identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
45458794FFAAFFE1A5DBFF75D86FFA81.text	45458794FFAAFFE1A5DBFF75D86FFA81.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Schuelkelia	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Schuelkelia gen. n.</p>
            <p> Type species:  Schuelkelia unicornis sp. n. (here designated). </p>
            <p>Diagnosis. Body slender, with distinct and relatively deep constrictions between head and pronotum and between pronotum and elytra; head capsule with occipital constriction only slightly narrower than vertex; tempora longer than eyes; mandible with several small mesal teeth and prostheca; antennae with indistinct club composed of three terminal antennomeres; submentum lacking lateral sutures; hypostomal ridges long and posteriorly reaching posterior tentorial pits; tempora and vertex lacking bristles; prothorax lacking lateral edges and pronotum lacking antebasal pits or groove; basisternal part of prosternum only slightly shorter than coxal part; procoxal sockets broadly closed; prosternal process weakly elevated and very narrow, with pointed tip; notosternal sutures and hypomeral ridges entire; sides of prothorax with dense bristles; mesoventral intercoxal process long, parallel-sided but weakly elevated, extending from anterior ridge of mesoventrite to posterior margins of mesocoxae; mesothorax with deep ventrolateral foveae; metaventral intercoxal process with pair of long spines; each elytron with two asetose and barely discernible rudiments of basal foveae; aedeagus asymmetrical, with large and complicated assemblage of darkly sclerotized internal structures and free parameres.</p>
            <p>Description. Body of male (Figs 1–2) elongate and slender, moderately convex, with deep constrictions between head and pronotum and between pronotum and elytra, brown and setose, BL about 1 mm.</p>
            <p>Head capsule (Figs 1–2, 3, 7–9) with anterior part (in front of occiput) approximately pentagonal in shape. Occipital constriction (Figs 3, 7; occ) only slightly narrower than vertex, extending dorsally and ventrally as transverse impressions separating anterior part of head capsule from 'neck' region (Figs 7–8; nr), which is subcylindrical and short. Tempora (Fig 7; tm) longer than eyes and rounded, weakly convergent posteromesally; vertex (Fig. 7; vt) strongly transverse; frons (Fig. 7; fr) subtrapezoidal, posteriorly confluent with vertex; frontoclypeal groove absent; anterior margin of clypeus (Fig. 7; cl) rounded; supraantennal tubercles weakly raised; antennal insertions broadly separated. Eyes bean-shaped (Fig. 8) and oblique in relation to long axis of head.</p>
            <p>Gular plate (Figs 3, 9; gp) very large and subtrapezoidal; gular sutures (Fig. 9; gs) superficial; posterior tentorial pits (Figs 3, 9; ptp) exposed, C-shaped and located slightly in front of transverse impression demarcating 'neck' region ventrally.</p>
            <p>Labrum (Fig. 10) transverse with indistinctly three-lobed anterior margin and two transverse rows of long dorsoanterior setae. Mandibles (Fig. 11) symmetrical, each subtriangular with broad base and curved, robust apical tooth (Fig. 11; ap), mesal margin with several small and sharp mesal teeth (Fig. 11; mt); prostheca (Fig. 11; pst) present, short and with dense setae along mesal margin. Each maxilla (Fig. 9) with subtriangular basistipes (Fig. 9; bst), broad mediostipes (Fig. 9; mst), elongate galea (Fig. 9; gal) and lacinia (Fig. 9; lac) and moderately long maxillary palp (Fig. 9; mxp) composed of distinctly elongate palpomere I, strongly elongate, pedunculate palpomere II, broad palpomere III broadest near distal third, and elongate and slender subconical and pointed palpomere IV. Labium (Fig. 9) with transverse submentum (Fig. 9; smn) not demarcated laterally from postcardinal parts of hypostomae; subrectangular mentum (Fig. 9; mn); and long prementum (Fig. 9; pmn) bearing small 3- segmented labial palps (Fig. 9; lp) and indistinctly demarcated median ligula with one pair of long setae. Hypostomal ridges (Fig. 9; hr) long and recurved, extending posteromesally up to posterior tentorial pits, not connected.</p>
            <p>Antennae (Figs 1–2) short, with small club composed of antennomeres IX–XI.</p>
            <p>Pronotum (Figs 1–2, 12) in dorsal view oval with arcuate anterior and posterior margins, weakly marked and obtuse-angled anterior and posterior corners and strongly rounded sides; lateral edges or carinae absent; antebasal pits and groove absent. Sides of pronotum and hypomera with long, dense bristles.</p>
            <p>Prosternum (Figs 4, 13) with moderately long basisternal part (Fig. 13; bst) only slightly shorter than coxal part and indistinctly demarcated from procoxal cavities (Fig. 13; pcc); median part of sternum with weakly elevated and very narrow subtriangular prosternal intercoxal process with pointed tip; procoxal sockets (Fig. 13; pcs) broadly closed by lateral lobes of prosternum; hypomera (Fig. 13; hy) elongate, each divided into broad lateral part confluent with side of pronotum and narrow inner (adcoxal) part; pronotosternal sutures (Fig. 13; nss) and hypomeral ridges (Fig. 13; hyr) complete; posterior margin of sternum with deep median subtriangular emargination.</p>
            <p>Mesonotum subtrapezoidal, with subtriangular and posteriorly pointed mesoscutellum, mesoscutoscutellar suture not visible in studied specimen. Mesoscutellum not visible between elytral bases in intact specimens.</p>
            <p>Mesoventrite (Figs 5, 14) with broad anterior ridge (Fig. 14; ar); mesoventral intercoxal process (Fig. 14; msvp) long and parallel-sided but moderately elevated (not keel-like), extending from anterior ridge of mesoventrite up to posterior margins of mesocoxae; lateral impressions functioning as procoxal rests (Fig. 14; pcr) present, separated at middle and asetose except posterior margins; ventrolateral foveae (Fig. 14; vlf) present and deep, dorsolateral foveae absent; mesocoxal projections (Fig 14; mcp) prominent, each with short and setose posterior lobe (Fig. 14; pl) and mesocoxal sockets (Fig. 14; mscs) located on their mesoventral surface; mesocoxal cavities (Fig. 14; mscc) posteriorly demarcated by dense setae.</p>
            <p>Metaventrite (Fig. 14; v3) subtrapezoidal, anteriorly fused with mesoventrite, posteriorly shallowly bisinuate and with moderately broad metaventral intercoxal process (Fig. 15; mtvp) bearing pair of long pointed spines. Metanepisterna and metepimera narrow.</p>
            <p>Metendosternite (= metafurca) (Fig. 14) with short and broad stalk and divergent lateral furcal arms (Fig. 15; lmfa).</p>
            <p>Elytra (Figs 1–2, 12) oval, with indistinct humeral calli and lacking basal impressions, each with two rudimentary and asetose basal foveae barely discernible even in transparent mounts; elytral apices rounded, unmodified.</p>
            <p>Hind wings absent.</p>
            <p>Legs (Figs 1–2, 14) moderately long and slender; pro- and mesocoxae oval, metacoxae strongly transverse; all trochanters short; all femora distinctly clavate; tibiae broadening distally; tarsi short and robust.</p>
            <p>Abdominal sternites (Fig. 6) unmodified, suture between VII and VIII distinct.</p>
            <p>Aedeagus (Figs 6, 15–16) very large, in repose occupying major part of abdomen between posterior margin of the first visible sternite and tip of abdomen; median lobe asymmetrical, stout; endophallus strongly asymmetrical and containing several large sclerites projecting distally from ostium; basal orifice located in submedian region of dorsal wall; parameres free (i.e., not fused with median lobe), each with single apical seta.</p>
            <p>Sexual dimorphism. Males with larger eyes, small horn-like posteromedian vertexal projection (Fig. 8) and strongly swollen profemora.</p>
            <p> Distribution and composition.  Schuelkelia is represented by one species occuring in Yunnan, China. </p>
            <p> Etymology. The name  Schuelkelia is dedicated to Michael Schülke, a distinguished specialist on  Staphylinidae , who collected this new genus. Gender feminine. </p>
            <p> Remarks. Among Palaearctic  Cyrtoscydmini ,  Schuelkelia can be readily distinguished from all other genera on the basis of oval pronotum lacking antebasal pits and groove, elytra lacking externally visible basal elytral foveae and impressions, eyes distant from the broad occipital constriction, long and parallel-sided but not keel-like mesoventral intercoxal process, the metaventral intercoxal process with a pair of long spines and a strongly asymmetrical and heavily sclerotized, complicated endophallus. As in many genera of  Scydmaeninae , this is a combination of synapomorphies that, separately or in different configurations, can be found in other taxa, but this particular set occurs together only in  Schuelkelia . The only character that seems to be an autapomorphy of  Schuelkelia is a row of small sharp mesal mandibular teeth; this feature is not known in any other world genus of  Cyrtoscydmini that has been revised so far. </p>
            <p> Schuelkelia belongs to a group of genera characterized by the submentum not demarcated laterally by sutures. Remaining Eurasian genera showing this character state are (in alphabetical order):  Elacatophora ,  Euconnus ,  Horaeomorphus ,  Leptocharis ,  Loeblites ,  Microscydmus, Nanoscydmus ,  Protoscydmus and  Syndicus . Comparative notes on this group are given below: </p>
            <p> Schuelkelia differs from  Elacatophora (Figs 18, 36, 51, 66) in the head capsule in lateral view only slightly higher near antennal insertions than at occipital constriction (head near antennal insertions about twice as high as at occipital constriction in  Elacatophora ), occipital constriction only slightly narrower than frons (as narrow as about half of frons or less in  Elacatophora , except one species,  E. euconnoides Jałoszyński, 2015b ), posterior ends of hypostomal ridges and posterior tentorial pits located clearly behind eyes (between eyes in  Elacatophora ), gular plate lacking long and narrow anterior part in front of the 'neck' region (in  Elacatophora gular plate with broad posterior part located on the 'neck' region and with much narrower, strongly elongate anterior part expanding onto the anterior part of head capsule and extending up to the level of eyes), maxillary palpomere IV subconical, with broad base (palpomere IV nearly rod-like in  Elacatophora ), pronotum lacking sublateral carinae and antebasal impressions (with carinae and impressions in  Elacatophora ), mesoventral intercoxal process weakly elevated (keel-like and strongly expanding ventrally in  Elacatophora ), metaventral intercoxal process with a pair of long spines (lacking spines in  Elacatophora ), elytra with rudimentary basal foveae (each elytron with one large and distinct fovea in  Elacatophora ). </p>
            <p> Schuelkelia differs from  Euconnus (Figs 19–22, 37, 52, 67) and  Protoscydmus in the lack of frontoclypeal groove (present in  Euconnus and  Protoscydmus ), basal elytral foveae visible only as indistinct rudiments in transparent mounts (two foveae on each elytron in  Euconnus , although in some subgenera, mainly from the Southern Hemisphere, and in  Protoscydmus rudimentary, but still marked on the surface of elytra), nearly contiguous metacoxae (moderately broadly to narrowly but always distinctly separated in  Euconnus and  Protoscydmus ; not separated in some Southern Hemisphere taxa currently placed as subgenera of  Euconnus , but their taxonomic status requires verification), metaventral intercoxal process with a pair of long spines (spines absent in  Euconnus and  Protoscydmus ). </p>
            <p> Schuelkelia differs from  Horaeomorphus (Figs 23, 38, 53, 68) in the pronotum lacking antebasal pits and groove (pits always present in  Horaeomorphus , often connected by a transverse groove), prosternum with weakly marked prosternal process with pointed posterior tip (in  Horaeomorphus prosternal process distinct, parallel-sided, lacking defined posterior tip), mesoventral intercoxal process parallel-sided and extending in front of mesocoxal cavities (in  Horaeomorphus process broad at base and narrowing caudad, not extending anteriorly beyond mesocoxal cavities), elytra with rudimentary basal foveae (two distinct foveae on each elytron in  Horaeomorphus ). </p>
            <p> Schuelkelia differs from  Leptocharis (Figs 24, 39, 54, 69) in rudimentary basal elytral foveae (one deep and distinct fovea and a hardly noticeable rudiment of another in  Leptocharis ), subtrapezoidal and broad mesoscutellum (strongly elongate with broadened posterior portion in  Leptocharis ), hypomeral ridges present (absent in  Leptocharis ), the mesoventral intercoxal process extending from the anterior ridge of mesoventrite up to posterior margins of mesocoxae (process in  Leptocharis extends from anterior to posterior margins of mesocoxae, intercoxal region in front of mesocoxae is convex but broad and not keel-like), mesothorax with a pair of deep ventrolateral foveae (absent in  Leptocharis ), the metaventral intercoxal process with spines (with a deep notch but lacking spines in  Leptocharis ). </p>
            <p> Schuelkelia differs from  Loeblites (Figs 25, 40, 55, 70) in short and clubbed antennae (very long, nearly filiform antennae in  Loeblites ), posterior tentorial pits exposed (hidden in a deep transverse groove in  Loeblites ), pronotum lacking antebasal pits (pits present in  Loeblites ), mesoventral intercoxal process extending from anterior ridge of mesoventrite to posterior margins of mesocoxae (process extending only from anterior to posterior margins of coxae in  Loeblites ), metaventral intercoxal process with a pair of long spines (lacking spines in  Loeblites ), elytra with rudimentary basal foveae (two deep and distinct foveae on each elytron in  Loeblites ), femora lacking dorsal grooves and glandular openings (grooves with glandular openings present on all femora in  Loeblites ). </p>
            <p> Schuelkelia differs from  Microscydmus (Figs 26, 41, 56, 71) in antennal insertions broadly separated (narrowly separated in  Microscydmus ), hypostomal ridges reaching posterior tentorial pits (ridges very short in  Microscydmus , not reaching posterior tentorial pits), notosternal sutures complete and visible on the surface (largely internalized in  Microscydmus ), pronotum lacking antebasal pits (pits present in  Microscydmus ), mesoventral intercoxal process long, connected with the anterior ridge of mesoventrite (process short, not connected with anterior ridge in  Microscydmus ); elytra with rudimentary foveae (one large and setose basal fovea on each elytron in  Microscydmus ). </p>
            <p> Schuelkelia differs from Nanoscydmus (Figs 27, 42, 57, 72) in the head capsule lacking median longitudinal groove and V-shaped carinae (both present in Nanoscydmus), tempora longer than eyes and therefore eyes distant from the occipital constriction (tempora much shorter than eyes and consequently eyes nearly adjacent to occipital constriction in Nanoscydmus), pronotum lacking antebasal pits and groove (with lateral pits connected by transverse groove in Nanoscydmus), present hypomeral ridges (absent in Nanoscydmus), mesoventral intercoxal process extending from anterior ridge of mesoventrite to posterior margins of mesocoxae (interrupted in front of mesocoxae in Nanoscydmus), metaventral intercoxal process with spines (lacking spines in Nanoscydmus). </p>
            <p> Schuelkelia differs from  Syndicus (Figs 34–35, 50, 65, 80) in the base of antennomere XI clearly separated from apex of antennomere XI by constriction (antennomeres X and XI forming a compact oval or at least tightly assembled, not separated in  Syndicus ), posterior tentorial pits exposed (hidden in a deep transverse groove in  Syndicus ); pronotum lacking antebasal pits (pits present in  Syndicus ), mesoventral intercoxal process extending from anterior ridge of mesoventrite to posterior margins of mesocoxae (process extending only from anterior to posterior margins of coxae in  Syndicus ), metaventral intercoxal process with a pair of long spines (lacking spines in  Syndicus ), elytra with rudimentary basal foveae (two deep and distinct foveae on each elytron in  Syndicus ), femora lacking dorsal grooves and glandular openings (grooves with glandular openings present on all femora in  Syndicus ). </p>
            <p> Among  Cyrtoscydmini known to occur in other regions and whose morphological details are sufficiently known, the submentum lacking lateral sutures can be found in sixteen genera; they differ from  Schuelkelia in characters listed below (only major features are mentioned): </p>
            <p> Schuelkelia differs from Amimoscydmus Jałoszyński, 2013a (Brazil) in hypostomal ridges posteriorly reaching posterior tentorial pits (ridges very short, not reaching tentorial pits in Amimoscydmus), long basisternal part of prosternum (rudimentary in Amimoscydmus), inner part of hypomera completely demarcated from prosternum (anteriorly and posteriorly fused with prosternum in Amimoscydmus), and pronotum lacking antebasal transverse groove (with groove in Amimoscydmus). </p>
            <p> Schuelkelia differs from  Archiconnus Franz, 1980 (Peru) in maxillary palpomere III slender (strongly swollen in  Archiconnus ), pronotum lacking antebasal pits (pits present in  Archiconnus ), prosternal process present (absent in  Archiconnus ), hypomeral ridges complete (anteriorly obliterated in  Archiconnus ), and metaventral intercoxal process with a pair of long spines (lacking spines in  Archiconnus ). </p>
            <p> Schuelkelia differs from  Euconnomorphus Franz, 1980 (Peru, Bolivia) in pronotum lacking antebasal pits (pits present in  Euconnomorphus ), prosternal process present (absent in  Euconnomorphus ), long basisternal part of prosternum (much shorter than coxal part in  Euconnomorphus ), mesoventral intercoxal process uninterrupted in front of mesocoxae (interrupted in  Euconnomorphus ), and metaventral intercoxal process with a pair of long spines (lacking spines in  Euconnomorphus ). </p>
            <p> Schuelkelia differs from  Heteroscydmus Franz, 1980 (Bolivia) in hypostomal ridges posteriorly reaching posterior tentorial pits (ridges very short, not reaching tentorial pits in  Heteroscydmus ), inner part of hypomera completely demarcated from prosternum (anteriorly fused with prosternum in  Heteroscydmus ), and pronotum lacking antebasal transverse groove (with groove in  Heteroscydmus ). </p>
            <p> Schuelkelia differs from Leascydmus Jałoszyński, 2014c (Australia) in the occipital constriction only slightly narrower than vertex (about half as broad as vertex in Leascydmus), broadly closed procoxal sockets (narrowly open in Leascydmus), absent anterior metaventral process (present in Leascydmus), contiguous metacoxae (broadly separated in Leascydmus), and metaventral intercoxal process with a pair of long spines (lacking spines in Leascydmus). </p>
            <p> Schuelkelia differs from  Microraphes Franz, 1980 (Brazil) in elongate head with long tempora (head short and with tempora much shorter than eyes in  Microraphes ), inner part of hypomera completely demarcated from prosternum (anteriorly fused with prosternum in  Microraphes ), basisternal part of prosternum shorter than coxal part (longer in  Microraphes ), prosternal process long and carinate (short and drop-shaped in  Microraphes ), and metaventral intercoxal process with a pair of long spines (lacking spines in  Microraphes ). </p>
            <p> Schuelkelia differs from  Mimoscydmus Franz, 1980 (Brazil) in elongate head with long tempora (head short and with tempora much shorter than eyes in  Mimoscydmus ), prosternal process lacking defined anterior tip (with anterior tip in  Mimoscydmus ), inner part of hypomera completely demarcated from prosternum (anteriorly fused with prosternum in  Mimoscydmus ), and pronotum lacking antebasal transverse groove (with groove in  Mimoscydmus ). </p>
            <p> Schuelkelia differs from  Parapseudoconnus Franz, 1980 (Brazil, Peru) in pronotum lacking antebasal pits (pits present in  Parapseudoconnus ), long basisternal part of prosternum (much shorter than coxal part in  Parapseudoconnus ), inner part of hypomera completely demarcated from prosternum (posteriorly fused with prosternum in  Parapseudoconnus ), elytra with rudiments of foveae (each elytron with one deep and setose basal fovea in  Parapseudoconnus ), and metaventral intercoxal process with a pair of long spines (lacking spines in  Parapseudoconnus ). </p>
            <p> Schuelkelia differs from Penicillidmus Jałoszyński, 2014b (Australia) in maxillary palpomere IV subconical with base about four times as broad as apical part (very narrow, with base only about twice as broad as apical part in Penicillidmus), head capsule lacking longitudinal and transverse carinae behind cardines (carinae present in Penicillidmus), pronotum lacking posterolateral rod-like penicilli (penicilli present in Penicillidmus), pronotum lacking antebasal pits (pits present in Penicillidmus), inner part of hypomera completely demarcated from prosternum (anteriorly fused with prosternum in Penicillidmus), and elytra with rudiments of basal foveae (each elytron with one large fovea located in a deep impression in Penicillidmus). </p>
            <p> Schuelkelia differs from  Sciacharis Broun, 1893 (Southern Hemisphere) in prothorax lacking antebasal pits (pits present in  Sciacharis ), elytra with rudiments of basal foveae (two distinct foveae on each elytron in  Sciacharis ), and the metaventral intercoxal process with a pair of long spines (lacking spines in  Sciacharis ). </p>
            <p> Schuelkelia differs from Sciacharoides Jałoszyński, 2014c (Australia) in prothorax lacking antebasal pits and grooves (pits and/or grooves present in Sciacharoides), mesocoxae distinctly separated by long mesoventral intercoxal process (process absent between coxae in Sciacharoides), and metaventral intercoxal process with a pair of long spines (lacking spines in Sciacharoides). </p>
            <p> Schuelkelia differs from Scydmaenozila Jałoszyński, 2014c (Australia) in prothorax lacking antebasal pits (pits present in Scydmaenozila), prosternal process present and carinate (indistinct in Scydmaenozila), complete hypomeral ridges (rudimentary in Scydmaenozila), mesoventral intercoxal process extending from anterior ridge of mesoventrite up to posterior margins of mesocoxae (process interrupted in front of mesocoxae in Scydmaenozila), and metaventral intercoxal process with a pair of long spines (lacking spines in Scydmaenozila). </p>
            <p> Schuelkelia differs from Scydmepitoxis Jałoszyński, 2014f (Australia) in the occipital constriction only slightly narrower than vertex (about half as broad as vertex in Scydmepitoxis), prothorax lacking deep posterior constriction and lacking antebasal pits (with these structures in Scydmepitoxis), the prosternal process present and carinate (absent in Scydmepitoxis), mesoventral intercoxal process long and distinct (indistinct, barely noticeable between mesocoxae in their anterior half in Scydmepitoxis), and metaventral intercoxal process with a pair of long spines (lacking spines in Scydmepitoxis). </p>
            <p> Schuelkelia differs from Spinosciacharis Jałoszyński, 2014c (Australia) in the occipital constriction only slightly narrower than vertex (about half as broad as vertex in Spinosciacharis), prothorax lacking antebasal pits and elytra with rudiments of basal foveae (pits and foveae present and distinct in Spinosciacharis), absent anterior metaventral process (present in Spinosciacharis), and metaventral intercoxal process with a pair of long spines (lacking spines in Spinosciacharis). </p>
            <p> Schuelkelia differs from  Stenichnoteras Scott, 1922 (Seychelles) in the prosternal process present and carinate (absent in  Stenichnoteras ), mesoventral intercoxal process extending from anterior ridge of mesoventrite up to posterior margins of mesocoxae (process interrupted in front of mesocoxae and posteriorly reaching middle of mesocoxae in  Stenichnoteras ), contiguous metacoxae (moderately broadly separated in  Stenichnoteras ), and metaventral intercoxal process with a pair of long spines (lacking spines in  Stenichnoteras ). </p>
            <p> Schuelkelia differs from  Venezolanoconnus Franz, 1988 (Venezuela) in maxillary palpomere III slender (strongly swollen in  Venezolanoconnus ), prosternal process present (absent in  Venezolanoconnus ), elytra lacking basal impressions and subhumeral lines (with distinct impressions and long subhumeral lines in  Venezolanoconnus ), and slender parameres (parameres extremely broad and hyaline in  Venezolanoconnus ). </p>
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	https://treatment.plazi.org/id/45458794FFAAFFE1A5DBFF75D86FFA81	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2015): Schuelkelia gen. n., a new eastern Palaearctic ant-like stone beetle, with synopsis of Eurasian genera of Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4007 (3): 343-369, DOI: 10.11646/zootaxa.4007.3.3
45458794FFAEFFE0A5DBFA64D8B7FB6B.text	45458794FFAEFFE0A5DBFA64D8B7FB6B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyrtoscydmini	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Key to Eurasian genera of  Cyrtoscydmini (  Protoscydmus Franz omitted) </p>
            <p> 1. Antenna lacking constriction between antennomeres X and XI which appear closely connected and antennae appear 10-segmented [subtropical Asia and Himalaya Mts]...................................  Syndicus Motschulsky (Figs 34–35) </p>
            <p>- Antennomere X and XI distinctly separated................................................................ 2</p>
            <p> 2. Antennae nearly as long as body and nearly filiform, very weakly and gradually thickened distally; femur of each leg with narrow longitudinal dorsal groove and small glandular opening [subtropical Asia]..................  Loeblites Franz (Fig. 25) </p>
            <p>- Antennae shorter than body and not filiform, either with club composed of 2–5 terminal antennomeres or distinctly gradually thickened distally; femora lacking dorsal grooves and glandular openings......................................... 3</p>
            <p>3. Distance between posterior margin of eye and occipital constriction (i.e., length of temple) comparable to length of eye or larger, eyes not adjacent to anterior margin of pronotum in intact specimens....................................... 4</p>
            <p>- Distance between posterior margin of eye and occipital constriction much shorter than length of eye, eyes adjacent or nearly adjacent to anterior margin of pronotum................................................................... 10</p>
            <p>4. Each elytron with two basal foveae distinct in dry-mounted and intact specimens.................................. 5</p>
            <p>- Each elytron with one basal fovea or rudiments of foveae not visible in dry-mounted and intact specimens.............. 6</p>
            <p>5. Mesoventral intercoxal process shorter than mesocoxae and less elevated than mesocoxae (in lateral view its ventral margin concealed by mesocoxae); metaventral intercoxal process with two long spines projecting posteriorly; pronotal base with distinct median pit in addition to lateral pits; head with small pit at posteromesal margin of each supraantennal tubercle; basal ely-</p>
            <p> tral foveae connected by U-shaped groove extending anteriorly [subtropical Asia and Himalaya Mts]...........................................................................................  Horaeomorphus Schaufuss (Fig. 23) - Mesoventral intercoxal process keel-like, longer than mesocoxae and more elevated than mesocoxae (its ventral margin visible in lateral view, not concealed by mesocoxae); metaventral intercoxal process lacking spines, with straight or concave posterior margin; pronotal base lacking median pit; head lacking pits near supraantennal tubercles; basal elytral foveae not connected [cosmopolitan]..............................................................  Euconnus Thomson (Figs 19–22) </p>
            <p> 6. Elytra with basal foveae not visible in dry-mounted specimens [China: Yunnan]........  Schuelkelia Jałoszyński (Figs 1–2) </p>
            <p>- Each elytron with one distinct fovea...................................................................... 7</p>
            <p>7. Pronotum in dorsal view broadest at or near base and narrowing anteriorly........................................ 8</p>
            <p>- Pronotum broadest near middle or in front of middle, narrowing both anteriorly and posteriorly........................ 9</p>
            <p> 8. Posterior margin of vertex with four distinct tufts or rosettes of dense bristles; maxillary palpomere III weakly narrowing in distal half; maxillary palpomere IV subconical [Thailand]..................................  Siamites Franz (Fig. 31) </p>
            <p> - Posterior margin of vertex with setae not arranged in paired tufts; maxillary palpomere III strongly narrowing in distal half; maxillary palpomere IV nearly rod-like, very slender and visible as terminal spine [Malaysia, Indonesia]............................................................................................  Elacatophora Schaufuss (Fig. 18) </p>
            <p> 9. Pronotum elongate, lacking paired basal pits; hypomeral ridges absent; metaventral intercoxal process notched at middle and lacking pair of long spines [Mediterranean]..........................................  Leptocharis Reitter (Fig. 24) </p>
            <p> - Pronotum about as long as broad, with paired basal pits; hypomeral ridges present and complete; metaventral intercoxal process with a pair of robust spines [cosmopolitan]........................  Microscydmus Saulcy &amp; Croissandeau (Fig. 26) </p>
            <p>10. Head with raised dorsal median triangular area pointing posteriorly and with median longitudinal groove [China, Nepal]............................................................................. Nanoscydmus Jałoszyński (Fig. 27)</p>
            <p>- Head lacking raised median triangular area and lacking longitudinal groove..................................... 11</p>
            <p> 11. Each elytron with one large basal fovea filled with dense setae [panpalaearctic]............  Neuraphes Thomson (Fig. 28) </p>
            <p>- Each elytron with variously visible but always asetose basal fovea.............................................. 12</p>
            <p>12. Pronotum with transverse antebasal groove................................................................ 13</p>
            <p>- Pronotum lacking transverse antebasal groove.............................................................. 14</p>
            <p> 13. Basal elytral fovea large and well-visible; mesoventral intercoxal process posteriorly reaching to middle of mesocoxa; anterior metaventral process present; aedeagus with parameres [panpalaearctic]..................  Scydmoraphes Reitter (Fig. 30) </p>
            <p> - Basal elytral fovea barely noticeable; mesoventral intercoxal process posteriorly reaching posterior margin of mesocoxa; anterior metaventral process absent; aedeagus lacking parameres [Sumatra, Sri Lanka]...............................................................................................  Stenichnodes (  Parastenichnaphes Franz ) (Fig. 32) </p>
            <p> 14. Pronotum with rounded lateral margins, lacking lateral edges; metaventral intercoxal process notched but lacking pair of spines; aedeagus with parameres [panpalaearctic]....................................  Stenichnus Thomson (Fig. 33) </p>
            <p>-. Pronotum with lateral edges in posterior half; metaventral intercoxal process with a pair of long spines; aedeagus lacking parameres [Japan: Shikoku].................................................... Rutaraphes Jałoszyński (Fig. 29)</p>
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	https://treatment.plazi.org/id/45458794FFAEFFE0A5DBFA64D8B7FB6B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2015): Schuelkelia gen. n., a new eastern Palaearctic ant-like stone beetle, with synopsis of Eurasian genera of Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4007 (3): 343-369, DOI: 10.11646/zootaxa.4007.3.3
45458794FFAFFFEFA5DBFAE6D8B7FE6E.text	45458794FFAFFFEFA5DBFAE6D8B7FE6E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyrtoscydmini	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Key to Chinese genera of  Cyrtoscydmini</p>
            <p> 1. Antenna lacking constriction between antennomeres X and XI which are closely connected and antennae appear 10-segmented.............................................................................  Syndicus Motschulsky</p>
            <p>- Antennomere X and XI distinctly separated................................................................. 2</p>
            <p> 2. Antennae nearly as long as body and nearly filiform, weakly and gradually thickened distally; femur of each leg with narrow longitudinal dorsal groove and small glandular opening...........................................  Loeblites Franz</p>
            <p>- Antennae shorter than body and not filiform, either with club composed of 2–5 terminal antennomeres or distinctly gradually thickened distally; femora lacking dorsal grooves and glandular openings.......................................... 3</p>
            <p>3. Distance between posterior margin of eye and occipital constriction (i.e., length of temple) comparable to length of eye or larger, eyes not adjacent to anterior margin of pronotum in intact specimens....................................... 4</p>
            <p>- Distance between posterior margin of eye and occipital constriction much shorter than length of eye, eyes adjacent or nearly adjacent to anterior margin of pronotum................................................................... 7</p>
            <p>4. Each elytron with two basal foveae distinct in dry-mounted specimens.......................................... 5</p>
            <p>- Each elytron with one basal fovea or rudiments of foveae not visible in dry-mounted specimens...................... 6</p>
            <p> 5. Mesoventral intercoxal process shorter than mesocoxae and less elevated than mesocoxae (in lateral view its ventral margin concealed by mesocoxae); metaventral intercoxal process with two long spines projecting posteriorly; pronotal base with distinct median pit in addition to lateral pits; head with small pit at posteromesal margin of each supraantennal tubercle; basal elytral foveae connected by U-shaped groove extending anteriorly............................  Horaeomorphus Schaufuss</p>
            <p> - Mesoventral intercoxal process keel-like, longer than mesocoxae and more elevated than mesocoxae (its ventral margin visible in lateral view, not concealed by mesocoxae); metaventral intercoxal process lacking spines, with straight or concave posterior margin; pronotal base lacking median pit; head lacking pits near supraantennal tubercles; basal elytral foveae not connected......................................................................................  Euconnus Thomson</p>
            <p> 6. Elytra with basal foveae not visible in dry-mounted specimens; pronotum lacking antebasal pits....  Schuelkelia Jałoszyński </p>
            <p> - Each elytron with one distinct fovea; pronotum with paired antebasal pits...........  Microscydmus Saulcy &amp; Croissandeau</p>
            <p>7. Head with raised dorsal median triangular area pointing posteriorly and frons with median longitudinal groove........................................................................................... Nanoscydmus Jałoszyński</p>
            <p>- Head lacking raised median triangular area and lacking longitudinal groove...................................... 8</p>
            <p> 8. Each elytron with one large basal fovea filled with dense setae.................................  Neuraphes Thomson</p>
            <p>- Each elytron with asetose basal fovea..................................................................... 9</p>
            <p> 9. Pronotum with lateral edges and transverse antebasal groove.................................  Scydmoraphes Reitter</p>
            <p> - Pronotum lacking lateral edges and lacking transverse antebasal groove..........................  Stenichnus Thomson</p>
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	https://treatment.plazi.org/id/45458794FFAFFFEFA5DBFAE6D8B7FE6E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2015): Schuelkelia gen. n., a new eastern Palaearctic ant-like stone beetle, with synopsis of Eurasian genera of Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4007 (3): 343-369, DOI: 10.11646/zootaxa.4007.3.3
45458794FFA1FFEBA5DBFF3FD857FD34.text	45458794FFA1FFEBA5DBFF3FD857FD34.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Schuelkelia unicornis	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Schuelkelia unicornis sp. n.</p>
            <p>(Figs 1–16)</p>
            <p> Type material. Holotype: CHINA (Yunnan Province): ♂, two labels: “ CHINA: Yunnan SE Pingbian, / 22º54'31''N, 103º41'44''E, 2100m, / primary subtropical broad-leaved / forest, litter sifted, 28.VIII.2014, / leg. M. Schülke [CH 14-22a]” [white, printed]; “  SCHUELKELIA /  unicornis m. / det. P. Jałoszyński, 2015 / HOLOTYPUS ” [red, printed] (cMS). Paratypes (8 exx.): 3 ♂ (one disarticulated), 5 ♀, same data as holotype, except for yellow “ paratypus ” label (cMS, cPJ). </p>
            <p> Diagnosis. This is the only known species of  Schuelkelia and can be identified on the basis of generic diagnostic characters and structures of the aedeagus (Figs 15–16). </p>
            <p>Description. Body of male (Figs 1–2) slender and moderately convex, brown, covered with setae slightly lighter than cuticle; BL 1.03–1.05 mm (mean 1.04 mm).</p>
            <p>Head (Figs 1–2, 3, 7–9) broadest at large, moderately convex and moderately coarsely faceted eyes, HL 0.18 mm, HW 0.18–0.19 mm (mean 0.19 mm); vertex and frons confluent, vertex laterally weakly convex, with small posteromedian horn-like projection (Figs 7–8), sides of frons between eyes slightly impressed; supraantennal tubercles feebly raised. Punctures on frons and vertex inconspicuous, fine; setae short, sparse and suberect. Antennae slender but short, AnL 0.38–0.40 mm (mean 0.35 mm), antennomeres I and II strongly elongate, III–V each about as long as broad, VI slightly transverse, VII as long as broad, VIII–X each distinctly transverse, XI about as long as broad, rounded at apex.</p>
            <p>Pronotum (Figs 1–2, 12) oval, broadest slightly in front of middle; PL 0.26–0.28 mm (mean 0.27 mm), PW 0.25–0.28 mm (mean 0.26 mm). Anterior margin nearly straight; anterior corners strongly obtuse-angled and blunt; sides strongly rounded; posterior corners indistinct, strongly obtuse-angled and broadly rounded; posterior margin arcuate with indistinct posteromedian expansion. Punctures on pronotal disc fine and inconspicuous; setae on dorsum sparse, moderately long and suberect, sides of pronotum with dense and long bristles with admixture of long, curved setae (Fig. 13).</p>
            <p>Elytra (Figs 1–2) oval, broadest at middle or slightly in front of middle; EL 0.58–0.60 mm (mean 0.59 mm), EW 0.38 mm, EI 1.53–1.60; humeral calli indistinct; basal impressions absent, apices rounded together. Punctures on elytra fine and inconspicuous; setae sparse, moderately long, suberect. Hind wings absent.</p>
            <p>Legs moderately long and slender, profemora (Fig. 1) distinctly swollen, much broader than meso- and metafemora.</p>
            <p>Aedeagus (Figs 15–16) asymmetrical and stout; AeL 0.28 mm; median lobe in ventral view broadest near base, with basal orifice located in sub-median region of dorsal wall; endophallus with several darkly sclerotized and robust asymmetrical sclerites, of which two long and slender and one broad and short protrude from apical ostium; parameres short, each with single long apical seta.</p>
            <p>Female. Similar to male but with unmodified vertex, slightly less swollen profemora, slightly smaller eyes, on average smaller body and shorter antennae; BL 0.91–0.98 mm (mean 0.95 mm); HL 0.16–0.18 mm (mean 0.17 mm), HW 0.16–0.18 mm (mean (0.17 mm), AnL 0.33–0.35 mm (mean 0.34 mm); PL 0.25 mm, PW 0.24–0.25 mm (mean 0.24 mm); EL 0.50–0.55 mm (mean 0.53 mm), EW 0.35–0.38 mm (mean 0.36 mm), EI 1.43–1.50.</p>
            <p>Distribution and habitat. Southern part of central China; all specimens were collected by sifting leaf litter in a subtropical forest (Fig. 17).</p>
            <p> Etymology. The name  unicornis refers to the tiny posteromedian horn-like projection of the vertex in male. </p>
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	https://treatment.plazi.org/id/45458794FFA1FFEBA5DBFF3FD857FD34	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jałoszyński, Paweł	Jałoszyński, Paweł (2015): Schuelkelia gen. n., a new eastern Palaearctic ant-like stone beetle, with synopsis of Eurasian genera of Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 4007 (3): 343-369, DOI: 10.11646/zootaxa.4007.3.3
