taxonID	type	description	language	source
B31220ED11E34B0E8DEB6744DCFF1612.taxon	materials_examined	Other material. One male, Yunnan Province, Guangnan County, Bamei Town, Ake Village, Miaopu Cave, 24 ° 14.767 ' N, 105 ° 05.384 ' E, alt. 690 m, 8 Jan. 2017, X. K. Jiang, H. M. Chen & X. Guo leg. (IBGAS).	en	Jiang, Xuankong, Guo, Xuan, Chen, Huiming, Xie, Zhicai (2018): Four new species of the Glyphiulusjavanicus group from southern China (Diplopoda, Spirostreptida, Cambalopsidae). ZooKeys 741: 155-179, DOI: http://dx.doi.org/10.3897/zookeys.741.23223, URL: http://dx.doi.org/10.3897/zookeys.741.23223
B31220ED11E34B0E8DEB6744DCFF1612.taxon	etymology	Etymology. This specific name is derived from the Latin word foetidus, meaning ' smelly' and refers to the extremely strong and unpleasant smell of the animals.	en	Jiang, Xuankong, Guo, Xuan, Chen, Huiming, Xie, Zhicai (2018): Four new species of the Glyphiulusjavanicus group from southern China (Diplopoda, Spirostreptida, Cambalopsidae). ZooKeys 741: 155-179, DOI: http://dx.doi.org/10.3897/zookeys.741.23223, URL: http://dx.doi.org/10.3897/zookeys.741.23223
B31220ED11E34B0E8DEB6744DCFF1612.taxon	diagnosis	Diagnosis. The new species can be diagnosed by the following combination of morphological characteristics: (1) all crests on collum complete and fully developed, carinotaxic formula I-III + P + M; (2) telopodite of male legs I strongly degenerated, bi-segmented, as high as coxal process; (3) coxosternal mesal process of anterior gonopod prolonged and subtriangular; (4) flagellum of posterior gonopod short, with multiple branches at inner margin. See also Key below.	en	Jiang, Xuankong, Guo, Xuan, Chen, Huiming, Xie, Zhicai (2018): Four new species of the Glyphiulusjavanicus group from southern China (Diplopoda, Spirostreptida, Cambalopsidae). ZooKeys 741: 155-179, DOI: http://dx.doi.org/10.3897/zookeys.741.23223, URL: http://dx.doi.org/10.3897/zookeys.741.23223
B31220ED11E34B0E8DEB6744DCFF1612.taxon	description	Description. Body segments with 53 - 67 p + 1 - 2 a + T (holotype 67 p + 1 a + T). Body size of ca. 45 - 63 mm long and 2.3 - 3.0 mm wide (holotype 62 and 2.7 mm, respectively). Colouration. Brown to dark brown in vivo (Fig. 1 A). In fixed condition, head red-brown with yellow dapples; collum yellow-brown, anterior and posterior margins and the crests red-brown; midbody red-brown, lateral crests, ozoporiferous tubercles and anterior rows of metatergal crests light yellow; antennae and legs pale to light yellow (Fig. 2). Head. Each eye patch with 30 - 45 pigmented ocelli arranged in five irregular vertical rows (Fig. 2 A, B). Antennae slender, 2.88 - 3.35 mm long. Terminal part of antennomeres V expanded (Fig. 2 B). Gnathochilarium with a separate promentum, polytrichous (Fig. 3 A). Collum. All crests on collum complete and fully developed, carinotaxic formula I-III + P + M (Fig. 2 A, B). Body segments. Postcollum constriction modest (Fig. 2 A). Metatergal crests well-developed (Fig. 2 A-E). Crests divided into two transverse rows of tubercles, carinotaxic formula 2 / 2 + I / i + 3 / 3 + I / i + 2 / 2. Anterior tubercle (except ozoporiferous one) small and round, posterior one strip-shaped (Fig. 2 A-E). Ozoporiferous tubercles round, wider than high, obviously larger than other tubercles (Fig. 3 E). Location of the tubercle behind ozopore relatively medial, set off from ozoporiferous tubercle (Figs 2 C-E, 3 E). Lateral crests rather small (Fig. 15). Midbody rings round in cross-section (Fig. 3 E), 2.02 - 2.44 mm high (vertical diameter) and 2.15 - 2.56 mm wide (horizontal diameter), the ratio of height to width 0.92 - 0.97. Telson. Epiproct simple, with a rounded caudal ridge and a strong dorsal tooth (Fig. 2 E). Paraprocts convex, polytrichous. Hypoproct crescent-shaped (Fig. 2 F). Walking legs. Slender, 2.71 - 3.15 mm long, obviously longer than body width (Fig. 3 E, F). Male sexual characters. Male legs I strongly degenerated, with a pair of bi-segmented telopodites and a pair of large, subdigitiform, coxal processes. Coxal processes contiguous medially and curved forward, with clusters of long and robust setae at base (Fig. 3 B). Male legs II normal. Penes trapeziform and small, each possessing three robust distolateral setae (Fig. 3 C). Male legs III modified, with coxa especially slender and elongated (Fig. 3 D). Femora VI and VII normal, not inflated. Anterior gonopods. Coxosterna shield-like, sunken medially. Coxosternal mesal processes prolonged, obviously higher than telopodites. Telopodites one-segmented, placed laterally, curved and moveable, with several distal setae and a field of microsetae at base (Figs 4 A, 5 A, 6 A). Posterior gonopods. Compact (Figs 4 B, 5 B, 6 B). Coxite with a medial lamelliform lobe and two rows of strong and curved setae at mediolateral margin. Flagella short with multiple branches at inner margin (Fig. 5 C). Lateral margin with a field of microsetae (Fig. 5 D).	en	Jiang, Xuankong, Guo, Xuan, Chen, Huiming, Xie, Zhicai (2018): Four new species of the Glyphiulusjavanicus group from southern China (Diplopoda, Spirostreptida, Cambalopsidae). ZooKeys 741: 155-179, DOI: http://dx.doi.org/10.3897/zookeys.741.23223, URL: http://dx.doi.org/10.3897/zookeys.741.23223
B31220ED11E34B0E8DEB6744DCFF1612.taxon	distribution	Distribution. Known only from the type locality, a cave in Xilin County, Guangxi, and another cave in Guangnan County, Yunnan. The two caves are ca. 35 kilometres apart.	en	Jiang, Xuankong, Guo, Xuan, Chen, Huiming, Xie, Zhicai (2018): Four new species of the Glyphiulusjavanicus group from southern China (Diplopoda, Spirostreptida, Cambalopsidae). ZooKeys 741: 155-179, DOI: http://dx.doi.org/10.3897/zookeys.741.23223, URL: http://dx.doi.org/10.3897/zookeys.741.23223
B20D5F093321C753F5A4B9D9ECBA62F1.taxon	etymology	Etymology. This specific name is derived from the Latin word calceus, meaning ' shoe' and refers to the shape of the coxosternal mesal process of the anterior gonopod.	en	Jiang, Xuankong, Guo, Xuan, Chen, Huiming, Xie, Zhicai (2018): Four new species of the Glyphiulusjavanicus group from southern China (Diplopoda, Spirostreptida, Cambalopsidae). ZooKeys 741: 155-179, DOI: http://dx.doi.org/10.3897/zookeys.741.23223, URL: http://dx.doi.org/10.3897/zookeys.741.23223
B20D5F093321C753F5A4B9D9ECBA62F1.taxon	diagnosis	Diagnosis. The new species can be diagnosed by the following combination of morphological characteristics: (1) all crests on collum complete and fully developed, carinotaxic formula I-III + P + M; (2) telopodite of male legs I bi-segmented, obviously shorter than coxal process; (3) coxosternal mesal process of anterior gonopod prolonged and shoe-shaped; (4) flagellum of posterior gonopod short and zigzag-shaped. See also Key below.	en	Jiang, Xuankong, Guo, Xuan, Chen, Huiming, Xie, Zhicai (2018): Four new species of the Glyphiulusjavanicus group from southern China (Diplopoda, Spirostreptida, Cambalopsidae). ZooKeys 741: 155-179, DOI: http://dx.doi.org/10.3897/zookeys.741.23223, URL: http://dx.doi.org/10.3897/zookeys.741.23223
B20D5F093321C753F5A4B9D9ECBA62F1.taxon	description	Description. Body segments with 58 - 67 p + 1 - 2 a + T (holotype with 67 p + 1 a + T). Body size of ca. 45 - 63 mm long and 2.6 - 3.1 mm wide (holotype 58 and 2.9 mm, respectively). Colouration. Brown to yellow brown in vivo (Fig. 1 B); brown to red-brown in fixed condition (Fig. 7 A-F). Head. Each eye patch with 8 - 15 pigmented ocelli, arranged in two irregular vertical rows (Fig. 7 B). Antennae slender, 2.90 - 3.28 mm long. Terminal part of antennomeres V expanded (Fig. 7 B). Gnathochilarium with a separate promentum, polytrichous (Fig. 8 A). Collum. All crests complete and obvious, carinotaxic formula I-III + P + M (Fig. 7 A, B). Body segments. Postcollum constriction obvious (Fig. 7 A). Metaterga strongly crested (Fig. 7 A-F). Crests with two transverse rows of tubercles, carinotaxic for mula 2 / 2 + I / i + 3 / 3 + I / i + 2 / 2. Anterior tubercle (except ozoporiferous one) small and upright, posterior one directed caudally, both with sharp tips (Fig. 7 A-F). Ozoporiferous tubercle round, higher than broad, obviously larger than other tubercles (Fig. 8 E). Location of the tubercle behind ozopore relatively medial, set off from ozoporiferous tubercle in caudal view (Figs 7 C, D, 8 E). Lateral crests rather small. Midbody rings round in cross-section (Fig. 8 E), 2.10 - 2.48 mm high (vertical diameter) and 2.19 - 2.59 mm wide (horizontal diameter), the ratio of height to width 0.95 - 0.98. Telson. Epiproct simple, with a rounded caudal ridge and a strong dorsal tooth. Paraprocts convex. Hypoproct crescent-shaped (Fig. 7 E, F). Walking legs. 3.17 - 3.67 mm long, obviously longer than body width (Fig. 8 E, F). Male sexual characters. Telopodite of male legs I strongly degenerated, bi-segmented. Coxal processes obviously longer than telopodites (Fig. 8 B). Penes broad, tongue-shaped (Fig. 8 C). Male legs III with slender and elongated coxa (Fig. 8 D). Femora VI and VII normal, not inflated. Anterior gonopods. Coxosternum shield-like, sunken medially. Coxosternal mesal processes of anterior gonopods elongated and shoe-shaped, obviously higher than telopodites. Telopodite one-segmented, curved and moveable, with round tip and a field of microsetae at base (Figs 9 A, 10 A, 11 A). Posterior gonopods. Mediolateral margins of coxite brush-like. Flagella short and zigzag-shaped (Fig. 10 C). A long seta at anterolateral margin (Figs 9 B, 11 B). Lateral margin with a field of microsetae (Fig. 10 D).	en	Jiang, Xuankong, Guo, Xuan, Chen, Huiming, Xie, Zhicai (2018): Four new species of the Glyphiulusjavanicus group from southern China (Diplopoda, Spirostreptida, Cambalopsidae). ZooKeys 741: 155-179, DOI: http://dx.doi.org/10.3897/zookeys.741.23223, URL: http://dx.doi.org/10.3897/zookeys.741.23223
B20D5F093321C753F5A4B9D9ECBA62F1.taxon	distribution	Distribution. Known only from the type locality, a cave in Tian'e County, Guangxi Zhuang Autonomous Region.	en	Jiang, Xuankong, Guo, Xuan, Chen, Huiming, Xie, Zhicai (2018): Four new species of the Glyphiulusjavanicus group from southern China (Diplopoda, Spirostreptida, Cambalopsidae). ZooKeys 741: 155-179, DOI: http://dx.doi.org/10.3897/zookeys.741.23223, URL: http://dx.doi.org/10.3897/zookeys.741.23223
A9BC06F8BB7CFCCED25F716BD6BADE66.taxon	etymology	Etymology. This specific name is derived from the type locality.	en	Jiang, Xuankong, Guo, Xuan, Chen, Huiming, Xie, Zhicai (2018): Four new species of the Glyphiulusjavanicus group from southern China (Diplopoda, Spirostreptida, Cambalopsidae). ZooKeys 741: 155-179, DOI: http://dx.doi.org/10.3897/zookeys.741.23223, URL: http://dx.doi.org/10.3897/zookeys.741.23223
A9BC06F8BB7CFCCED25F716BD6BADE66.taxon	diagnosis	Diagnosis. The new species can be diagnosed by the following combination of morphological characteristics: (1) all crests on collum fully developed, carinotaxic formula 1 a + 2 c + III-IV + 5 c + 6 a + pc + ma + pc + 6 a + 5 c + IV-III + 2 c + 1 a; (2) metatergal crests not divided, carinotaxic formula 2 + I / i + 3 + I / i + 2 (3) telopodite of male legs I complete, not degenerated, five-segmented; (4) anterior gonopod possessing a coxosternal mesal process and a coxosternal lateral process, coxosternal mesal process with a long and sharp tip, coxosternal lateral process with a blunt tip; (5) flagellum of posterior gonopod extremely long and smooth, slightly curved. See also Key below.	en	Jiang, Xuankong, Guo, Xuan, Chen, Huiming, Xie, Zhicai (2018): Four new species of the Glyphiulusjavanicus group from southern China (Diplopoda, Spirostreptida, Cambalopsidae). ZooKeys 741: 155-179, DOI: http://dx.doi.org/10.3897/zookeys.741.23223, URL: http://dx.doi.org/10.3897/zookeys.741.23223
A9BC06F8BB7CFCCED25F716BD6BADE66.taxon	description	Description. Body segments with 56 - 73 p + 1 a + T (holotype 73 p + 1 a + T). Body size of ca. 38 - 55 mm long and 2.0 - 2.3 mm wide (holotype 54 and 2.3 mm, respectively). Colouration. Brown to dark brown in vivo (Fig. 1 C); taupe to red-brown in fixed condition (Fig. 12 A-F). Head. Each eye patch with 9 - 12 pigmented ocelli arranged in 2 - 3 irregular vertical rows (Fig. 12 B). Antennae slender, 2.20 - 2.38 mm long. Terminal part of antennomeres V slightly expanded (Fig. 12 B). Gnathochilarium with a separate promentum, polytrichous (Fig. 13 A). Collum. All crests developed, carinotaxic formula 1 a + 2 c + III-IV + 5 c + 6 a + pc + ma + pc + 6 a + 5 c + IV-III + 2 c + 1 a (Fig. 12 A, B). Body segments. Postcollum constriction modest (Fig. 12 A). Metaterga strongly crested (Fig. 12 A-F). All metatergal crests undivided (Fig. 12 A-D, F), carinotaxic formula 2 + I / i + 3 + I / i + 2. Anterior part of crest round and broad, posterior part strip-shaped. Ozoporiferous tubercles large and round, as high as broad. Lateral crests fully developed. Midbody rings round in cross-section (Fig. 13 E), 1.70 - 2.01 mm high (vertical diameter) and 1.74 - 2.08 mm wide (horizontal diameter), the ratio of height to width 0.96 - 0.99. Telson. Epiproct with a rounded caudal ridge and an evident, axial, dorsal rib (Fig. 12 F). Paraproct convex, with an evident depression near caudal edge, polytrichous. Hypoproct crescent-shaped (Fig. 12 E, F). Walking legs. 2.64 - 2.80 mm long, obviously longer than body width (Fig. 13 E, F). Male sexual characters. Telopodite of male legs I complete, five-segmented (Fig. 13 B). Penes rather broad and round (Fig. 13 C). Male legs II and III modified as usual (Fig. 13 C, D). Femora VI and VII normal, not inflated. Anterior gonopods. Coxosternum shield-like, sunken medially. Distal part of coxosternum with a deep indentation, the latter separating a mesal process and a lateral process. Coxosternal mesal process digitiform, obviously higher than telopodite. Coxosternal lateral process broad, with a blunt tip, nearly as high as telopodite. Telopodite short, one-segmented with thin and round tip and a field of microsetae at base (Figs 14 A, 15 A, 16 A). Posterior gonopods. Mediolateral margins of coxite brush-like. Flagella smooth, curved and extremely long. Lateral margin with a field of microsetae (Figs 14 B, 15 B, 16 B).	en	Jiang, Xuankong, Guo, Xuan, Chen, Huiming, Xie, Zhicai (2018): Four new species of the Glyphiulusjavanicus group from southern China (Diplopoda, Spirostreptida, Cambalopsidae). ZooKeys 741: 155-179, DOI: http://dx.doi.org/10.3897/zookeys.741.23223, URL: http://dx.doi.org/10.3897/zookeys.741.23223
A9BC06F8BB7CFCCED25F716BD6BADE66.taxon	distribution	Distribution. Known only from the type locality, a cave in Guangnan County, Yunnan Province.	en	Jiang, Xuankong, Guo, Xuan, Chen, Huiming, Xie, Zhicai (2018): Four new species of the Glyphiulusjavanicus group from southern China (Diplopoda, Spirostreptida, Cambalopsidae). ZooKeys 741: 155-179, DOI: http://dx.doi.org/10.3897/zookeys.741.23223, URL: http://dx.doi.org/10.3897/zookeys.741.23223
BF9AC4608DC9DEBBD73E5ABCE8BB9CAE.taxon	materials_examined	Other material examined. Seven males, 6 females and 4 juveniles, Lingyun County, Sicheng Town, Shuiyuan Cave 24 ° 21.992 ' N, 106 ° 34.670 ' E, alt. 450 m, 3 Jan. 2011, H. M. Chen leg. (IBGAS); 17 males and 14 females, Lingyun County, Sicheng Town, Naling Cave 24 ° 21.926 ' N, 106 ° 33.911 ' E, alt. 500 m, 4 Jan. 2011, H. M. Chen leg. (IBGAS); 3 males and 1 female, Fengshan County, Yuanyang Cave 24 ° 32.518 ' N, 107 ° 03.768 ' E, alt. 640 m, 3 Jan. 2017, X. K. Jiang, H. M. Chen & X. Guo leg. (IBGAS); 1 male, 2 females and 1 juvenile, Nandan County, Bachuan Cave 25 ° 03.966 ' N, 107 ° 37.392 ' E, 31 Jan. 2017, H. M. Chen & C. Chen leg. (IBGAS); 17 males, 24 females and 24 juveniles, Donglan County, Xinyan Village, Qiumotun, Ganma Cave 24 ° 26.784 ' N, 107 ° 20.584 ' E, alt. 320 m, 2 Feb. 2017, H. M. Chen & C. Chen leg. (IBGAS).	en	Jiang, Xuankong, Guo, Xuan, Chen, Huiming, Xie, Zhicai (2018): Four new species of the Glyphiulusjavanicus group from southern China (Diplopoda, Spirostreptida, Cambalopsidae). ZooKeys 741: 155-179, DOI: http://dx.doi.org/10.3897/zookeys.741.23223, URL: http://dx.doi.org/10.3897/zookeys.741.23223
BF9AC4608DC9DEBBD73E5ABCE8BB9CAE.taxon	etymology	Etymology. This specific name is derived from the Latin word impletus, meaning ' plentiful', referring to the large number of specimens of the new species in our collections.	en	Jiang, Xuankong, Guo, Xuan, Chen, Huiming, Xie, Zhicai (2018): Four new species of the Glyphiulusjavanicus group from southern China (Diplopoda, Spirostreptida, Cambalopsidae). ZooKeys 741: 155-179, DOI: http://dx.doi.org/10.3897/zookeys.741.23223, URL: http://dx.doi.org/10.3897/zookeys.741.23223
BF9AC4608DC9DEBBD73E5ABCE8BB9CAE.taxon	diagnosis	Diagnosis. The new species can be diagnosed by the following combination of morphological characteristics: (1) all crests on collum complete and fully developed, carinotaxic formula I-III + P + M; (2) telopodite of male leg I bi-segmented, shorter than coxal process; (3) coxosternal mesal process of anterior gonopod slender and strongly prolonged; (4) flagellum of posterior gonopod short and zigzag-shaped. See also Key below.	en	Jiang, Xuankong, Guo, Xuan, Chen, Huiming, Xie, Zhicai (2018): Four new species of the Glyphiulusjavanicus group from southern China (Diplopoda, Spirostreptida, Cambalopsidae). ZooKeys 741: 155-179, DOI: http://dx.doi.org/10.3897/zookeys.741.23223, URL: http://dx.doi.org/10.3897/zookeys.741.23223
BF9AC4608DC9DEBBD73E5ABCE8BB9CAE.taxon	description	Description. Body segments with 71 - 82 p + 1 a + T (holotype 73 p + 1 a + T). Body ca. 51 - 66 mm long and 2.3 - 3.2 mm wide (holotype 64 mm and 3.0 mm, respectively). Colouration. Brown to dark brown in vivo (Fig. 1 D). In fixed condition, yellow-brown to red-brown, tergal crests dark red-brown to castaneous brown (Fig. 17 A-F). Head. Each eye patch with 7 - 20 pigmented ocelli arranged in 1 - 3 irregular vertical rows (Fig. 17 A, B). Antennae slender, 2.34 - 3.31 mm long. Terminal part of antennomeres V obviously expanded (Fig. 17 B). Gnathochilarium with a separate promentum, polytrichous (Fig. 18 A). Collum. All crests on collum complete and fully developed, carinotaxic formula I-III + P + M (Fig. 17 A, B). Body segments. Postcollum constriction modest (Fig. 17 A). Metaterga strongly crested (Fig. 17 A-E). Metatergal crests divided into two transverse rows of tubercles, carinotaxic formula 2 / 2 + I / i + 3 / 3 + I / i + 2 / 2. Anterior tubercle (except ozoporiferous one) small and upright, posterior one directed caudally, both tubercles with sharp tips (Fig. 17 A-E). Ozoporiferous tubercle round, higher than broad, obviously larger than other tubercles (Fig. 18 E). Location of the tubercle behind ozopore relatively medial, set off from ozoporiferous tubercle in caudal view (Figs 17 B-E, 18 E). Lateral crests well developed. Midbody rings round in cross-section (Fig. 18 E), 1.88 - 2.42 mm high (vertical diameter) and 1.91 - 2.42 mm wide (horizontal diameter), the ratio of height to width 0.95 - 1.00. Telson. Epiproct simple, with a rounded caudal ridge and a strong dorsal tooth. Paraprocts convex, polytrichous. Hypoproct crescent-shaped (Fig. 17 E, F). Walking legs. Slender, 2.52 - 3.41 mm long, longer than body width (Fig. 18 E, F). Male sexual characters. Telopodite of male legs I strongly degraded, bi-segmented (Fig. 18 B). Penes rather small and oval (Fig. 18 C). Male legs II and III modified as usual (Fig. 18 C, D). Femora VI and VII normal, not inflated. Anterior gonopods. Coxosternum shield-like, sunken medially. Coxosternal mesal process slender and strongly prolonged. Telopodite thin, curved with a rounded tip, and a field of microsetae at base (Figs 19 A, 20 A, 21 A). Posterior gonopods. Mediolateral margins of coxite brush-like. Flagella short and zigzag-shaped. Lateral margin with a field of microsetae (Figs 19 B, 20 B, 21 B).	en	Jiang, Xuankong, Guo, Xuan, Chen, Huiming, Xie, Zhicai (2018): Four new species of the Glyphiulusjavanicus group from southern China (Diplopoda, Spirostreptida, Cambalopsidae). ZooKeys 741: 155-179, DOI: http://dx.doi.org/10.3897/zookeys.741.23223, URL: http://dx.doi.org/10.3897/zookeys.741.23223
BF9AC4608DC9DEBBD73E5ABCE8BB9CAE.taxon	distribution	Distribution. Known from the type locality and several caves scattered in northwestern Guangxi.	en	Jiang, Xuankong, Guo, Xuan, Chen, Huiming, Xie, Zhicai (2018): Four new species of the Glyphiulusjavanicus group from southern China (Diplopoda, Spirostreptida, Cambalopsidae). ZooKeys 741: 155-179, DOI: http://dx.doi.org/10.3897/zookeys.741.23223, URL: http://dx.doi.org/10.3897/zookeys.741.23223
