taxonID	type	description	language	source
45458794FFAAFFE1A5DBFF75D86FFA81.taxon	materials_examined	Type species: Schuelkelia unicornis sp. n. (here designated).	en	Jałoszyński, Paweł (2015): Schuelkelia gen. n., a new eastern Palaearctic ant-like stone beetle, with synopsis of Eurasian genera of Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4007 (3): 343-369, DOI: 10.11646/zootaxa.4007.3.3
45458794FFAAFFE1A5DBFF75D86FFA81.taxon	diagnosis	Diagnosis. Body slender, with distinct and relatively deep constrictions between head and pronotum and between pronotum and elytra; head capsule with occipital constriction only slightly narrower than vertex; tempora longer than eyes; mandible with several small mesal teeth and prostheca; antennae with indistinct club composed of three terminal antennomeres; submentum lacking lateral sutures; hypostomal ridges long and posteriorly reaching posterior tentorial pits; tempora and vertex lacking bristles; prothorax lacking lateral edges and pronotum lacking antebasal pits or groove; basisternal part of prosternum only slightly shorter than coxal part; procoxal sockets broadly closed; prosternal process weakly elevated and very narrow, with pointed tip; notosternal sutures and hypomeral ridges entire; sides of prothorax with dense bristles; mesoventral intercoxal process long, parallel-sided but weakly elevated, extending from anterior ridge of mesoventrite to posterior margins of mesocoxae; mesothorax with deep ventrolateral foveae; metaventral intercoxal process with pair of long spines; each elytron with two asetose and barely discernible rudiments of basal foveae; aedeagus asymmetrical, with large and complicated assemblage of darkly sclerotized internal structures and free parameres.	en	Jałoszyński, Paweł (2015): Schuelkelia gen. n., a new eastern Palaearctic ant-like stone beetle, with synopsis of Eurasian genera of Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4007 (3): 343-369, DOI: 10.11646/zootaxa.4007.3.3
45458794FFAAFFE1A5DBFF75D86FFA81.taxon	description	Description. Body of male (Figs 1 – 2) elongate and slender, moderately convex, with deep constrictions between head and pronotum and between pronotum and elytra, brown and setose, BL about 1 mm. Head capsule (Figs 1 – 2, 3, 7 – 9) with anterior part (in front of occiput) approximately pentagonal in shape. Occipital constriction (Figs 3, 7; occ) only slightly narrower than vertex, extending dorsally and ventrally as transverse impressions separating anterior part of head capsule from ' neck' region (Figs 7 – 8; nr), which is subcylindrical and short. Tempora (Fig 7; tm) longer than eyes and rounded, weakly convergent posteromesally; vertex (Fig. 7; vt) strongly transverse; frons (Fig. 7; fr) subtrapezoidal, posteriorly confluent with vertex; frontoclypeal groove absent; anterior margin of clypeus (Fig. 7; cl) rounded; supraantennal tubercles weakly raised; antennal insertions broadly separated. Eyes bean-shaped (Fig. 8) and oblique in relation to long axis of head. Gular plate (Figs 3, 9; gp) very large and subtrapezoidal; gular sutures (Fig. 9; gs) superficial; posterior tentorial pits (Figs 3, 9; ptp) exposed, C-shaped and located slightly in front of transverse impression demarcating ' neck' region ventrally. Labrum (Fig. 10) transverse with indistinctly three-lobed anterior margin and two transverse rows of long dorsoanterior setae. Mandibles (Fig. 11) symmetrical, each subtriangular with broad base and curved, robust apical tooth (Fig. 11; ap), mesal margin with several small and sharp mesal teeth (Fig. 11; mt); prostheca (Fig. 11; pst) present, short and with dense setae along mesal margin. Each maxilla (Fig. 9) with subtriangular basistipes (Fig. 9; bst), broad mediostipes (Fig. 9; mst), elongate galea (Fig. 9; gal) and lacinia (Fig. 9; lac) and moderately long maxillary palp (Fig. 9; mxp) composed of distinctly elongate palpomere I, strongly elongate, pedunculate palpomere II, broad palpomere III broadest near distal third, and elongate and slender subconical and pointed palpomere IV. Labium (Fig. 9) with transverse submentum (Fig. 9; smn) not demarcated laterally from postcardinal parts of hypostomae; subrectangular mentum (Fig. 9; mn); and long prementum (Fig. 9; pmn) bearing small 3 - segmented labial palps (Fig. 9; lp) and indistinctly demarcated median ligula with one pair of long setae. Hypostomal ridges (Fig. 9; hr) long and recurved, extending posteromesally up to posterior tentorial pits, not connected. Antennae (Figs 1 – 2) short, with small club composed of antennomeres IX – XI. Pronotum (Figs 1 – 2, 12) in dorsal view oval with arcuate anterior and posterior margins, weakly marked and obtuse-angled anterior and posterior corners and strongly rounded sides; lateral edges or carinae absent; antebasal pits and groove absent. Sides of pronotum and hypomera with long, dense bristles. Prosternum (Figs 4, 13) with moderately long basisternal part (Fig. 13; bst) only slightly shorter than coxal part and indistinctly demarcated from procoxal cavities (Fig. 13; pcc); median part of sternum with weakly elevated and very narrow subtriangular prosternal intercoxal process with pointed tip; procoxal sockets (Fig. 13; pcs) broadly closed by lateral lobes of prosternum; hypomera (Fig. 13; hy) elongate, each divided into broad lateral part confluent with side of pronotum and narrow inner (adcoxal) part; pronotosternal sutures (Fig. 13; nss) and hypomeral ridges (Fig. 13; hyr) complete; posterior margin of sternum with deep median subtriangular emargination. Mesonotum subtrapezoidal, with subtriangular and posteriorly pointed mesoscutellum, mesoscutoscutellar suture not visible in studied specimen. Mesoscutellum not visible between elytral bases in intact specimens. Mesoventrite (Figs 5, 14) with broad anterior ridge (Fig. 14; ar); mesoventral intercoxal process (Fig. 14; msvp) long and parallel-sided but moderately elevated (not keel-like), extending from anterior ridge of mesoventrite up to posterior margins of mesocoxae; lateral impressions functioning as procoxal rests (Fig. 14; pcr) present, separated at middle and asetose except posterior margins; ventrolateral foveae (Fig. 14; vlf) present and deep, dorsolateral foveae absent; mesocoxal projections (Fig 14; mcp) prominent, each with short and setose posterior lobe (Fig. 14; pl) and mesocoxal sockets (Fig. 14; mscs) located on their mesoventral surface; mesocoxal cavities (Fig. 14; mscc) posteriorly demarcated by dense setae. Metaventrite (Fig. 14; v 3) subtrapezoidal, anteriorly fused with mesoventrite, posteriorly shallowly bisinuate and with moderately broad metaventral intercoxal process (Fig. 15; mtvp) bearing pair of long pointed spines. Metanepisterna and metepimera narrow. Metendosternite (= metafurca) (Fig. 14) with short and broad stalk and divergent lateral furcal arms (Fig. 15; lmfa). Elytra (Figs 1 – 2, 12) oval, with indistinct humeral calli and lacking basal impressions, each with two rudimentary and asetose basal foveae barely discernible even in transparent mounts; elytral apices rounded, unmodified. Hind wings absent. Legs (Figs 1 – 2, 14) moderately long and slender; pro- and mesocoxae oval, metacoxae strongly transverse; all trochanters short; all femora distinctly clavate; tibiae broadening distally; tarsi short and robust. Abdominal sternites (Fig. 6) unmodified, suture between VII and VIII distinct. Aedeagus (Figs 6, 15 – 16) very large, in repose occupying major part of abdomen between posterior margin of the first visible sternite and tip of abdomen; median lobe asymmetrical, stout; endophallus strongly asymmetrical and containing several large sclerites projecting distally from ostium; basal orifice located in submedian region of dorsal wall; parameres free (i. e., not fused with median lobe), each with single apical seta. Sexual dimorphism. Males with larger eyes, small horn-like posteromedian vertexal projection (Fig. 8) and strongly swollen profemora.	en	Jałoszyński, Paweł (2015): Schuelkelia gen. n., a new eastern Palaearctic ant-like stone beetle, with synopsis of Eurasian genera of Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4007 (3): 343-369, DOI: 10.11646/zootaxa.4007.3.3
45458794FFAAFFE1A5DBFF75D86FFA81.taxon	distribution	Distribution and composition. Schuelkelia is represented by one species occuring in Yunnan, China.	en	Jałoszyński, Paweł (2015): Schuelkelia gen. n., a new eastern Palaearctic ant-like stone beetle, with synopsis of Eurasian genera of Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4007 (3): 343-369, DOI: 10.11646/zootaxa.4007.3.3
45458794FFAAFFE1A5DBFF75D86FFA81.taxon	etymology	Etymology. The name Schuelkelia is dedicated to Michael Schülke, a distinguished specialist on Staphylinidae, who collected this new genus. Gender feminine.	en	Jałoszyński, Paweł (2015): Schuelkelia gen. n., a new eastern Palaearctic ant-like stone beetle, with synopsis of Eurasian genera of Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4007 (3): 343-369, DOI: 10.11646/zootaxa.4007.3.3
45458794FFAAFFE1A5DBFF75D86FFA81.taxon	discussion	Remarks. Among Palaearctic Cyrtoscydmini, Schuelkelia can be readily distinguished from all other genera on the basis of oval pronotum lacking antebasal pits and groove, elytra lacking externally visible basal elytral foveae and impressions, eyes distant from the broad occipital constriction, long and parallel-sided but not keel-like mesoventral intercoxal process, the metaventral intercoxal process with a pair of long spines and a strongly asymmetrical and heavily sclerotized, complicated endophallus. As in many genera of Scydmaeninae, this is a combination of synapomorphies that, separately or in different configurations, can be found in other taxa, but this particular set occurs together only in Schuelkelia. The only character that seems to be an autapomorphy of Schuelkelia is a row of small sharp mesal mandibular teeth; this feature is not known in any other world genus of Cyrtoscydmini that has been revised so far. Schuelkelia belongs to a group of genera characterized by the submentum not demarcated laterally by sutures. Remaining Eurasian genera showing this character state are (in alphabetical order): Elacatophora, Euconnus, Horaeomorphus, Leptocharis, Loeblites, Microscydmus, Nanoscydmus, Protoscydmus and Syndicus. Comparative notes on this group are given below:	en	Jałoszyński, Paweł (2015): Schuelkelia gen. n., a new eastern Palaearctic ant-like stone beetle, with synopsis of Eurasian genera of Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4007 (3): 343-369, DOI: 10.11646/zootaxa.4007.3.3
45458794FFA1FFEBA5DBFF3FD857FD34.taxon	materials_examined	Type material. Holotype: CHINA (Yunnan Province): ♂, two labels: “ CHINA: Yunnan SE Pingbian, / 22 º 54 ' 31 '' N, 103 º 41 ' 44 '' E, 2100 m, / primary subtropical broad-leaved / forest, litter sifted, 28. VIII. 2014, / leg. M. Schülke [CH 14 - 22 a] ” [white, printed]; “ SCHUELKELIA / unicornis m. / det. P. Jałoszyński, 2015 / HOLOTYPUS ” [red, printed] (cMS). Paratypes (8 exx.): 3 ♂ (one disarticulated), 5 ♀, same data as holotype, except for yellow “ paratypus ” label (cMS, cPJ).	en	Jałoszyński, Paweł (2015): Schuelkelia gen. n., a new eastern Palaearctic ant-like stone beetle, with synopsis of Eurasian genera of Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4007 (3): 343-369, DOI: 10.11646/zootaxa.4007.3.3
45458794FFA1FFEBA5DBFF3FD857FD34.taxon	diagnosis	Diagnosis. This is the only known species of Schuelkelia and can be identified on the basis of generic diagnostic characters and structures of the aedeagus (Figs 15 – 16).	en	Jałoszyński, Paweł (2015): Schuelkelia gen. n., a new eastern Palaearctic ant-like stone beetle, with synopsis of Eurasian genera of Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4007 (3): 343-369, DOI: 10.11646/zootaxa.4007.3.3
45458794FFA1FFEBA5DBFF3FD857FD34.taxon	description	Description. Body of male (Figs 1 – 2) slender and moderately convex, brown, covered with setae slightly lighter than cuticle; BL 1.03 – 1.05 mm (mean 1.04 mm). Head (Figs 1 – 2, 3, 7 – 9) broadest at large, moderately convex and moderately coarsely faceted eyes, HL 0.18 mm, HW 0.18 – 0.19 mm (mean 0.19 mm); vertex and frons confluent, vertex laterally weakly convex, with small posteromedian horn-like projection (Figs 7 – 8), sides of frons between eyes slightly impressed; supraantennal tubercles feebly raised. Punctures on frons and vertex inconspicuous, fine; setae short, sparse and suberect. Antennae slender but short, AnL 0.38 – 0.40 mm (mean 0.35 mm), antennomeres I and II strongly elongate, III – V each about as long as broad, VI slightly transverse, VII as long as broad, VIII – X each distinctly transverse, XI about as long as broad, rounded at apex. Pronotum (Figs 1 – 2, 12) oval, broadest slightly in front of middle; PL 0.26 – 0.28 mm (mean 0.27 mm), PW 0.25 – 0.28 mm (mean 0.26 mm). Anterior margin nearly straight; anterior corners strongly obtuse-angled and blunt; sides strongly rounded; posterior corners indistinct, strongly obtuse-angled and broadly rounded; posterior margin arcuate with indistinct posteromedian expansion. Punctures on pronotal disc fine and inconspicuous; setae on dorsum sparse, moderately long and suberect, sides of pronotum with dense and long bristles with admixture of long, curved setae (Fig. 13). Elytra (Figs 1 – 2) oval, broadest at middle or slightly in front of middle; EL 0.58 – 0.60 mm (mean 0.59 mm), EW 0.38 mm, EI 1.53 – 1.60; humeral calli indistinct; basal impressions absent, apices rounded together. Punctures on elytra fine and inconspicuous; setae sparse, moderately long, suberect. Hind wings absent. Legs moderately long and slender, profemora (Fig. 1) distinctly swollen, much broader than meso- and metafemora. Aedeagus (Figs 15 – 16) asymmetrical and stout; AeL 0.28 mm; median lobe in ventral view broadest near base, with basal orifice located in sub-median region of dorsal wall; endophallus with several darkly sclerotized and robust asymmetrical sclerites, of which two long and slender and one broad and short protrude from apical ostium; parameres short, each with single long apical seta. Female. Similar to male but with unmodified vertex, slightly less swollen profemora, slightly smaller eyes, on average smaller body and shorter antennae; BL 0.91 – 0.98 mm (mean 0.95 mm); HL 0.16 – 0.18 mm (mean 0.17 mm), HW 0.16 – 0.18 mm (mean (0.17 mm), AnL 0.33 – 0.35 mm (mean 0.34 mm); PL 0.25 mm, PW 0.24 – 0.25 mm (mean 0.24 mm); EL 0.50 – 0.55 mm (mean 0.53 mm), EW 0.35 – 0.38 mm (mean 0.36 mm), EI 1.43 – 1.50.	en	Jałoszyński, Paweł (2015): Schuelkelia gen. n., a new eastern Palaearctic ant-like stone beetle, with synopsis of Eurasian genera of Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4007 (3): 343-369, DOI: 10.11646/zootaxa.4007.3.3
45458794FFA1FFEBA5DBFF3FD857FD34.taxon	distribution	Distribution and habitat. Southern part of central China; all specimens were collected by sifting leaf litter in a subtropical forest (Fig. 17).	en	Jałoszyński, Paweł (2015): Schuelkelia gen. n., a new eastern Palaearctic ant-like stone beetle, with synopsis of Eurasian genera of Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4007 (3): 343-369, DOI: 10.11646/zootaxa.4007.3.3
45458794FFA1FFEBA5DBFF3FD857FD34.taxon	etymology	Etymology. The name unicornis refers to the tiny posteromedian horn-like projection of the vertex in male.	en	Jałoszyński, Paweł (2015): Schuelkelia gen. n., a new eastern Palaearctic ant-like stone beetle, with synopsis of Eurasian genera of Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4007 (3): 343-369, DOI: 10.11646/zootaxa.4007.3.3
