identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
410D8786A3634854FF6FAB1FF48EFB2A.text	410D8786A3634854FF6FAB1FF48EFB2A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoribates chistyakovi	<div><p>Protoribates chistyakovi sp. nov.</p><p>(Figs 1–14)</p><p>Diagnosis. Body size: 630–680 × 431–498. Rostral, lamellar and interlamellar setae long, setiform, barbed, in specifically curved mediodistally. Bothridial setae long, setiform, ciliate. Notogastral setae minute. Distance between porose areas A1–A1 shorter than A2–A2. Subcapitular setae h longest, m shortest and thinnest. Epimeral setal formula: 3–1–3–3; setae 3с longest. Circumpedal carinae short. Adanal setae ad 1 and ad 2 of medium size, thickened. Legs tridactylous. Femora II undulate ventroanteriorly. Trochanters IV with two or three teeth dorsoanteriorly. Setae l’ on femora I, II and d on femora III and some ventral setae on tarsi thick.</p><p>Description. Measurements. Body length: 680 (holotype, female), 630, 680 (two paratypes: two females); notogaster width: 498 (holotype), 431, 498 (two paratypes).</p><p>Integument. Body color brown. Body surface (including genital and anal plates, subcapitular mentum and genae and leg segments) densely microporose (visible under high magnification, ×1000). Macrofoveolae absent. Lateral parts of prodorsum between sublamellae and acetabula I, II microgranulate. Femora II distinctly striate antiaxially.</p><p>Prodorsum (Figs 1, 3). Rostrum broadly rounded. Lamellae (lam) located dorso-laterally, half as long as prodorsum (measured in lateral view). Prolamellae absent. Sublamellae (slam) about half as long as lamellae. Sublamellar porose areas (Al) oval (16–20 × 10–12), located very near to sublamellae. Rostral (ro, 61–69), lamellar (le, 102–114) and interlamellar (in, 123–131) setae setiform, barbed, le inserted on the lamellar ends, in specifically curved mediodistally. Exobothridial setae (ex, 16) thin, slightly barbed. Bothridial setae (bs, 155–159) setiform, ciliate. Sejugal porose areas band-like, diffuse. Tutoria (tu) ridge-like, straight. Lateral ridges (r) present, thin.</p><p>Notogaster (Figs 1, 3–5). Anterior notogastral margin convex medially. Dorsophragmata (D) comparatively short, longitudinally elongated, directed posteromedially. Pteromorphs with distinct hinges. Ten pairs of notogastral setae minute (8), thin, smooth. Four pairs of porose areas rounded, Aa (16) larger than A1, A2, A3 (10– 12). Distance A1–A1 shorter than A2–A2. Setae lp inserted posteriorly or posteromedially to A1, setae h 3 inserted far anteriorly to A2. Lyrifissures (ia, im, ip, ih, ips) and opisthonotal gland openings (gla) clearly visible, ip located posteriorly to A3.</p><p>Gnathosoma (Figs 6–8). Subcapitulum longer than wide (151–155 × 106–114). Subcapitular setae setiform, h longest (45–49), barbed, a (24–28) barbed mediodistally, m shortest (12–16), slightly barbed, thinnest. Two pairs of adoral setae (or 1, or 2, 20) setiform, heavily barbed. Palps (length 94–98) with setation 0–2–1–3–9(+ω). Postpalpal setae (ep, 6) spiniform. Chelicerae (length 164–172) with two barbed setae, cha (53–57) longer than chb (28–32). Trägårdh’s organ (Tg) long, tapered.</p><p>Epimeral and lateral podosomal regions (Figs 2, 3). Epimeral setal formula: 3–1–3–3. Setae setiform, thin, slightly barbed, 3с (45–49) longer than 1b (28–32) and others (14–16). Pedotecta I (Pd I) and II (Pd II) lamina-like, Pd II represented by two connected parts, trapezoid in ventral view, rounded anteriorly. Discidia (dis) triangular, broadly rounded. Custodia absent. Circumpedal carinae (cp) short, directed to acetabula IV.</p><p>Anogenital region (Figs 2–5, 9). Five pairs of genital (g 1, 20; g 2, 16; g 3– g 5, 12–14), one pair of aggenital (ag, 14–16), two pairs of anal (an 1, an 2, 32–36) setae and adanal setae ad 3 (28–32) setiform, slightly barbed. Adanal setae ad 1 and ad 2 of medium size, similar in length (61–69), thickened, erect, barbed. Setae ad 3 inserted anterior to anterior margin of anal aperture. Adanal lyrifissures (iad) located close and parallel to anal plates. Marginoventral porose area (Amar) semioval, interrupted posteromedially. Ovipositor elongated (245 × 36), blades (102) shorter than length of distal section (beyond middle fold; 143). Each of three blades with four straight, smooth setae; ψ1 = τ1 (36–41) longer than ψ2 = τ a = τ b = τ c (16). Six coronal setae minute (4).</p><p>Legs (Figs 10–14). Tridactylous, median claw thicker than laterals, all barbed dorsally. Tibiae I and II with one tubercle ventrobasally. Trochanters IV with two (rarely three) teeth dorsoanteriorly. Femora II undulate ventroanteriorly. Dorsoparaxial porose areas (p.a.) on femora and trochanters III, IV, posteroventral porose areas on tarsi and anteroventral porose areas on tibiae well visible. Formulas of leg setation and solenidia: I (1–5–3–4– 20) [1–2–2], II (1–5–3–4–15) [1–1–2], III (2–3–1–3–15) [1–1–0], IV (1–2–2–3–12) [0–1–0]; homology of setae and solenidia indicated in Table 1. Solenidia ω1 on tarsi I, ω1 and ω2 on tarsi II and σ on genua III bacilliform, other solenidia setiform. Famuli (ɛ) short, thin, straight, indistinctly dilated and truncated distally, inserted posteriorly to ω2. Setae l’ on femora I, II and d on femora III and some ventral setae on tarsi thick.</p><p>Roman letters refer to normal setae, Greek letters to solenidia (except ɛ = famulus). Single prime (’) marks setae on the anterior and double prime (”) setae on the posterior side of a given leg segment. Parentheses refer to a pair of setae. Trtrochanter, Fe—femur, Ge—genu, Ti—Tibia, Ta—tarsus.</p><p>*—s on tarsi IV absent in P. shvanderovi sp. nov.</p><p>Material examined. Holotype (female): Northern Vietnam, Tam Dao National Park, 950 m a.s.l., meadow, sample (No. VIE-125) of grass rhizosphere, 17.X.1988 (collected by J. Starý &amp; Nguen Tri Tien) . One paratype (one female): Northern Vietnam, Tam Dao National Park, 930 m a.s.l., sample (No. VIE-129) of decaying grass on the soil surface from batata field, 17.X.1988 (collected by J. Starý &amp; Nguen Tri Tien) . One paratype (one female): Northern Vietnam, Tam Dao National Park, 1150 m a.s.l., secondary submontane foggy forest, leaf litter sample (No. VIE-193), 20.X.1988 (collected by J. Starý &amp; Nguen Tri Tien) .</p><p>Type deposition. The holotype is deposited in the collection of the Senckenberg Institute, Görlitz, Germany ; two paratypes are deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia .</p><p>Etymology. The species epithet honors Dr. Mikhail P. Chistyakov (1937–2006)—acarologist from Nizhniy Novgorod, Russia—on the 80th anniversary of his birth.</p><p>Remarks. Protoribates chistyakovi sp. nov. is morphologically most similar to P. sangumburiensis (Choi, 1996) from Korea in having setiform bothridial setae, long prodorsal setae and adanal setae ad 1 and ad 2 and tridactylous legs. However, the new species differs from the latter by the larger body size (630–680 × 431–498 vs. 532–592 × 280–320), localization of lamellar setae on lamellar ends (vs. on prodorsum, medially to lamellar ends), broadly rounded rostrum (vs. clearly protruding), thin and rather short epimeral setae 1b (vs. thick and comparatively long) and curved interlamellar setae in medio-distal parts (vs. straight, not curved).</p></div>	https://treatment.plazi.org/id/410D8786A3634854FF6FAB1FF48EFB2A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ermilov, Sergey G.;Starý, Josef	Ermilov, Sergey G., Starý, Josef (2017): New Protoribates (Acari, Oribatida, Haplozetidae) from Vietnam. Zootaxa 4258 (6): 501-524, DOI: 10.11646/zootaxa.4258.6.1
410D8786A36A4855FF6FADC4F44FF8AD.text	410D8786A36A4855FF6FADC4F44FF8AD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoribates shaldybinae	<div><p>Protoribates shaldybinae sp. nov.</p><p>(Figs 15–24)</p><p>Diagnosis. Body size: 647–680 × 415–431. Body surface distinctly foveolate. Rostral, lamellar and interlamellar setae long, setiform, barbed. Bothridial setae long, with heads slightly dilated and barbed unilaterally. Notogastral setae minute. Distance between porose areas A1–A1 similar to A2–A2. Subcapitular setae h longest, m shortest and thinnest. Epimeral setal formula: 3–1–3–2; setae 3с longest, 4c absent. Pedotecta II dentate anteriorly. Custodia developed. Circumpedal carinae long. Adanal setae ad 1 and ad 2 of medium size, thickened. Legs I monodactylous, legs II–IV tridactylous, lateral claws with one ventrodistal tooth. Tarsi I and II with two teeth dorsally. Femora II pointed ventroanteriorly.</p><p>Description. Measurements. Body length: 680 (holotype, female), 647–680 (three paratypes: three females); notogaster width: 415 (holotype), 415–431 (three paratypes).</p><p>Integument (Figs 15, 16, 18, 19). Body color brown. Body surface (including genital and anal plates, subcapitular mentum and genae and leg segments) densely microporose (visible under high magnification, ×1000) and sparsely macrofoveolate (diameter of foveolae up to 2). Lateral parts of prodorsum between sublamellae and acetabula I, II microgranulate.</p><p>Prodorsum (Figs 15, 17). Rostrum broadly rounded. Lamellae located dorso-laterally, half as long as prodorsum (measured in lateral view). Prolamellae absent. Sublamellae about one third of lamellae. Sublamellar porose areas oval (18–24 × 10–12), located near to sublamellae. Rostral (77–82), lamellar (123–127) and interlamellar (155–172) setae setiform, barbed, le inserted on the lamellar ends. Exobothridial setae (36) thin, slightly barbed. Bothridial setae (131–143) with long, barbed unilaterally stalks and heads short, slightly dilated and barbed unilaterally, pointed distally. Sejugal porose areas band-like, diffuse. Tutoria ridge-like, slightly curved mediobasally. Lateral ridges present, thin.</p><p>Notogaster (Figs 15, 17–19). Anterior notogastral margin convex medially. Dorsophragmata comparatively short, longitudinally elongated, directed posteromedially. Pteromorphs with distinct hinges. Ten pairs of notogastral setae minute (8), thin, smooth. Four pairs of porose areas rounded, Aa largest (10–12), A1, A2 and A3 small (4–6). Distance A1–A1 similar to A2–A2. Setae lp inserted anteromedially to A1, setae h 3 inserted far anteriorly to A2. Lyrifissures and opisthonotal gland openings clearly visible, ip located posterolaterally to A3.</p><p>Gnathosoma. Generally, similar to P. chistyakovi sp. nov. Subcapitulum longer than wide (151–155 × 102– 110). Subcapitular setae setiform, h longest (49–53), barbed, a (20–24) slightly barbed, m shortest (12–16), slightly barbed, thinnest. Two pairs of adoral setae (20) setiform, heavily barbed. Palps (length 94–98) with setation 0–2– 1–3–9(+ω). Postpalpal setae (6) spiniform. Chelicerae (length 168–172) with two barbed setae, cha (61–65) longer than chb (32–36). Trägårdh’s organ long, tapered.</p><p>Epimeral and lateral podosomal regions (Figs 16, 17). Epimeral setal formula: 3–1–3–2. Setae 4c and their alveoli absent. Setae setiform, thin, slightly barbed, 3с (41–45) longer than 1b (22–24) and others (12–16). Pedotecta I and II lamina-like, Pd II represented by two connected parts, trapezoid in ventral view, distinctly dentate anteriorly. Custodia present, elongate triangular. Circumpedal carinae long, directed to Pd II.</p><p>Anogenital region (Figs 16–19). Five pairs of genital (g 1, 20; g 2– g 5, 16), one pair of aggenital (14–16) and two pairs of anal (32–36) setae and adanal setae ad 3 (24) setiform, slightly barbed. Adanal setae ad 1 (98–110) and ad 2 (82–90) of medium size, thickened, erect, barbed. Setae ad 3 inserted anterior to anterior margin of anal aperture. Adanal lyrifissures located close and parallel to anal plates. Marginoventral porose area semioval, interrupted posteromedially. Ovipositor, generally, similar to P. chistyakovi sp. nov., elongated (221 × 41), blades (90) shorter than length of distal section (beyond middle fold; 131). Each of three blades with four straight, smooth setae; ψ1 = τ1 (36–41) longer than ψ2 = τ a = τ b = τ c (16). Six coronal setae minute (4).</p><p>Legs (Figs 20–24). Legs I monodactylous, legs II–IV tridactylous, median claw thicker than laterals, all barbed dorsally. Lateral claws with one ventrodistal tooth. Tarsi I and II with two strong teeth dorsally. Tibiae I and II with one tubercle ventrobasally. Femora II pointed ventroanteriorly. Dorsoparaxial porose areas on femora and trochanters III, IV, posteroventral porose areas on tarsi and anteroventral porose areas on tibiae well visible. Formulas of leg setation and solenidia: I (1–5–3–4–20) [1–2–2], II (1–5–3–4–15) [1–1–2], III (2–3–1–3–15) [1–1– 0], IV (1–2–2–3–12) [0–1–0]; homology of setae and solenidia indicated in Table 1. Solenidia ω1 on tarsi I, ω1 and ω2 on tarsi II and σ on genua III bacilliform, other solenidia setiform. Famuli short, thin, straight, indistinctly dilated and truncated distally, inserted posteriorly to ω2.</p><p>Material examined. Holotype (female) and one paratype (female): Northern Vietnam, Tam Dao National Park, 950 m a.s.l., meadow, sample (No. VIE-125) of grass rhizosphere, 17.X.1988 (collected by J. Starý &amp; Nguen Tri Tien) . Two paratypes (two females): Northern Vietnam, Tam Dao National Park, 1020 m a.s.l., secondary mixed forest, sample (No. VIE-179) of leaf litter, 19.X.1988 (collected by J. Starý &amp; Nguen Tri Tien) .</p><p>Type deposition. The holotype is deposited in the collection of the Senckenberg Institute, Görlitz, Germany ; three paratypes are deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia .</p><p>Etymology. The species is named after the late Dr. Elena S. Shaldybina, the acarologist from Nizhniy Novgorod, Russia, for her significant contribution to our knowledge of oribatid mites.</p><p>Remarks. Protoribates shaldybinae sp. nov. is morphologically most similar to P. dentatus (Berlese, 1883) (see Weigmann et al. 1993) from the Holarctic and Oriental regions and Fiji in having bothridial setae with heads slightly dilated unilaterally and pointed distally, long prodorsal setae, custodia and adanal setae ad 1, minute notogastral setae and monodactylous legs I and tridactylous legs II–IV. However, the new species differs from the latter by the macrofoveolate body surface (vs. not macrofoveolate), dentate pedotecta II (vs. not dentate), long adanal setae ad 2 (vs. minute) and dorsal teeth on tarsi I and II (vs. without teeth).</p></div>	https://treatment.plazi.org/id/410D8786A36A4855FF6FADC4F44FF8AD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ermilov, Sergey G.;Starý, Josef	Ermilov, Sergey G., Starý, Josef (2017): New Protoribates (Acari, Oribatida, Haplozetidae) from Vietnam. Zootaxa 4258 (6): 501-524, DOI: 10.11646/zootaxa.4258.6.1
410D8786A36D4849FF6FA9A9F576FE22.text	410D8786A36D4849FF6FA9A9F576FE22.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoribates shvanderovi	<div><p>Protoribates shvanderovi sp. nov.</p><p>(Figs 25–34)</p><p>Diagnosis. Body size: 448–464 × 249–265. Rostral, lamellar and interlamellar setae long, setiform, barbed. Bothridial setae long, with heads strongly dilated and barbed unilaterally. Notogastral setae minute. Distance between porose areas A1–A1 shorter than A2–A2. Subcapitular setae h longest, m shortest and thinnest. Epimeral setal formula: 3–1–3–3; setae 3с longest. Circumpedal carinae long. Adanal setae ad 1 and ad 2 of medium size, thickened. Legs monodactylous. Femora II rounded ventroanteriorly.</p><p>Description. Measurements. Body length: 464 (holotype, male), 448 (one paratype: one male); notogaster width: 249 (holotype), 265 (one paratype).</p><p>Integument. Body color light brownish. Body surface (including genital and anal plates, subcapitular mentum and genae and leg segments) densely microporose (visible under high magnification, ×1000) and sparsely macrofoveolate, but macrofoveolae (their diameter up to 2) very poorly visible. Lateral parts of prodorsum between sublamellae and acetabula I, II microgranulate.</p><p>Prodorsum (Figs 25, 27). Rostrum broadly rounded. Lamellae located dorso-laterally, half as long as prodorsum (measured in lateral view). Prolamellae absent. Sublamellae about two third of lamellae. Sublamellar porose areas oval (12 × 6–8), located very near to sublamellae. Rostral (49–57), lamellar (69–73) and interlamellar (94–98) setae setiform, barbed, le inserted on the lamellar ends. Exobothridial setae (12) thin, slightly barbed. Bothridial setae (69–77) with long, barbed unilaterally stalks and heads short, strongly dilated and barbed unilaterally, pointed distally. Sejugal porose areas band-like, diffuse. Tutoria ridge-like, straight. Lateral ridges present, thin.</p><p>Notogaster (Figs 25, 27–29). Anterior notogastral margin convex medially. Dorsophragmata comparatively short, longitudinally elongated, directed posteromedially. Pteromorphs with distinct hinges. Ten pairs of notogastral setae minute (4), thin, smooth. Four pairs of porose areas rounded, Aa (8–10) larger than others (A1, A2, A3, 6). Distance A1–A1 shorter than A2–A2. Setae lp inserted posteriorly to A1, setae h 3 inserted far anteriorly to A2. Lyrifissures and opisthonotal gland openings clearly visible, ip located posterolaterally to A3.</p><p>Gnathosoma (Fig. 27). Morphology similar to P. chistyakovi sp. nov. Subcapitulum longer than wide (110 × 77). Subcapitular setae setiform, slightly barbed, h longest (28–32) longer than a (18–20), m shortest (12) and thinnest. Two pairs of adoral setae (12) setiform, heavily barbed. Palps (length 65) with setation 0–2–1–3–9(+ω). Postpalpal setae (4) spiniform. Chelicerae (length 123) with two barbed setae, cha (57) longer than chb (28). Trägårdh’s organ long, tapered.</p><p>Epimeral and lateral podosomal regions (Figs 26, 27). Epimeral setal formula: 3–1–3–3. Setae setiform, thin, slightly barbed, 3с (22–24) longer than 1b (18–20) and others (10–12). Pedotecta I and II lamina-like, Pd II represented by two connected parts, trapezoid in ventral view, rounded anteriorly. Custodia absent. Circumpedal carinae long, directed to Pd II.</p><p>Anogenital region (Figs 26–29). Five pairs of genital (g 1, 18–20; g 2– g 5, 10–12), one pair of aggenital (10–12) and two pairs of anal (20–24) setae and adanal setae ad 3 (16) setiform, slightly barbed. Adanal setae ad 1 and ad 2 of medium size, similar in length (49–57), thickened, erect, barbed. Setae ad 3 inserted laterally to anterior margin of anal aperture. Adanal lyrifissures located close and parallel to anal plates. Marginoventral porose area semioval, interrupted posteromedially.</p><p>Legs (Figs 30–34). Legs monodactylous, claws thick, barbed dorsally. Tibiae I and II without clear tubercles ventrobasally. Femora II rounded ventroanteriorly. Dorsoparaxial porose areas on femora and trochanters III, IV, posteroventral porose areas on tarsi and anteroventral porose areas on tibiae poorly visible. Formulas of leg setation and solenidia: I (1–5–3–4–20) [1–2–2], II (1–5–3–4–15) [1–1–2], III (2–3–1–3–15) [1–1–0], IV (1–2–2–3–11) [0– 1–0]; homology of setae and solenidia indicated in Table 1. Solenidia ω1 on tarsi I, ω1 and ω2 on tarsi II and σ on genua III bacilliform, other solenidia setiform. Famuli short, thin, straight, indistinctly dilated and truncated distally, inserted posteriorly to ω2. Setae s absent on tarsi IV.</p><p>Material examined. Holotype (male) and one paratype (male): Northern Vietnam, Tam Dao National Park, 950 m a.s.l., meadow, sample (No. VIE-125) of grass rhizosphere, 17.X.1988 (collected by J. Starý &amp; Nguen Tri Tien) .</p><p>Type deposition. The holotype is deposited in the collection of the Senckenberg Institute, Görlitz, Germany ; one paratype is deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia .</p><p>Etymology. The species is named after the late Dr. Fridrikh A. Shvanderov, the acarologist from Nizhniy Novgorod, Russia, for his significant contribution to our knowledge of eriophyoid mites.</p><p>Remarks. Protoribates shvanderovi sp. nov. is morphologically most similar to P. gracilis (Aoki, 1982) from Japan and Vietnam in having long bothridial setae with heads strongly dilated unilaterally and pointed distally, long adanal setae ad 1 and ad 2 and monodactylous legs. However, the new species differs from the latter by the larger body size (448–464 × 249–265 vs. 340 × 180), localization of lamellar setae on the lamellar ends (vs. on prodorsum, medially to lamellar ends), relative lengths of lamellar, interlamellar and bothridial setae, viz., le longer than ro, in longer than bs (vs. le shorter than ro, in shorter than bs), and short dorsophragmata (vs. long).</p></div>	https://treatment.plazi.org/id/410D8786A36D4849FF6FA9A9F576FE22	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ermilov, Sergey G.;Starý, Josef	Ermilov, Sergey G., Starý, Josef (2017): New Protoribates (Acari, Oribatida, Haplozetidae) from Vietnam. Zootaxa 4258 (6): 501-524, DOI: 10.11646/zootaxa.4258.6.1
410D8786A377484AFF6FAB3DF5CFFEB2.text	410D8786A377484AFF6FAB3DF5CFFEB2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protoribates	<div><p>Key to the Vietnamese species of the genus Protoribates</p><p>1. Bothridial setae setiform............................................................................... 2</p><p>- Bothridial setae with developed (dilated unilaterally) heads................................................... 6</p><p>2. Legs bidactylous; rostral and lamellar setae and epimeral setae 3b and 4a bacilliform; body size: 381–431 × 182–249......................... P. heterodactylus Ermilov &amp; Anichkin, 2011 (see Ermilov &amp; Anichkin 2011 a). Distribution: Vietnam.</p><p>- Legs mono- or tridactylous; rostral and lamellar setae and epimeral setae 3b and 4a setiform........................ 3</p><p>3. Legs monodactylous................................................................................... 4</p><p>- Legs tridactylous......................................................................................5</p><p>4. Lamellar setae minute, inserted medially to the lamellar ends; adanal setae ad 1 and ad 2 short, similar to anal setae in length; distance between notogastral porose areas A1–A1 longer than A2–A2; body size: 315–348 × 140–166......................................... P. cattienensis Ermilov &amp; Anichkin, 2011 (see Ermilov &amp; Anichkin 2011 b). Distribution: Vietnam.</p><p>- Lamellar setae long, inserted on the lamellar ends; adanal setae ad 1 and ad 2 distinctly longer than anal setae; distance between notogastral porose areas A1–A1 shorter than A2–A2; body size: 525–565 × 300–330................................................................ P. lophotrichus (Berlese, 1904) (see Miko et al. 1994). Distribution: semicosmopolitan.</p><p>5. Adanal setae ad 1 and ad 2 of medium size, similar in length; interlamellar setae specifically curved mediodistally; dorsophragmata longitudinally elongated; body size: 630–680 × 431–498.............. P. chistyakovi sp. nov. Distribution: Vietnam.</p><p>- Adanal setae ad 1 of medium size, ad 2 short; interlamellar setae straight; dorsophragmata semi-oval; body length: 490–680........................................................ P. duosetus (Hammer, 1979) . Distribution: Oriental region.</p><p>6. Legs I monodactylous, legs II–IV tridactylou1s............................................................. 7</p><p>- All legs monodactylous or tridactylous.....................................................................8</p><p>7. Adanal setae ad 1 and ad 2 of medium size, similar in length; body surface distinctly foveolate; pedotecta II dentate anteriorly; body size: 647–680 × 415–431...................................... P. shaldybinae sp. nov. Distribution: Vietnam.</p><p>- Adanal setae ad 1 of medium size, ad 2 short; body surface not distinctly foveolate; pedotecta II rounded anteriorly; body length: 470–555............................... P. dentatus (Berlese, 1883) (see Weigmann et al. 1993). Distribution: Vietnam.</p><p>8. Legs monodactylous.................................................................................. 9</p><p>- Legs tridactylous................................................................................... 11</p><p>9. Heads of bothridial narrowly dilated; adanal setae ad 1 and ad 2 short, slightly differs from anal setae in length; body size: 320– 440 × 238–266.......................................................................................... P. capucinus Berlese, 190 2 8 (see Hammer 1961; Pérez-Íñigo 1974; Weigmann 2006; Bayartogtokh 2010). Distribution: cosmopolitan.</p><p>- Heads of bothridial setae broadly dilated; adanal setae ad 1 and ad 2 of medium size, distinctly longer than anal setae..... 10</p><p>10. Lamellar setae inserted on the lamellar ends; lamellar setae longer than rostral setae, interlamellar setae longer than bothridial setae; dorsophragmata comparatively short; body size: 448–464 × 249–265.. P. shvanderovi sp. nov. Distribution: Vietnam.</p><p>- Lamellar setae inserted on the prodorsum, medially to the lamellar ends; lamellar setae shorter than rostral setae, interlamellar setae shorter than bothridial setae; dorsophragmata comparatively long; body size: 340 × 180................................................................................ P. gracilis (Aoki, 1982) . Distribution: Japan and Vietnam.</p><p>11. Adanal setae ad 1 and ad 2 short, similar in length; interlamellar setae not longer than lamellar setae; body length: 680............................................................... P. acutus (Hammer, 1979) . Distribution: Oriental region.</p><p>- Adanal setae ad 1 of medium size, ad 2 short; interlamellar setae distinctly longer than lamellar setae................... 12</p><p>1. The typical case is given. However, sometimes the number of leg claws in P. dentatus is variable (see Weigmann et al. 1993).</p><p>2. Mahunka (1988 b) described P. paracapucinus from Malaysia. At present, it was recorded from the Palaearctic, Oriental, Neotropical and Ethiopian regions and Vanuatu (Subías 2004, online version 2016; Weigmann &amp; Ermilov 2016). This species differs from P. capucinus by the presence of 10 pairs of notogastral setae (vs. 11) and 12 barbs on bothridial setae (vs. more than 20), and the localization of notogastral porose areas A3 (between h 1 and p 1 vs. p 1 and p 2) (see Mahunka 1988 b). However, the literature analysis (e.g. Hammer 1961; Pérez-Íñigo 1974; Weigmann 2006; Bayartogtokh 2010) shows that some populations of P. capucinus also have ten pairs of notogastral setae, numerous barbs on bothridial setae and localization of porose areas A3 between h 1 and p 1. Thus, P. capucinus and P. paracapucinus are morphologically very similar and the clear differences between them are not visible, therefore, potentially, the latter is a junior synonym of P. capucinus, but the additional research is necessary.</p><p>12. Body surface distinctly foveolate; distance between notogastral porose areas A1–A1 shorter than A2–A2; femora II rounded ventroanteriorly; body size: 448–798 × 266–532..................................................................... P. bisculpturatus (Mahunka, 1988) (see Mahunka 1988 b and also Corpuz-Raros 2013). Distribution: Oriental region.</p><p>- Body surface not distinctly foveolate; distance between notogastral porose areas A1–A1 and A2–A2 similar; femora II pointed ventroanteriorly; body size: 836–976 × 582–664......................................................................................... P. maximus (Mahunka, 1988) (see Mahunka 1988 a). Distribution: Oriental region and Iran.</p></div>	https://treatment.plazi.org/id/410D8786A377484AFF6FAB3DF5CFFEB2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ermilov, Sergey G.;Starý, Josef	Ermilov, Sergey G., Starý, Josef (2017): New Protoribates (Acari, Oribatida, Haplozetidae) from Vietnam. Zootaxa 4258 (6): 501-524, DOI: 10.11646/zootaxa.4258.6.1
