taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
385C87B6FFF30E50FCF9FABCFAD6F950.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927025/files/figure.png	https://doi.org/10.5281/zenodo.14927025	Figure 4. Aedeagi and vesici in Canthophorus spp. A, B, C. dubius dubius (Scopoli, 1763), Russia: Krasnaya Polyana, paratype; C–G, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Russia: Pukhlyakovskiy (C, D), Ukraine: Opolznevoe (E), Greece: Lake Kremasta (F), Tunisia: Sidi el Hani (G); H, C. melanopterus contrarius (Wagner, 1956). Aedeagus in lateral view (A) and in dorsal view (B, C); vesica in dorsoposterior view (D) and in lateral view (E–H). a.l – apical lobes of conjunctiva; dl.l – dorsolateral lobes of conjunctiva; l.b – lateral branch of vesica; sp – spicule of dorsolateral lobes; v.l – ventral lobe of conjunctiva; vs – vesica. Scale bares: 0.25 mm.	Figure 4. Aedeagi and vesici in Canthophorus spp. A, B, C. dubius dubius (Scopoli, 1763), Russia: Krasnaya Polyana, paratype; C–G, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Russia: Pukhlyakovskiy (C, D), Ukraine: Opolznevoe (E), Greece: Lake Kremasta (F), Tunisia: Sidi el Hani (G); H, C. melanopterus contrarius (Wagner, 1956). Aedeagus in lateral view (A) and in dorsal view (B, C); vesica in dorsoposterior view (D) and in lateral view (E–H). a.l – apical lobes of conjunctiva; dl.l – dorsolateral lobes of conjunctiva; l.b – lateral branch of vesica; sp – spicule of dorsolateral lobes; v.l – ventral lobe of conjunctiva; vs – vesica. Scale bares: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFF30E50FCF9FABCFAD6F950.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927029/files/figure.png	https://doi.org/10.5281/zenodo.14927029	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFF30E50FCF9FABCFAD6F950.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927033/files/figure.png	https://doi.org/10.5281/zenodo.14927033	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFF30E50FCF9FABCFAD6F950.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927035/files/figure.png	https://doi.org/10.5281/zenodo.14927035	Figure 9. Female terminalia in Canthophorus spp. A, B, C. dubius (Scopoli, 1763), Russia: Krasnaya Polyana (A), Italy: Lecco (B); C, C. melanopterus melanopterus (Herrich- Schaeffer, 1835), Azerbaijan: Ganja; D, C. melanopterus mariae n. ssp., Tajikistan: Kondara Gorge, paratype. Gynatrial sac, dorsal view (B), capsule of spermatheca (C, D). Scale bars: 0.25 mm.	Figure 9. Female terminalia in Canthophorus spp. A, B, C. dubius (Scopoli, 1763), Russia: Krasnaya Polyana (A), Italy: Lecco (B); C, C. melanopterus melanopterus (Herrich- Schaeffer, 1835), Azerbaijan: Ganja; D, C. melanopterus mariae n. ssp., Tajikistan: Kondara Gorge, paratype. Gynatrial sac, dorsal view (B), capsule of spermatheca (C, D). Scale bars: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFF30E50FCF9FABCFAD6F950.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927037/files/figure.png	https://doi.org/10.5281/zenodo.14927037	Figure 10. Female terminalia in Canthophorus spp. A, B, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (A), Russia: Krasnoyarsk Terr. (B); C, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; D, E, C. wagneri Asanova, 1964, Russia: Anapa (D), Armenia: Sevan (E). Gynatrial sac (A, D) and its posterior part (B, C, E) in dorsal view. Scale bar: 0.25 mm.	Figure 10. Female terminalia in Canthophorus spp. A, B, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (A), Russia: Krasnoyarsk Terr. (B); C, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; D, E, C. wagneri Asanova, 1964, Russia: Anapa (D), Armenia: Sevan (E). Gynatrial sac (A, D) and its posterior part (B, C, E) in dorsal view. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFF30E50FCF9FABCFAD6F950.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927039/files/figure.png	https://doi.org/10.5281/zenodo.14927039	Figure 11. Female terminalia in Canthophorus spp. A, B, C. mixtus mixtus Asanova, 1964, Ukraine: Aleshki, paratype (A), Azerbaijan: Kars (B); C, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht, paratype; D, C. niveimarginatus Scott, 1874; Russia: Kamen’-Rybolov; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Gynatrial sac (A, C–E) and its posterior part (B) in dorsal view. Scale bar: 0.25 mm.	Figure 11. Female terminalia in Canthophorus spp. A, B, C. mixtus mixtus Asanova, 1964, Ukraine: Aleshki, paratype (A), Azerbaijan: Kars (B); C, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht, paratype; D, C. niveimarginatus Scott, 1874; Russia: Kamen’-Rybolov; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Gynatrial sac (A, C–E) and its posterior part (B) in dorsal view. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFF30E50FCF9FABCFAD6F950.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927021/files/figure.png	https://doi.org/10.5281/zenodo.14927021	Figure 2. Pygophores in Canthophorus and Adomerus spp. A, B, C. melanopterus (Herrich-Schaeffer, 1835); C, D, A. aeneus (Walker, 1867) n. comb.; E, F, A. maculipes (Mulsant & Rey, 1852) n. comb. Ventral view (A, C, E) and posterodorsal view (B, D, F). In A, C, E, subgenital plate is shown in dotted lines. d.inf – dorsal infolding; l.r – lateral ridge of subgenital plate; l. inf – lateral infolding; p.r – posterior ridge of subgenital plate; v.inf – ventral infolding. Scale bar: 0.25 mm.	Figure 2. Pygophores in Canthophorus and Adomerus spp. A, B, C. melanopterus (Herrich-Schaeffer, 1835); C, D, A. aeneus (Walker, 1867) n. comb.; E, F, A. maculipes (Mulsant & Rey, 1852) n. comb. Ventral view (A, C, E) and posterodorsal view (B, D, F). In A, C, E, subgenital plate is shown in dotted lines. d.inf – dorsal infolding; l.r – lateral ridge of subgenital plate; l. inf – lateral infolding; p.r – posterior ridge of subgenital plate; v.inf – ventral infolding. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFF30E50FCF9FABCFAD6F950.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927023/files/figure.png	https://doi.org/10.5281/zenodo.14927023	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFF30E50FCF9FABCFAD6F950.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927027/files/figure.png	https://doi.org/10.5281/zenodo.14927027	Figure 5. Aedeagi of Canthophorus spp. in dorsal view. A, C. impressus (Horváth, 1880), Armenia: Gekhmakhan Mt.; B, C. wagneri Asanova, 1964, Russia: Makhachkala; C, Canthophorus mixtus Asanova, 1964, Armenia: Kipchakh, 1964; D, C. niveimarginatus Scott, 1874, Mongolia: Ulaangom; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	Figure 5. Aedeagi of Canthophorus spp. in dorsal view. A, C. impressus (Horváth, 1880), Armenia: Gekhmakhan Mt.; B, C. wagneri Asanova, 1964, Russia: Makhachkala; C, Canthophorus mixtus Asanova, 1964, Armenia: Kipchakh, 1964; D, C. niveimarginatus Scott, 1874, Mongolia: Ulaangom; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFF30E50FCF9FABCFAD6F950.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927053/files/figure.png	https://doi.org/10.5281/zenodo.14927053	Figure 18. Aedeagus of Adomerus aeneus (Walker, 1867) n. comb. A, lateral; B, dorsal; and C, ventral views. d.l – dorsal lobes of conjunctiva; da.l – dorsoapical lobe; dl.l – dorsolateral lobes; vl.l – ventrolateral lobes; vs – vesica. Scale bar: 0.25 mm.	Figure 18. Aedeagus of Adomerus aeneus (Walker, 1867) n. comb. A, lateral; B, dorsal; and C, ventral views. d.l – dorsal lobes of conjunctiva; da.l – dorsoapical lobe; dl.l – dorsolateral lobes; vl.l – ventrolateral lobes; vs – vesica. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFF30E50FCF9FABCFAD6F950.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927055/files/figure.png	https://doi.org/10.5281/zenodo.14927055	Figure 19. Aedeagus of Adomerus maculipes (Mulsant & Rey, 1852) n. comb., A, lateral; B, dorsal; and C, ventral views. d.l – dorsal lobes of conjunctiva; da.l – dorsoapical lobe; dl.l – dorsolateral lobes; s.g – secondary gonopore; vl.l – ventrolateral lobes; vs – vesica. Scale bar: 0.25 mm.	Figure 19. Aedeagus of Adomerus maculipes (Mulsant & Rey, 1852) n. comb., A, lateral; B, dorsal; and C, ventral views. d.l – dorsal lobes of conjunctiva; da.l – dorsoapical lobe; dl.l – dorsolateral lobes; s.g – secondary gonopore; vl.l – ventrolateral lobes; vs – vesica. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFF30E50FCF9FABCFAD6F950.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927031/files/figure.png	https://doi.org/10.5281/zenodo.14927031	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFF30E50FCF9FABCFAD6F950.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927057/files/figure.png	https://doi.org/10.5281/zenodo.14927057	Figure 20. Female terminalia in Adomerus spp. A, B, A. aeneus (Walker, 1867) n. comb.; C, D, A. maculipes (Mulsant & Rey, 1852) n. comb. External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A, C); gynatrial sac, dorsal view (B, D). a.p – anterior pouches of gynatrium; d.f – dividing fold in cavity of anterior pouch; f.r – first rami; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; s.r – second rami; tr – torus. Scale bar: 0.25 mm.	Figure 20. Female terminalia in Adomerus spp. A, B, A. aeneus (Walker, 1867) n. comb.; C, D, A. maculipes (Mulsant & Rey, 1852) n. comb. External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A, C); gynatrial sac, dorsal view (B, D). a.p – anterior pouches of gynatrium; d.f – dividing fold in cavity of anterior pouch; f.r – first rami; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; s.r – second rami; tr – torus. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFF30E50FCF9FABCFAD6F950.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927019/files/figure.png	https://doi.org/10.5281/zenodo.14927019	Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency.	Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFFE0E52FC07F944FA84FEF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927019/files/figure.png	https://doi.org/10.5281/zenodo.14927019	Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency.	Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFFE0E52FC07F944FA84FEF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927023/files/figure.png	https://doi.org/10.5281/zenodo.14927023	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFFE0E52FC07F944FA84FEF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927025/files/figure.png	https://doi.org/10.5281/zenodo.14927025	Figure 4. Aedeagi and vesici in Canthophorus spp. A, B, C. dubius dubius (Scopoli, 1763), Russia: Krasnaya Polyana, paratype; C–G, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Russia: Pukhlyakovskiy (C, D), Ukraine: Opolznevoe (E), Greece: Lake Kremasta (F), Tunisia: Sidi el Hani (G); H, C. melanopterus contrarius (Wagner, 1956). Aedeagus in lateral view (A) and in dorsal view (B, C); vesica in dorsoposterior view (D) and in lateral view (E–H). a.l – apical lobes of conjunctiva; dl.l – dorsolateral lobes of conjunctiva; l.b – lateral branch of vesica; sp – spicule of dorsolateral lobes; v.l – ventral lobe of conjunctiva; vs – vesica. Scale bares: 0.25 mm.	Figure 4. Aedeagi and vesici in Canthophorus spp. A, B, C. dubius dubius (Scopoli, 1763), Russia: Krasnaya Polyana, paratype; C–G, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Russia: Pukhlyakovskiy (C, D), Ukraine: Opolznevoe (E), Greece: Lake Kremasta (F), Tunisia: Sidi el Hani (G); H, C. melanopterus contrarius (Wagner, 1956). Aedeagus in lateral view (A) and in dorsal view (B, C); vesica in dorsoposterior view (D) and in lateral view (E–H). a.l – apical lobes of conjunctiva; dl.l – dorsolateral lobes of conjunctiva; l.b – lateral branch of vesica; sp – spicule of dorsolateral lobes; v.l – ventral lobe of conjunctiva; vs – vesica. Scale bares: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFFE0E52FC07F944FA84FEF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927029/files/figure.png	https://doi.org/10.5281/zenodo.14927029	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFFE0E52FC07F944FA84FEF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927031/files/figure.png	https://doi.org/10.5281/zenodo.14927031	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFFE0E52FC07F944FA84FEF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927033/files/figure.png	https://doi.org/10.5281/zenodo.14927033	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFFE0E52FC07F944FA84FEF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927035/files/figure.png	https://doi.org/10.5281/zenodo.14927035	Figure 9. Female terminalia in Canthophorus spp. A, B, C. dubius (Scopoli, 1763), Russia: Krasnaya Polyana (A), Italy: Lecco (B); C, C. melanopterus melanopterus (Herrich- Schaeffer, 1835), Azerbaijan: Ganja; D, C. melanopterus mariae n. ssp., Tajikistan: Kondara Gorge, paratype. Gynatrial sac, dorsal view (B), capsule of spermatheca (C, D). Scale bars: 0.25 mm.	Figure 9. Female terminalia in Canthophorus spp. A, B, C. dubius (Scopoli, 1763), Russia: Krasnaya Polyana (A), Italy: Lecco (B); C, C. melanopterus melanopterus (Herrich- Schaeffer, 1835), Azerbaijan: Ganja; D, C. melanopterus mariae n. ssp., Tajikistan: Kondara Gorge, paratype. Gynatrial sac, dorsal view (B), capsule of spermatheca (C, D). Scale bars: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFFE0E52FC07F944FA84FEF1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927041/files/figure.png	https://doi.org/10.5281/zenodo.14927041	Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm.	Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFFC0E53FC3CFEE4FAD2FF10.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927019/files/figure.png	https://doi.org/10.5281/zenodo.14927019	Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency.	Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFFC0E53FC3CFEE4FAD2FF10.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927023/files/figure.png	https://doi.org/10.5281/zenodo.14927023	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFFC0E53FC3CFEE4FAD2FF10.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927029/files/figure.png	https://doi.org/10.5281/zenodo.14927029	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFFC0E53FC3CFEE4FAD2FF10.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927031/files/figure.png	https://doi.org/10.5281/zenodo.14927031	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFFC0E53FC3CFEE4FAD2FF10.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927033/files/figure.png	https://doi.org/10.5281/zenodo.14927033	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFFC0E53FC3CFEE4FAD2FF10.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927041/files/figure.png	https://doi.org/10.5281/zenodo.14927041	Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm.	Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFFD0E4AFC16FE84FDBDFCFE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927019/files/figure.png	https://doi.org/10.5281/zenodo.14927019	Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency.	Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFFD0E4AFC16FE84FDBDFCFE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927023/files/figure.png	https://doi.org/10.5281/zenodo.14927023	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFFD0E4AFC16FE84FDBDFCFE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927025/files/figure.png	https://doi.org/10.5281/zenodo.14927025	Figure 4. Aedeagi and vesici in Canthophorus spp. A, B, C. dubius dubius (Scopoli, 1763), Russia: Krasnaya Polyana, paratype; C–G, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Russia: Pukhlyakovskiy (C, D), Ukraine: Opolznevoe (E), Greece: Lake Kremasta (F), Tunisia: Sidi el Hani (G); H, C. melanopterus contrarius (Wagner, 1956). Aedeagus in lateral view (A) and in dorsal view (B, C); vesica in dorsoposterior view (D) and in lateral view (E–H). a.l – apical lobes of conjunctiva; dl.l – dorsolateral lobes of conjunctiva; l.b – lateral branch of vesica; sp – spicule of dorsolateral lobes; v.l – ventral lobe of conjunctiva; vs – vesica. Scale bares: 0.25 mm.	Figure 4. Aedeagi and vesici in Canthophorus spp. A, B, C. dubius dubius (Scopoli, 1763), Russia: Krasnaya Polyana, paratype; C–G, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Russia: Pukhlyakovskiy (C, D), Ukraine: Opolznevoe (E), Greece: Lake Kremasta (F), Tunisia: Sidi el Hani (G); H, C. melanopterus contrarius (Wagner, 1956). Aedeagus in lateral view (A) and in dorsal view (B, C); vesica in dorsoposterior view (D) and in lateral view (E–H). a.l – apical lobes of conjunctiva; dl.l – dorsolateral lobes of conjunctiva; l.b – lateral branch of vesica; sp – spicule of dorsolateral lobes; v.l – ventral lobe of conjunctiva; vs – vesica. Scale bares: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFFD0E4AFC16FE84FDBDFCFE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927029/files/figure.png	https://doi.org/10.5281/zenodo.14927029	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFFD0E4AFC16FE84FDBDFCFE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927031/files/figure.png	https://doi.org/10.5281/zenodo.14927031	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFFD0E4AFC16FE84FDBDFCFE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927033/files/figure.png	https://doi.org/10.5281/zenodo.14927033	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFFD0E4AFC16FE84FDBDFCFE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927035/files/figure.png	https://doi.org/10.5281/zenodo.14927035	Figure 9. Female terminalia in Canthophorus spp. A, B, C. dubius (Scopoli, 1763), Russia: Krasnaya Polyana (A), Italy: Lecco (B); C, C. melanopterus melanopterus (Herrich- Schaeffer, 1835), Azerbaijan: Ganja; D, C. melanopterus mariae n. ssp., Tajikistan: Kondara Gorge, paratype. Gynatrial sac, dorsal view (B), capsule of spermatheca (C, D). Scale bars: 0.25 mm.	Figure 9. Female terminalia in Canthophorus spp. A, B, C. dubius (Scopoli, 1763), Russia: Krasnaya Polyana (A), Italy: Lecco (B); C, C. melanopterus melanopterus (Herrich- Schaeffer, 1835), Azerbaijan: Ganja; D, C. melanopterus mariae n. ssp., Tajikistan: Kondara Gorge, paratype. Gynatrial sac, dorsal view (B), capsule of spermatheca (C, D). Scale bars: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFFD0E4AFC16FE84FDBDFCFE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927021/files/figure.png	https://doi.org/10.5281/zenodo.14927021	Figure 2. Pygophores in Canthophorus and Adomerus spp. A, B, C. melanopterus (Herrich-Schaeffer, 1835); C, D, A. aeneus (Walker, 1867) n. comb.; E, F, A. maculipes (Mulsant & Rey, 1852) n. comb. Ventral view (A, C, E) and posterodorsal view (B, D, F). In A, C, E, subgenital plate is shown in dotted lines. d.inf – dorsal infolding; l.r – lateral ridge of subgenital plate; l. inf – lateral infolding; p.r – posterior ridge of subgenital plate; v.inf – ventral infolding. Scale bar: 0.25 mm.	Figure 2. Pygophores in Canthophorus and Adomerus spp. A, B, C. melanopterus (Herrich-Schaeffer, 1835); C, D, A. aeneus (Walker, 1867) n. comb.; E, F, A. maculipes (Mulsant & Rey, 1852) n. comb. Ventral view (A, C, E) and posterodorsal view (B, D, F). In A, C, E, subgenital plate is shown in dotted lines. d.inf – dorsal infolding; l.r – lateral ridge of subgenital plate; l. inf – lateral infolding; p.r – posterior ridge of subgenital plate; v.inf – ventral infolding. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFFD0E4AFC16FE84FDBDFCFE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927041/files/figure.png	https://doi.org/10.5281/zenodo.14927041	Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm.	Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFFD0E4AFC16FE84FDBDFCFE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927043/files/figure.png	https://doi.org/10.5281/zenodo.14927043	Figure 13. Posterior part of gynatrial sac in Canthophorus melanopterus melanopterus (Herrich-Schaeffer, 1835), dorsal view. A, Ukraine: Kamenets-Podol’skiy; B, Slovenia, Krajina; C, France: Nice; D, Russia: 30 km NW Rutul; E, Ukraine: Alma Riv.; F, Armenia: ravine of Dzhervezh; G, Armenia: Dzhrvezh O Yerevan; H, Morocco: Aourir; I, “Egypt” [coll. Eversmann]; J, Algérie [coll. A.F Morawitz]. Scale bar: 0.25 mm.	Figure 13. Posterior part of gynatrial sac in Canthophorus melanopterus melanopterus (Herrich-Schaeffer, 1835), dorsal view. A, Ukraine: Kamenets-Podol’skiy; B, Slovenia, Krajina; C, France: Nice; D, Russia: 30 km NW Rutul; E, Ukraine: Alma Riv.; F, Armenia: ravine of Dzhervezh; G, Armenia: Dzhrvezh O Yerevan; H, Morocco: Aourir; I, “Egypt” [coll. Eversmann]; J, Algérie [coll. A.F Morawitz]. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFFD0E4AFC16FE84FDBDFCFE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927045/files/figure.png	https://doi.org/10.5281/zenodo.14927045	Figure 14. Posterior part of gynatrial sac in Canthophorus melanopterus (Herrich-Schaeffer, 1835), dorsal view. A, Tunisia: Sfax; B, Tunisia: Sidi el Hani; C, Libya: Tripoli; D, Algeria: Sidi-bel-Abbes; E, Tunisia [N.L. Pastukhov]; F, Tunisia: Al Hushayshinah; G, Egypt: “Aegypt.”; H, C. melanopterus contrarius (Wagner, 1956); I, C. melanopterus mariae n. ssp. [holotype]. Scale bar: 0.25 mm.	Figure 14. Posterior part of gynatrial sac in Canthophorus melanopterus (Herrich-Schaeffer, 1835), dorsal view. A, Tunisia: Sfax; B, Tunisia: Sidi el Hani; C, Libya: Tripoli; D, Algeria: Sidi-bel-Abbes; E, Tunisia [N.L. Pastukhov]; F, Tunisia: Al Hushayshinah; G, Egypt: “Aegypt.”; H, C. melanopterus contrarius (Wagner, 1956); I, C. melanopterus mariae n. ssp. [holotype]. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFE40E47FF05FCF2FC3AFDD0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927019/files/figure.png	https://doi.org/10.5281/zenodo.14927019	Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency.	Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFE40E47FF05FCF2FC3AFDD0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927023/files/figure.png	https://doi.org/10.5281/zenodo.14927023	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFE40E47FF05FCF2FC3AFDD0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927025/files/figure.png	https://doi.org/10.5281/zenodo.14927025	Figure 4. Aedeagi and vesici in Canthophorus spp. A, B, C. dubius dubius (Scopoli, 1763), Russia: Krasnaya Polyana, paratype; C–G, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Russia: Pukhlyakovskiy (C, D), Ukraine: Opolznevoe (E), Greece: Lake Kremasta (F), Tunisia: Sidi el Hani (G); H, C. melanopterus contrarius (Wagner, 1956). Aedeagus in lateral view (A) and in dorsal view (B, C); vesica in dorsoposterior view (D) and in lateral view (E–H). a.l – apical lobes of conjunctiva; dl.l – dorsolateral lobes of conjunctiva; l.b – lateral branch of vesica; sp – spicule of dorsolateral lobes; v.l – ventral lobe of conjunctiva; vs – vesica. Scale bares: 0.25 mm.	Figure 4. Aedeagi and vesici in Canthophorus spp. A, B, C. dubius dubius (Scopoli, 1763), Russia: Krasnaya Polyana, paratype; C–G, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Russia: Pukhlyakovskiy (C, D), Ukraine: Opolznevoe (E), Greece: Lake Kremasta (F), Tunisia: Sidi el Hani (G); H, C. melanopterus contrarius (Wagner, 1956). Aedeagus in lateral view (A) and in dorsal view (B, C); vesica in dorsoposterior view (D) and in lateral view (E–H). a.l – apical lobes of conjunctiva; dl.l – dorsolateral lobes of conjunctiva; l.b – lateral branch of vesica; sp – spicule of dorsolateral lobes; v.l – ventral lobe of conjunctiva; vs – vesica. Scale bares: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFE40E47FF05FCF2FC3AFDD0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927029/files/figure.png	https://doi.org/10.5281/zenodo.14927029	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFE40E47FF05FCF2FC3AFDD0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927031/files/figure.png	https://doi.org/10.5281/zenodo.14927031	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFE40E47FF05FCF2FC3AFDD0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927033/files/figure.png	https://doi.org/10.5281/zenodo.14927033	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFE40E47FF05FCF2FC3AFDD0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927035/files/figure.png	https://doi.org/10.5281/zenodo.14927035	Figure 9. Female terminalia in Canthophorus spp. A, B, C. dubius (Scopoli, 1763), Russia: Krasnaya Polyana (A), Italy: Lecco (B); C, C. melanopterus melanopterus (Herrich- Schaeffer, 1835), Azerbaijan: Ganja; D, C. melanopterus mariae n. ssp., Tajikistan: Kondara Gorge, paratype. Gynatrial sac, dorsal view (B), capsule of spermatheca (C, D). Scale bars: 0.25 mm.	Figure 9. Female terminalia in Canthophorus spp. A, B, C. dubius (Scopoli, 1763), Russia: Krasnaya Polyana (A), Italy: Lecco (B); C, C. melanopterus melanopterus (Herrich- Schaeffer, 1835), Azerbaijan: Ganja; D, C. melanopterus mariae n. ssp., Tajikistan: Kondara Gorge, paratype. Gynatrial sac, dorsal view (B), capsule of spermatheca (C, D). Scale bars: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFE40E47FF05FCF2FC3AFDD0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927041/files/figure.png	https://doi.org/10.5281/zenodo.14927041	Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm.	Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFE40E47FF05FCF2FC3AFDD0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927043/files/figure.png	https://doi.org/10.5281/zenodo.14927043	Figure 13. Posterior part of gynatrial sac in Canthophorus melanopterus melanopterus (Herrich-Schaeffer, 1835), dorsal view. A, Ukraine: Kamenets-Podol’skiy; B, Slovenia, Krajina; C, France: Nice; D, Russia: 30 km NW Rutul; E, Ukraine: Alma Riv.; F, Armenia: ravine of Dzhervezh; G, Armenia: Dzhrvezh O Yerevan; H, Morocco: Aourir; I, “Egypt” [coll. Eversmann]; J, Algérie [coll. A.F Morawitz]. Scale bar: 0.25 mm.	Figure 13. Posterior part of gynatrial sac in Canthophorus melanopterus melanopterus (Herrich-Schaeffer, 1835), dorsal view. A, Ukraine: Kamenets-Podol’skiy; B, Slovenia, Krajina; C, France: Nice; D, Russia: 30 km NW Rutul; E, Ukraine: Alma Riv.; F, Armenia: ravine of Dzhervezh; G, Armenia: Dzhrvezh O Yerevan; H, Morocco: Aourir; I, “Egypt” [coll. Eversmann]; J, Algérie [coll. A.F Morawitz]. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFE40E47FF05FCF2FC3AFDD0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927045/files/figure.png	https://doi.org/10.5281/zenodo.14927045	Figure 14. Posterior part of gynatrial sac in Canthophorus melanopterus (Herrich-Schaeffer, 1835), dorsal view. A, Tunisia: Sfax; B, Tunisia: Sidi el Hani; C, Libya: Tripoli; D, Algeria: Sidi-bel-Abbes; E, Tunisia [N.L. Pastukhov]; F, Tunisia: Al Hushayshinah; G, Egypt: “Aegypt.”; H, C. melanopterus contrarius (Wagner, 1956); I, C. melanopterus mariae n. ssp. [holotype]. Scale bar: 0.25 mm.	Figure 14. Posterior part of gynatrial sac in Canthophorus melanopterus (Herrich-Schaeffer, 1835), dorsal view. A, Tunisia: Sfax; B, Tunisia: Sidi el Hani; C, Libya: Tripoli; D, Algeria: Sidi-bel-Abbes; E, Tunisia [N.L. Pastukhov]; F, Tunisia: Al Hushayshinah; G, Egypt: “Aegypt.”; H, C. melanopterus contrarius (Wagner, 1956); I, C. melanopterus mariae n. ssp. [holotype]. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFE40E47FF05FCF2FC3AFDD0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927049/files/figure.png	https://doi.org/10.5281/zenodo.14927049	Figure 16. Geographical variability of male and female terminalia in Canthophorus melanopterus (Herrich-Schaeffer, 1835). 1–6 in legend correspond to numbers of forms 1–6 by female terminalia in C. m. melanopterus; 7, females of C. m. mariae n. ssp.; 8, males with long lateral branch of vesica; 9, males with short lateral branch of vesica.	Figure 16. Geographical variability of male and female terminalia in Canthophorus melanopterus (Herrich-Schaeffer, 1835). 1–6 in legend correspond to numbers of forms 1–6 by female terminalia in C. m. melanopterus; 7, females of C. m. mariae n. ssp.; 8, males with long lateral branch of vesica; 9, males with short lateral branch of vesica.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFE90E40FCF8FDC4FF73FC7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927019/files/figure.png	https://doi.org/10.5281/zenodo.14927019	Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency.	Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFE90E40FCF8FDC4FF73FC7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927025/files/figure.png	https://doi.org/10.5281/zenodo.14927025	Figure 4. Aedeagi and vesici in Canthophorus spp. A, B, C. dubius dubius (Scopoli, 1763), Russia: Krasnaya Polyana, paratype; C–G, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Russia: Pukhlyakovskiy (C, D), Ukraine: Opolznevoe (E), Greece: Lake Kremasta (F), Tunisia: Sidi el Hani (G); H, C. melanopterus contrarius (Wagner, 1956). Aedeagus in lateral view (A) and in dorsal view (B, C); vesica in dorsoposterior view (D) and in lateral view (E–H). a.l – apical lobes of conjunctiva; dl.l – dorsolateral lobes of conjunctiva; l.b – lateral branch of vesica; sp – spicule of dorsolateral lobes; v.l – ventral lobe of conjunctiva; vs – vesica. Scale bares: 0.25 mm.	Figure 4. Aedeagi and vesici in Canthophorus spp. A, B, C. dubius dubius (Scopoli, 1763), Russia: Krasnaya Polyana, paratype; C–G, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Russia: Pukhlyakovskiy (C, D), Ukraine: Opolznevoe (E), Greece: Lake Kremasta (F), Tunisia: Sidi el Hani (G); H, C. melanopterus contrarius (Wagner, 1956). Aedeagus in lateral view (A) and in dorsal view (B, C); vesica in dorsoposterior view (D) and in lateral view (E–H). a.l – apical lobes of conjunctiva; dl.l – dorsolateral lobes of conjunctiva; l.b – lateral branch of vesica; sp – spicule of dorsolateral lobes; v.l – ventral lobe of conjunctiva; vs – vesica. Scale bares: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFE90E40FCF8FDC4FF73FC7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927031/files/figure.png	https://doi.org/10.5281/zenodo.14927031	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFE90E40FCF8FDC4FF73FC7F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927045/files/figure.png	https://doi.org/10.5281/zenodo.14927045	Figure 14. Posterior part of gynatrial sac in Canthophorus melanopterus (Herrich-Schaeffer, 1835), dorsal view. A, Tunisia: Sfax; B, Tunisia: Sidi el Hani; C, Libya: Tripoli; D, Algeria: Sidi-bel-Abbes; E, Tunisia [N.L. Pastukhov]; F, Tunisia: Al Hushayshinah; G, Egypt: “Aegypt.”; H, C. melanopterus contrarius (Wagner, 1956); I, C. melanopterus mariae n. ssp. [holotype]. Scale bar: 0.25 mm.	Figure 14. Posterior part of gynatrial sac in Canthophorus melanopterus (Herrich-Schaeffer, 1835), dorsal view. A, Tunisia: Sfax; B, Tunisia: Sidi el Hani; C, Libya: Tripoli; D, Algeria: Sidi-bel-Abbes; E, Tunisia [N.L. Pastukhov]; F, Tunisia: Al Hushayshinah; G, Egypt: “Aegypt.”; H, C. melanopterus contrarius (Wagner, 1956); I, C. melanopterus mariae n. ssp. [holotype]. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFEE0E40FF50FC60FB5DF96B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927031/files/figure.png	https://doi.org/10.5281/zenodo.14927031	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFEE0E40FF50FC60FB5DF96B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927035/files/figure.png	https://doi.org/10.5281/zenodo.14927035	Figure 9. Female terminalia in Canthophorus spp. A, B, C. dubius (Scopoli, 1763), Russia: Krasnaya Polyana (A), Italy: Lecco (B); C, C. melanopterus melanopterus (Herrich- Schaeffer, 1835), Azerbaijan: Ganja; D, C. melanopterus mariae n. ssp., Tajikistan: Kondara Gorge, paratype. Gynatrial sac, dorsal view (B), capsule of spermatheca (C, D). Scale bars: 0.25 mm.	Figure 9. Female terminalia in Canthophorus spp. A, B, C. dubius (Scopoli, 1763), Russia: Krasnaya Polyana (A), Italy: Lecco (B); C, C. melanopterus melanopterus (Herrich- Schaeffer, 1835), Azerbaijan: Ganja; D, C. melanopterus mariae n. ssp., Tajikistan: Kondara Gorge, paratype. Gynatrial sac, dorsal view (B), capsule of spermatheca (C, D). Scale bars: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFEE0E40FF50FC60FB5DF96B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927045/files/figure.png	https://doi.org/10.5281/zenodo.14927045	Figure 14. Posterior part of gynatrial sac in Canthophorus melanopterus (Herrich-Schaeffer, 1835), dorsal view. A, Tunisia: Sfax; B, Tunisia: Sidi el Hani; C, Libya: Tripoli; D, Algeria: Sidi-bel-Abbes; E, Tunisia [N.L. Pastukhov]; F, Tunisia: Al Hushayshinah; G, Egypt: “Aegypt.”; H, C. melanopterus contrarius (Wagner, 1956); I, C. melanopterus mariae n. ssp. [holotype]. Scale bar: 0.25 mm.	Figure 14. Posterior part of gynatrial sac in Canthophorus melanopterus (Herrich-Schaeffer, 1835), dorsal view. A, Tunisia: Sfax; B, Tunisia: Sidi el Hani; C, Libya: Tripoli; D, Algeria: Sidi-bel-Abbes; E, Tunisia [N.L. Pastukhov]; F, Tunisia: Al Hushayshinah; G, Egypt: “Aegypt.”; H, C. melanopterus contrarius (Wagner, 1956); I, C. melanopterus mariae n. ssp. [holotype]. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFEE0E40FF50FC60FB5DF96B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927049/files/figure.png	https://doi.org/10.5281/zenodo.14927049	Figure 16. Geographical variability of male and female terminalia in Canthophorus melanopterus (Herrich-Schaeffer, 1835). 1–6 in legend correspond to numbers of forms 1–6 by female terminalia in C. m. melanopterus; 7, females of C. m. mariae n. ssp.; 8, males with long lateral branch of vesica; 9, males with short lateral branch of vesica.	Figure 16. Geographical variability of male and female terminalia in Canthophorus melanopterus (Herrich-Schaeffer, 1835). 1–6 in legend correspond to numbers of forms 1–6 by female terminalia in C. m. melanopterus; 7, females of C. m. mariae n. ssp.; 8, males with long lateral branch of vesica; 9, males with short lateral branch of vesica.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFEE0E41FC2FF979FBD4FC8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927019/files/figure.png	https://doi.org/10.5281/zenodo.14927019	Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency.	Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFEE0E41FC2FF979FBD4FC8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927023/files/figure.png	https://doi.org/10.5281/zenodo.14927023	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFEE0E41FC2FF979FBD4FC8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927027/files/figure.png	https://doi.org/10.5281/zenodo.14927027	Figure 5. Aedeagi of Canthophorus spp. in dorsal view. A, C. impressus (Horváth, 1880), Armenia: Gekhmakhan Mt.; B, C. wagneri Asanova, 1964, Russia: Makhachkala; C, Canthophorus mixtus Asanova, 1964, Armenia: Kipchakh, 1964; D, C. niveimarginatus Scott, 1874, Mongolia: Ulaangom; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	Figure 5. Aedeagi of Canthophorus spp. in dorsal view. A, C. impressus (Horváth, 1880), Armenia: Gekhmakhan Mt.; B, C. wagneri Asanova, 1964, Russia: Makhachkala; C, Canthophorus mixtus Asanova, 1964, Armenia: Kipchakh, 1964; D, C. niveimarginatus Scott, 1874, Mongolia: Ulaangom; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFEE0E41FC2FF979FBD4FC8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927029/files/figure.png	https://doi.org/10.5281/zenodo.14927029	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFEE0E41FC2FF979FBD4FC8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927031/files/figure.png	https://doi.org/10.5281/zenodo.14927031	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFEE0E41FC2FF979FBD4FC8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927033/files/figure.png	https://doi.org/10.5281/zenodo.14927033	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFEE0E41FC2FF979FBD4FC8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927037/files/figure.png	https://doi.org/10.5281/zenodo.14927037	Figure 10. Female terminalia in Canthophorus spp. A, B, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (A), Russia: Krasnoyarsk Terr. (B); C, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; D, E, C. wagneri Asanova, 1964, Russia: Anapa (D), Armenia: Sevan (E). Gynatrial sac (A, D) and its posterior part (B, C, E) in dorsal view. Scale bar: 0.25 mm.	Figure 10. Female terminalia in Canthophorus spp. A, B, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (A), Russia: Krasnoyarsk Terr. (B); C, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; D, E, C. wagneri Asanova, 1964, Russia: Anapa (D), Armenia: Sevan (E). Gynatrial sac (A, D) and its posterior part (B, C, E) in dorsal view. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFEE0E41FC2FF979FBD4FC8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927041/files/figure.png	https://doi.org/10.5281/zenodo.14927041	Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm.	Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFEF0E42FCEAFC19FB64FAFF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927019/files/figure.png	https://doi.org/10.5281/zenodo.14927019	Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency.	Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFEF0E42FCEAFC19FB64FAFF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927023/files/figure.png	https://doi.org/10.5281/zenodo.14927023	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFEF0E42FCEAFC19FB64FAFF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927029/files/figure.png	https://doi.org/10.5281/zenodo.14927029	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFEF0E42FCEAFC19FB64FAFF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927031/files/figure.png	https://doi.org/10.5281/zenodo.14927031	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFEF0E42FCEAFC19FB64FAFF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927033/files/figure.png	https://doi.org/10.5281/zenodo.14927033	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFEF0E42FCEAFC19FB64FAFF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927037/files/figure.png	https://doi.org/10.5281/zenodo.14927037	Figure 10. Female terminalia in Canthophorus spp. A, B, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (A), Russia: Krasnoyarsk Terr. (B); C, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; D, E, C. wagneri Asanova, 1964, Russia: Anapa (D), Armenia: Sevan (E). Gynatrial sac (A, D) and its posterior part (B, C, E) in dorsal view. Scale bar: 0.25 mm.	Figure 10. Female terminalia in Canthophorus spp. A, B, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (A), Russia: Krasnoyarsk Terr. (B); C, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; D, E, C. wagneri Asanova, 1964, Russia: Anapa (D), Armenia: Sevan (E). Gynatrial sac (A, D) and its posterior part (B, C, E) in dorsal view. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFEF0E42FCEAFC19FB64FAFF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927041/files/figure.png	https://doi.org/10.5281/zenodo.14927041	Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm.	Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFEC0E43FC3DFAEAFB49F9DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927019/files/figure.png	https://doi.org/10.5281/zenodo.14927019	Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency.	Figure 1. Hemelytral membranes in Canthophorus spp. A, C. dubius dubius (Scopoli, 1763), France: Roquefort-les-Pins; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Ascania Nova; D, C. melanopterus contrarius (Wagner, 1956), Cyprus: Château de Buffavento; E, C. impressus impressus (Horváth, 1880), Russia: Khodzhal-Makhi; F, C. impressus hajastanicus n. ssp., Georgia: gorge of Paravanichet Riv., paratype. Abdomens removed to show membrane colour in transparency.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFEC0E43FC3DFAEAFB49F9DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927023/files/figure.png	https://doi.org/10.5281/zenodo.14927023	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFEC0E43FC3DFAEAFB49F9DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927027/files/figure.png	https://doi.org/10.5281/zenodo.14927027	Figure 5. Aedeagi of Canthophorus spp. in dorsal view. A, C. impressus (Horváth, 1880), Armenia: Gekhmakhan Mt.; B, C. wagneri Asanova, 1964, Russia: Makhachkala; C, Canthophorus mixtus Asanova, 1964, Armenia: Kipchakh, 1964; D, C. niveimarginatus Scott, 1874, Mongolia: Ulaangom; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	Figure 5. Aedeagi of Canthophorus spp. in dorsal view. A, C. impressus (Horváth, 1880), Armenia: Gekhmakhan Mt.; B, C. wagneri Asanova, 1964, Russia: Makhachkala; C, Canthophorus mixtus Asanova, 1964, Armenia: Kipchakh, 1964; D, C. niveimarginatus Scott, 1874, Mongolia: Ulaangom; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFEC0E43FC3DFAEAFB49F9DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927031/files/figure.png	https://doi.org/10.5281/zenodo.14927031	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFEC0E43FC3DFAEAFB49F9DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927033/files/figure.png	https://doi.org/10.5281/zenodo.14927033	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFEC0E43FC3DFAEAFB49F9DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927037/files/figure.png	https://doi.org/10.5281/zenodo.14927037	Figure 10. Female terminalia in Canthophorus spp. A, B, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (A), Russia: Krasnoyarsk Terr. (B); C, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; D, E, C. wagneri Asanova, 1964, Russia: Anapa (D), Armenia: Sevan (E). Gynatrial sac (A, D) and its posterior part (B, C, E) in dorsal view. Scale bar: 0.25 mm.	Figure 10. Female terminalia in Canthophorus spp. A, B, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (A), Russia: Krasnoyarsk Terr. (B); C, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; D, E, C. wagneri Asanova, 1964, Russia: Anapa (D), Armenia: Sevan (E). Gynatrial sac (A, D) and its posterior part (B, C, E) in dorsal view. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFEC0E43FC3DFAEAFB49F9DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927041/files/figure.png	https://doi.org/10.5281/zenodo.14927041	Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm.	Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFED0E7CFC19F9C5FB41FF11.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927023/files/figure.png	https://doi.org/10.5281/zenodo.14927023	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFED0E7CFC19F9C5FB41FF11.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927027/files/figure.png	https://doi.org/10.5281/zenodo.14927027	Figure 5. Aedeagi of Canthophorus spp. in dorsal view. A, C. impressus (Horváth, 1880), Armenia: Gekhmakhan Mt.; B, C. wagneri Asanova, 1964, Russia: Makhachkala; C, Canthophorus mixtus Asanova, 1964, Armenia: Kipchakh, 1964; D, C. niveimarginatus Scott, 1874, Mongolia: Ulaangom; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	Figure 5. Aedeagi of Canthophorus spp. in dorsal view. A, C. impressus (Horváth, 1880), Armenia: Gekhmakhan Mt.; B, C. wagneri Asanova, 1964, Russia: Makhachkala; C, Canthophorus mixtus Asanova, 1964, Armenia: Kipchakh, 1964; D, C. niveimarginatus Scott, 1874, Mongolia: Ulaangom; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFED0E7CFC19F9C5FB41FF11.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927029/files/figure.png	https://doi.org/10.5281/zenodo.14927029	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFED0E7CFC19F9C5FB41FF11.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927031/files/figure.png	https://doi.org/10.5281/zenodo.14927031	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFED0E7CFC19F9C5FB41FF11.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927033/files/figure.png	https://doi.org/10.5281/zenodo.14927033	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFED0E7CFC19F9C5FB41FF11.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927037/files/figure.png	https://doi.org/10.5281/zenodo.14927037	Figure 10. Female terminalia in Canthophorus spp. A, B, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (A), Russia: Krasnoyarsk Terr. (B); C, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; D, E, C. wagneri Asanova, 1964, Russia: Anapa (D), Armenia: Sevan (E). Gynatrial sac (A, D) and its posterior part (B, C, E) in dorsal view. Scale bar: 0.25 mm.	Figure 10. Female terminalia in Canthophorus spp. A, B, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (A), Russia: Krasnoyarsk Terr. (B); C, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; D, E, C. wagneri Asanova, 1964, Russia: Anapa (D), Armenia: Sevan (E). Gynatrial sac (A, D) and its posterior part (B, C, E) in dorsal view. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFED0E7CFC19F9C5FB41FF11.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927041/files/figure.png	https://doi.org/10.5281/zenodo.14927041	Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm.	Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD20E7DFCCEFE84FC8DF929.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927023/files/figure.png	https://doi.org/10.5281/zenodo.14927023	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD20E7DFCCEFE84FC8DF929.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927027/files/figure.png	https://doi.org/10.5281/zenodo.14927027	Figure 5. Aedeagi of Canthophorus spp. in dorsal view. A, C. impressus (Horváth, 1880), Armenia: Gekhmakhan Mt.; B, C. wagneri Asanova, 1964, Russia: Makhachkala; C, Canthophorus mixtus Asanova, 1964, Armenia: Kipchakh, 1964; D, C. niveimarginatus Scott, 1874, Mongolia: Ulaangom; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	Figure 5. Aedeagi of Canthophorus spp. in dorsal view. A, C. impressus (Horváth, 1880), Armenia: Gekhmakhan Mt.; B, C. wagneri Asanova, 1964, Russia: Makhachkala; C, Canthophorus mixtus Asanova, 1964, Armenia: Kipchakh, 1964; D, C. niveimarginatus Scott, 1874, Mongolia: Ulaangom; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD20E7DFCCEFE84FC8DF929.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927029/files/figure.png	https://doi.org/10.5281/zenodo.14927029	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD20E7DFCCEFE84FC8DF929.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927031/files/figure.png	https://doi.org/10.5281/zenodo.14927031	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD20E7DFCCEFE84FC8DF929.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927033/files/figure.png	https://doi.org/10.5281/zenodo.14927033	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD20E7DFCCEFE84FC8DF929.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927037/files/figure.png	https://doi.org/10.5281/zenodo.14927037	Figure 10. Female terminalia in Canthophorus spp. A, B, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (A), Russia: Krasnoyarsk Terr. (B); C, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; D, E, C. wagneri Asanova, 1964, Russia: Anapa (D), Armenia: Sevan (E). Gynatrial sac (A, D) and its posterior part (B, C, E) in dorsal view. Scale bar: 0.25 mm.	Figure 10. Female terminalia in Canthophorus spp. A, B, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (A), Russia: Krasnoyarsk Terr. (B); C, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; D, E, C. wagneri Asanova, 1964, Russia: Anapa (D), Armenia: Sevan (E). Gynatrial sac (A, D) and its posterior part (B, C, E) in dorsal view. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD20E7DFCCEFE84FC8DF929.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927041/files/figure.png	https://doi.org/10.5281/zenodo.14927041	Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm.	Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD30E7EFF58F8BFFC8CFDCC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927029/files/figure.png	https://doi.org/10.5281/zenodo.14927029	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD30E7EFF58F8BFFC8CFDCC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927031/files/figure.png	https://doi.org/10.5281/zenodo.14927031	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD00E7EFEB3FDD8FB78FA21.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927023/files/figure.png	https://doi.org/10.5281/zenodo.14927023	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD00E7EFEB3FDD8FB78FA21.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927027/files/figure.png	https://doi.org/10.5281/zenodo.14927027	Figure 5. Aedeagi of Canthophorus spp. in dorsal view. A, C. impressus (Horváth, 1880), Armenia: Gekhmakhan Mt.; B, C. wagneri Asanova, 1964, Russia: Makhachkala; C, Canthophorus mixtus Asanova, 1964, Armenia: Kipchakh, 1964; D, C. niveimarginatus Scott, 1874, Mongolia: Ulaangom; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	Figure 5. Aedeagi of Canthophorus spp. in dorsal view. A, C. impressus (Horváth, 1880), Armenia: Gekhmakhan Mt.; B, C. wagneri Asanova, 1964, Russia: Makhachkala; C, Canthophorus mixtus Asanova, 1964, Armenia: Kipchakh, 1964; D, C. niveimarginatus Scott, 1874, Mongolia: Ulaangom; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD00E7EFEB3FDD8FB78FA21.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927029/files/figure.png	https://doi.org/10.5281/zenodo.14927029	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD00E7EFEB3FDD8FB78FA21.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927031/files/figure.png	https://doi.org/10.5281/zenodo.14927031	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD00E7EFEB3FDD8FB78FA21.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927033/files/figure.png	https://doi.org/10.5281/zenodo.14927033	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD00E7EFEB3FDD8FB78FA21.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927039/files/figure.png	https://doi.org/10.5281/zenodo.14927039	Figure 11. Female terminalia in Canthophorus spp. A, B, C. mixtus mixtus Asanova, 1964, Ukraine: Aleshki, paratype (A), Azerbaijan: Kars (B); C, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht, paratype; D, C. niveimarginatus Scott, 1874; Russia: Kamen’-Rybolov; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Gynatrial sac (A, C–E) and its posterior part (B) in dorsal view. Scale bar: 0.25 mm.	Figure 11. Female terminalia in Canthophorus spp. A, B, C. mixtus mixtus Asanova, 1964, Ukraine: Aleshki, paratype (A), Azerbaijan: Kars (B); C, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht, paratype; D, C. niveimarginatus Scott, 1874; Russia: Kamen’-Rybolov; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Gynatrial sac (A, C–E) and its posterior part (B) in dorsal view. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD00E7EFEB3FDD8FB78FA21.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927041/files/figure.png	https://doi.org/10.5281/zenodo.14927041	Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm.	Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD00E78FC3FF9B7FB3CFACA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927023/files/figure.png	https://doi.org/10.5281/zenodo.14927023	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD00E78FC3FF9B7FB3CFACA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927027/files/figure.png	https://doi.org/10.5281/zenodo.14927027	Figure 5. Aedeagi of Canthophorus spp. in dorsal view. A, C. impressus (Horváth, 1880), Armenia: Gekhmakhan Mt.; B, C. wagneri Asanova, 1964, Russia: Makhachkala; C, Canthophorus mixtus Asanova, 1964, Armenia: Kipchakh, 1964; D, C. niveimarginatus Scott, 1874, Mongolia: Ulaangom; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	Figure 5. Aedeagi of Canthophorus spp. in dorsal view. A, C. impressus (Horváth, 1880), Armenia: Gekhmakhan Mt.; B, C. wagneri Asanova, 1964, Russia: Makhachkala; C, Canthophorus mixtus Asanova, 1964, Armenia: Kipchakh, 1964; D, C. niveimarginatus Scott, 1874, Mongolia: Ulaangom; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD00E78FC3FF9B7FB3CFACA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927029/files/figure.png	https://doi.org/10.5281/zenodo.14927029	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD00E78FC3FF9B7FB3CFACA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927031/files/figure.png	https://doi.org/10.5281/zenodo.14927031	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD00E78FC3FF9B7FB3CFACA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927033/files/figure.png	https://doi.org/10.5281/zenodo.14927033	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD00E78FC3FF9B7FB3CFACA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927039/files/figure.png	https://doi.org/10.5281/zenodo.14927039	Figure 11. Female terminalia in Canthophorus spp. A, B, C. mixtus mixtus Asanova, 1964, Ukraine: Aleshki, paratype (A), Azerbaijan: Kars (B); C, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht, paratype; D, C. niveimarginatus Scott, 1874; Russia: Kamen’-Rybolov; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Gynatrial sac (A, C–E) and its posterior part (B) in dorsal view. Scale bar: 0.25 mm.	Figure 11. Female terminalia in Canthophorus spp. A, B, C. mixtus mixtus Asanova, 1964, Ukraine: Aleshki, paratype (A), Azerbaijan: Kars (B); C, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht, paratype; D, C. niveimarginatus Scott, 1874; Russia: Kamen’-Rybolov; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Gynatrial sac (A, C–E) and its posterior part (B) in dorsal view. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD00E78FC3FF9B7FB3CFACA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927041/files/figure.png	https://doi.org/10.5281/zenodo.14927041	Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm.	Figure 12. Posterior pouches of gynatrial sac in Canthophorus spp., posterio-inner view. A, C. dubius (Scopoli, 1763), Italy: Lecco; B, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv.; C, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme”; D, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; E, C. wagneri Asanova, 1964, Russia: Anapa (some diagnostic characters is indicated by arrows); F, C. mixtus Asanova, 1964, Ukraine: Aleshki, paratype (pouch is shown straightened). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD60E7AFCF9FADEFD6AFEE8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927023/files/figure.png	https://doi.org/10.5281/zenodo.14927023	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD60E7AFCF9FADEFD6AFEE8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927031/files/figure.png	https://doi.org/10.5281/zenodo.14927031	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD60E7AFCF9FADEFD6AFEE8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927033/files/figure.png	https://doi.org/10.5281/zenodo.14927033	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD60E7AFCF9FADEFD6AFEE8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927039/files/figure.png	https://doi.org/10.5281/zenodo.14927039	Figure 11. Female terminalia in Canthophorus spp. A, B, C. mixtus mixtus Asanova, 1964, Ukraine: Aleshki, paratype (A), Azerbaijan: Kars (B); C, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht, paratype; D, C. niveimarginatus Scott, 1874; Russia: Kamen’-Rybolov; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Gynatrial sac (A, C–E) and its posterior part (B) in dorsal view. Scale bar: 0.25 mm.	Figure 11. Female terminalia in Canthophorus spp. A, B, C. mixtus mixtus Asanova, 1964, Ukraine: Aleshki, paratype (A), Azerbaijan: Kars (B); C, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht, paratype; D, C. niveimarginatus Scott, 1874; Russia: Kamen’-Rybolov; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Gynatrial sac (A, C–E) and its posterior part (B) in dorsal view. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD60E7AFCF9FADEFD6AFEE8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927051/files/figure.png	https://doi.org/10.5281/zenodo.14927051	Figure 17. Comparison of body measurements in Canthophorus mixtus mixtus Asanova, 1964 (1, 2) and C. mixtus hissaricus Asanova, 1964 n. stat. (3). 2, specimens from Trans-Ili and Alatau Khozretishi Range. BL – body length; BW – body width; PW – pronotum width.	Figure 17. Comparison of body measurements in Canthophorus mixtus mixtus Asanova, 1964 (1, 2) and C. mixtus hissaricus Asanova, 1964 n. stat. (3). 2, specimens from Trans-Ili and Alatau Khozretishi Range. BL – body length; BW – body width; PW – pronotum width.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD40E7AFF53FEFCFA99FC1A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927023/files/figure.png	https://doi.org/10.5281/zenodo.14927023	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD40E7AFF53FEFCFA99FC1A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927027/files/figure.png	https://doi.org/10.5281/zenodo.14927027	Figure 5. Aedeagi of Canthophorus spp. in dorsal view. A, C. impressus (Horváth, 1880), Armenia: Gekhmakhan Mt.; B, C. wagneri Asanova, 1964, Russia: Makhachkala; C, Canthophorus mixtus Asanova, 1964, Armenia: Kipchakh, 1964; D, C. niveimarginatus Scott, 1874, Mongolia: Ulaangom; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	Figure 5. Aedeagi of Canthophorus spp. in dorsal view. A, C. impressus (Horváth, 1880), Armenia: Gekhmakhan Mt.; B, C. wagneri Asanova, 1964, Russia: Makhachkala; C, Canthophorus mixtus Asanova, 1964, Armenia: Kipchakh, 1964; D, C. niveimarginatus Scott, 1874, Mongolia: Ulaangom; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD40E7AFF53FEFCFA99FC1A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927029/files/figure.png	https://doi.org/10.5281/zenodo.14927029	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD40E7AFF53FEFCFA99FC1A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927031/files/figure.png	https://doi.org/10.5281/zenodo.14927031	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	Figure 7. Spicules of conjunctival dorsolateral lobes in Canthophorus spp.A, C. dubius dubius (Scopoli, 1763), France:Forêt des Morières; B, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; C, D, C. melanopterus melanopterus (Herrich-Schaeffer, 1835), Ukraine: Opolznevoe (C), Tunisia, Sidi el Hani (D); E, C. melanopterus contrarius (Wagner, 1956); F, C.melanopterus mariae n. ssp., Tajikistan:Kondara Gorge, paratype; G, C. impressus impressus (Horváth, 1880), Ukraine:Demir-kapu; H, C. impressus hajastanicus n. ssp., Armenia:Gekhmakhan Mt., paratype; I, C. wagneri wagneri Asanova, 1964, Russia: Il’ Mt. SE Vladikavkaz; J, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly, paratype; K–Q, C. mixtus mixtus Asanova, 1964, Kazakhstan: Chagan-Aba (K), Kazakhstan: Yanvartsevo (L); Armenia: Kipchakh (M), Kazakhstan:Khanskaya stavka(N),Tajikistan:KhozretishiRange[Gurjeva] (O,P), Kazakhstan:Trans-Ili Alatau,gorge Levyy Talgar(Q);R,S, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype (R), Tajikistan: Khozretishi Range, 75 km NE Muminabad (S); T, U, C. niveimarginatus Scott, 1874, Russia: Kyzyl (T), Mongolia: Stariy Tsakhir (U); V, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD40E7AFF53FEFCFA99FC1A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927033/files/figure.png	https://doi.org/10.5281/zenodo.14927033	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD40E7AFF53FEFCFA99FC1A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927039/files/figure.png	https://doi.org/10.5281/zenodo.14927039	Figure 11. Female terminalia in Canthophorus spp. A, B, C. mixtus mixtus Asanova, 1964, Ukraine: Aleshki, paratype (A), Azerbaijan: Kars (B); C, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht, paratype; D, C. niveimarginatus Scott, 1874; Russia: Kamen’-Rybolov; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Gynatrial sac (A, C–E) and its posterior part (B) in dorsal view. Scale bar: 0.25 mm.	Figure 11. Female terminalia in Canthophorus spp. A, B, C. mixtus mixtus Asanova, 1964, Ukraine: Aleshki, paratype (A), Azerbaijan: Kars (B); C, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht, paratype; D, C. niveimarginatus Scott, 1874; Russia: Kamen’-Rybolov; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Gynatrial sac (A, C–E) and its posterior part (B) in dorsal view. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD40E7BFC13FB8FFE54FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927023/files/figure.png	https://doi.org/10.5281/zenodo.14927023	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD40E7BFC13FB8FFE54FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927029/files/figure.png	https://doi.org/10.5281/zenodo.14927029	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	Figure 6. Ventral lobes of conjunctiva in Canthophorus spp. A–C, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; D–I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype (D–F), Russia: Tuapse (G–I); J–M, C. melanopterus (Herrich- Schaeffer, 1835), Ukraine: Yalta (J, K), Armenia: Tigranashen (L, M); N, O, C. impressus (Horváth, 1880), Ukraine: Velikiy Anadol; P, Q, C. wagneri wagneri Asanova, 1964, Russia: Akhty; R, S, C. wagneri hyrcanicus n. ssp., Turkmenistan: gorge Khozly; T, U, C. mixtus Asanova, 1964, Kazakhstan: Dzhanibek; V, W, C. niveimarginatus Scott, 1874, Russia: Kyzyl; X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Ventral view (A, D, G, J, L, N, P, R, T, V, X), lateral view (B, E, H, K, M, O, Q, S, U, W, Y), and anterior view (C, F, I). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD40E7BFC13FB8FFE54FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927033/files/figure.png	https://doi.org/10.5281/zenodo.14927033	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	Figure 8. Female terminalia in Canthophorus spp. A–G, C. melanopterus (Herrich-Schaeffer, 1835), Ukraine: Alma Riv. (A), Azerbaijan: Ganja (B), Ukraine: Kamenets-Podol’skiy (C), Armenia: Dzhervezh (D), Ukraine: Khadzhibeyskiy liman (E), Morocco: Aourir (F); Egypt: “Aegipt.” (G); H, C. dubius dubius (Scopoli, 1763), France: Forêt des Morières; I, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; J, K, C. impressus impressus (Horváth, 1880), Ukraine: “yayla Besh-tekme” (J), Russia: Beshtau Mt. (K); L, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; M, N, C. wagneri wagneri Asanova, 1964, Russia: Anapa (M), Armenia: Sevan Lake (N); O–S, C. mixtus mixtus Asanova, 1964, Ukraine: Kremenets Mt. nr Izyum (O), Armenia: Spitak (P), Ukraine: Aleshki, paratype (Q), Ukraine: Nyrkovo – Vyemka, paratype (R), Azerbaijan: Kars (S); T, U, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht [paratypes]; V, W, C. niveimarginatus Scott, 1874; Russia: Kosh-Agach (V), Russia: Kamen’-Rybolov (W); X, Y, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek (X), Kyrgyzstan: Berk-Suu Riv (Y). External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A); gynatrial sac, dorsal view (B); shape of right first ramus (C–Y). a.g – anterior part of gynatrial sac; d.d – distal part of spermathecal duct; f.r – first rami; g.c – gynatrial cone; gn – gonangulum; gp.I – gonapophyses I; gp.II – gonapophyses II; gx.II – gonocoxites II; m.d – median part of spermathecal duct; p.d – proximal part of spermathecal duct; p.g – posterior part of gynatrial sac; p.m – posterior margin of gynatrial sac; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; tr – triangulum. Scale bars: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD40E7BFC13FB8FFE54FBC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927039/files/figure.png	https://doi.org/10.5281/zenodo.14927039	Figure 11. Female terminalia in Canthophorus spp. A, B, C. mixtus mixtus Asanova, 1964, Ukraine: Aleshki, paratype (A), Azerbaijan: Kars (B); C, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht, paratype; D, C. niveimarginatus Scott, 1874; Russia: Kamen’-Rybolov; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Gynatrial sac (A, C–E) and its posterior part (B) in dorsal view. Scale bar: 0.25 mm.	Figure 11. Female terminalia in Canthophorus spp. A, B, C. mixtus mixtus Asanova, 1964, Ukraine: Aleshki, paratype (A), Azerbaijan: Kars (B); C, C. mixtus hissaricus Asanova, 1964; n. stat., Tajikistan: urochishche Ruydasht, paratype; D, C. niveimarginatus Scott, 1874; Russia: Kamen’-Rybolov; E, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek. Gynatrial sac (A, C–E) and its posterior part (B) in dorsal view. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD50E7BFC26FF7FFA2CFD13.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927021/files/figure.png	https://doi.org/10.5281/zenodo.14927021	Figure 2. Pygophores in Canthophorus and Adomerus spp. A, B, C. melanopterus (Herrich-Schaeffer, 1835); C, D, A. aeneus (Walker, 1867) n. comb.; E, F, A. maculipes (Mulsant & Rey, 1852) n. comb. Ventral view (A, C, E) and posterodorsal view (B, D, F). In A, C, E, subgenital plate is shown in dotted lines. d.inf – dorsal infolding; l.r – lateral ridge of subgenital plate; l. inf – lateral infolding; p.r – posterior ridge of subgenital plate; v.inf – ventral infolding. Scale bar: 0.25 mm.	Figure 2. Pygophores in Canthophorus and Adomerus spp. A, B, C. melanopterus (Herrich-Schaeffer, 1835); C, D, A. aeneus (Walker, 1867) n. comb.; E, F, A. maculipes (Mulsant & Rey, 1852) n. comb. Ventral view (A, C, E) and posterodorsal view (B, D, F). In A, C, E, subgenital plate is shown in dotted lines. d.inf – dorsal infolding; l.r – lateral ridge of subgenital plate; l. inf – lateral infolding; p.r – posterior ridge of subgenital plate; v.inf – ventral infolding. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD50E7BFC26FF7FFA2CFD13.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927023/files/figure.png	https://doi.org/10.5281/zenodo.14927023	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD50E7BFC26FF7FFA2CFD13.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927053/files/figure.png	https://doi.org/10.5281/zenodo.14927053	Figure 18. Aedeagus of Adomerus aeneus (Walker, 1867) n. comb. A, lateral; B, dorsal; and C, ventral views. d.l – dorsal lobes of conjunctiva; da.l – dorsoapical lobe; dl.l – dorsolateral lobes; vl.l – ventrolateral lobes; vs – vesica. Scale bar: 0.25 mm.	Figure 18. Aedeagus of Adomerus aeneus (Walker, 1867) n. comb. A, lateral; B, dorsal; and C, ventral views. d.l – dorsal lobes of conjunctiva; da.l – dorsoapical lobe; dl.l – dorsolateral lobes; vl.l – ventrolateral lobes; vs – vesica. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD50E7BFC26FF7FFA2CFD13.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927057/files/figure.png	https://doi.org/10.5281/zenodo.14927057	Figure 20. Female terminalia in Adomerus spp. A, B, A. aeneus (Walker, 1867) n. comb.; C, D, A. maculipes (Mulsant & Rey, 1852) n. comb. External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A, C); gynatrial sac, dorsal view (B, D). a.p – anterior pouches of gynatrium; d.f – dividing fold in cavity of anterior pouch; f.r – first rami; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; s.r – second rami; tr – torus. Scale bar: 0.25 mm.	Figure 20. Female terminalia in Adomerus spp. A, B, A. aeneus (Walker, 1867) n. comb.; C, D, A. maculipes (Mulsant & Rey, 1852) n. comb. External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A, C); gynatrial sac, dorsal view (B, D). a.p – anterior pouches of gynatrium; d.f – dividing fold in cavity of anterior pouch; f.r – first rami; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; s.r – second rami; tr – torus. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD50E7BFCE6FC81FB6DFACE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927021/files/figure.png	https://doi.org/10.5281/zenodo.14927021	Figure 2. Pygophores in Canthophorus and Adomerus spp. A, B, C. melanopterus (Herrich-Schaeffer, 1835); C, D, A. aeneus (Walker, 1867) n. comb.; E, F, A. maculipes (Mulsant & Rey, 1852) n. comb. Ventral view (A, C, E) and posterodorsal view (B, D, F). In A, C, E, subgenital plate is shown in dotted lines. d.inf – dorsal infolding; l.r – lateral ridge of subgenital plate; l. inf – lateral infolding; p.r – posterior ridge of subgenital plate; v.inf – ventral infolding. Scale bar: 0.25 mm.	Figure 2. Pygophores in Canthophorus and Adomerus spp. A, B, C. melanopterus (Herrich-Schaeffer, 1835); C, D, A. aeneus (Walker, 1867) n. comb.; E, F, A. maculipes (Mulsant & Rey, 1852) n. comb. Ventral view (A, C, E) and posterodorsal view (B, D, F). In A, C, E, subgenital plate is shown in dotted lines. d.inf – dorsal infolding; l.r – lateral ridge of subgenital plate; l. inf – lateral infolding; p.r – posterior ridge of subgenital plate; v.inf – ventral infolding. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD50E7BFCE6FC81FB6DFACE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927023/files/figure.png	https://doi.org/10.5281/zenodo.14927023	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	Figure 3. Right parameres in Canthophorus and Adomerus spp. A, B, C. dubius dubius (Scopoli, 1763), France: Seine-et-Marne; C, D, C. dubius sanigarum n. ssp., Russia: Krasnaya Polyana, paratype; E, C. melanopterus (Herrich-Schaeffer, 1835), Russia: Krasnaya Polyana; F, C. impressus impressus (Horváth, 1880), Ukraine: Demir-kapu; G, C. impressus hajastanicus n. ssp., Armenia: Gekhmakhan Mt., paratype; H, C. wagneri Asanova, 1964, Russia: Makhachkala; I, C. mixtus mixtus Asanova, 1964, Armenia: Kipchakh; J, C. mixtus mixtus Asanova, 1964, Armenia: Gyumri; K, C. mixtus hissaricus Asanova, 1964 n. stat., Tajikistan: urochishche Ruydasht, paratype; L, C. niveimarginatus Scott, 1874, Russia: Kyzyl; M, C. coeruleus (Reuter, 1902), Kyrgyzstan: urochishche Ak-terek; N, O, A. aeneus (Walker, 1867) n. comb.; P, Q, A. maculipes (Mulsant & Rey, 1852) n. comb. Outer-ventral view (A, C, E, F–M), inner view (N, P), and dorsal view (B, D, O, Q). Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD50E7BFCE6FC81FB6DFACE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927055/files/figure.png	https://doi.org/10.5281/zenodo.14927055	Figure 19. Aedeagus of Adomerus maculipes (Mulsant & Rey, 1852) n. comb., A, lateral; B, dorsal; and C, ventral views. d.l – dorsal lobes of conjunctiva; da.l – dorsoapical lobe; dl.l – dorsolateral lobes; s.g – secondary gonopore; vl.l – ventrolateral lobes; vs – vesica. Scale bar: 0.25 mm.	Figure 19. Aedeagus of Adomerus maculipes (Mulsant & Rey, 1852) n. comb., A, lateral; B, dorsal; and C, ventral views. d.l – dorsal lobes of conjunctiva; da.l – dorsoapical lobe; dl.l – dorsolateral lobes; s.g – secondary gonopore; vl.l – ventrolateral lobes; vs – vesica. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
385C87B6FFD50E7BFCE6FC81FB6DFACE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14927057/files/figure.png	https://doi.org/10.5281/zenodo.14927057	Figure 20. Female terminalia in Adomerus spp. A, B, A. aeneus (Walker, 1867) n. comb.; C, D, A. maculipes (Mulsant & Rey, 1852) n. comb. External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A, C); gynatrial sac, dorsal view (B, D). a.p – anterior pouches of gynatrium; d.f – dividing fold in cavity of anterior pouch; f.r – first rami; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; s.r – second rami; tr – torus. Scale bar: 0.25 mm.	Figure 20. Female terminalia in Adomerus spp. A, B, A. aeneus (Walker, 1867) n. comb.; C, D, A. maculipes (Mulsant & Rey, 1852) n. comb. External terminalia (genital plates) and vestibulum of gynatrium, ventral view (A, C); gynatrial sac, dorsal view (B, D). a.p – anterior pouches of gynatrium; d.f – dividing fold in cavity of anterior pouch; f.r – first rami; p.p – posterior pouches of gynatrial sac; r.s – ring sclerites; s.r – second rami; tr – torus. Scale bar: 0.25 mm.	2018-05-31	Gapon, Dmitry A.		Zenodo	biologists	Gapon, Dmitry A.			
