taxonID	type	description	language	source
410487DCB7191D7AFF23520AFB4F54BD.taxon	diagnosis	Diagnosis. Carapace with flat dorsal surface, regions distinct, median groove shallow. Merus of third maxilliped with external margin broadly rounded, shallow notch on anterior margin rounded; ischium trapezoidal, becoming broader distally, ratio exopod / ischium 0.5. Male first gonopod latero-mesially compressed, slightly arched in caudal and cephalic views, of same thickness along its length; distal processes well developed. In mesial view, marginal suture straight along most of gonopod, slightly concave in the middle. Marginal plate between caudal and mesial surfaces abruptly widening distally, forming triangular or rectangular apical projection overreaching cephalic spine, lying on caudal margin of mesial surface, ending in acute distal margin. Mesial surface ending distally in tongue-like apical projection, disto-cephalically directed, longer than apical projection of marginal plate, distal margin broad, rounded, sculptured with shallow longitudinal grooves and scattered subacute granules or thin and divided with acute and robust spines. Mesial process as triangular spine, stout and acute, cephalad or laterally directed, forming 90 º angle with respect to principal axis of gonopod. Lateral suture visible through proximal twothirds, separating marginal plate from caudal surface. In cephalic view, mesial process conical, stout and acute. Lateral surface with 1 or 2 small, triangular, distomedial tubercles; distal third of laterocaudal portion, forming a high border, rounded, with a mid-triangular and low prominence or a distocaudal spine, strong, conical and acute. Apical projection of mesial surface, tongue-like, with rounded, broad, granulated margin or armed with acute spines. In lateral view, caudal margin with wide and shallow or deep and narrow lateral notch, located at midlength. Lateral surface widening caudally in distal third into a semirounded plate, with 1 or 2 small triangular, distomedial tubercles, caudal border high, laterally directed with triangular and low prominence or distocaudal spine. Deep suture in the base of apical projection of mesial surface, marking separation of lateral and mesial plates. In caudal view, gonopod slightly arched laterally. Distal half with lateral margin expanded, armed with triangular prominence or acute and conical spine. In apical view, apical projection of marginal plate ending in acute tip or thin border armed with spines. Apical projection of mesial surface broad and rounded, with acute tubercles or thin and divided with acute, robust spines. Apical cavity covered by hood formed by crest of lateral plate or open surrounded by crests of mesial and lateral surfaces. Caudal border of lateral surface ending distally in triangular prominence or in acute, conical, strong spine. Opening of spermatic channel at the base of apical projection of mesial surface.	en	Villalobos, José Luis, Álvarez, Fernando (2013): Two new genera and three new species of freshwater crabs (Crustacea: Pseudothelphusidae: Potamocarcinini) from Chiapas, Mexico. Zootaxa 3599 (5): 457-470, DOI: 10.11646/zootaxa.3599.5.3
410487DCB7191D7AFF23520AFB4F54BD.taxon	materials_examined	Type species. Sylvathelphusa kalebi n. sp. Other species included. Sylvathelphusa cavernicola n. sp.	en	Villalobos, José Luis, Álvarez, Fernando (2013): Two new genera and three new species of freshwater crabs (Crustacea: Pseudothelphusidae: Potamocarcinini) from Chiapas, Mexico. Zootaxa 3599 (5): 457-470, DOI: 10.11646/zootaxa.3599.5.3
410487DCB7191D7AFF23520AFB4F54BD.taxon	etymology	Etymology. The generic name is derived from the Latin sylva (forest) and Thelphusa, the generic name for freshwater crabs.	en	Villalobos, José Luis, Álvarez, Fernando (2013): Two new genera and three new species of freshwater crabs (Crustacea: Pseudothelphusidae: Potamocarcinini) from Chiapas, Mexico. Zootaxa 3599 (5): 457-470, DOI: 10.11646/zootaxa.3599.5.3
410487DCB7191D7AFF23520AFB4F54BD.taxon	distribution	Distribution. Southern and central portions of Chiapas, Mexico.	en	Villalobos, José Luis, Álvarez, Fernando (2013): Two new genera and three new species of freshwater crabs (Crustacea: Pseudothelphusidae: Potamocarcinini) from Chiapas, Mexico. Zootaxa 3599 (5): 457-470, DOI: 10.11646/zootaxa.3599.5.3
410487DCB7191D7AFF23520AFB4F54BD.taxon	discussion	Remarks. Based on the morphology of the male gonopod, Sylvathelphusa n. gen. belongs in the tribe Potamocarcinini and is closely related to Potamocarcinus. A conical mesial process, with a stout and acute spine, cephalad or laterad directed, forming a 90 º angle with respect to the principal axis of the gonopod, and a marginal plate placed between the caudal and mesial surfaces, and overreaching the cephalic spine, are traits found in the species of both genera. The gonopod morphology of Potamocarcinus and Sylvathelphusa n. gen. differ, however, in the form, ornamentation and position of the apical projections of the marginal and mesial plates, in the cover of the apical cavity by the apical crests, and in the position of the aperture of the spermatic channel.	en	Villalobos, José Luis, Álvarez, Fernando (2013): Two new genera and three new species of freshwater crabs (Crustacea: Pseudothelphusidae: Potamocarcinini) from Chiapas, Mexico. Zootaxa 3599 (5): 457-470, DOI: 10.11646/zootaxa.3599.5.3
410487DCB7191D7EFF235423FB8453C0.taxon	materials_examined	Type material. Holotype: male, cb 17.3 mm, cl 10.5 mm; allotype: female, cb 14.8 mm, cl 9.6 mm; paratype: male, cb 15.1 mm, cl 9.4 mm; La Pera nature reserve, 12 km NE of Berriozábal (16 ° 49 ' 53 " N, 93 ° 17 ' 42 " W; 1178 m), Municipio de Tuxtla Gutiérrez, Chiapas, Mexico; 12 Sep 2008; colls: K. Zárate, I. A. Córdoba, A. J. Flores, C. A. Pérez, G. Salinas; CNCR 26179. Other material examined (non-types). 5 males, cb 12.0 - 15.5 mm, cl 7.8 - 10.5 mm; 2 females 15.0 mm both, cl 9.2 - 9.5 mm; same locality as holotype; 27 Nov 2008; colls: J. L. Villalobos, C. Hernández, K. Zárate, A. García, C. Enríquez, E. Torres; CNCR 26181.	en	Villalobos, José Luis, Álvarez, Fernando (2013): Two new genera and three new species of freshwater crabs (Crustacea: Pseudothelphusidae: Potamocarcinini) from Chiapas, Mexico. Zootaxa 3599 (5): 457-470, DOI: 10.11646/zootaxa.3599.5.3
410487DCB7191D7EFF235423FB8453C0.taxon	description	Description. Small sized crabs (cb <17.3 mm). In dorsal view, carapace oval shaped, widest in the middle; dorsal surface flat, punctated, regions faintly indicated; pair of gastric pits barely distinct, oblique, elongated, close to each other on metagastric region; cervical grooves narrow, shallow, straight, not reaching anterolateral margin of carapace (Fig. 2 a). Postfrontal lobes distinct, low, separated by median groove. Frontal region granulated, depressed with respect to carapace surface; inferior frontal border granulated, projected, bilobed, with deep medial notch; superior frontal border straight, bilobed in dorsal view, lobes defined by row of sharp granules separated by shallow mesial notch (Fig. 2 a). Superior margin of orbit formed by granules, continuous with inferior frontal border; lower orbital margin marked by row of granules; exorbital angle projected, narrow, acute. Lateral margins of carapace serrated, anterior half with sharp granules, posterior half smooth. Posterior margin of carapace slightly concave. Eyes with cornea pigmented, faintly faceted, not filling orbits (Fig. 2 d). Epistome narrow, epistomial tooth low, triangular, slightly deflexed, borders smooth (Fig. 2 c). Suborbital, subhepatic and pterygostomian regions pubescent. Third maxilliped with ratio exopod / ischium 0.47, ischium with outer margin slightly sinuous, inner margin straight, external margin of merus widely curved (Fig. 2 c). Aperture of efferent branchial channel subrectangular. Chelipeds subequal in size and shape; merus subtriangular in cross section, 4.3 times as long as wide, superior internal border with row of conical and acute spines, increasing in size distally, alternating with small spinules. Carpus smooth, prominent and acute subdistal spine on inner side. Chela slender (length / height 3.6), surface smooth, with faint minute teeth on upper and lower rounded borders. Fingers slender, as long as palm, not gaping, tips crossing; both fingers with sharp triangular teeth on cutting edges, becoming smaller distally; dactylus with alternating large and small teeth (Fig. 2 b). Thoracic sternites of third maxillipeds and first pereiopods completely fused, except for small notches at lateral edges of sternum; sternal sulci s 4 / s 5, s 5 / s 6 and s 6 / s 7 distinct, just failing to reach midline of thoracic sternum. All abdominal segments free. Telson subtriangular, narrow; lateral margins distinctly straight, tip rounded. Male gonopod not completely chitinized, latero-mesially compressed, slightly arched in caudal and cephalic views, distal half wider. In mesial view, marginal suture almost straight along most of gonopod, slightly concave in middle (Fig. 3 a). Marginal plate between caudal and mesial surfaces, without marginal setae basally; distal half widening distally, ending in triangular apical projection overreaching mesial process, lying on caudal margin of apical projection, ending in acute distal margin with triangular tubercle (Fig. 3 a). Caudal portion of marginal plate limited by lateral suture, distal third rounded. Mesial surface ending in tongue-like apical projection, distocephalically directed, longer than apical projection of marginal plate, distal margin broad, rounded, sculptured with shallow longitudinal grooves and scattered subacute granules (Fig. 3 a). Mesial process as triangular, stout, acute spine, directed cephalad, forming 90 º angle with respect to principal axis of gonopod (Fig. 3 a). Caudal surface with medial shallow, rounded notch. Lateral suture visible through proximal two thirds, apically separating marginal plate from caudal surface. In cephalic view, mesial process conical, stout, acute (Fig. 3 b). Lateral surface with small triangular distomedial tubercle (Fig. 3 b); distal third of laterocaudal portion forming high, rounded border, with central depression (Fig. 3 b). Distal end of marginal plate subacute. In lateral view, caudal surface with medial constriction (Fig. 3 c). Lateral surface becoming broader caudally, forming on distal third ample plate with high caudal margin, laterally directed (Fig. 3 c). Deep suture at base of apical projection of mesial surface, separating lateral and mesial plates (Fig. 3 c). In caudal view, gonopod slightly arched laterally (Fig. 3 d). Distal half with lateral margin expanded, with low, distal, triangular prominence (Fig. 3 d). In apical view, apical portion of marginal plate ending in acute tip (Fig. 2 e). Mesial surface with broad, rounded distal margin, with few granules (Fig. 2 e). Mesial process conical, stout, acute, directed cephalically (Fig. 2 e). Apical cavity covered by hood formed by distal crest of lateral surface. Opening of spermatic channel located caudally (Fig. 2 e). Color in life. Carapace dark olive or gray, chelipeds white and ambulatory legs light brown, thoracic sternites and abdomen light yellow.	en	Villalobos, José Luis, Álvarez, Fernando (2013): Two new genera and three new species of freshwater crabs (Crustacea: Pseudothelphusidae: Potamocarcinini) from Chiapas, Mexico. Zootaxa 3599 (5): 457-470, DOI: 10.11646/zootaxa.3599.5.3
410487DCB7191D7EFF235423FB8453C0.taxon	etymology	Etymology. The species is named after Kaleb Zárate, biologist and speleologist, who first discovered this population.	en	Villalobos, José Luis, Álvarez, Fernando (2013): Two new genera and three new species of freshwater crabs (Crustacea: Pseudothelphusidae: Potamocarcinini) from Chiapas, Mexico. Zootaxa 3599 (5): 457-470, DOI: 10.11646/zootaxa.3599.5.3
410487DCB7191D7EFF235423FB8453C0.taxon	discussion	Remarks. This is the first record of a completely terrestrial pseudothelphusid crab in Mexico, although Pseudothelphusa parabelliana Álvarez, 1989, Odontothelphusa maxillipes (Rathbun, 1898), Potamocarcinus magnus (Rathbun, 1895) and Raddaus bocourti (A. Milne-Edwards, 1866), have been observed at considerable distances from the nearest stream. The crabs representing the new species were found in a tropical rainforest NE of the town of Berriozábal, at altitudes ranging from 950 to 1200 m. They were found under limestone in crevices, where the soil is humid and contains abundant organic matter composed of leaf litter in different stages of decomposition. No streams, rivers or other aquatic bodies were observed nearby.	en	Villalobos, José Luis, Álvarez, Fernando (2013): Two new genera and three new species of freshwater crabs (Crustacea: Pseudothelphusidae: Potamocarcinini) from Chiapas, Mexico. Zootaxa 3599 (5): 457-470, DOI: 10.11646/zootaxa.3599.5.3
410487DCB71D1D71FF2352B6FC6650C3.taxon	materials_examined	Type material. Holotype: male, cb 32.8 mm, cl 20.3 mm; female allotype, cb 46.6 mm, cl 28.6 mm; Cueva de las Arañas, near the town of San Fernando (16 ° 51 ' 26.0 " N, 93 ° 16 ' 01.5 " W; 1140 m), Municipio de San Fernando, Chiapas, Mexico; 5 Sep 2008; colls: F. Álvarez, E. Torres, K. Zárate, G. Armendáriz, A. García and J. L. Villalobos; CNCR 25324. Paratypes: 3 males, cb 28.6 - 24.1 mm, cl 17.1 - 14.4 mm; 1 female, cb 17.0 mm, cl 11.0 mm; same locality, date and collectors as holotype, CNCR 27210.	en	Villalobos, José Luis, Álvarez, Fernando (2013): Two new genera and three new species of freshwater crabs (Crustacea: Pseudothelphusidae: Potamocarcinini) from Chiapas, Mexico. Zootaxa 3599 (5): 457-470, DOI: 10.11646/zootaxa.3599.5.3
410487DCB71D1D71FF2352B6FC6650C3.taxon	description	Description. Crabs of moderate size (cb <47 mm). In dorsal view, carapace oval, widest in the middle (cb / cl average = 1.62); dorsal surface flat, punctated, regions faintly indicated, pair of gastric pits barely distinct. Cervical grooves shallow, straight, not reaching anterolateral margin (Fig. 4 a). Postfrontal lobes indistinct, separated by shallow median groove. Frontal region granulated, slightly curved towards superior frontal margin; inferior border faintly granulated, projected, divided into two convex lobes; superior border bilobed with shallow mesial notch. Superior orbital margin carinate, continuous with inferior frontal border; inferior orbital margin marked by row of granules; exorbital angle projected, narrow and acute (Fig. 4 d). Anterolateral margins serrated with sharp granules along anterior half, posterior half almost smooth. Posterior margin concave. Eyes with cornea pigmented, faintly faceted, not filling orbits (Fig. 4 d). Epistome narrow, epistomial tooth low, triangular, slightly deflexed, borders smooth. Pterygostomial region covered by dense pubescence around mouthparts. Endopod of third maxilliped with outer margin of ischium sinuous, inner margin straight; external margin of merus widely curved; ratio exopod / ischium 0.52 (Fig. 4 c). Aperture of efferent branchial channel subrectangular. Chelipeds similar in shape, subequal in length, same color as walking legs. Major cheliped with merus subtriangular in cross section; 3.3 times as long as wide; dorsal suface smooth, internal dorsal margin with row of conical, acute spines, increasing in size distally alternating with small spinules; external margin granulated. Carpus smooth, prominent and acute subdistal spine on inner surface, preceded by two acute spinules. Palm high, compressed, surface smooth, with minute squamous granules on dorsal and ventral margins. Fingers slender, shorter than palm, not gaping, tips crossing; both fingers with triangular sharp teeth on cutting edges, smaller distally; dactylus with smaller teeth sometimes interspersed with larger ones (Fig. 4 b). Thoracic sternites of third maxillipeds and first pereiopods completely fused, except for small notches at lateral edges of sternum; sternal sulci s 4 / s 5, s 5 / s 6 and s 6 / s 7 distinct, just failing to reach midline of thoracic sternum. All abdominal segments free. Telson subtriangular, narrower than sixth abdominal segment; lateral margins straigth, tip acute. Male gonopod latero-mesially compressed, slightly arched in caudal and cephalic views, distal half wider. In mesial view, marginal suture almost straight along most of gonopod, slightly concave in the middle (Fig. 5 a). Marginal plate between caudal and mesial surfaces, proximal half narrow, with few long marginal setae at base; distal half wider, forming rectangular projection, distal margin rounded with prominent spine (Fig. 5 a). Caudal portion of marginal plate limited by straight and incipient lateral suture. Mesial surface ending apically in tonguelike apical projection, disto-cephalically directed and longer than apical projection of marginal plate, distal margin with acute and robust spines. Mesial process as a triangular, stout, acute spine, laterad directed, forming an 85 º angle with respect to principal axis of gonopod. Caudal surface with narrow, deep medial notch. Lateral suture visible through proximal two thirds; on distal third interrupted and reappearing separating marginal plate from caudal surface. In cephalic view, mesial process conical, stout, acute (Fig. 5 b). Lateral surface broad, semicircular, with one slender, acute distomedial tubercle. Caudal margin with acute spinules and distal triangular, acute spine. Apical projection of mesial surface tongue-like, with thin serrate margin. In lateral view, distal margin of mesial surface with 7 strong teeth, spine on lateral surface acute, constriction of gonopod at middle portion evident (Fig. 5 c). In caudal view, gonopod straight. Distal half with lateral margin expanded as rounded plate with spinular edge and distal, acute spine (Fig. 5 d). In apical view, apical projection of mesial surface with thin distal margin serrate (Fig. 4 e), caudal surface concave, with few short setae in the apical cavity. Mesial process conical, stout directed laterad. Apex cavity elongated, partially covered by hood formed by distal crest of lateral surface, with few setae. Caudal border of lateral plate ending distally in acute spine (Fig. 4 e).	en	Villalobos, José Luis, Álvarez, Fernando (2013): Two new genera and three new species of freshwater crabs (Crustacea: Pseudothelphusidae: Potamocarcinini) from Chiapas, Mexico. Zootaxa 3599 (5): 457-470, DOI: 10.11646/zootaxa.3599.5.3
410487DCB71D1D71FF2352B6FC6650C3.taxon	discussion	Remarks. Sylvathelphusa cavernicola n. sp. represents the third stygobitic freshwater crab of the family Pseudothelphusidae recorded in Chiapas after Typhlopseudothelphusa mocinoi Rioja, 1952 and T. hyba Rodríguez & Hobbs, 1989. Interestingly, two different lineages represented by these two genera have colonized the karstic environment in a relatively reduced area in central Chiapas. Morphologically the gonopod of S. cavernicola n. sp. is different from those of the species of Typhlopseudothelphusa in that in the latter it is bent mesially, and there are no supra-apical processes, being more similar to Potamocarcinus where the gonopod is straight and stout. The most evident differences with Typhlopseudothelphusa are the stoutness of the gonopod, with the marginal plate and the distal crest of the mesial surface distally projected beyond the mesial process.	en	Villalobos, José Luis, Álvarez, Fernando (2013): Two new genera and three new species of freshwater crabs (Crustacea: Pseudothelphusidae: Potamocarcinini) from Chiapas, Mexico. Zootaxa 3599 (5): 457-470, DOI: 10.11646/zootaxa.3599.5.3
410487DCB7121D72FF2351BAFB8F5293.taxon	diagnosis	Diagnosis. Carapace with dorsal surface convex, regions discernible, median groove narrow. Exopod of third maxilliped subequal in length to ischium. First male gonopod with distal third strongly bent laterally, compressed lateromesially, distal half slightly narrower, mesial margin widely curved. In mesial view, marginal suture straight along proximal two-thirds of gonopod, laterally bent on distal third (Fig. 7 a). Marginal plate between caudal and mesial surfaces. Mesial process as triangular, stout, acute spine, cephalad directed, forming 90 º angle with respect to principal axis of gonopod (Fig. 7 a). Caudal surface straight, without medial notch. Lateral suture visible through almost entire length of gonopod. In cephalic view, distal portion of gonopod laterally directed, proximal two-thirds thicker, distal one compressed, separated from medial third by deep, wide, rounded constriction (Fig. 7 b). Apical projection of mesial surface digitiform, with transverse rows of spines. In lateral view, gonopod of uniform thickness, surface punctated with scarce short setae, apical elements and apex cavity visible (Fig. 7 c). Mesial surface of distal crest, thin, high, bordering inner surface of apical cavity. Distal crest of lateral surface blunt, cephalically reaching proximal half of mesial process, limiting apical cavity. Apical cavity narrow, elongated along a cephalo-caudal axis, limited cephalically by irregular laminar projection of lateral crest and caudally by marginal plate; opening of spermatic channel subcircular, in caudal position (Fig. 7 c). In caudal view, distal portion of gonopod laterally directed, lateral suture visible along proximal two-thirds (Fig. 7 d). Distal notch separating marginal plate from lateral surface obliterated. Distal end of marginal plate with acute spinules.	en	Villalobos, José Luis, Álvarez, Fernando (2013): Two new genera and three new species of freshwater crabs (Crustacea: Pseudothelphusidae: Potamocarcinini) from Chiapas, Mexico. Zootaxa 3599 (5): 457-470, DOI: 10.11646/zootaxa.3599.5.3
410487DCB7121D72FF2351BAFB8F5293.taxon	materials_examined	Type species. Tzotzilthelphusa villarosalensis n. sp.	en	Villalobos, José Luis, Álvarez, Fernando (2013): Two new genera and three new species of freshwater crabs (Crustacea: Pseudothelphusidae: Potamocarcinini) from Chiapas, Mexico. Zootaxa 3599 (5): 457-470, DOI: 10.11646/zootaxa.3599.5.3
410487DCB7121D72FF2351BAFB8F5293.taxon	etymology	Etymology. The generic name is derived from Tzotzil the name of the native people that inhabit this region of Chiapas and Thelphusa, the generic name for freshwater crabs.	en	Villalobos, José Luis, Álvarez, Fernando (2013): Two new genera and three new species of freshwater crabs (Crustacea: Pseudothelphusidae: Potamocarcinini) from Chiapas, Mexico. Zootaxa 3599 (5): 457-470, DOI: 10.11646/zootaxa.3599.5.3
410487DCB7121D72FF2351BAFB8F5293.taxon	distribution	Distribution. Known only from a small area south of the town of Teopisca in central Chiapas, Mexico.	en	Villalobos, José Luis, Álvarez, Fernando (2013): Two new genera and three new species of freshwater crabs (Crustacea: Pseudothelphusidae: Potamocarcinini) from Chiapas, Mexico. Zootaxa 3599 (5): 457-470, DOI: 10.11646/zootaxa.3599.5.3
410487DCB7121D72FF2351BAFB8F5293.taxon	discussion	Remarks. Based on the morphology of the male gonopod which does not show any torsion, although the distal third is inclined, Tzotzilthelphusa n. gen. is placed in the tribe Potamocarcinini. The new genus appears morphologically similar to Phrygiopilus in that the gonopods of both genera show a distal third inclined laterally, a laterally oriented apical cavity and a conical mesial process produced as a robust spine cephalically oriented at a 90 º angle with respect to the longitudinal axis of the gonopod. The most important difference between Tzotzilthelphusa n. gen. and Phrygiopilus is the extent of development of the apical processes of the gonopod. In Phrygiopilus the supra-apical process is a large cap-like process developed on top of the apical cavity, while in Tzotzilthelphusa n. gen. the crest deriving from the mesial surface develops into a lobular projection with spinules. Phrygiopilus is also found in Chiapas, in the Montebello Lakes district with two species (P. montebelloensis Álvarez & Villalobos, 1998 and P. yoshibensis Álvarez & Villalobos, 1998) and in northern Guatemala (Álvarez & Villalobos, 1998).	en	Villalobos, José Luis, Álvarez, Fernando (2013): Two new genera and three new species of freshwater crabs (Crustacea: Pseudothelphusidae: Potamocarcinini) from Chiapas, Mexico. Zootaxa 3599 (5): 457-470, DOI: 10.11646/zootaxa.3599.5.3
410487DCB7111D72FF23524AFB7454FD.taxon	materials_examined	Type material. Holotype: male, cb 31.2 mm, cl 20.9 mm; ditch along highway 101 near Villa de las Rosas (16 º 21 ' 0.5 " N, 92 º 23 ' 59.07 " W; 1152 m), Municipio de Villa de las Rosas, Chiapas, Mexico; 3 Dec. 2007; coll. F. Álvarez, J. L. Villalobos, A. Botello, C. Hernández; CNCR 26192. Allotype: female, cb 24.8 mm, cl 16.8 mm,; same locality as holotype, 30 Apr 2009; coll. F. Álvarez, J. L. Villalobos, G. Armendáriz, E. Torres; CNCR 25385. Paratypes: 4 males, cb 32.5 - 21.0 mm, cl 21.0 - 18.7 mm; 3 females, cb 21.9 - 19.7 mm, cl 15.0 - 13.4 mm; same locality, date and collectors as allotype; CNCR 27339.	en	Villalobos, José Luis, Álvarez, Fernando (2013): Two new genera and three new species of freshwater crabs (Crustacea: Pseudothelphusidae: Potamocarcinini) from Chiapas, Mexico. Zootaxa 3599 (5): 457-470, DOI: 10.11646/zootaxa.3599.5.3
410487DCB7111D72FF23524AFB7454FD.taxon	description	Description. In dorsal view carapace moderately oval, widest in middle (cb / cl = 1.63); dorsal surface convex, punctated, regions faintly indicated, cephalic and branchial somewhat elevated; pair of gastric pits barely distinct, oblique, narrowly elongated; cervical grooves wide, shallow, arched, not reaching anterolateral margin (Fig. 6 a). Postfrontal lobes and median groove discernible. Frontal region curved downwards to frontal border; superior frontal border absent, inferior one smooth, projected, divided into 2 low convex lobes, separated by rounded medial depression (Fig 6 d). Superior orbital margin continuous with frontal border; inferior orbital margin marked by row of rounded granules; exorbital angle flat. Anterolateral margin smooth along anterior third, granulated on branchial region, straight through posterior half. Posterior margin concave. Eyes with cornea normally faceted, pigmented, filling orbits. Epistome narrow; epistomial tooth low, triangular, deflexed, borders smooth. Suborbital and subhepatic regions of carapace smooth, punctated; pterygostomial region pubescent around mouthparts. Third maxilliped with ischium rectangular, merus with external margin widely rounded, ratio exopod / ischium 0.9 (Fig. 6 c). Aperture of efferent branchial channel subquadrate. Chelipeds different in shape and size. Major cheliped with merus subtriangular in cross section, short, thick, shorter than carapace length, dorsal surface smooth; internal dorsal margin with row of acute granules; external margin delimited by rounded granules on proximal two-thirds. Carpus smooth with small, low tubercles on internal margin. Palm compressed, high (length / high = 1.3); fingers longer than palm, stout, smooth, not gaping, with low, triangular teeth on cutting edges (Fig. 6 b). Thoracic sternites of third maxilliped and first pereiopod completely fused, except for short fissures at lateral edges of sternum; sternal sulci s 4 / s 5, s 5 / s 6 and s 6 / s 7 distinct, just failing to reach midline of thoracic sternum; sternal sulcus s 7 / s 8 reaching midline. All abdominal segments free. Telson subtriangular, lateral margins straight, tip rounded. First male gonopod as described under genus diagnosis.	en	Villalobos, José Luis, Álvarez, Fernando (2013): Two new genera and three new species of freshwater crabs (Crustacea: Pseudothelphusidae: Potamocarcinini) from Chiapas, Mexico. Zootaxa 3599 (5): 457-470, DOI: 10.11646/zootaxa.3599.5.3
410487DCB7111D72FF23524AFB7454FD.taxon	etymology	Etymology. The specific name is derived from " Villa de las Rosas " the name of the closest town to the type locality in Chiapas, Mexico.	en	Villalobos, José Luis, Álvarez, Fernando (2013): Two new genera and three new species of freshwater crabs (Crustacea: Pseudothelphusidae: Potamocarcinini) from Chiapas, Mexico. Zootaxa 3599 (5): 457-470, DOI: 10.11646/zootaxa.3599.5.3
410487DCB7111D72FF23524AFB7454FD.taxon	discussion	Remarks. Tzotzilthelphusa villarosalensis n. sp. has a gonopod with the distal third bent laterally, but without torsion. The apical cavity of the gonopod is oblique to the principal axis of the gonopod due to the elevation of the mesial surface. The gonopod morphology of the new species is similar to that of Typhlopseudothelphusa hyba, raising an interesting problem because that species was placed in Typhlopseudothelphusa due to its external morphology, which is very similar to that of T. mocinoi. However, considering the gonopod morphology T. hyba could be placed in Tzotzilthelphusa n. gen., a problem that will have to be addressed in the future. Geographically, T. hyba and T. villarosalensis n. sp. are only 40 km apart to the south of San Cristóbal de las Casas.	en	Villalobos, José Luis, Álvarez, Fernando (2013): Two new genera and three new species of freshwater crabs (Crustacea: Pseudothelphusidae: Potamocarcinini) from Chiapas, Mexico. Zootaxa 3599 (5): 457-470, DOI: 10.11646/zootaxa.3599.5.3
