identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
3B0087B31444FF8CFF3AFA21FF21428F.text	3B0087B31444FF8CFF3AFA21FF21428F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copelatus alternatus Sharp 1882	<div><p>Copelatus alternatus Sharp, 1882</p><p>Source of material. All instar larvae studied were associated with adults collected at the following locality: Argentina, Tucumán Province. El Cochuna Provincial Park, 3/ 9-XII-2007, roadside pond; M. C. Michat leg. The identification is firm as C. alternatus is the only Copelatus species known from that locality.</p><p>Diagnosis (instar III). The third instar of Copelatus alternatus can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 0.90 mm; L3 &lt;2.00 mm; U &lt;0.40 mm; head capsule rounded, slightly constricted at level of occipital region (Fig. 15) with scale-like sculpticels present over frontoclypeus and parietale; anterior margin of frontoclypeus narrowly convex, extending mesally at about level of adnasalia (Fig. 15); adnasalia margined with a mix of bluntly rounded and truncated teeth (Fig. 63); MP2/MP1&gt; 1.50; GA/MP1&gt; 1.50; 1.30 &lt;LP2/LP1 &lt;1.90; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with less than 3 PV and 4 AV secondary setae; metafemur with less than 3 AV secondary setae; urogomphus composed of one urogomphomere; U/HW &lt;0.40; U/LAS &lt;0.50; Neotropical.</p><p>Description, instar I (Figs 1–14)</p><p>Body: Measurements and ratios aimed to characterize body shape are shown in Table 2.</p><p>(1) Michat &amp; Torres (2009)</p><p>(1) Michat &amp; Torres (2009)</p><p>Head (Figs 1–9): HL = 0.45–0.47 mm; Head capsule rounded, lacking a clear constriction at level of occiput; occipital suture absent; anterior margin of frontoclypeus lightly convex, extending mesally at about level of adnasalia; A/MP = 1.48–1.60; MP/LP = 1.70–1.79; MP2/MP1 = 1.96–2.17; GA/MP1 = 2.50–3.04; LP2/LP1 = 2.27–2.50; MNL/MNW = 2.75–2.88.</p><p>Thorax (Figs 10–11): L3 = 0.83–0.88 mm.</p><p>Abdomen (Figs 12–14): LAS = 0.30–0.32 mm; LAS subcylindrical, not constricted posteriorly at point of insertion of urogomphi. Urogomphus: U = 0.18–0.19 mm, composed of one urogomphomere; U/HW = 0.36–0.39; U/LAS = 0.56–0.65.</p><p>Chaetotaxy (Figs 1–14): Position and number of additional setae on legs are shown in Table 3 and Figs 10– 11.</p><p>Description, instar II. As first-instar larva except as follows:</p><p>Body: Measurements and ratios aimed to characterize body shape are shown in Table 4.</p><p>Head: HL = 0.65 mm; anterior margin of frontoclypeus more narrowly convex; scale-like sculpticels welldeveloped over frontoclypeus and parietale; adnasalia margined with short bluntly rounded and truncated teeth (see Fig. 63); occipital suture present; A/MP = 1.59–1.64; MP/LP = 1.64–1.76; MP2/MP1 = 2.00–2.14; GA/MP1 = 2.38–2.57; LP2/LP1 = 1.78–1.89; MNL/MNW = 2.60–2.78.</p><p>Thorax: L3 = 1.32–1.34 mm.</p><p>Abdomen: LAS = 0.43–0.44 mm; LAS subcylindrical to subconical, less abruptly converging posteriorly at point of insertion of urogomphi. Urogomphus: U = 0.23–0.24 mm, composed of one urogomphomere; U/HW = 0.31–0.32; U/LAS = 0.54–0.56.</p><p>Chaetotaxy: Parietale with 3–5 temporal spines; position and number of secondary setae on legs are shown in Table 5.</p><p>Description, instar III (Figs 15–16, 63). As second-instar larva except as follows:</p><p>Body: Measurements and ratios aimed to characterize body shape are shown in Table 6.</p><p>Head (Figs 15, 63): HL = 0.88–0.89 mm; A/MP = 1.52–1.56; MP/LP = 1.77–1.90; MP2/MP1 = 1.58–1.75; GA/MP1 = 1.83–2.00; LP2/LP1 = 1.42–1.58; MNL/MNW = 2.43–2.83.</p><p>Thorax: L3 = 1.74–1.81 mm.</p><p>Abdomen (Fig. 16): LAS = 0.68–0.73 mm; U = 0.27–0.30 mm; U/HW = 0.27–0.32; U/LAS = 0.37–0.44.</p><p>Chaetotaxy: Parietale with 3–7 temporal spines. Position and number of secondary setae on legs are shown in Table 7.</p></div>	https://treatment.plazi.org/id/3B0087B31444FF8CFF3AFA21FF21428F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Alarie, Yves;Michat, Mariano C.;Watanabe, Kohei;Shaverdo, Helena;Wang, Liang-Jong;Watts, Chris H. S.	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31448FF97FF3AFA8DFEDD442B.text	3B0087B31448FF97FF3AFA8DFEDD442B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copelatus caelatipennis subsp. princeps Young 1963	<div><p>Copelatus caelatipennis princeps Young, 1963</p><p>Source of material. The two larvae studied (one instar II and one instar III) were associated with adults collected at the following locality: USA. Florida, Orlando. Junction International Drive &amp; Hwy 523, 26–28.IX.2016. 28°53.83’N 81°37.92’W; Y. Alarie leg. The identification is firm as C. caelatipennis princeps is the only Copelatus species known from that locality.</p><p>Diagnosis (instar III). The third instar of Copelatus c. princeps can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 0.80 mm; L3 &lt;2.00 mm; U &lt;0.40 mm; head capsule rounded to subquadrate, strongly constricted at level of occipital region (Fig. 17), scale-like sculpticels present over frontoclypeus and parietale; anterior margin of frontoclypeus narrowly convex, extending mesally well beyond level of adnasalia (Fig. 17); adnasalia margined with short bluntly rounded and truncated teeth (Fig. 63); MP2/MP1&gt; 1.50; GA/MP1&gt; 1.50; 1.30 &lt;LP2/LP1 &lt;1.90; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with less than 3 PV and 4 AV secondary setae; metafemur with less than 3 AV secondary setae; urogomphus composed of one urogomphomere; U/HW = 0.40; U/LAS = 0.50; Nearctic and Neotropical.</p><p>Description, instar II</p><p>Body: Measurements and ratios aimed to characterize body shape are shown in Table 4.</p><p>Head: Head capsule rounded to subquadrate, strongly constricted at level of occiput; anterior margin of frontoclypeus narrowly convex, extending well beyond level of adnasalia; adnasalia margined with short bluntly rounded and truncated teeth (see Fig. 63); scale-like sculpticels well-developed over frontoclypeus and parietale; HL = 0.58 mm; A/MP = 1.41; MP/LP = 1.85; MP2/MP1 = 1.73; GA/MP1 = 2.00; LP2/LP1 = 1.73; MNL/MNW = 2.72.</p><p>Thorax: L3 = 1.05 mm.</p><p>Abdomen: LAS = 0.39 mm; LAS subconical, not constricted posteriorly at point of insertion of urogomphi. Urogomphus: U = 0.22 mm, composed of one urogomphomere; U/HW = 0.38; U/LAS = 0.58.</p><p>Chaetotaxy: Parietale with 7 temporal spines; position and number of secondary setae on legs are shown in Table 5.</p><p>Description, instar III (Figs 17–18, 63). As second-instar larva except as follows:</p><p>Body: Measurements and ratios aimed to characterize body shape are shown in Table 6.</p><p>Head (Figs 18, 63): HL = 0.81 mm; A/MP = 1.50; MP/LP = 1.98; MP2/MP1 = 1.67; GA/MP1 = 2.00; LP2/LP1 = 1.51; MNL/MNW = 2.58.</p><p>Thorax: L3 = 1.58 mm.</p><p>Abdomen (Fig. 18): LAS = 0.64 mm; U = 0.34 mm; U/HW = 0.43; U/LAS = 0.52.</p><p>Chaetotaxy: Parietale with 10–12 temporal spines; position and number of secondary setae on legs are shown in Table 7.</p></div>	https://treatment.plazi.org/id/3B0087B31448FF97FF3AFA8DFEDD442B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Alarie, Yves;Michat, Mariano C.;Watanabe, Kohei;Shaverdo, Helena;Wang, Liang-Jong;Watts, Chris H. S.	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31453FF97FF3AFD29FF21403F.text	3B0087B31453FF97FF3AFD29FF21403F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copelatus glyphicus (Say 1823) NA	<div><p>Copelatus glyphicus (Say, 1823)</p><p>Source of material. The five instar III larvae studied were associated with adults collected at the following locality: Canada. Québec, Trois-Rivières. Ephemeral pond in red maple forest at Chemin des Oblats, 06.VIII.1986, 08.VIII.1986, 12.VIII.1986; Y. Alarie leg. The identification is firm as C. glyphicus is the only Copelatus species known from that region.</p><p>Diagnosis (instar III). The third instar of Copelatus glyphicus can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 0.80–0.90 mm; L3 &lt;2.00 mm; U &lt;0.40 mm; head capsule rounded, strongly constricted at level of occipital region (Fig. 19), scale-like sculpticels present over frontoclypeus and parietale; anterior margin of frontoclypeus broadly convex, extending mesally beyond level of adnasalia (Fig. 19); adnasalia margined with short bluntly rounded and truncated teeth (Fig. 63); MP2/MP1&gt; 1.50; GA/MP1&gt; 1.50; 1.30 &lt;LP2/LP1 &lt;1.90; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with less than 3 PV and 4 AV secondary setae; metafemur with less than 3 AV secondary setae; urogomphus composed of one urogomphomere; U/HW &lt;0.40; U/LAS &lt;0.50; Nearctic.</p><p>Description, instar III (Figs 19–20, 63)</p><p>Body: Measurements and ratios aimed to characterize body shape are shown in Table 6.</p><p>Head (Figs 19, 63): Head capsule rounded, strongly constricted at level of occiput; anterior margin of frontoclypeus broadly convex, extending beyond level of adnasalia; scale-like sculpticels well-developed both on frontoclypeus and parietale; adnasalia margined with short bluntly rounded and truncated teeth (Fig. 63); HL = 0.81–0.94 mm; A/MP = 1.34–1.38; MP/LP = 2.12–2.17; MP2/MP1 = 1.50–1.73; GA/MP1 = 1.79–1.88; LP2/LP1 = 1.40–1.56; MNL/MNW = 2.85–3.04.</p><p>Thorax: L3 = 1.60–1.74 mm.</p><p>Abdomen (Fig. 20): LAS = 0.65–0.74 mm; LAS subcylindrical, not constricted posteriorly at point of insertion of urogomphi. Urogomphus: U = 0.28–0.32 mm, composed of one urogomphomere; U/HW = 0.31–0.33; U/LAS = 0.42–0.44.</p><p>Chaetotaxy: Parietale with 6–9 temporal spines; position and number of secondary setae on legs are shown in Table 7.</p></div>	https://treatment.plazi.org/id/3B0087B31453FF97FF3AFD29FF21403F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Alarie, Yves;Michat, Mariano C.;Watanabe, Kohei;Shaverdo, Helena;Wang, Liang-Jong;Watts, Chris H. S.	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31453FF94FF3AF95AFE7444BB.text	3B0087B31453FF94FF3AF95AFE7444BB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copelatus japonicus Sharp 1884	<div><p>Copelatus japonicus Sharp, 1884</p><p>Source of material. The four instar III larvae studied were associated with adults collected at the following locality: Taiwan. Taipei, Beitou, 24.VIII.1994; L.J. Wang leg. The identification is firm as C. japonicus is the only Copelatus species collected at this locality.</p><p>Diagnosis (instar III). The third instar of Copelatus japonicus can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 0.90 mm; L3 &lt;2.00 mm; U &lt;0.40 mm; head capsule rounded, strongly constricted at level of occipital region (Fig. 21), scale-like sculpticels present over frontoclypeus and parietale; anterior margin of frontoclypeus narrowly convex, extending mesally at about level of adnasalia (Fig. 21); adnasalia margined with bluntly rounded and truncated teeth (Fig. 63); MP2/MP1&gt; 1.50; GA/ MP1&gt; 1.50; 1.30 &lt;LP2/LP1 &lt;1.90; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with less than 3 PV and 4 AV secondary setae; metafemur with less than 3 AV secondary setae; urogomphus composed of two urogomphomeres; U/HW &lt;0.40; U/LAS &lt;0.50; Palaearctic.</p><p>Description, instar III (Figs 21–24, 63)</p><p>Body: Measurements and ratios aimed to characterize body shape are shown in Table 6.</p><p>Head (Figs 21, 63): Head capsule rounded, strongly constricted at level of occiput; anterior margin of frontoclypeus narrowly convex, extending mesally at about level of adnasalia; scale-like sculpticels well-developed over frontoclypeus and parietale; adnasalia margined with bluntly rounded and truncated teeth (Fig. 63); HL = 0.86–0.90 mm; A/MP = 1.51–1.60; MP/LP = 1.77–1.88; MP2/MP1 = 1.57–1.82; GA/MP1 = 1.67–1.85; LP2/LP1 = 1.28–1.61; MNL/MNW = 2.50–2.72.</p><p>Thorax (Figs 23–24): L3 = 1.84–1.92 mm.</p><p>Abdomen (Fig. 22): LAS = 0.66–0.72 mm; LAS subconical, not constricted posteriorly at point of insertion of urogomphi. Urogomphus, U = 0.28–0.30 mm, composed of two urogomphomeres; U/HW = 0.28–0.30; U/LAS = 0.42–0.43.</p><p>Chaetotaxy: Parietale with 4–6 temporal spines; position and number of secondary setae on legs are shown in Figs 23–24 and Table 7.</p></div>	https://treatment.plazi.org/id/3B0087B31453FF94FF3AF95AFE7444BB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Alarie, Yves;Michat, Mariano C.;Watanabe, Kohei;Shaverdo, Helena;Wang, Liang-Jong;Watts, Chris H. S.	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31450FF94FF3AFCACFF2141BD.text	3B0087B31450FF94FF3AFCACFF2141BD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copelatus masculinus Regimbart 1899	<div><p>Copelatus masculinus Régimbart, 1899</p><p>Source of material. The five instar II and five instar III larvae studied were reared ex ovo from adults collected at the following locality: Japan. Tarumai-shitsugen, Yonaguni-cho Yonaguni-jima Island, Okinawa Prefecture, 24°26’31.2’’N 122°58’06.2’’E, 17.VI.2018; K. Watanabe leg.</p><p>Diagnosis (instar III). The third instar of Copelatus masculinus can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 1.00 mm; L3 &lt;2.00 mm; U &lt;0.40 mm; head capsule rounded, slightly constricted at level of occipital region (Fig. 25), scale-like sculpticels present over frontoclypeus and parietale; anterior margin of frontoclypeus narrowly convex, extending mesally slightly beyond adnasalia (Fig. 25); adnasalia margined with short bluntly rounded teeth (Fig. 64); MP2/MP1&gt; 1.50; GA/MP1&gt; 1.50; LP2/LP1 &lt;1.10; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with less than 3 PV and 4 AV secondary setae; metafemur with less than 3 AV secondary setae; urogomphus composed of two urogomphomeres; U/HW &lt;0.40; U/LAS &lt;0.50; Oriental and Palaearctic.</p><p>Description, instar II</p><p>Body: Measurements and ratios aimed to characterize body shape are shown in Table 4.</p><p>Head: Head capsule rounded, slightly constricted at level of occiput; anterior margin of frontoclypeus narrowly convex, extending mesally slightly beyond level of adnasalia; scale-like sculpticels well-developed over frontoclypeus and parietale; adnasalia margined with short bluntly rounded teeth (see Fig. 64); HL = 0.68–0.72 mm; A/MP = 1.33–1.36; MP/LP = 1.99–2.07; MP2/MP1 = 1.62–1.72; GA/MP1 = 1.92–2.13; LP2/LP1 = 1.33–1.55; MNL/MNW = 2.83–3.33.</p><p>Thorax: L3 = 1.32–1.42 mm.</p><p>Abdomen: LAS = 0.46–0.54 mm; LAS subcylindrical to subconical, not constricted posteriorly at point of insertion of urogomphi. Urogomphus: U = 0.23–0.25 mm, composed of two urogomphomeres; U/HW = 0.32–0.34; U/LAS = 0.46–0.51.</p><p>Chaetotaxy: Parietale with 5–7 temporal spines; position and number of secondary setae on legs are shown in Table 5.</p><p>Description, instar III (Figs 25–27, 64). As second-instar larva except as follows:</p><p>Body: Measurements and ratios aimed to characterize body shape are shown in Table 6.</p><p>Head (Figs 25, 27, 64): HL = 0.97–0.99 mm; A/MP = 1.34–1.37; MP/LP = 2.04–2.14; MP2/MP1 = 1.51–1.65; GA/MP1 = 1.61–1.85; LP2/LP1 = 1.06–1.12; MNL/MNW = 2.84–2.92.</p><p>Thorax: L3 = 1.90–2.00 mm.</p><p>Abdomen (Fig. 26): LAS = 0.75–0.77 mm; U = 0.31–0.32 mm, U/HW = 0.29–0.31; U/LAS = 0.40–0.43.</p><p>Chaetotaxy: Parietale with 7–9 temporal spines; position and number of secondary setae on legs are shown in Table 7.</p></div>	https://treatment.plazi.org/id/3B0087B31450FF94FF3AFCACFF2141BD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Alarie, Yves;Michat, Mariano C.;Watanabe, Kohei;Shaverdo, Helena;Wang, Liang-Jong;Watts, Chris H. S.	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B3145AFF9EFF3AFF0CFEDD4344.text	3B0087B3145AFF9EFF3AFF0CFEDD4344.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copelatus nakamurai Gueorguiev 1970	<div><p>Copelatus nakamurai Guéorguiev, 1970</p><p>Source of material. The two instar II and five instar III larvae studied were reared ex ovo from adults collected at the following locality: Japan. Kasamatshu-cho, Mudoji, Gifu Prefecture, 4.V.2018. Y. Yoshimura, leg.</p><p>Diagnosis (instar III). The third instar of Copelatus nakamurai can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 0.80 mm; L3 &lt;2.00 mm; U &lt;0.40 mm; head capsule subquadrate, strongly constricted at level of occipital region (Fig. 28), scale-like sculpticels lacking over frontoclypeus, present over parietale; anterior margin of frontoclypeus narrowly convex, extending mesally well beyond adnasalia (Fig. 28); adnasalia margined with elongate acute teeth (Fig. 64); MP2/MP1&gt; 1.50; GA/MP1&gt; 1.50; 1.30 &lt;LP2/LP1 &lt;1.90; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with less than 3 PV and 4 AV secondary setae; metafemur with less than 3 AV secondary setae; urogomphus composed of one urogomphomere; U/HW &lt;0.40; U/LAS &lt;0.50; Palaearctic.</p><p>Description, instar II</p><p>Body: Measurements and ratios aimed to characterize body shape are shown in Table 4.</p><p>Head: Head capsule subquadrate, strongly constricted at level of occiput; anterior margin of frontoclypeus narrowly convex, extending well beyond level of adnasalia; scale-like sculpticels lacking over frontoclypeus, welldeveloped over parietale; adnasalia margined with elongate acute teeth (see Fig. 64); HL = 0.61 mm; A/MP = 1.49; MP/LP = 2.06; MP2/MP1 = 1.60; GA/MP1 = 1.91; LP2/LP1 = 2.24; MNL/MNW = 3.11.</p><p>Thorax: L3 = 1.23 mm.</p><p>Abdomen: LAS = 0.45 mm; LAS subcylindrical, slightly constricted posteriorly at point of insertion of urogomphi. Urogomphus: U = 0.26 mm, composed of one urogomphomere; U/HW = 0.41; U/LAS = 0.58.</p><p>Chaetotaxy: Parietale with 8 temporal spines; position and number of secondary setae on legs are shown in Table 5.</p><p>Description, instar III (Figs 28–29, 64). As second-instar larva except as follows:</p><p>Body: Measurements and ratios aimed to characterize body shape are shown in Table 6.</p><p>Head (Figs 28, 64): HL = 0.81–0.84 mm; A/MP = 1.42–1.46; MP/LP = 2.02–2.25; MP2/MP1 = 1.51–1.74; GA/MP1 = 1.75–1.99; LP2/LP1 = 1.55–1.78; MNL/MNW = 2.85–2.98.</p><p>Thorax: L3 = 1.66–1.72 mm.</p><p>Abdomen (Fig 29): LAS = 0.66–0.69 mm; Urogomphus, U = 0.31–0.33 mm, composed of one urogomphomere; U/HW = 0.35–0.38; U/LAS = 0.45–0.49.</p><p>Chaetotaxy: Parietale with 9–10 temporal spines; position and number of secondary setae on legs are shown in Table 7.</p></div>	https://treatment.plazi.org/id/3B0087B3145AFF9EFF3AFF0CFEDD4344	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Alarie, Yves;Michat, Mariano C.;Watanabe, Kohei;Shaverdo, Helena;Wang, Liang-Jong;Watts, Chris H. S.	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B3145AFF9CFF3AFAC2FE74442B.text	3B0087B3145AFF9CFF3AFAC2FE74442B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copelatus oblitus Sharp 1882	<div><p>Copelatus oblitus Sharp, 1882</p><p>Source of material. The five instar II and five instar III larvae studied were reared ex ovo from adults collected at the following locality: Japan. Mantaburu, Yonaguni-cho, Yonaguni-jima Island, Okinawa Prefecture. 24°27’04.1”N 122°57’36.8”E, 13VI.2019; K. Watanabe leg.</p><p>Diagnosis (instar III). The third instar of Copelatus oblitus can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 0.80 mm; L3 &lt;2.00 mm; U &lt;0.30 mm; head capsule ovate, not constricted at level of occipital region (Fig. 30), scale-like sculpticels present over frontoclypeus and parietale; anterior margin of frontoclypeus narrowly convex, extending mesally well below level of adnasalia (Fig. 30); adnasalia margined with short bluntly rounded teeth (Fig. 64); MP2/MP1&gt; 1.50; GA/MP1&gt; 1.50; 1.30 &lt;LP2/LP1 &lt;1.90; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with less than 3 PV and 4 AV secondary setae; metafemur with less than 3 AV secondary setae; urogomphus composed of two urogomphomeres; U/HW &lt;0.40; U/LAS &lt;0.50; Oriental and Palaearctic.</p><p>Description, instar II</p><p>Body: Measurements and ratios aimed to characterize body shape are shown in Table 4.</p><p>Head: Head capsule ovate, not constricted at level of occiput; anterior margin of frontoclypeus narrowly convex, extending mesally below level of adnasalia; scale-like sculpticels present over frontoclypeus and parietale; adnasalia margined with short bluntly rounded teeth (see Fig. 64); HL = 0.55–0.58 mm; A/MP = 1.39–1.51; MP/LP = 1.70–1.82; MP2/MP1 = 1.67–1.95; GA/MP1 = 1.85–2.32; LP2/LP1 = 1.56–1.89; MNL/MNW = 2.41–2.61.</p><p>Thorax: L3 = 1.10–1.13 mm.</p><p>Abdomen: LAS = 0.37–0.39 mm; LAS subconical, not constricted posteriorly at point of insertion of urogomphi. Urogomphus: U = 0.17–0.19 mm, composed of two urogomphomeres; U/HW = 0.26–0.29; U/LAS = 0.45–0.51.</p><p>Chaetotaxy: Parietale with 3–5 temporal spines; position and number of secondary setae on legs are shown in Table 5.</p><p>Description, instar III (Figs 30–33, 64). As second-instar larva except as follows:</p><p>Body: Measurements and ratios aimed to characterize body shape are shown in Table 6.</p><p>Head (Figs 30, 64): HL = 0.74–0.80 mm; A/MP = 1.50–1.60; MP/LP = 1.65–1.82; MP2/MP1 = 1.44–1.67; GA/MP1 = 1.59–1.83; LP2/LP1 = 1.31–1.47; MNL/MNW = 2.40–2.60.</p><p>Thorax (Figs 32–33): L3 = 1.57–1.64 mm.</p><p>Abdomen (Fig 31): LAS = 0.54–0.60 mm; U = 0.20–0.25 mm, U/HW = 0.22–0.27; U/LAS = 0.37–0.47.</p><p>Chaetotaxy: Parietale with 3–4 temporal spines; position and number of secondary setae on legs are shown in Figs 32–33 and Table 7.</p></div>	https://treatment.plazi.org/id/3B0087B3145AFF9CFF3AFAC2FE74442B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Alarie, Yves;Michat, Mariano C.;Watanabe, Kohei;Shaverdo, Helena;Wang, Liang-Jong;Watts, Chris H. S.	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31458FFA5FF3AFD29FE7447C3.text	3B0087B31458FFA5FF3AFD29FE7447C3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copelatus parallelus Zimmermann 1920	<div><p>Copelatus parallelus Zimmermann, 1920</p><p>Source of material. All larvae studied were reared ex ovo from were adults collected at the following locality: Japan. Kyoto Prefecture. Uji-gawa, Fushimi-ku, Kyoto-shi, Kyoto Prefecture, 34°55’08.8”N 135°45’28.3”E, 6.XI.2016; K. Watanabe leg.</p><p>Diagnosis (instar III). The third instar of Copelatus parallelus can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 0.80 mm; L3 &lt;2.00 mm; U = 0.40 mm; head capsule subquadrate, strongly constricted at level of occipital region (Fig. 48), scale-like sculpticels present over frontoclypeus and parietale; anterior margin of frontoclypeus broadly convex, extending mesally at about level of adnasalia (Fig. 48); adnasalia margined with strong acute teeth (Fig. 64); MP2/MP1&gt; 1.50; GA/MP1&gt; 1.50; 1.30 &lt;LP2/LP1 &lt;1.90; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with less than 3 PV and 4 AV secondary setae; metafemur with less than 3 AV secondary setae; urogomphus composed of one urogomphomere; U/HW&gt; 0.40; U/LAS&gt; 0.60; Palaearctic.</p><p>Description, instar I (Figs 34–47)</p><p>Body: Measurements and ratios aimed to characterize body shape are shown in Table 2.</p><p>Head (Figs 34–42): HL = 0.43 mm; Head capsule subquadrate, strongly constricted at level of occiput; occipital suture absent; anterior margin of frontoclypeus broadly convex extending beyond level of adnasalia mesally; scale-like sculpticels well-developed over frontoclypeus and parietale; adnasalia margined with strong acute teeth; occipital carina absent; A/MP = 1.27; MP/LP = 2.30; MP2/MP1 = 2.17; GA/MP1 = 2.60; LP2/LP1 = 2.74; MNL/ MNW = 3.66.</p><p>Thorax (Figs 43–44): L3 = 0.80 mm.</p><p>Abdomen (Figs 45–47): LAS = 0.32 mm; LAS subconical, slightly constricted posteriorly at point of insertion of urogomphi. Urogomphus: U = 0.17 mm, composed of one urogomphomere; U/HW = 0.36; U/LAS = 0.52.</p><p>Chaetotaxy (Figs 34–47): Position and number of additional setae on legs are shown in Table 3 and Figs 43–44.</p><p>Description, instar II. As for first-instar larva except as follows:</p><p>Body: Measurements and ratios aimed to characterize body shape are shown in Table 4.</p><p>Head: Anterior margin of frontoclypeus extending mesally at about same level of adnasalia; adnasalia margined with strong acute teeth (see Fig. 64); occipital suture present; HL = 0.56 mm; A/MP = 1.39; MP/LP = 2.24; MP2/ MP1 = 1.86; GA/MP1 = 2.44; LP2/LP1 = 2.12; MNL/MNW = 3.55.</p><p>Thorax: L3 = 1.07 mm.</p><p>Abdomen: LAS = 0.41 mm. Urogomphus: U = 0.18 mm, composed of one urogomphomere; U/HW = 0.30; U/LAS = 0.45.</p><p>Chaetotaxy: Parietale with 7 temporal spines; position and number of secondary setae on legs are shown in Table 5.</p><p>Description, instar III (Figs 48–51, 64). As second-instar larva except as follows:</p><p>Body: Measurements and ratios aimed to characterize body shape are shown in Table 6.</p><p>Head (Figs 48, 64): HL = 0.78 mm; A/MP = 1.29; MP/LP = 2.38; MP2/MP1 = 1.69; GA/MP1 = 1.82; LP2/LP1 = 1.67; MNL/MNW = 3.07.</p><p>Thorax (Figs 50–51): L3 = 1.47 mm.</p><p>Abdomen (Fig. 49): LAS = 0.61 mm; U = 0.40 mm; U/HW = 0.50; U/LAS = 0.65.</p><p>Chaetotaxy: Parietale with 7 temporal spines. Position and number of secondary setae on legs are shown in Figs 50–51 and Table 7.</p></div>	https://treatment.plazi.org/id/3B0087B31458FFA5FF3AFD29FE7447C3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Alarie, Yves;Michat, Mariano C.;Watanabe, Kohei;Shaverdo, Helena;Wang, Liang-Jong;Watts, Chris H. S.	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31466FFA2FF3AFF0CFF2145A4.text	3B0087B31466FFA2FF3AFF0CFF2145A4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copelatus tenebrosus Regimbart 1880	<div><p>Copelatus tenebrosus Régimbart, 1880</p><p>Source of material. The single instar III larvae studied was reared ex ovo from were adults collected at the following locality: Japan. Mantaburu, Yonaguni-cho, Yonaguni-jima Island, Okinawa Prefecture, 24°27’04.1”N 122°57’36.8”E, 13.VI.2019; E. K. Watanabe leg.</p><p>Diagnosis (instar III). The third instar of Copelatus tenebrosus can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 0.80 mm; L3 &lt;2.00 mm; U &lt;0.40 mm; head capsule rounded, slightly constricted at level of occipital region (Fig. 52), scale-like sculpticels present over frontoclypeus and parietale; anterior margin of frontoclypeus narrowly convex, extending mesally well beyond level of adnasalia (Fig. 52); adnasalia margined with mostly short bluntly rounded teeth (Fig. 65); MP2/MP1&gt; 1.50; GA/MP1&gt; 1.50; 1.30 &lt;LP2/LP1 &lt;1.90; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with less than 3 PV and 4 AV secondary setae; metafemur with less than 3 AV secondary setae; urogomphus composed of two urogomphomeres; U/HW &lt;0.40; U/LAS &lt;0.50; Australia, Oriental, Palaearctic.</p><p>Description, instar III (Figs 52–53, 65)</p><p>Body: Measurements and ratios aimed to characterize body shape are shown in Table 6.</p><p>Head (Figs 52, 65): Head capsule rounded, slightly constricted at level of occiput; anterior margin of frontoclypeus narrowly convex, extending mesally well beyond level of adnasalia; scale-like sculpticels present over frontoclypeus and parietale; adnasalia margined with mostly short bluntly rounded teeth (Fig. 65); HL = 0.84 mm; A/MP = 1.40; MP/LP = 2.03; MP2/MP1 = 1.64; GA/MP1 = 1.85; LP2/LP1 = 1.43; MNL/MNW = 2.87.</p><p>Thorax: L3 = 1.73 mm.</p><p>Abdomen (Fig. 53): LAS = 0.63 mm; LAS subconical, not constricted posteriorly at point of insertion of urogomphi. Urogomphus, U = 0.30 mm, composed of two urogomphomeres; U/HW = 0.33; U/LAS = 0.47.</p><p>Chaetotaxy: Parietale with 5–6 temporal spines; position and number of secondary setae on legs are shown in Table 7.</p></div>	https://treatment.plazi.org/id/3B0087B31466FFA2FF3AFF0CFF2145A4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Alarie, Yves;Michat, Mariano C.;Watanabe, Kohei;Shaverdo, Helena;Wang, Liang-Jong;Watts, Chris H. S.	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31466FFA2FF3AFBA4FEDD41F0.text	3B0087B31466FFA2FF3AFBA4FEDD41F0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Exocelina australiae (Clark 1863) AU	<div><p>Exocelina australiae (Clark, 1863)</p><p>Source of material. The three instar III larvae studied were collected in association with adults at the following locality: South Australia. Ten kilometers North of Forreston, 9.VIII.1999; C. H. S. Watts leg.</p><p>Diagnosis (instar III). The third instar of Exocelina australiae can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 1.85–1.90 mm; L3&gt; 2.90 mm; U&gt; 0.50 mm; head capsule subquadrate, strongly constricted at level of occipital region (Fig. 54), scale-like sculpticels present over frontoclypeus and parietale; anterior margin of frontoclypeus broadly convex, extending mesally at level of adnasalia (Fig. 54); adnasalia margined with mostly short bluntly rounded teeth (Fig. 65); MP2/MP1&gt; 1.50; GA/MP1&gt; 1.50; LP2/LP1&gt; 2.00; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with more than 5 PV and 5 AV secondary setae; metafemur with more than 4 AV secondary setae; urogomphus composed of one urogomphomere; U/HW &lt;0.40; U/LAS &lt;0.50; Australian.</p><p>Description, instar III (Figs 54–55, 65)</p><p>Body: Measurements and ratios aimed to characterize body shape are shown in Table 6.</p><p>Head (Figs 54, 65): Head capsule subquadrate, strongly constricted at level of occiput; anterior margin of frontoclypeus broadly convex, extending mesally at about level of adnasalia; scale-like sculpticels present over frontoclypeus and parietale; adnasalia margined with mostly short bluntly rounded teeth (Fig. 65); HL = 1.85–1.90 mm; A/MP = 1.37–1.41; MP/LP = 2.36–2.41; MP2/MP1 = 1.56–1.67; GA/MP1 = 1.53–1.73; LP2/LP1 = 2.00–2.67; MNL/MNW = 2.72–2.96.</p><p>Thorax: L3 = 3.71–3.87 mm.</p><p>Abdomen (Fig. 55): LAS = 1.33–1.63 mm; LAS subcylindrical, not constricted posteriorly at point of insertion of urogomphi. Urogomphus, U = 0.56–0.58 mm, composed of one urogomphomere; U/HW = 0.25–0.26; U/LAS = 0.35–0.44.</p><p>Chaetotaxy: Parietale with 9–10 temporal spines; position and number of secondary setae on legs are shown in Table 7.</p></div>	https://treatment.plazi.org/id/3B0087B31466FFA2FF3AFBA4FEDD41F0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Alarie, Yves;Michat, Mariano C.;Watanabe, Kohei;Shaverdo, Helena;Wang, Liang-Jong;Watts, Chris H. S.	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31468FFACFF3AFF0CFF214344.text	3B0087B31468FFACFF3AFF0CFF214344.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Exocelina ferruginea (Sharp 1882) AU	<div><p>Exocelina ferruginea (Sharp, 1882)</p><p>Source of material. The three instar II and three instar III larvae studied were collected in association with adults at the following locality: South Australia. One kilometer North of Forreston, 2.IX.2000; C. H. S. Watts leg.</p><p>Diagnosis (instar III). The third instar of Exocelina ferruginea can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 1.10–1.20 mm; L3 = 2.20–2.50 mm; U&gt; 0.50 mm; head capsule subquadrate, strongly constricted at level of occipital region (Fig. 56), scale-like sculpticels present over frontoclypeus and parietale; anterior margin of frontoclypeus broadly convex, extending mesally at level of adnasalia (Fig. 54); adnasalia margined with short acute teeth (Fig. 65); MP2/MP1&gt; 1.50; GA/ MP1&gt; 1.50; LP2/LP1&gt; 2.00; inner margin of stipes lacking a dorsal linear row of spinulae; profemur with less than 3 PV and more than 5 AV secondary setae; metafemur with less than 3 AV secondary setae; urogomphus composed of one urogomphomere; U/HW&gt; 0.40; U/LAS&gt; 0.50; Australian.</p><p>Description, instar II</p><p>Body: Measurements and ratios aimed to characterize body shape are shown in Table 4.</p><p>Head: Head capsule subquadrate, strongly constricted at level of occiput; anterior margin of frontoclypeus broadly convex, extending mesally at level of adnasalia; scale-like sculpticels present over frontoclypeus and parietale; adnasalia margined with short acute teeth (see Fig. 65); HL = 0.85–0.90 mm; A/MP = 1.33–1.38; MP/LP = 2.52–3.00; MP2/MP1 = 1.83; GA/MP1 = 2.00–2.33; LP2/LP1 = 3.17–3.80; MNL/MNW = 2.77–3.18.</p><p>Thorax: L3 = 1.70–1.84 mm.</p><p>Abdomen: LAS = 0.54–0.63 mm; LAS subcylindrical, not constricted posteriorly at point of insertion of urogomphi. Urogomphus: U = 0.46–0.51 mm, composed of one urogomphomere; U/HW = 0.50–0.53; U/LAS = 0.74–0.96.</p><p>Chaetotaxy: Parietale with 4–6 temporal spines; position and number of secondary setae on legs are shown in Table 5.</p><p>Description, instar III (Figs 56–57, 65). As second-instar larva except as follows:</p><p>Body: Measurements and ratios aimed to characterize body shape are shown in Table 6.</p><p>Head (Figs 56, 65): HL = 1.08–1.23 mm; A/MP = 1.27–1.33; MP/LP = 2.79–3.27; MP2/MP1 = 1.55–1.82; GA/MP1 = 1.65–2.06; LP2/LP1 = 2.22–2.50; MNL/MNW = 2.88–2.94.</p><p>Thorax: L3 = 2.19–2.44 mm.</p><p>Abdomen (Fig. 57): LAS = 0.79–0.85 mm; U = 0.44–0.71 mm, U/HW = 0.38–0.57; U/LAS = 0.56–0.88.</p><p>Chaetotaxy: Parietale with 4–7 temporal spines; position and number of secondary setae on legs are shown in Table 7.</p></div>	https://treatment.plazi.org/id/3B0087B31468FFACFF3AFF0CFF214344	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Alarie, Yves;Michat, Mariano C.;Watanabe, Kohei;Shaverdo, Helena;Wang, Liang-Jong;Watts, Chris H. S.	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
3B0087B31468FFA8FF3AFACFFE11472F.text	3B0087B31468FFA8FF3AFACFFE11472F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liopterus haemorrhoidalis (Fabricius 1787)	<div><p>Liopterus haemorrhoidalis (Fabricius, 1787)</p><p>Source of material. The four instar III larvae were collected in association with adults at the following locality: Belarus, Vicebsk Region, area near upy Village, wetland of evinka River, Carex, moss. 12.VIII.1998;H.Shaverdo leg.</p><p>Diagnosis (instar III). The third instar of Liopterus haemorrhoidalis can easily be distinguished from that of the other species studied in this paper by the following combination of characters: HL = 1.40 mm; L3&gt; 2.90 mm; U&gt; 0.50 mm; head capsule subquadrate, strongly constricted at level of occipital region (Fig. 58), scale-like sculpticels lacking over frontoclypeus and parietale; anterior margin of frontoclypeus broadly convex, extending mesally at level of adnasalia (Fig. 58); adnasalia margined with short bluntly rounded teeth (Fig. 65); MP2/MP1 &lt;1.30; GA/MP1 &lt;1.50; 1.30 &lt;LP2/LP1 &lt;1.90; inner margin of stipes with a dorsal linear row of spinulae (Fig. 60); profemur with less than 3 PV and more than 5 AV secondary setae; metafemur with more than 4 AV secondary setae (Fig. 61); urogomphus composed of one urogomphomere; U/HW = 0.40; U/LAS&gt; 0.50; Palaearctic.</p><p>Description, instar III (Figs 58–62, 65)</p><p>Body: Measurements and ratios aimed to characterize body shape are shown in Table 6.</p><p>Head (Figs 58, 60, 65): Head capsule subquadrate, strongly constricted at level of occiput; anterior margin of frontoclypeus broadly convex, extending mesally at about level of adnasalia; scale-like sculpticels lacking over frontoclypeus and parietale; adnasalia margined with mostly short bluntly rounded teeth (Fig. 65); HL = 1.36–1.39 mm; A/MP = 1.16–1.27; MP/LP = 1.67–1.74; MP2/MP1 = 1.15–1.28; GA/MP1 = 1.27–1.49; LP2/LP1 = 1.54–1.71; MNL/MNW = 2.63–2.71.</p><p>Thorax (Figs 61–62): L3 = 2.86–3.06 mm.</p><p>Abdomen (Fig. 59): LAS = 1.09–1.17 mm; LAS subconical, constricted posteriorly at point of insertion of urogomphi. Urogomphus, U = 0.60–0.63 mm, composed of one urogomphomere; U/HW = 0.38–0.40; U/LAS = 0.54–0.57.</p><p>Chaetotaxy: Parietale with 7–10 temporal spines; position and number of secondary setae on legs are shown in Figs 61–62 and Table 7.</p></div>	https://treatment.plazi.org/id/3B0087B31468FFA8FF3AFACFFE11472F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Alarie, Yves;Michat, Mariano C.;Watanabe, Kohei;Shaverdo, Helena;Wang, Liang-Jong;Watts, Chris H. S.	Alarie, Yves, Michat, Mariano C., Watanabe, Kohei, Shaverdo, Helena, Wang, Liang-Jong, Watts, Chris H. S. (2022): An outlook on larval morphology of Copelatinae diving beetles with phylogenetic considerations (Coleoptera: Adephaga, Dytiscidae). Zootaxa 5175 (2): 151-205, DOI: 10.11646/zootaxa.5175.2.1
