taxonID	type	description	language	source
455A87DB446BFFA72C92210D8E52FBF2.taxon	description	urn: lsid: zoobank. org: act: C 9107 B 76 - 69 A 3 - 4304 - AA 05 - 43 D 5542 E 6 ADF	en	Gnezdilov, Vladimir M., Malenovský, Igor (2023): First record of the planthopper tribe Adenissini (Hemiptera: Caliscelidae) from Socotra Island, with description of a new genus and two new species. European Journal of Taxonomy 888 (1): 46-63, DOI: 10.5852/ejt.2023.888.2207, URL: http://dx.doi.org/10.5852/ejt.2023.888.2207
455A87DB446BFFA72C92210D8E52FBF2.taxon	type_taxon	Type species Oceatra litoralis gen. et sp. nov., designated here.	en	Gnezdilov, Vladimir M., Malenovský, Igor (2023): First record of the planthopper tribe Adenissini (Hemiptera: Caliscelidae) from Socotra Island, with description of a new genus and two new species. European Journal of Taxonomy 888 (1): 46-63, DOI: 10.5852/ejt.2023.888.2207, URL: http://dx.doi.org/10.5852/ejt.2023.888.2207
455A87DB446BFFA72C92210D8E52FBF2.taxon	diagnosis	Diagnosis Metope with median and sublateral carinae; median carina running to clypeus, but far not reaching upper margin of metope. Forewings with hypocostal plate and reticulate venation (with many transverse veins). Cubitus posterior (CuP) of forewings indistinct. Phallobase with ventral lobe narrow, without denticles, and dorso-lateral lobes each with mushroom-shaped outgrowth above ventral lobe. Style with large, curved capitulum, without neck, bearing small lateral tooth or lacking it. Gonoplacs short, rounded, with dorso-caudal angle not protruding beyond the apex of anal tube (in lateral view).	en	Gnezdilov, Vladimir M., Malenovský, Igor (2023): First record of the planthopper tribe Adenissini (Hemiptera: Caliscelidae) from Socotra Island, with description of a new genus and two new species. European Journal of Taxonomy 888 (1): 46-63, DOI: 10.5852/ejt.2023.888.2207, URL: http://dx.doi.org/10.5852/ejt.2023.888.2207
455A87DB446BFFA72C92210D8E52FBF2.taxon	etymology	Etymology The name of the genus is a combination from the Latin noun ‘ oceanus ’ (= ‘ ocean’, ‘ sea’, ‘ large body of water’) and Socotra, referring to the geographical isolation of the newly described planthopper taxa on Socotra Island in the Indian Ocean. Gender feminine.	en	Gnezdilov, Vladimir M., Malenovský, Igor (2023): First record of the planthopper tribe Adenissini (Hemiptera: Caliscelidae) from Socotra Island, with description of a new genus and two new species. European Journal of Taxonomy 888 (1): 46-63, DOI: 10.5852/ejt.2023.888.2207, URL: http://dx.doi.org/10.5852/ejt.2023.888.2207
455A87DB446AFFA22C46236F8AECFA0E.taxon	description	urn: lsid: zoobank. org: act: C 3 F 31 EA 9 - 7579 - 425 A-BA 43 - C 2 A 916 A 36 F 78 Figs 1 – 2, 4 B – C, 5 A – C, 6 – 7	en	Gnezdilov, Vladimir M., Malenovský, Igor (2023): First record of the planthopper tribe Adenissini (Hemiptera: Caliscelidae) from Socotra Island, with description of a new genus and two new species. European Journal of Taxonomy 888 (1): 46-63, DOI: 10.5852/ejt.2023.888.2207, URL: http://dx.doi.org/10.5852/ejt.2023.888.2207
455A87DB446AFFA22C46236F8AECFA0E.taxon	diagnosis	Diagnosis See the identification key to species of Oceatra gen. nov. above.	en	Gnezdilov, Vladimir M., Malenovský, Igor (2023): First record of the planthopper tribe Adenissini (Hemiptera: Caliscelidae) from Socotra Island, with description of a new genus and two new species. European Journal of Taxonomy 888 (1): 46-63, DOI: 10.5852/ejt.2023.888.2207, URL: http://dx.doi.org/10.5852/ejt.2023.888.2207
455A87DB446AFFA22C46236F8AECFA0E.taxon	etymology	Etymology From the Latin adjective ‘ litoralis ’ (= ‘ of the seashore’, ‘ littoral’), referring to the habitat of the new species in the coastal salt marshes.	en	Gnezdilov, Vladimir M., Malenovský, Igor (2023): First record of the planthopper tribe Adenissini (Hemiptera: Caliscelidae) from Socotra Island, with description of a new genus and two new species. European Journal of Taxonomy 888 (1): 46-63, DOI: 10.5852/ejt.2023.888.2207, URL: http://dx.doi.org/10.5852/ejt.2023.888.2207
455A87DB446AFFA22C46236F8AECFA0E.taxon	materials_examined	Type material Holotype YEMEN • ♂; Socotra Island, ca 3 km NE of Shuab; 12 ° 34.1 ′ N, 53 ° 23.9 ′ E; 3 m a. s. l.; 20 – 21 Jun. 2012; I. Malenovský, P. Kment, J. Bezděk, J. Hájek, V. Hula, J. Niedobová and L. Purchart leg.; sweeping and suction sampling in Avicennia marina mangrove, salt marsh and sand dunes; MMBC. Paratypes YEMEN – Socotra Island • 1 ♀; same collection data as for holotype; MMBC • 1 ♂, 1 ♀; same collection data as for holotype; ZIN.	en	Gnezdilov, Vladimir M., Malenovský, Igor (2023): First record of the planthopper tribe Adenissini (Hemiptera: Caliscelidae) from Socotra Island, with description of a new genus and two new species. European Journal of Taxonomy 888 (1): 46-63, DOI: 10.5852/ejt.2023.888.2207, URL: http://dx.doi.org/10.5852/ejt.2023.888.2207
455A87DB446AFFA22C46236F8AECFA0E.taxon	description	Description MEASUREMENTS. Total body length: males – 3.0 mm, females – 4.0 mm. COLORATION. Male (Fig. 1). General coloration light ochreous. Upper half of metope dark brown to black, carinae light ochreous (Fig. 1 C). Postclypeus with dark brown dots and spots laterally. Anteclypeus dark brown. Rostrum dark brown ventrally, black apically. Pedicel brown to dark brown. Coryphe with dark brown dots (Fig. 1 A). Preocular fields dark brown. Paranotal lobes of pronotum with dark brown dots. Mesonotum with dark brown spot in each anterior angle. Forewing with dark brown pattern concentrated along outer clavus margin between CuP and Pcu and in cells of corium in its apical half (Fig. 1 B). Hind coxae brown posteriorly. Femora and tibiae with dark brown bands. Third tarsomeres with dark brown apical halves. Claws and dorso-lateral plates of pretarsus dark brown. Apices of spines on legs black. Each laterotergite with large dark brown to black spot. Style light yellow, with dark brown capitulum. Penis dark brown (Fig. 1 D). Female (Fig. 2) similar to male. Forewing with dark brown cells along outer clavus margin and in posterior half of corium (Fig. 2 A) or only with sparse dark brown spots scattered across whole wing. Visible abdominal tergites VI – VIII with dark brown spots. Gonoplacs brown (Fig. 2 C – D). Anal tube with black median stripe running and expanding from paraproct to its hind margin. EXTERNAL STRUCTURE. Metope (Figs 1 C, 2 B, 5 B) about as wide as long, with lateral margins obtusely angular below eyes; upper margin sharply trapezoidal, concave. Median carina distinct from the level of eye middle through whole postclypeus, interrupted by metopoclypeal suture; sublateral carinae extending from upper margin of metope almost to metopoclypeal suture. Metopoclypeal suture distinct. Coryphe transverse, nearly twice as wide as long medially (Fig. 5 A). Ocelli absent. Pedicel globular. Rostrum long, reaching sternite VI, not narrowing apically; second segment 0.25 times as long as third one (Fig. 5 C). Pronotum wide, depressed in the middle, without carinae or with very short median carina at its hind margin (Fig. 5 A). Paradiscal fields of pronotum very narrow, not visible behind eyes. Paranotal lobes nearly rectangular. Mesonotum as long as pronotum medially, with median and lateral carinae. Forewing (Fig. 4 B – C) widely truncate apically, reaching hind margin of pygofer in male and surpassing hind margin of tergite VII in female. Hypocostal plate relatively wide and long, extending along three quarters of wing length. Radius and median veins running from basal cell in one short stem. R, M, CuA, and Pcu furcating apically. Hind tibia with two lateral spines in apical half and seven apical spines. First metatarsomere 1.5 times as long as second one, with two latero-apical and five intermediate spines arranged in arc. Second metatarsomere with only two latero-apical spines. Claws, in dorsal view, nearly twice as long as arolium of pretarsus, each with a single long seta. Dorso-lateral plates of pretarsus narrow. MALE TERMINALIA (Figs 1 D, 6). Anal tube, in dorsal view, wide, 0.7 times as wide as long at midline, slightly widening apically, with slightly obtusely angulate anterior margin (Fig. 6 C). Anal column short and wide. Pygofer, in lateral view, narrow, with hind margins strongly convex in upper half (Fig. 6 A – B). Anal tube and phallobase with well-sclerotised connection (Fig. 6 F). Phallobase, in lateral view, wide, slightly enlarged apically (Fig. 6 A, F). Dorso-lateral lobes of phallobase fused dorsally in one lobe with strongly convex margin (Fig. 6 I); each dorso-lateral lobe with mushroom-shaped outgrowth above ventral phallobase lobe (Fig. 6 F, mot). Ventral phallobase lobe short and narrow, narrowing apically (Fig. 6 G, vl). Apical processes of aedeagus extending beyond phallobase margin, long and wide, each with one long spiny and several small teeth apically (Fig. 6 F, aep). Ventral aedeagal hooks arising at aedeagal apex, almost as long as aedeagus, directed basally, acute apically (Fig. 6 F – G, vh). Connective with large, widely opened cup. Style massive, with large curved capitulum without neck, hind margin of style straight, caudal angle rounded (Fig. 6 D). Capitulum, in dorsal view, with small apical tooth and weak lateral tooth (Fig. 6 E). FEMALE TERMINALIA (Figs 2 C – D, 7). Hind margin of sternite VII with two short teeth medially (Fig. 7 C). Anal tube, in dorsal view, as wide as long, with hind margin concave (Fig. 7 A); in lateral view, ventral part of anal tube convex (Fig. 7 B). Anal column short and wide.	en	Gnezdilov, Vladimir M., Malenovský, Igor (2023): First record of the planthopper tribe Adenissini (Hemiptera: Caliscelidae) from Socotra Island, with description of a new genus and two new species. European Journal of Taxonomy 888 (1): 46-63, DOI: 10.5852/ejt.2023.888.2207, URL: http://dx.doi.org/10.5852/ejt.2023.888.2207
455A87DB446AFFA22C46236F8AECFA0E.taxon	distribution	Distribution and ecology So far only known from the western coast of Socotra Island, close to the cape and village of Shuab (Fig. 9). The type series was collected in coastal sand dunes and salt marshes in vegetation dominated by the semi-succulent plant species Limonium sokotranum (Vierh.) Radcl. - Sm. (Plumbaginaceae), Arthrocnemum macrostachyum (Moric.) K. Koch (Amaranthaceae) and Zygophyllum qatarense Hadidi (Zygophyllaceae) (Fig. 10 A). Most specimens were swept from L. sokotranum which might be a host plant for O. litoralis gen. et sp. nov. Another planthopper species, Haloflata arthrocnemi Świerczewski, Malenovský & Stroiński, 2017 from the family of Flatidae, co-occurred at the site (see Świerczewski et al. 2017 for details).	en	Gnezdilov, Vladimir M., Malenovský, Igor (2023): First record of the planthopper tribe Adenissini (Hemiptera: Caliscelidae) from Socotra Island, with description of a new genus and two new species. European Journal of Taxonomy 888 (1): 46-63, DOI: 10.5852/ejt.2023.888.2207, URL: http://dx.doi.org/10.5852/ejt.2023.888.2207
455A87DB446FFFAF2C4921428804FE51.taxon	description	urn: lsid: zoobank. org: act: 1 A 305662 - 5 F 2 E- 4191 - BE 76 - 45 A 01 CC 1 B 2 F 3 Figs 3, 5 D – E, 8	en	Gnezdilov, Vladimir M., Malenovský, Igor (2023): First record of the planthopper tribe Adenissini (Hemiptera: Caliscelidae) from Socotra Island, with description of a new genus and two new species. European Journal of Taxonomy 888 (1): 46-63, DOI: 10.5852/ejt.2023.888.2207, URL: http://dx.doi.org/10.5852/ejt.2023.888.2207
455A87DB446FFFAF2C4921428804FE51.taxon	diagnosis	Diagnosis See the identification key to species of Oceatra gen. nov. above.	en	Gnezdilov, Vladimir M., Malenovský, Igor (2023): First record of the planthopper tribe Adenissini (Hemiptera: Caliscelidae) from Socotra Island, with description of a new genus and two new species. European Journal of Taxonomy 888 (1): 46-63, DOI: 10.5852/ejt.2023.888.2207, URL: http://dx.doi.org/10.5852/ejt.2023.888.2207
455A87DB446FFFAF2C4921428804FE51.taxon	etymology	Etymology Derived from the present participle of the Latin verb ‘ scandō ’ (= ‘ to climb’, ‘ ascend’, ‘ mount’), referring to the type locality of the new species close to the top of the highest peak of Socotra Island, Mt Scand.	en	Gnezdilov, Vladimir M., Malenovský, Igor (2023): First record of the planthopper tribe Adenissini (Hemiptera: Caliscelidae) from Socotra Island, with description of a new genus and two new species. European Journal of Taxonomy 888 (1): 46-63, DOI: 10.5852/ejt.2023.888.2207, URL: http://dx.doi.org/10.5852/ejt.2023.888.2207
455A87DB446FFFAF2C4921428804FE51.taxon	materials_examined	Type material Holotype YEMEN • ♂; Socotra Island, Hagher Mountains, Mt Scand env.; 12 ° 34.6 ′ N, 54 ° 01.5 ′ E; 1450 m a. s. l.; 16 – 18 Jun. 2012; I. Malenovský, P. Kment, J. Bezděk, J. Hájek, V. Hula, J. Niedobová and L. Purchart leg.; sweeping and suction sampling in montane evergreen woodland; MMBC.	en	Gnezdilov, Vladimir M., Malenovský, Igor (2023): First record of the planthopper tribe Adenissini (Hemiptera: Caliscelidae) from Socotra Island, with description of a new genus and two new species. European Journal of Taxonomy 888 (1): 46-63, DOI: 10.5852/ejt.2023.888.2207, URL: http://dx.doi.org/10.5852/ejt.2023.888.2207
455A87DB446FFFAF2C4921428804FE51.taxon	description	Description MEASUREMENTS. Total body length: male – 3.8 mm. COLORATION (Fig. 3). General coloration dark brown. Metope, genae and postclypeus dark brown to black, with light ochreous transverse band above metopoclypeal suture and light ochreous dots and pustules (traces of larval sensory pits) between lateral margins and sublateral carinae, genae with light ochreous spot above each pedicel; median carina light ochreous, sublateral carinae brown (Fig. 3 C). Anteclypeus dark brown to black including median carina. Rostrum dark brown, black apically. Scapus and pedicel black. Preocular fields dark brown to black, each with two light ochreous spots at anterior angles (Fig. 3 B). Coryphe and pronotum with paranotal lobes dark brown, with light ochreous dots (Fig. 3 A). Mesonotum dark brown, with light ochreous dots and carinae. Scutellum and tegulae light ochreous. Forewings brown to dark brown, each with three large light areas: basally, around basal cell and basal portions of main veins; a band in the middle across of corium; a subapical patch in apical fourth of wing at wing margin (Fig. 3 A – B). Veins and hypocostal plate light brownish. Episternae and epimerae black. Fore and middle coxae light ochreous, with brown areas. Legs largely dark brown to black, with light ochreous bands. Abdominal tergites black, with ochreous spots and stripe along midline. Laterotergites black. Abdominal sternites dark brown to black, with light ochreous hind margins; sternites III – VI each with pair of large brown ochreous lateral patches. Fore and middle legs with second tarsomeres light ochreous dorsally; third tarsomeres and claws black. First metatarsomere black basally and apically, light ochreous medially; third metatarsomere light ochreous basally and black apically. Apices of spines on legs black. Anal tube light ochreous. Pygofer dark brown to black, except light ochreous near anal tube and ventral edge. Styles dark brown to black, with large light yellow patch on the plate. EXTERNAL STRUCTURE. Generally similar to O. litoralis gen. et sp. nov. Metope 0.8 times as long as wide at midline below the eyes (Figs 3 C, 5 E). Coryphe 2.5 times as wide as long medially (Figs 3 A, 5 D). Pronotum depressed in the middle, with very short median carina at hind margin (Fig. 5 D). Forewing reaching middle of tergite VI. First metatarsomere with two latero-apical and six intermediate spines arranged in arc. Claws, in dorsal view, 0.3 times as long as arolium of pretarsus. Arolium of pretarsus, in dorsal view, with straight hind margin. Dorso-lateral plates of pretarsus narrow. MALE TERMINALIA (Fig. 8). In general as described for O. litoralis gen. et sp. nov. Anal tube, in dorsal view, 0.8 times as wide as long medially, parallel-sided, anterior margin rounded (Fig. 8 B). Style with hind margin convex, caudal angle obtuse (Fig. 8 E); in dorsal view, style with distinct pointed lateral tooth (Fig. 8 F, lt). Phallobase with dorso-lateral lobes fused dorsally in one lobe with notched margin (Fig. 8 D). Aedeagus with each apical process with two long spiny teeth (Fig. 8 A, aep).	en	Gnezdilov, Vladimir M., Malenovský, Igor (2023): First record of the planthopper tribe Adenissini (Hemiptera: Caliscelidae) from Socotra Island, with description of a new genus and two new species. European Journal of Taxonomy 888 (1): 46-63, DOI: 10.5852/ejt.2023.888.2207, URL: http://dx.doi.org/10.5852/ejt.2023.888.2207
455A87DB446FFFAF2C4921428804FE51.taxon	distribution	Distribution and ecology So far only known from the highest elevations of the Hagher mountains in the central part of the Socotra Island (Fig. 9). The holotype was collected by sweeping on low shrubs in quite open, montane evergreen woodland (Fig. 10 B). The local low shrub plant community was dominated by Hypericum scopulorum Balf. f. (Hypericaceae), Leucas hagghierensis Al-Gifri & Cortés-Burns (Lamiaceae), and Euryops arabicus Steud. ex Jaub. & Spach. (Asteraceae). Oceatra scandens gen. et sp. nov. was found syntopic with the flatid planthoppers Socoflata aurolineata Stroiński, Malenovský & Świerczewski, 2018, S. histrionica Stroiński, Malenovský & Świerczewski, 2018, and Kirkamflata socotrana Świerczewski, Malenovský & Stroiński, 2014 (see Świerczewski et al. 2014 and Stroiński et al. 2018 for more details including a more explicit description of the habitat).	en	Gnezdilov, Vladimir M., Malenovský, Igor (2023): First record of the planthopper tribe Adenissini (Hemiptera: Caliscelidae) from Socotra Island, with description of a new genus and two new species. European Journal of Taxonomy 888 (1): 46-63, DOI: 10.5852/ejt.2023.888.2207, URL: http://dx.doi.org/10.5852/ejt.2023.888.2207
