Moreover, P. bafutensis differs from P. proselytus by a wider head, longer antennae and sparser punctation of the scutellum.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD8FF8D55741524FE4CFDCE.taxon	distribution	Distribution. Cameroon and Niger.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFDAFF82554B1624FDF1FC0E.taxon	description	(Figs. 6 – 10)	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFDAFF82554B1624FDF1FC0E.taxon	materials_examined	Type locality. Cameroon, Lolodorf [= village of Lolo]. Type material examined. Philonthus conradti: HOLOTYPE: ♀, ‛ S. O. Kamerun: Lolodorf, L. Conradt, 1895. Chicago NHMus, M. Bernhauer collection // Philonthus conradti Bernhauer, Type [ochre oblong label, handwritten] ’ (FMNH). Hesperus longicornis: COTYPE: 1 ♀, ʻANGOLA, Ngauyanga // Hesperus longicornis COTYPE Cameron, M. Cameron-Bequest B. M. 1955 - 147 ʼ (BMNH). Additional material examined. ANGOLA: 1 spec., Ngauyanga (BMNH). NIGERIA: 1 spec., Ibadan, 1. v. 1956 (NMUK), 1 spec., Aroche, viii. 1945 (NMUK). ZIMBABWE: 3 spec., Kutsaga, near Harare airport, 18. vi. 1997, W. Rossi (LHPC),	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFDAFF82554B1624FDF1FC0E.taxon	description	Redescription. Body length 7.9 mm, length of fore body (to end of elytra) 3.8 mm. Colouration. Head black, pronotum, scutellum, elytra and abdomen black-brown, posterior margin of all tergites narrowly brown-red, maxillary palpi, labial palpi and antennomeres 1 and 2 yellow-brown, remaining antennomeres black-brown, mandibles brown, femora and tarsi yellow-brown, tibiae darker. Scutellum bluish iridescent, abdomen slightly golden-red iridescent. Head transverse, wider than long (ratio 27: 20), inconspicuously narrowed posteriad. Posterior angles obtusely rounded with two long black bristles. Eyes longer than temples (ratio 9: 7). Four coarse punctures present between eyes, distance between medial interocular punctures three times as long as distance between medial and lateral puncture, median punctures slightly shifted anteriad. Inner margin of eyes and entire temporal area with variably large punctures, vertex of head largely impunctate. Dorsal surface without appreciable microsculpture. Antennae slender and long, exceeding posterior margin of pronotum by antennomeres 10 and 11 when reclined. All antennomeres longer than wide. Relative lengths of antennomeres 1 – 11: 1 = 11 units, 2 = 5 units, 3 = 6 units, 4 = 5 units, 5 – 7 = 6 units, 8 = 5.5 units and 9 – 11 = 5 units. Pronotum wider than long (ratio 32: 30), parallel-sided, each anterior angle distinctly deflexed, vaguely obtusely rounded, posterior angles conspicuously rounded. Wide area along midline impunctate, each dorsal row consisting of four coarse and large, irregularly placed punctures and width many smaller punctures, lateral portions of pronotum with scattered punctures. Surface without microsculpture. Scutellum densely and finely punctate. Punctures as large as eye-facets, separated by one puncture diameter in transverse direction. Elytra combined wider than long (ratio 42: 38), hardly widened posteriad. Punctation fine and dense. Punctures somewhat larger than eye-facets, distance between punctures mostly smaller than their diameter. Surface without microsculpture; setation brown-yellow. Legs. Metatarsus as long as metatibia. Relative length of metatarsomeres 1 – 5: 1 = 7 units, 2 = 4 units, 3 = 3.5 units, 4 = 3 units and 5 = 7 units. Abdomen wide, gradually narrowed towards apex from visible tergite III. First three tergites with two basal lines, elevated area between lines impunctate. Punctation at base of all tergites slightly coarser than that on elytra, gradually becoming finer and sparser towards posterior margin of each tergite. Surface between punctures without microsculpture; setation similar to that on elytra. Male. Protarsomeres 1 – 4 simple, with few modified pale setae laterally. Sternite VIII (Fig. 9), sternite IX (Fig. 10), aedeagus (Figs. 6 – 8). Female. Protarsomeres 1 – 3 simple, each covered with modified pale setae ventrally.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFDAFF82554B1624FDF1FC0E.taxon	diagnosis	Differential diagnosis. Pseudohesperus conradti is similar to P. apsilus sp. nov., but may be distinguished from the latter by shorter antennae and eyes, paler antennomere 1 and a different shape of the aedeagus; it differs from P. proselytus by a wider head, longer antennae, paler antennomere 1 and denser punctation of the scutellum, pronotal sides and abdomen.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFDAFF82554B1624FDF1FC0E.taxon	distribution	Distribution. Cameroon and Zimbabwe.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD5FF80555C105CFDF9FBCE.taxon	description	(Figs. 33 – 36)	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD5FF80555C105CFDF9FBCE.taxon	materials_examined	Type locality. Republic of South Africa, Natal. Type material examined. HOLOTYPE: J, ‘ Natal, Sharp Coll. 1905 - 313, // Philonthus eccoptomus Tottenham, TYPE [ochre oblong label, handwritten] // Holotype: J, Philonthus eccoptomus Tottenham, det. 1962 R. G., Booth, 2009 [white oblong label, handwritten] ’ (BMNH).	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD5FF80555C105CFDF9FBCE.taxon	description	Redescription. Body length 9.1 mm, length of fore body (to end of elytra) 4.7 mm. Colouration. Head, pronotum and abdomen black, scutellum and elytra dark brown-violet, maxillary, labial palpi, mandibles and legs yellow-brown, antennomeres 1 – 3 dark brown, remaining antennomeres black. Abdomen with very conspicuous, golden to rainbow-coloured iridescence. Head transverse, distinctly wider than long (ratio 32: 26), posterior angles rounded, bearing one long black bristle. Four coarse punctures present between eyes, distance between median interocular punctures three times as large as distance between medial and lateral puncture. Medial punctures shifted anteriad. Eyes flat, longer than temples (ratio 11: 8). Surface except wide area along midline and clypeus finely and irregularly punctate. Diameter of punctures markedly larger than eye-facets, distance between punctures irregular. Surface without microsculpture. Antennae slender and long, all antennomeres longer than wide, exceeding posterior margin of pronotum by antennomere 11 when reclined. Relative lengths of antennomeres 1 – 11: 1 = 11 units, 2 = 6.5 units, 3 = 7.5 units, 4 – 8 = 5.5 units, 9 and 10 = 5 units and 11 = 5.5 units. Pronotum highly convex, parallel-sided, wider than long (ratio 37: 35), anterior angles conspicuously deflexed, vaguely obtusely rounded, with several punctures, posterior angles markedly rounded. Wide midline impunctate, each side along midline with many variably large punctures. Surface without microsculpture. Scutellum very finely and densely punctate, punctures as large as eye-facets, separated by one puncture diameter or slightly smaller distance. Elytra wider than long (ratio 37: 35), slightly widened posteriad. Punctation fine and dense, punctures larger than those on scutellum, separated by distance mostly smaller than one puncture diameter in transverse direction. Surface without microsculpture; setation brown. Legs. Metatarsus longer than metatibia (ratio 29: 27), relative lengths of metatarsomeres 1 – 5: 1 = 9 units, 2 = 5 units, 3 = 4 units, 4 = 3 units and 5 = 7 units. Abdomen wide, very gradually narrowed from visible tergite III towards apex. First three visible tergites with two basal lines, elevated area between lines with scattered punctures. Punctation at base of all tergites coarser and sparser than that on elytra, becoming finer and sparser towards posterior margin of each tergite. Surface without microsculpture; setation similar to that on elytra. Male. Protarsomeres 1 – 4 simple, each with few modified pale setae laterally. Sternite IX (Fig. 36), aedeagus (Figs. 33 – 35). Female. Unknown.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD5FF80555C105CFDF9FBCE.taxon	diagnosis	Differential diagnosis. Pseudohesperus eccoptomus is quite similar to P. tyto sp. nov., but differs from the latter by longer eyes, coarser punctation of abdomen and a different shape of the aedeagus. Moreover, it may be distinguished from P. proselytus by longer eyes and a very conspicuous, golden to rainbow-coloured iridescence of the abdomen.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD5FF80555C105CFDF9FBCE.taxon	distribution	Distribution. Republic of South Africa.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD7FF8155521024FD98FA04.taxon	description	(Figs. 37 – 40)	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD7FF8155521024FD98FA04.taxon	materials_examined	Type locality. Natal. Type material examined. HOLOTYPE: J, ‘ Natal, 1914, // TYPUS Hesperus natalensis O. Scheerpeltz [red oblong label, handwritten], E. Ross – Berlin donarit 1934, ’ (NHMW).	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD7FF8155521024FD98FA04.taxon	description	Redescription. Body length 9.4 mm, length of fore body (to end of elytra) 3.9 mm. Colouration. Head and pronotum black, with very feeble bluish tinge, scutellum and elytra dark black-violet, abdomen black, very strongly golden-violet-greenish iridescent, maxillary, labial palpi and mandibles yellow-brown, antennae dark brown, legs brown yellow, tarsi paler distally. Head wider than long (ratio 31: 25), parallel-sided, posterior angles obtusely rounded, midline wide impunctate, each side with many varying large punctures. Temporal area with many punctures. Four coarse punctures present between eyes, distance between medial punctures three times as large as distance between medial and lateral puncture, medial punctures markedly shifted anteriad. Eyes large, longer than temples (ratio 12: 10). Surface without microsculpture. Antennae incomplete in the holotype: left antenna with only six antennomeres, right antennae with only two antennomeres. Antennnomere 1 as long as antennomeres 5 and 6 combined. Pronotum almost as long as wide, parallel-sided, anterior angles rounded, with several small punctures, posterior angles markedly rounded. Midline with wide impunctate area, punctures near midline coarse and dense, finer and sparser towards sides, diameter of smaller punctures mostly slightly larger than eye-facets, separated by 1 – 2 puncture diameters. Surface without microsculpture. Scutellum densely and finely punctate, punctures as large as those on pronotum, separated by one puncture diameter in transverse direction. Elytra wider than long (ratio 49: 42), widened posteriad. Punctation very fine and dense. Diameter of punctures as large as smaller punctures on elytra, separated by one puncture diameter in transverse direction. Surface without microsculpture; setation brown. Legs. Metatibia as long as metatarsus, metatarsomere 1 longer than metatarsomere 5, almost as long as metatarsomeres 2 – 4 combined. Abdomen wide, very gradually narrowed posteriad, first three visible tergites with two basal lines, elevated area between lines punctate. Punctation at base of all tergites dense, most punctures droplet-shaped, sparser towards posterior margin of each tergite. Surface without microsculpture, setation similar to that on elytra. Male. Protarsomeres 1 – 3 not dilated, protarsomere 1 almost as long as protarsomeres 2 – 4 combined, longer than protarsomere 5. Sternite IX (Fig. 40), aedeagus (Figs. 37 – 39). Female. Unknown.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD7FF8155521024FD98FA04.taxon	diagnosis	Differential diagnosis. Pseudohesperus natalensis may be distinguished from the similar P. bafutensis by a different colouration of the pronotum and abdomen, wider elytra, paler legs and a different shape of the aedeagus.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD7FF8155521024FD98FA04.taxon	distribution	Distribution. Republic of South Africa.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD1FF87552514A4FD3AFD89.taxon	description	(Figs. 11 ‒ 12)	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD1FF87552514A4FD3AFD89.taxon	materials_examined	Type locality. Democratic Republic of the Congo, Kivu, Tshibinda. Type material examined. HOLOTYPE: ♀, ‘ Democratic Republic of the Congo, Kivu: Tshibinda, xi, 1932, L Burgeon, Musée du Congo, // Philonthus tamulus, Tottenham, Type [yellow square label, handwritten] ’ (MRAC).	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD1FF87552514A4FD3AFD89.taxon	description	Redescription. Body length 7.5 mm, length of fore body (to end of elytra) 3.4 mm. Colouration. Head black, pronotum black-brown, elytra including suture black, abdomen pitchy brown, golden-bluish iridescent, maxillary palpi and labial palpi yellow-brown, mandibles brown, slightly paler distally, antennae black, legs pitchy brown with femora vaguely lighter. Head wider than long (ratio 24: 20), sides very slightly rounded, posterior angles obtusely rounded, base straight. Four coarse punctures present between eyes, distance between medial interocular punctures four times as long as distance between medial and lateral puncture, medial punctures slightly shifted anteriad, outer punctures situated halfway between inner punctures and marginal punctures of eyes, one or two much smaller shallow punctures present behind these; temples with several variably large punctures. Eyes flat, slightly shorter than temples (ratio 7: 8). Surface with very fine, almost inconspicuous microsculpture. Antennae slender and long, all antennomeres longer than wide, exceeding posterior margin of pronotum by length of antennomere 2 when reclined. Relative lengths of antennomeres 1 – 11: 1 = 7 units, 2 and 3 = 5 units, 4 and 5 = 4.5 units, 6 and 7 = 4 units, 8 – 10 = 3.5 units and 11 = 6 units. Pronotum highly convex, as wide as long, vaguely narrowed anteriad, anterior angles conspicuously deflexed, obtusely rounded, posterior angles markedly rounded. Each dorsal row irregular, consisting of about four coarse punctures and 5 – 6 very small but distinct punctures (number of punctures in dorsal row difficult to state owing to its irregular nature), other similar small and 1 – 2 larger punctures scattered over whole lateral area, surface with microsculpture similar to that on head. Scutellum sparsely and finely punctate, punctures smaller than eye-facets, separated by 1.5 – 2 puncture diameters in transverse direction. Elytra combined wider than long (ratio 39: 36), slightly widened posteriad, very finely and densely punctate, with several distinctly larger punctures in indistinct longitudinal rows. Legs. Metatarsus shorter than metatibia (ratio 21: 23), relative lengths of metatarsomeres 1 – 5: 1 = 6 units, 2 = 3.5 units, 3 = 3 units, 4 = 2.5 units and 5 = 5 units. Abdomen wide, very slightly and gradually narrowed towards apex. First four visible tergites with two basal lines, elevated area between lines with scattered punctures. Punctation at base of all tergites coarser and sparser than on elytra, gradually becoming finer and sparser towards posterior margin of each tergite. Surface without microsculpture; setation similar to that on elytra. Male. Unknown. Female. Protarsomeres 1 – 4 simple, each with few modified pale setae laterally. Tergite X (Fig. 11), gonocoxite of female genital segment (Fig. 12).	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD1FF87552514A4FD3AFD89.taxon	diagnosis	Differential diagnosis. Pseudohesperus proselytus may be distinguished from P. apsilus sp. nov. by a narrower head, shorter antennae and sparser punctation of the abdomen, from P. bafutensis by a narrower head, longer antennae and sparser punctation of the scutellum, from P. conradti by a narrower head, shorter antennae, darker antennomere 1 and sparser punctation of the scutellum, pronotal sides and abdomen, from P. eccoptomus by shorter eyes and the lack of rainbow-coloured iridescence of the abdomen, from P. tauraco sp. nov. by a narrower head, shorter antennae and denser punctation of the scutellum, from P. tyto sp. nov. by a narrower head, shorter antennae, darker antennomere 1 and sparser punctation of the scutellum and abdomen, and from P. varanus sp. nov. by a narrower head and shorter eyes and antennae.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD1FF87552514A4FD3AFD89.taxon	distribution	Distribution. Democratic Republic of the Congo.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD0FF8455CC16E4FD10FB69.taxon	description	(Figs. 13 – 17)	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD0FF8455CC16E4FD10FB69.taxon	materials_examined	Type locality. Democratic Republic of the Congo, Massif Ruwenzori, Kyandolire, 1700 m a. s. l. Type material. HOLOTYPE: J, ‘ Democratic Republic of the Congo [‛ Congo Belge’], Massif Ruwenzori, Kyandolire, 1700 m, Camp des Garde (terreau), 7. - 9. x. 1952, P. Vanschuytbroeck & J. Kekenbosch 1136 - 39 // Holotypus, Pseudohesperus tauraco sp. nov., Hromádka det. 2010 [red oblong label, printed] ’ (MRAC). PARATYPES: DEMOCRATIC REPUBLIC OF THE CONGO: ♀, Massif Ruwenzori, Kyandolire, 1.750 m, Riv. Mulaku, Kakalari (terreau), [‛ Congo Belge’]: P. N. A., 16. X. 1952, P. Vanschuytbroeck & J. Kekenbosch, 1302 (MRAC); ♀, Massif Ruwenzori, Kalonge, 2.060 m, Ruiss, Kalambure affl. Katauleko, [‘ Congo Belge’]: P. N. A. 30. i. - 21. ii. 1953, P. Vanschuytbroeck & J. Kekenbosch 2225 - 59 (MRAC), ♀, Massif Ruwenzori, Kalonge, 2.210 m (terreau), [‛ Congo Belge’]: P. N. A., 4. ix. 1952, P. Vanschuytbroeck & J. Kekenbosch 934 (MRAC), J, Kivu: T. Kalohe, Bltale, N. O. Kahusi, 1600 m, 29. vi. 1951, N. Leleup, Récolté dans l’humus (LHPC); ♀, Massif Ruwenzori, Kalonge 2.010 m Riv. Nyamwamba, affl. Butahu, [‛ Congo Belge’]: P. N. A., 2. - 3. ii. 1953, P. Vanschuytbroeck & J. Kekenbosch 2214 - 21 (MRAC); J, Massif Ruwenzori 1.780 m, Kyandolire, 1780 m, R. Bwisonge, affl. Butahu (terreau), [‛ Congo Belge’]: P. N. A. 12. x. 1952, P. Vanschuytbroeck & J. Kekenbosch 1253 - 57 (MRAC); J, Massif Ruwenzori, Kalonge, 2.480 m. (ét bambus), R. Nyamwamba-Ihongero, [‛ Congo Belge’]: P. N. A., 26 - 27. viii. 1952, P. Vanschuytbroeck & J. Kekenbosch (MRAC); ♀, Massif Ruwenzori, Kyandolire, 1750 m, Riv. Mulaku af Kakalari (terreau), [‛ Congo Belge’]: P. N. A., 14. x. 1952, P. Vanschuytbroeck & J. Kekenbosch (LHPC). Each paratype is provided with a red printed paratype label.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD0FF8455CC16E4FD10FB69.taxon	description	Description. Body length 6.1 mm, length of fore body (to end of elytra) 3.5 mm. Colouration. Head black, pronotum, scutellum, elytra and abdomen black-brown, mandibles, maxillary and labial palpi brown, apex of labial and maxillary palpomeres 3 yellowbrown, antennae black, femora and tarsomeres 5 of all tarsi brown-yellow, tibiae and remaining tarsomeres dark brown. Head transverse, markedly wider than long (ratio 27: 20), parallel-sided. Posterior angles obtusely rounded. Eyes flat, slightly longer than temples (ratio 8: 7). Four coarse punctures present between eyes, distance between medial interocular punctures four times as large as distance between medial and lateral puncture. Medial punctures distinctly shifted anteriad. Middle portion of head with wide impunctate median area, each side and temple area with several variably large punctures. Posterior margin of each eye with two coarse punctures. Surface without microsculpture. Antennae very long, exceeding posterior margin of pronotum by antennomeres 10 and 11 when reclined. All antennomeres longer than wide. Relative lengths of antennomeres 1 – 11: 1 = 10 units, 2 and 3 = 7 units, 4 = 6 units, 5 and 6 = 5.5 units, 7 – 10 = 5 units and 11 = 6 units. Pronotum highly convex, wider than long (ratio 30: 28), anterior angles conspicuously deflexed, vaguely obtusely rounded, posterior angles markedly rounded. Each dorsal row with 4 – 5 large punctures, and 9 – 10 small punctures, middle portion of pronotum with wide impunctate longitudinal band. Each side with many irregularly arranged small punctures and several large punctures between them. Surface without microsculpture. Scutellum very finely punctate, punctures smaller than eye-facets and separated by two puncture diameters in transverse direction; setation black. Elytra combined wider than long (ratio 38: 33), hardly widened posteriad. Punctation very fine and sparse, punctures smaller than eye-facets, separated by 2 – 3 puncture diameters. Surface without microsculpture; setation dark. Legs. Metatarsus about as long as metatibia. Relative lengths of metatarsomeres 1 – 5: 1 = 10 units, 2 = 5 units, 3 = 4.5 units, 4 = 4 units and 5 = 7 units. Abdomen wide, very slightly narrowed towards apex. First three visible tergites with two basal lines, elevated area between lines with scattered punctures. Punctation of visible tergites finer and much sparser than that on elytra, becoming sparser towards posterior margin of each tergite. Surface between punctures without microsculpture. Setation similar to that on elytra. Male. Protarsomeres 1 – 4 simple, each with few modified pale setae laterally. Sternite VIII (Fig. 16), sternite IX (Fig. 17), aedeagus (Figs. 13 – 15). Female. Unknown.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD0FF8455CC16E4FD10FB69.taxon	diagnosis	Differential diagnosis. Pseudohesperus tauraco sp. nov. may be distinguished from P. tyto sp. nov. by a narrower and sparser punctation of the head and a different shape of the aedeagus and from P. proselytus by a wider head, shorter antennae and denser punctation of the scutellum.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD0FF8455CC16E4FD10FB69.taxon	etymology	Etymology. The name of this species, a noun in apposition, is the Latin generic name of the African red-crested turaco Tauraco erythrolophus (Vieillot, 1819).	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD0FF8455CC16E4FD10FB69.taxon	distribution	Distribution. Democratic Republic of the Congo.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD3FF9A558810C4FBBFFBAB.taxon	description	(Figs. 23 – 27)	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD3FF9A558810C4FBBFFBAB.taxon	materials_examined	Type locality. Ethiopia, Shoa, Sodere. Type material. HOLOTYPE: J, ‘ ETHIOPIA: Shoa, Sodere, 1971 // HOLOTYPE Pseudohesperus tyto sp. nov. Hromádka det. [red oblong label, printed] ’ (MRAC). PARATYPES: ETHIOPIA: 3 JJ, Abessin, Maraco, 5. v. 1914 (LHPC); 1 J, Abyssinie // Tottenham megalopoides Tottenham, TYPE [ochre oblong label, handwritten] (IRSB). DEMOCRATIC REPUBLIC OF THE CONGO: 1 J, Congo Belge, Libenge, 29. iv. 1948, R. Cremer – M. Neuman (IRSB).	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD3FF9A558810C4FBBFFBAB.taxon	description	Description. Body length 8.3 – 8.9 mm, length of fore body (to end of elytra) 4.2 – 4.6 mm. Colouration. Head and pronotum black, scutellum slightly pitchy, abdomen black, conspicuously blue-golden iridescent, mandibles, maxillary and labial palpi brown, anterior half of palpomere 3 yellow-brown, ventral side of antennomere 1 yellow-brown, dorsal side and antennomeres 2 and 3 dark brown, remaining antennomeres black, legs pitchy with femora hardly lighter. Head markedly transverse, wider than long (ratio 32: 22), parallel-sided, posterior angles obtusely rounded, bearing one long black bristle each. Eyes flat, hardly longer than temples (ratio 10: 9), base straight. Four coarse punctures present between eyes, distance between medial interocular punctures 2.5 times as long as distance between medial and lateral puncture. Middle portion of head with relatively wide impunctate longitudinal band. Each side and temporal area with many punctures of variable size. Surface with very fine microsculpture. Antennae very long, exceeding posterior margin of pronotum by the length of antennomere 1 when reclined. All antennomeres longer than wide. Relative lengths of antennomeres 1 – 11: 1 = 10 units, 2 = 5 units, 3 = 6 units, 4 – 9 = 5 units, 10 = 4.5 units and 11 = 5 units. Pronotum highly convex, hardly wider than long (ratio 33: 31), each anterior angle strongly deflexed, obtusely rounded, posterior angles conspicuously rounded. Middle portion of pronotum with wide impunctate longitudinal band. Each dorsal row irregular, consisting of four large and 5 – 6 small but distinct punctures, number of punctures difficult to state owing to its irregularity; other similarly small and 1 – 2 larger punctures diffusely scattered over whole lateral area. Surface with microsculpture similar to that on head. Scutellum finely punctate, punctures as large as eye-facets, separated by one puncture diameter in transverse direction; setation black. Elytra vaguely wider than long (ratio 42: 40), very slightly widened posteriad. Punctation fine and dense, diameter of punctures somewhat larger than those on scutellum, separated by 1 – 1.5 puncture diameters. Surface without microsculpture; setation greyish. Legs. Metatarsus as long as metatibia. Metatarsomere 1 longer than metatarsomere 5. Relative lengths of metatarsomeres 1 – 5: 1 = 8 units, 2 = 4 units, 3 = 3.5 units, 4 = 3 units and 5 = 6 units. Abdomen slightly narrowed from visible tergite III towards apex. First three visible tergites with two basal lines, elevated area between lines sparsely punctate. Base of each tergite finely and densely punctate, gradually becoming finer and much sparser towards posterior margin of tergite. Surface between tergites without microsculpture; setation similar to that on elytra. Male. Protarsomeres 1 – 4 simple, each with few modified pale setae laterally. Sternite VIII (Fig. 26), sternite IX (Fig. 27), aedeagus (Figs. 23 – 25). Female. Protarsomeres 1 – 4 of the same shape as in male.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD3FF9A558810C4FBBFFBAB.taxon	diagnosis	Differential diagnosis. Pseudohesperus tyto sp. nov. may be distinguished from P. tauraco sp. nov. by a wider and densely punctate head and different shape of the aedeagus, from P. eccoptomus by shorter eyes, finer punctation of the abdomen and a different shape of the aedeagus, and from P. proselytus by a wider head, longer antennae, paler antennomere 1 and denser punctation of the scutellum and abdomen. Note. I have found this new species under the name Philonthus megalopoides identified by Tottenham in the collection of IRSB, but the species has never been formally described.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD3FF9A558810C4FBBFFBAB.taxon	etymology	Etymology. The name of this species, a noun in apposition, is the Latin generic name of the African grass-owl Tyto capensis (Smith, 1834).	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFD3FF9A558810C4FBBFFBAB.taxon	distribution	Distribution. Democratic Republic of the Congo, Ethiopia. Fig. 41. Distribution of the Afrotropical Pseudohesperus. ● – P. apsilus sp. nov.; ♣ – P. bafutensis (Levasseur, 1967); ♥ – P. conradti (Bernhauer, 1912); ▼ – P. eccoptomus (Tottenham, 1962); × – P. natalensis (Scheerpeltz, 1956); ○ – P. proselytus (Herman, 2001); ◊ – P. tauraco sp. nov.; * – P. tyto sp. nov.; + – P. varanus sp. nov.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFCDFF9B55AF10DFFD3AFB09.taxon	description	(Figs. 28 – 32)	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFCDFF9B55AF10DFFD3AFB09.taxon	materials_examined	Type locality. Democratic Republic of the Congo, Kibali-Ituri, Blukwa 1820 - 2100 m a. s. l. Type material examined. HOLOTYPE: J, ‘ Democratic Republic of the Congo, Kibali-Ituri 1820 - 2100 m, (Dans humus en forêt) i. 1954, N. Leleup // HOLOTYPE, Pseudohesperus varanus sp. nov., Hromádka det. 2009 [red oblong label, printed] ’ (IRSB). PARATYPES: DEMOCRATIC REPUBLIC OF THE CONGO: 1 J 1 ♀, Kibali-Ituri: Terr. Djugu, Mt. Aboro, 2200 m, 1.1954, H. Leleup (IRSB, LHPC).	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFCDFF9B55AF10DFFD3AFB09.taxon	description	Description. Body length 7.5 mm, length of fore body (to end of elytra) 3.3 mm. Colouration. Head black, pronotum black-brown, elytra and abdomen chestnut brown, scutellum and suture of elytra brown-yellow, abdomen bluish iridescent, maxillary palpi, labial palpi, femora and tarsi yellow-brown, mandibles and tibiae darker. Head vaguely wider than long (ratio 32: 30), distinctly narrowed posteriad, arching from posterior margin of eyes towards neck, posterior angles rounded, bearing one long black bristle each. Eyes flat, slightly shorter than temples (ratio 8: 9). Four coarse punctures present between eyes, distance between medial interocular punctures four times as large as distance between medial and lateral puncture. Posterior margin of eye with two coarse punctures, temporal area with several variably large punctures. Surface without microsculpture. Antennae slender and long, exceeding posterior margin of pronotum by antennomere 11 when reclined, all antennomeres longer than wide, relative lengths of antennomeres 1 – 11: 1 = 9 units, 2 and 3 = 6 units, 4 – 7 = 5 units, 8 and 9 = 4 units, 10 = 3.5 units and 11 = 5 units. Pronotum highly convex, almost as wide as long, parallel-sided, anterior angle with several variably large bristles, posterior angles markedly rounded. Sides bearing one long black bristle in anterior third. Each dorsal row with four coarse punctures and many small, but distinct punctures, each sublateral row with two coarse punctures, second puncture distinctly shifted laterad. Each side with several scattered punctures. Surface without microsculpture. Scutellum finely and densely punctate, punctures as large as eye-facets, separated by one puncture diameter in transverse direction. Elytra combined wider than long (ratio 40: 35), hardly wider posteriorly. Punctation slightly denser than that on scutellum. Surface between punctures without microsculpture; setation short and yellowish-brown. Legs. Metatarsus as long as metatibia, relative lengths of metatarsomeres 1 – 5: 1 = 7 units, 2 = 4 units, 3 = 3.5 units, 4 = 3 units and 5 = 6 units. Abdomen wide, very gradually narrowed towards apex. First three visible tergites with two basal lines, elevated area between basal lines with scattered punctures. Punctation of visible tergites coarser and much sparser than that on elytra. Surface without microsculpture, very shiny; setation longer, yellowish-brown. Male. Protarsomeres 1 – 4 simple, each with few modified pale setae laterally. Sternite VIII (Fig. 31), sternite IX (Fig. 32), aedeagus (Figs. 28 – 30). Female. Unknown.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFCDFF9B55AF10DFFD3AFB09.taxon	diagnosis	Differential diagnosis. Pseudohesperus varanus sp. nov. can be separated from P. proselytus by a wider head, longer eyes and antennae and different shape of the aedeagus, from P. bafutensis by a finer punctation of the elytra, sparser punctation of the abdomen and different shape of the aedeagus and from P. proselytus by a wider head and longer eyes and antennae.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFCDFF9B55AF10DFFD3AFB09.taxon	etymology	Etymology. The name of this species, a noun in apposition, is the Latin generic name of the African rock monitor Varanus albigularis Daudin, 1802.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFCDFF9B55AF10DFFD3AFB09.taxon	distribution	Distribution. Democratic Republic of the Congo.	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFCCFF9B550511E7FEF7F9CC.taxon	materials_examined	Material examined: P. ichor: PARATYPE: J, Senegal, Bad: Wassadou (IRSB). P. parasanguineus: HOLOTYPE: J, Senegal, Thies (MNHN).	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFCFFF9855FC1797FD13FB7C.taxon	materials_examined	Material examined. G. maritimus: J, Ethiopia, Bale 8 km W of Dinshu (LHPC). P. meridioafricanus: SYNTYPE: J SW Afr. Kaokoveld Anabib 100 miles W Ohopoho (NHMW).	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFCFFF98550114A4FC6BFE8E.taxon	materials_examined	Material examined. P. rubrovittatus: HOLOTYPE: ♀, Belgian Congo [= Democratic Republic of the Congo], Kivu: Lulenga (MRAC). P. nimeaglius: PARATYPE: J, Kenya, Mau Escarpment 2150 - 2200 m (BMNH).	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFCFFF9855EF15E3FE52FDC8.taxon	materials_examined	Material examined. P. taterae: HOLOTYPE: J, South Africa, Vredefort (NHMW). P. discoideisimilis: ‘ COTYPE’: J, Cape: Prov.: Near Areb Mnt. (NHMW).	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFCFFF985511162BFD16FD10.taxon	materials_examined	Material examined. P. ventralis: J, Ethiopia, Bele 8 km. W of Dinshu (LHPC). P. flandrianus: PARATYPE: J, Democratic Republic of the Congo: Tshuapa: Flandra (IRSB).	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFCFFF9855FC176AFE27FC34.taxon	materials_examined	Material examined. P. wittei: HOLOTYPE: J, Belgian Congo [= Democratic Republic of the Congo], Katanga: Kansenia (FMNH). P captivus: HOLOTYPE: J, Belgian Congo [= Democratic Republic of the Congo], Reg. Kolwezi, Katanga, Musonoie (MNHN).	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFCEFF99553D14A4FE8DFEA4.taxon	materials_examined	Material examined. HOLOTYPE: ♀, Belgian Congo [= Democratic Republic of the Congo], vers Rweru, volc. Mikeno, alt. 2400 m. (MRAC).	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFCEFF995524150FFC36FE29.taxon	materials_examined	Material examined. HOLOTYPE: ♀, Ostafrika, Katamayu, sept. 1934, A. F. J. Gerte (FMNH).	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFCEFF9955251582FF79FD87.taxon	materials_examined	Material examined. HOLOTYPE: J, Belgian Congo [= Democratic Republic of the Congo], Haut-Uele: Moto (FMNH).	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFCEFF99553D16DCFC24FCFB.taxon	materials_examined	Material examined. HOLOTYPE: J, [Belgian Congo], Manyema, Sibatawa Kilengwe (FMNH).	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFCEFF9955201750FCAEFC6E.taxon	materials_examined	Material examined. HOLOTYPE: J, [Belgian Congo], Kivu: N’Gama (MRAC).	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFCEFF99552917C4FEF2FBC4.taxon	materials_examined	Material examined. HOLOTYPE: ♀, D. Ostafrika [= Deutsche Ostafrika, i. e. German East Africa, referring to Tanzania], Kahama (FMNH).	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFCEFF995538101DFE3BFB1D.taxon	materials_examined	Material examined. PARATYPE: ♀, Congo Belge [= Democratic Republic of the Congo], Nyasheke (Nyamuragira, 1820 m) [Congo Belge] (IRSB).	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFCEFF9955471176FC0EFA91.taxon	materials_examined	Material examined. PARATYPE: ♀, [Belgian Congo], Mt. Sesero, près Bitashimwa, 200 m (IRSB).	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
4A5A3E7BFFCEFF99557B11EAFC09FA05.taxon	materials_examined	Material examined. PARATYPE: ♀, Belgian Congo, forest near Mont Kamatembe, 2200 m (IRSB).	en	Hromádka, Lubomír (2010): Revision of the Afrotropical species of the genus Pseudohesperus, with taxonomic notes on other Afrotropical species of the subtribe Philonthina (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae 50 (2): 495-516, DOI: 10.5281/zenodo.5326048
