taxonID	type	description	language	source
3B0087B31444FF8CFF3AFA21FF21428F.taxon	materials_examined	Source of material. All instar larvae studied were associated with adults collected at the following locality: Argentina, Tucumán Province. El Cochuna Provincial Park, 3 / 9 - XII- 2007, roadside pond; M. C. Michat leg. The identification is firm as C. alternatus is the only Copelatus species known from that locality.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31444FF8CFF3AFA21FF21428F.taxon	diagnosis	Diagnosis (instar III). The third instar of Copelatus alternatus can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 0.90 mm; L 3 <2.00 mm; U <0.40 mm; head capsule rounded, slightly constricted at level of occipital region (Fig. 15) with scale-like sculpticels present over frontoclypeus and parietale; anterior margin of frontoclypeus narrowly convex, extending mesally at about level of adnasalia (Fig. 15); adnasalia margined with a mix of bluntly rounded and truncated teeth (Fig. 63); MP 2 / MP 1> 1.50; GA / MP 1> 1.50; 1.30 <LP 2 / LP 1 <1.90; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with less than 3 PV and 4 AV secondary setae; metafemur with less than 3 AV secondary setae; urogomphus composed of one urogomphomere; U / HW <0.40; U / LAS <0.50; Neotropical.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31444FF8CFF3AFA21FF21428F.taxon	description	Description, instar I (Figs 1 – 14) Body: Measurements and ratios aimed to characterize body shape are shown in Table 2. Head (Figs 1 – 9): HL = 0.45 – 0.47 mm; Head capsule rounded, lacking a clear constriction at level of occiput; occipital suture absent; anterior margin of frontoclypeus lightly convex, extending mesally at about level of adnasalia; A / MP = 1.48 – 1.60; MP / LP = 1.70 – 1.79; MP 2 / MP 1 = 1.96 – 2.17; GA / MP 1 = 2.50 – 3.04; LP 2 / LP 1 = 2.27 – 2.50; MNL / MNW = 2.75 – 2.88. Thorax (Figs 10 – 11): L 3 = 0.83 – 0.88 mm. Abdomen (Figs 12 – 14): LAS = 0.30 – 0.32 mm; LAS subcylindrical, not constricted posteriorly at point of insertion of urogomphi. Urogomphus: U = 0.18 – 0.19 mm, composed of one urogomphomere; U / HW = 0.36 – 0.39; U / LAS = 0.56 – 0.65. Chaetotaxy (Figs 1 – 14): Position and number of additional setae on legs are shown in Table 3 and Figs 10 – 11. Description, instar II. As first-instar larva except as follows: Body: Measurements and ratios aimed to characterize body shape are shown in Table 4. Head: HL = 0.65 mm; anterior margin of frontoclypeus more narrowly convex; scale-like sculpticels welldeveloped over frontoclypeus and parietale; adnasalia margined with short bluntly rounded and truncated teeth (see Fig. 63); occipital suture present; A / MP = 1.59 – 1.64; MP / LP = 1.64 – 1.76; MP 2 / MP 1 = 2.00 – 2.14; GA / MP 1 = 2.38 – 2.57; LP 2 / LP 1 = 1.78 – 1.89; MNL / MNW = 2.60 – 2.78. Thorax: L 3 = 1.32 – 1.34 mm. Abdomen: LAS = 0.43 – 0.44 mm; LAS subcylindrical to subconical, less abruptly converging posteriorly at point of insertion of urogomphi. Urogomphus: U = 0.23 – 0.24 mm, composed of one urogomphomere; U / HW = 0.31 – 0.32; U / LAS = 0.54 – 0.56. Chaetotaxy: Parietale with 3 – 5 temporal spines; position and number of secondary setae on legs are shown in Table 5. Description, instar III (Figs 15 – 16, 63). As second-instar larva except as follows: Body: Measurements and ratios aimed to characterize body shape are shown in Table 6. Head (Figs 15, 63): HL = 0.88 – 0.89 mm; A / MP = 1.52 – 1.56; MP / LP = 1.77 – 1.90; MP 2 / MP 1 = 1.58 – 1.75; GA / MP 1 = 1.83 – 2.00; LP 2 / LP 1 = 1.42 – 1.58; MNL / MNW = 2.43 – 2.83. Thorax: L 3 = 1.74 – 1.81 mm. Abdomen (Fig. 16): LAS = 0.68 – 0.73 mm; U = 0.27 – 0.30 mm; U / HW = 0.27 – 0.32; U / LAS = 0.37 – 0.44. Chaetotaxy: Parietale with 3 – 7 temporal spines. Position and number of secondary setae on legs are shown in Table 7.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31448FF97FF3AFA8DFEDD442B.taxon	materials_examined	Source of material. The two larvae studied (one instar II and one instar III) were associated with adults collected at the following locality: USA. Florida, Orlando. Junction International Drive & Hwy 523, 26 – 28. IX. 2016. 28 ° 53.83 ’ N 81 ° 37.92 ’ W; Y. Alarie leg. The identification is firm as C. caelatipennis princeps is the only Copelatus species known from that locality.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31448FF97FF3AFA8DFEDD442B.taxon	diagnosis	Diagnosis (instar III). The third instar of Copelatus c. princeps can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 0.80 mm; L 3 <2.00 mm; U <0.40 mm; head capsule rounded to subquadrate, strongly constricted at level of occipital region (Fig. 17), scale-like sculpticels present over frontoclypeus and parietale; anterior margin of frontoclypeus narrowly convex, extending mesally well beyond level of adnasalia (Fig. 17); adnasalia margined with short bluntly rounded and truncated teeth (Fig. 63); MP 2 / MP 1> 1.50; GA / MP 1> 1.50; 1.30 <LP 2 / LP 1 <1.90; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with less than 3 PV and 4 AV secondary setae; metafemur with less than 3 AV secondary setae; urogomphus composed of one urogomphomere; U / HW = 0.40; U / LAS = 0.50; Nearctic and Neotropical.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31448FF97FF3AFA8DFEDD442B.taxon	description	Description, instar II Body: Measurements and ratios aimed to characterize body shape are shown in Table 4. Head: Head capsule rounded to subquadrate, strongly constricted at level of occiput; anterior margin of frontoclypeus narrowly convex, extending well beyond level of adnasalia; adnasalia margined with short bluntly rounded and truncated teeth (see Fig. 63); scale-like sculpticels well-developed over frontoclypeus and parietale; HL = 0.58 mm; A / MP = 1.41; MP / LP = 1.85; MP 2 / MP 1 = 1.73; GA / MP 1 = 2.00; LP 2 / LP 1 = 1.73; MNL / MNW = 2.72. Thorax: L 3 = 1.05 mm. Abdomen: LAS = 0.39 mm; LAS subconical, not constricted posteriorly at point of insertion of urogomphi. Urogomphus: U = 0.22 mm, composed of one urogomphomere; U / HW = 0.38; U / LAS = 0.58. Chaetotaxy: Parietale with 7 temporal spines; position and number of secondary setae on legs are shown in Table 5. Description, instar III (Figs 17 – 18, 63). As second-instar larva except as follows: Body: Measurements and ratios aimed to characterize body shape are shown in Table 6. Head (Figs 18, 63): HL = 0.81 mm; A / MP = 1.50; MP / LP = 1.98; MP 2 / MP 1 = 1.67; GA / MP 1 = 2.00; LP 2 / LP 1 = 1.51; MNL / MNW = 2.58. Thorax: L 3 = 1.58 mm. Abdomen (Fig. 18): LAS = 0.64 mm; U = 0.34 mm; U / HW = 0.43; U / LAS = 0.52. Chaetotaxy: Parietale with 10 – 12 temporal spines; position and number of secondary setae on legs are shown in Table 7.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31453FF97FF3AFD29FF21403F.taxon	materials_examined	Source of material. The five instar III larvae studied were associated with adults collected at the following locality: Canada. Québec, Trois-Rivières. Ephemeral pond in red maple forest at Chemin des Oblats, 06. VIII. 1986, 08. VIII. 1986, 12. VIII. 1986; Y. Alarie leg. The identification is firm as C. glyphicus is the only Copelatus species known from that region.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31453FF97FF3AFD29FF21403F.taxon	diagnosis	Diagnosis (instar III). The third instar of Copelatus glyphicus can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 0.80 – 0.90 mm; L 3 <2.00 mm; U <0.40 mm; head capsule rounded, strongly constricted at level of occipital region (Fig. 19), scale-like sculpticels present over frontoclypeus and parietale; anterior margin of frontoclypeus broadly convex, extending mesally beyond level of adnasalia (Fig. 19); adnasalia margined with short bluntly rounded and truncated teeth (Fig. 63); MP 2 / MP 1> 1.50; GA / MP 1> 1.50; 1.30 <LP 2 / LP 1 <1.90; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with less than 3 PV and 4 AV secondary setae; metafemur with less than 3 AV secondary setae; urogomphus composed of one urogomphomere; U / HW <0.40; U / LAS <0.50; Nearctic.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31453FF97FF3AFD29FF21403F.taxon	description	Description, instar III (Figs 19 – 20, 63) Body: Measurements and ratios aimed to characterize body shape are shown in Table 6. Head (Figs 19, 63): Head capsule rounded, strongly constricted at level of occiput; anterior margin of frontoclypeus broadly convex, extending beyond level of adnasalia; scale-like sculpticels well-developed both on frontoclypeus and parietale; adnasalia margined with short bluntly rounded and truncated teeth (Fig. 63); HL = 0.81 – 0.94 mm; A / MP = 1.34 – 1.38; MP / LP = 2.12 – 2.17; MP 2 / MP 1 = 1.50 – 1.73; GA / MP 1 = 1.79 – 1.88; LP 2 / LP 1 = 1.40 – 1.56; MNL / MNW = 2.85 – 3.04. Thorax: L 3 = 1.60 – 1.74 mm. Abdomen (Fig. 20): LAS = 0.65 – 0.74 mm; LAS subcylindrical, not constricted posteriorly at point of insertion of urogomphi. Urogomphus: U = 0.28 – 0.32 mm, composed of one urogomphomere; U / HW = 0.31 – 0.33; U / LAS = 0.42 – 0.44. Chaetotaxy: Parietale with 6 – 9 temporal spines; position and number of secondary setae on legs are shown in Table 7.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31453FF94FF3AF95AFE7444BB.taxon	materials_examined	Source of material. The four instar III larvae studied were associated with adults collected at the following locality: Taiwan. Taipei, Beitou, 24. VIII. 1994; L. J. Wang leg. The identification is firm as C. japonicus is the only Copelatus species collected at this locality.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31453FF94FF3AF95AFE7444BB.taxon	diagnosis	Diagnosis (instar III). The third instar of Copelatus japonicus can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 0.90 mm; L 3 <2.00 mm; U <0.40 mm; head capsule rounded, strongly constricted at level of occipital region (Fig. 21), scale-like sculpticels present over frontoclypeus and parietale; anterior margin of frontoclypeus narrowly convex, extending mesally at about level of adnasalia (Fig. 21); adnasalia margined with bluntly rounded and truncated teeth (Fig. 63); MP 2 / MP 1> 1.50; GA / MP 1> 1.50; 1.30 <LP 2 / LP 1 <1.90; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with less than 3 PV and 4 AV secondary setae; metafemur with less than 3 AV secondary setae; urogomphus composed of two urogomphomeres; U / HW <0.40; U / LAS <0.50; Palaearctic.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31453FF94FF3AF95AFE7444BB.taxon	description	Description, instar III (Figs 21 – 24, 63) Body: Measurements and ratios aimed to characterize body shape are shown in Table 6. Head (Figs 21, 63): Head capsule rounded, strongly constricted at level of occiput; anterior margin of frontoclypeus narrowly convex, extending mesally at about level of adnasalia; scale-like sculpticels well-developed over frontoclypeus and parietale; adnasalia margined with bluntly rounded and truncated teeth (Fig. 63); HL = 0.86 – 0.90 mm; A / MP = 1.51 – 1.60; MP / LP = 1.77 – 1.88; MP 2 / MP 1 = 1.57 – 1.82; GA / MP 1 = 1.67 – 1.85; LP 2 / LP 1 = 1.28 – 1.61; MNL / MNW = 2.50 – 2.72. Thorax (Figs 23 – 24): L 3 = 1.84 – 1.92 mm. Abdomen (Fig. 22): LAS = 0.66 – 0.72 mm; LAS subconical, not constricted posteriorly at point of insertion of urogomphi. Urogomphus, U = 0.28 – 0.30 mm, composed of two urogomphomeres; U / HW = 0.28 – 0.30; U / LAS = 0.42 – 0.43. Chaetotaxy: Parietale with 4 – 6 temporal spines; position and number of secondary setae on legs are shown in Figs 23 – 24 and Table 7.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31450FF94FF3AFCACFF2141BD.taxon	materials_examined	Source of material. The five instar II and five instar III larvae studied were reared ex ovo from adults collected at the following locality: Japan. Tarumai-shitsugen, Yonaguni-cho Yonaguni-jima Island, Okinawa Prefecture, 24 ° 26 ’ 31.2 ’’ N 122 ° 58 ’ 06.2 ’’ E, 17. VI. 2018; K. Watanabe leg.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31450FF94FF3AFCACFF2141BD.taxon	diagnosis	Diagnosis (instar III). The third instar of Copelatus masculinus can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 1.00 mm; L 3 <2.00 mm; U <0.40 mm; head capsule rounded, slightly constricted at level of occipital region (Fig. 25), scale-like sculpticels present over frontoclypeus and parietale; anterior margin of frontoclypeus narrowly convex, extending mesally slightly beyond adnasalia (Fig. 25); adnasalia margined with short bluntly rounded teeth (Fig. 64); MP 2 / MP 1> 1.50; GA / MP 1> 1.50; LP 2 / LP 1 <1.10; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with less than 3 PV and 4 AV secondary setae; metafemur with less than 3 AV secondary setae; urogomphus composed of two urogomphomeres; U / HW <0.40; U / LAS <0.50; Oriental and Palaearctic.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31450FF94FF3AFCACFF2141BD.taxon	description	Description, instar II Body: Measurements and ratios aimed to characterize body shape are shown in Table 4. Head: Head capsule rounded, slightly constricted at level of occiput; anterior margin of frontoclypeus narrowly convex, extending mesally slightly beyond level of adnasalia; scale-like sculpticels well-developed over frontoclypeus and parietale; adnasalia margined with short bluntly rounded teeth (see Fig. 64); HL = 0.68 – 0.72 mm; A / MP = 1.33 – 1.36; MP / LP = 1.99 – 2.07; MP 2 / MP 1 = 1.62 – 1.72; GA / MP 1 = 1.92 – 2.13; LP 2 / LP 1 = 1.33 – 1.55; MNL / MNW = 2.83 – 3.33. Thorax: L 3 = 1.32 – 1.42 mm. Abdomen: LAS = 0.46 – 0.54 mm; LAS subcylindrical to subconical, not constricted posteriorly at point of insertion of urogomphi. Urogomphus: U = 0.23 – 0.25 mm, composed of two urogomphomeres; U / HW = 0.32 – 0.34; U / LAS = 0.46 – 0.51. Chaetotaxy: Parietale with 5 – 7 temporal spines; position and number of secondary setae on legs are shown in Table 5. Description, instar III (Figs 25 – 27, 64). As second-instar larva except as follows: Body: Measurements and ratios aimed to characterize body shape are shown in Table 6. Head (Figs 25, 27, 64): HL = 0.97 – 0.99 mm; A / MP = 1.34 – 1.37; MP / LP = 2.04 – 2.14; MP 2 / MP 1 = 1.51 – 1.65; GA / MP 1 = 1.61 – 1.85; LP 2 / LP 1 = 1.06 – 1.12; MNL / MNW = 2.84 – 2.92. Thorax: L 3 = 1.90 – 2.00 mm. Abdomen (Fig. 26): LAS = 0.75 – 0.77 mm; U = 0.31 – 0.32 mm, U / HW = 0.29 – 0.31; U / LAS = 0.40 – 0.43. Chaetotaxy: Parietale with 7 – 9 temporal spines; position and number of secondary setae on legs are shown in Table 7.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B3145AFF9EFF3AFF0CFEDD4344.taxon	materials_examined	Source of material. The two instar II and five instar III larvae studied were reared ex ovo from adults collected at the following locality: Japan. Kasamatshu-cho, Mudoji, Gifu Prefecture, 4. V. 2018. Y. Yoshimura, leg.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B3145AFF9EFF3AFF0CFEDD4344.taxon	diagnosis	Diagnosis (instar III). The third instar of Copelatus nakamurai can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 0.80 mm; L 3 <2.00 mm; U <0.40 mm; head capsule subquadrate, strongly constricted at level of occipital region (Fig. 28), scale-like sculpticels lacking over frontoclypeus, present over parietale; anterior margin of frontoclypeus narrowly convex, extending mesally well beyond adnasalia (Fig. 28); adnasalia margined with elongate acute teeth (Fig. 64); MP 2 / MP 1> 1.50; GA / MP 1> 1.50; 1.30 <LP 2 / LP 1 <1.90; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with less than 3 PV and 4 AV secondary setae; metafemur with less than 3 AV secondary setae; urogomphus composed of one urogomphomere; U / HW <0.40; U / LAS <0.50; Palaearctic.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B3145AFF9EFF3AFF0CFEDD4344.taxon	description	Description, instar II Body: Measurements and ratios aimed to characterize body shape are shown in Table 4. Head: Head capsule subquadrate, strongly constricted at level of occiput; anterior margin of frontoclypeus narrowly convex, extending well beyond level of adnasalia; scale-like sculpticels lacking over frontoclypeus, welldeveloped over parietale; adnasalia margined with elongate acute teeth (see Fig. 64); HL = 0.61 mm; A / MP = 1.49; MP / LP = 2.06; MP 2 / MP 1 = 1.60; GA / MP 1 = 1.91; LP 2 / LP 1 = 2.24; MNL / MNW = 3.11. Thorax: L 3 = 1.23 mm. Abdomen: LAS = 0.45 mm; LAS subcylindrical, slightly constricted posteriorly at point of insertion of urogomphi. Urogomphus: U = 0.26 mm, composed of one urogomphomere; U / HW = 0.41; U / LAS = 0.58. Chaetotaxy: Parietale with 8 temporal spines; position and number of secondary setae on legs are shown in Table 5. Description, instar III (Figs 28 – 29, 64). As second-instar larva except as follows: Body: Measurements and ratios aimed to characterize body shape are shown in Table 6. Head (Figs 28, 64): HL = 0.81 – 0.84 mm; A / MP = 1.42 – 1.46; MP / LP = 2.02 – 2.25; MP 2 / MP 1 = 1.51 – 1.74; GA / MP 1 = 1.75 – 1.99; LP 2 / LP 1 = 1.55 – 1.78; MNL / MNW = 2.85 – 2.98. Thorax: L 3 = 1.66 – 1.72 mm. Abdomen (Fig 29): LAS = 0.66 – 0.69 mm; Urogomphus, U = 0.31 – 0.33 mm, composed of one urogomphomere; U / HW = 0.35 – 0.38; U / LAS = 0.45 – 0.49. Chaetotaxy: Parietale with 9 – 10 temporal spines; position and number of secondary setae on legs are shown in Table 7.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B3145AFF9CFF3AFAC2FE74442B.taxon	materials_examined	Source of material. The five instar II and five instar III larvae studied were reared ex ovo from adults collected at the following locality: Japan. Mantaburu, Yonaguni-cho, Yonaguni-jima Island, Okinawa Prefecture. 24 ° 27 ’ 04.1 ” N 122 ° 57 ’ 36.8 ” E, 13 VI. 2019; K. Watanabe leg.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B3145AFF9CFF3AFAC2FE74442B.taxon	diagnosis	Diagnosis (instar III). The third instar of Copelatus oblitus can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 0.80 mm; L 3 <2.00 mm; U <0.30 mm; head capsule ovate, not constricted at level of occipital region (Fig. 30), scale-like sculpticels present over frontoclypeus and parietale; anterior margin of frontoclypeus narrowly convex, extending mesally well below level of adnasalia (Fig. 30); adnasalia margined with short bluntly rounded teeth (Fig. 64); MP 2 / MP 1> 1.50; GA / MP 1> 1.50; 1.30 <LP 2 / LP 1 <1.90; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with less than 3 PV and 4 AV secondary setae; metafemur with less than 3 AV secondary setae; urogomphus composed of two urogomphomeres; U / HW <0.40; U / LAS <0.50; Oriental and Palaearctic.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B3145AFF9CFF3AFAC2FE74442B.taxon	description	Description, instar II Body: Measurements and ratios aimed to characterize body shape are shown in Table 4. Head: Head capsule ovate, not constricted at level of occiput; anterior margin of frontoclypeus narrowly convex, extending mesally below level of adnasalia; scale-like sculpticels present over frontoclypeus and parietale; adnasalia margined with short bluntly rounded teeth (see Fig. 64); HL = 0.55 – 0.58 mm; A / MP = 1.39 – 1.51; MP / LP = 1.70 – 1.82; MP 2 / MP 1 = 1.67 – 1.95; GA / MP 1 = 1.85 – 2.32; LP 2 / LP 1 = 1.56 – 1.89; MNL / MNW = 2.41 – 2.61. Thorax: L 3 = 1.10 – 1.13 mm. Abdomen: LAS = 0.37 – 0.39 mm; LAS subconical, not constricted posteriorly at point of insertion of urogomphi. Urogomphus: U = 0.17 – 0.19 mm, composed of two urogomphomeres; U / HW = 0.26 – 0.29; U / LAS = 0.45 – 0.51. Chaetotaxy: Parietale with 3 – 5 temporal spines; position and number of secondary setae on legs are shown in Table 5. Description, instar III (Figs 30 – 33, 64). As second-instar larva except as follows: Body: Measurements and ratios aimed to characterize body shape are shown in Table 6. Head (Figs 30, 64): HL = 0.74 – 0.80 mm; A / MP = 1.50 – 1.60; MP / LP = 1.65 – 1.82; MP 2 / MP 1 = 1.44 – 1.67; GA / MP 1 = 1.59 – 1.83; LP 2 / LP 1 = 1.31 – 1.47; MNL / MNW = 2.40 – 2.60. Thorax (Figs 32 – 33): L 3 = 1.57 – 1.64 mm. Abdomen (Fig 31): LAS = 0.54 – 0.60 mm; U = 0.20 – 0.25 mm, U / HW = 0.22 – 0.27; U / LAS = 0.37 – 0.47. Chaetotaxy: Parietale with 3 – 4 temporal spines; position and number of secondary setae on legs are shown in Figs 32 – 33 and Table 7.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31458FFA5FF3AFD29FE7447C3.taxon	materials_examined	Source of material. All larvae studied were reared ex ovo from were adults collected at the following locality: Japan. Kyoto Prefecture. Uji-gawa, Fushimi-ku, Kyoto-shi, Kyoto Prefecture, 34 ° 55 ’ 08.8 ” N 135 ° 45 ’ 28.3 ” E, 6. XI. 2016; K. Watanabe leg.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31458FFA5FF3AFD29FE7447C3.taxon	diagnosis	Diagnosis (instar III). The third instar of Copelatus parallelus can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 0.80 mm; L 3 <2.00 mm; U = 0.40 mm; head capsule subquadrate, strongly constricted at level of occipital region (Fig. 48), scale-like sculpticels present over frontoclypeus and parietale; anterior margin of frontoclypeus broadly convex, extending mesally at about level of adnasalia (Fig. 48); adnasalia margined with strong acute teeth (Fig. 64); MP 2 / MP 1> 1.50; GA / MP 1> 1.50; 1.30 <LP 2 / LP 1 <1.90; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with less than 3 PV and 4 AV secondary setae; metafemur with less than 3 AV secondary setae; urogomphus composed of one urogomphomere; U / HW> 0.40; U / LAS> 0.60; Palaearctic.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31458FFA5FF3AFD29FE7447C3.taxon	description	Description, instar I (Figs 34 – 47) Body: Measurements and ratios aimed to characterize body shape are shown in Table 2. Head (Figs 34 – 42): HL = 0.43 mm; Head capsule subquadrate, strongly constricted at level of occiput; occipital suture absent; anterior margin of frontoclypeus broadly convex extending beyond level of adnasalia mesally; scale-like sculpticels well-developed over frontoclypeus and parietale; adnasalia margined with strong acute teeth; occipital carina absent; A / MP = 1.27; MP / LP = 2.30; MP 2 / MP 1 = 2.17; GA / MP 1 = 2.60; LP 2 / LP 1 = 2.74; MNL / MNW = 3.66. Thorax (Figs 43 – 44): L 3 = 0.80 mm. Abdomen (Figs 45 – 47): LAS = 0.32 mm; LAS subconical, slightly constricted posteriorly at point of insertion of urogomphi. Urogomphus: U = 0.17 mm, composed of one urogomphomere; U / HW = 0.36; U / LAS = 0.52. Chaetotaxy (Figs 34 – 47): Position and number of additional setae on legs are shown in Table 3 and Figs 43 – 44. Description, instar II. As for first-instar larva except as follows: Body: Measurements and ratios aimed to characterize body shape are shown in Table 4. Head: Anterior margin of frontoclypeus extending mesally at about same level of adnasalia; adnasalia margined with strong acute teeth (see Fig. 64); occipital suture present; HL = 0.56 mm; A / MP = 1.39; MP / LP = 2.24; MP 2 / MP 1 = 1.86; GA / MP 1 = 2.44; LP 2 / LP 1 = 2.12; MNL / MNW = 3.55. Thorax: L 3 = 1.07 mm. Abdomen: LAS = 0.41 mm. Urogomphus: U = 0.18 mm, composed of one urogomphomere; U / HW = 0.30; U / LAS = 0.45. Chaetotaxy: Parietale with 7 temporal spines; position and number of secondary setae on legs are shown in Table 5. Description, instar III (Figs 48 – 51, 64). As second-instar larva except as follows: Body: Measurements and ratios aimed to characterize body shape are shown in Table 6. Head (Figs 48, 64): HL = 0.78 mm; A / MP = 1.29; MP / LP = 2.38; MP 2 / MP 1 = 1.69; GA / MP 1 = 1.82; LP 2 / LP 1 = 1.67; MNL / MNW = 3.07. Thorax (Figs 50 – 51): L 3 = 1.47 mm. Abdomen (Fig. 49): LAS = 0.61 mm; U = 0.40 mm; U / HW = 0.50; U / LAS = 0.65. Chaetotaxy: Parietale with 7 temporal spines. Position and number of secondary setae on legs are shown in Figs 50 – 51 and Table 7.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31466FFA2FF3AFF0CFF2145A4.taxon	materials_examined	Source of material. The single instar III larvae studied was reared ex ovo from were adults collected at the following locality: Japan. Mantaburu, Yonaguni-cho, Yonaguni-jima Island, Okinawa Prefecture, 24 ° 27 ’ 04.1 ” N 122 ° 57 ’ 36.8 ” E, 13. VI. 2019; E. K. Watanabe leg.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31466FFA2FF3AFF0CFF2145A4.taxon	diagnosis	Diagnosis (instar III). The third instar of Copelatus tenebrosus can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 0.80 mm; L 3 <2.00 mm; U <0.40 mm; head capsule rounded, slightly constricted at level of occipital region (Fig. 52), scale-like sculpticels present over frontoclypeus and parietale; anterior margin of frontoclypeus narrowly convex, extending mesally well beyond level of adnasalia (Fig. 52); adnasalia margined with mostly short bluntly rounded teeth (Fig. 65); MP 2 / MP 1> 1.50; GA / MP 1> 1.50; 1.30 <LP 2 / LP 1 <1.90; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with less than 3 PV and 4 AV secondary setae; metafemur with less than 3 AV secondary setae; urogomphus composed of two urogomphomeres; U / HW <0.40; U / LAS <0.50; Australia, Oriental, Palaearctic.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31466FFA2FF3AFF0CFF2145A4.taxon	description	Description, instar III (Figs 52 – 53, 65) Body: Measurements and ratios aimed to characterize body shape are shown in Table 6. Head (Figs 52, 65): Head capsule rounded, slightly constricted at level of occiput; anterior margin of frontoclypeus narrowly convex, extending mesally well beyond level of adnasalia; scale-like sculpticels present over frontoclypeus and parietale; adnasalia margined with mostly short bluntly rounded teeth (Fig. 65); HL = 0.84 mm; A / MP = 1.40; MP / LP = 2.03; MP 2 / MP 1 = 1.64; GA / MP 1 = 1.85; LP 2 / LP 1 = 1.43; MNL / MNW = 2.87. Thorax: L 3 = 1.73 mm. Abdomen (Fig. 53): LAS = 0.63 mm; LAS subconical, not constricted posteriorly at point of insertion of urogomphi. Urogomphus, U = 0.30 mm, composed of two urogomphomeres; U / HW = 0.33; U / LAS = 0.47. Chaetotaxy: Parietale with 5 – 6 temporal spines; position and number of secondary setae on legs are shown in Table 7.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31466FFA2FF3AFBA4FEDD41F0.taxon	materials_examined	Source of material. The three instar III larvae studied were collected in association with adults at the following locality: South Australia. Ten kilometers North of Forreston, 9. VIII. 1999; C. H. S. Watts leg.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31466FFA2FF3AFBA4FEDD41F0.taxon	diagnosis	Diagnosis (instar III). The third instar of Exocelina australiae can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 1.85 – 1.90 mm; L 3> 2.90 mm; U> 0.50 mm; head capsule subquadrate, strongly constricted at level of occipital region (Fig. 54), scale-like sculpticels present over frontoclypeus and parietale; anterior margin of frontoclypeus broadly convex, extending mesally at level of adnasalia (Fig. 54); adnasalia margined with mostly short bluntly rounded teeth (Fig. 65); MP 2 / MP 1> 1.50; GA / MP 1> 1.50; LP 2 / LP 1> 2.00; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with more than 5 PV and 5 AV secondary setae; metafemur with more than 4 AV secondary setae; urogomphus composed of one urogomphomere; U / HW <0.40; U / LAS <0.50; Australian.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31466FFA2FF3AFBA4FEDD41F0.taxon	description	Description, instar III (Figs 54 – 55, 65) Body: Measurements and ratios aimed to characterize body shape are shown in Table 6. Head (Figs 54, 65): Head capsule subquadrate, strongly constricted at level of occiput; anterior margin of frontoclypeus broadly convex, extending mesally at about level of adnasalia; scale-like sculpticels present over frontoclypeus and parietale; adnasalia margined with mostly short bluntly rounded teeth (Fig. 65); HL = 1.85 – 1.90 mm; A / MP = 1.37 – 1.41; MP / LP = 2.36 – 2.41; MP 2 / MP 1 = 1.56 – 1.67; GA / MP 1 = 1.53 – 1.73; LP 2 / LP 1 = 2.00 – 2.67; MNL / MNW = 2.72 – 2.96. Thorax: L 3 = 3.71 – 3.87 mm. Abdomen (Fig. 55): LAS = 1.33 – 1.63 mm; LAS subcylindrical, not constricted posteriorly at point of insertion of urogomphi. Urogomphus, U = 0.56 – 0.58 mm, composed of one urogomphomere; U / HW = 0.25 – 0.26; U / LAS = 0.35 – 0.44. Chaetotaxy: Parietale with 9 – 10 temporal spines; position and number of secondary setae on legs are shown in Table 7.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31468FFACFF3AFF0CFF214344.taxon	materials_examined	Source of material. The three instar II and three instar III larvae studied were collected in association with adults at the following locality: South Australia. One kilometer North of Forreston, 2. IX. 2000; C. H. S. Watts leg.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31468FFACFF3AFF0CFF214344.taxon	diagnosis	Diagnosis (instar III). The third instar of Exocelina ferruginea can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 1.10 – 1.20 mm; L 3 = 2.20 – 2.50 mm; U> 0.50 mm; head capsule subquadrate, strongly constricted at level of occipital region (Fig. 56), scale-like sculpticels present over frontoclypeus and parietale; anterior margin of frontoclypeus broadly convex, extending mesally at level of adnasalia (Fig. 54); adnasalia margined with short acute teeth (Fig. 65); MP 2 / MP 1> 1.50; GA / MP 1> 1.50; LP 2 / LP 1> 2.00; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with less than 3 PV and more than 5 AV secondary setae; metafemur with less than 3 AV secondary setae; urogomphus composed of one urogomphomere; U / HW> 0.40; U / LAS> 0.50; Australian.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31468FFACFF3AFF0CFF214344.taxon	description	Description, instar II Body: Measurements and ratios aimed to characterize body shape are shown in Table 4. Head: Head capsule subquadrate, strongly constricted at level of occiput; anterior margin of frontoclypeus broadly convex, extending mesally at level of adnasalia; scale-like sculpticels present over frontoclypeus and parietale; adnasalia margined with short acute teeth (see Fig. 65); HL = 0.85 – 0.90 mm; A / MP = 1.33 – 1.38; MP / LP = 2.52 – 3.00; MP 2 / MP 1 = 1.83; GA / MP 1 = 2.00 – 2.33; LP 2 / LP 1 = 3.17 – 3.80; MNL / MNW = 2.77 – 3.18. Thorax: L 3 = 1.70 – 1.84 mm. Abdomen: LAS = 0.54 – 0.63 mm; LAS subcylindrical, not constricted posteriorly at point of insertion of urogomphi. Urogomphus: U = 0.46 – 0.51 mm, composed of one urogomphomere; U / HW = 0.50 – 0.53; U / LAS = 0.74 – 0.96. Chaetotaxy: Parietale with 4 – 6 temporal spines; position and number of secondary setae on legs are shown in Table 5. Description, instar III (Figs 56 – 57, 65). As second-instar larva except as follows: Body: Measurements and ratios aimed to characterize body shape are shown in Table 6. Head (Figs 56, 65): HL = 1.08 – 1.23 mm; A / MP = 1.27 – 1.33; MP / LP = 2.79 – 3.27; MP 2 / MP 1 = 1.55 – 1.82; GA / MP 1 = 1.65 – 2.06; LP 2 / LP 1 = 2.22 – 2.50; MNL / MNW = 2.88 – 2.94. Thorax: L 3 = 2.19 – 2.44 mm. Abdomen (Fig. 57): LAS = 0.79 – 0.85 mm; U = 0.44 – 0.71 mm, U / HW = 0.38 – 0.57; U / LAS = 0.56 – 0.88. Chaetotaxy: Parietale with 4 – 7 temporal spines; position and number of secondary setae on legs are shown in Table 7.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31468FFA8FF3AFACFFE11472F.taxon	materials_examined	Source of material. The four instar III larvae were collected in association with adults at the following locality: Belarus, Vicebsk Region, area near upy Village, wetland of evinka River, Carex, moss. 12. VIII. 1998; H. Shaverdo leg.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31468FFA8FF3AFACFFE11472F.taxon	diagnosis	Diagnosis (instar III). The third instar of Liopterus haemorrhoidalis can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 1.40 mm; L 3> 2.90 mm; U> 0.50 mm; head capsule subquadrate, strongly constricted at level of occipital region (Fig. 58), scale-like sculpticels lacking over frontoclypeus and parietale; anterior margin of frontoclypeus broadly convex, extending mesally at level of adnasalia (Fig. 58); adnasalia margined with short bluntly rounded teeth (Fig. 65); MP 2 / MP 1 <1.30; GA / MP 1 <1.50; 1.30 <LP 2 / LP 1 <1.90; inner margin of stipes with a dorsal linear row of spinulae (Fig. 60); profemur with less than 3 PV and more than 5 AV secondary setae; metafemur with more than 4 AV secondary setae (Fig. 61); urogomphus composed of one urogomphomere; U / HW = 0.40; U / LAS> 0.50; Palaearctic.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31468FFA8FF3AFACFFE11472F.taxon	description	Description, instar III (Figs 58 – 62, 65) Body: Measurements and ratios aimed to characterize body shape are shown in Table 6. Head (Figs 58, 60, 65): Head capsule subquadrate, strongly constricted at level of occiput; anterior margin of frontoclypeus broadly convex, extending mesally at about level of adnasalia; scale-like sculpticels lacking over frontoclypeus and parietale; adnasalia margined with mostly short bluntly rounded teeth (Fig. 65); HL = 1.36 – 1.39 mm; A / MP = 1.16 – 1.27; MP / LP = 1.67 – 1.74; MP 2 / MP 1 = 1.15 – 1.28; GA / MP 1 = 1.27 – 1.49; LP 2 / LP 1 = 1.54 – 1.71; MNL / MNW = 2.63 – 2.71. Thorax (Figs 61 – 62): L 3 = 2.86 – 3.06 mm. Abdomen (Fig. 59): LAS = 1.09 – 1.17 mm; LAS subconical, constricted posteriorly at point of insertion of urogomphi. Urogomphus, U = 0.60 – 0.63 mm, composed of one urogomphomere; U / HW = 0.38 – 0.40; U / LAS = 0.54 – 0.57. Chaetotaxy: Parietale with 7 – 10 temporal spines; position and number of secondary setae on legs are shown in Figs 61 – 62 and Table 7.	en	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
