taxonID	type	description	language	source
3601F35E9E529332A3F8FC90FDCBDE04.taxon	materials_examined	Type species: Ceratoderus graniger Mulsant, 1851 (by monotypy).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E549334A3F8FB16FC96DDFC.taxon	description	Figs. 11 (habitus), 15 (aedeagus), 128 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E549334A3F8FB16FC96DDFC.taxon	materials_examined	Type Material. Holotype (male): Peru: Pantiacolla Lodge, Monk Saki Trail, Alto Madre de Dios River, at black light, elev. 400 m, 12 ° 39 ' S, 71 ° 13 ' W, 25 x 2000, R. Brooks (PERU 1 B 00 098 A) (SEMC). Paratypes (42): Peru: Same data as holotype (32 SEMC); Madre de Dios, Hostel Erica (near Salvacion), elev. 550 m, 12 ° 53 ' S, 71 ° 14 ' W, 3 – 5 ix 1989, R. A. Faitoute et al. (10 USNM).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E549334A3F8FB16FC96DDFC.taxon	diagnosis	Differential Diagnosis. The habitus of this species (Fig. 11) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus could be compared with that of E. transversus (Figs. 12, 15); however, the median lobe is shaped differently, considerably smaller in E. transversoides, and the internal sac is different in extent and sculpture in the two species, being narrower and with denser spicules in E. transversoides.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E549334A3F8FB16FC96DDFC.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 1.36 / 0.69; head (width) 0.41; pronotum 0.52 / 0.51; elytra 0.89 / 0.69. Habitus and sculpture as illustrated (Fig. 11). Head black, dorsum brown to reddish brown, venter and coxae dark brown. Eye with ca. 4 – 5 facets between canthus and posterior margin. Elytra with elongate granules linking punctures. Metaventral depression moderately deep, ca. eight granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E549334A3F8FB16FC96DDFC.taxon	etymology	Etymology. Named in reference to the similarity of the aedeagus to that of E. transversus.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E549334A3F8FB16FC96DDFC.taxon	distribution	Distribution. Currently known only from Peru (Fig. 128).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E549334A3F8FDC0FB89DEF3.taxon	description	Figs. 8 (habitus), 12 (aedeagus), 127 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E549334A3F8FDC0FB89DEF3.taxon	materials_examined	Type Material. Holotype (male): Bolivia: La Paz, Alcoche, lamping, elev. 600 m, 15 ° 45 ' S, 67 ° 40 ' W, 19 xii 1966, Balough, Mahunka, Zicsi (493 Soil Zoological Exp.) (HNHM). Paratypes: Same data as holotype (12 HNHM).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E549334A3F8FDC0FB89DEF3.taxon	diagnosis	Differential Diagnosis. The habitus of this species (Fig. 8) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus (Fig. 12) has a median lobe somewhat similar in general shape to that of E. transversoides (Fig. 15), but it is much longer and apically is shaped differently. The parameres are similar in shape in the two species, but in E. transversus they are longer compared to the length of the basal piece.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E549334A3F8FDC0FB89DEF3.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 1.65 / 0.81; head (width) 0.44; pronotum 0.59 / 0.60; elytra 1.10 / 0.81. Habitus and sculpture as illustrated (Fig. 8). Head black, dorsum red, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 3 – 4 facets between canthus and posterior margin. Protibiae slightly arcuate. Elytral punctures linked by elongate granules. Metaventral depression moderately deep, ca. eight granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E549334A3F8FDC0FB89DEF3.taxon	etymology	Etymology. Named in reference to the transverse apex of the median lobe of the aedeagus.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E549334A3F8FDC0FB89DEF3.taxon	distribution	Distribution. Currently known only from the type locality in Bolivia (Fig. 127).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E579336A3F8F92BFE66D87A.taxon	description	Figs. 9 (habitus), 13 (aedeagus), 127 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E579336A3F8F92BFE66D87A.taxon	materials_examined	Type material: Holotype (male): " Type [red disc]; Yungas Valley Bolivia. 12 – 13. vi. 1937; H. E. Hinton Collector; Epimetopus hintoni Type! J. Balfour-Browne det.; Hinton Coll. B. M. 1939 - 583 " (BMNH). New collection record: Bolivia: Santa Cruz, Amboro National Park, Los Volcanes, MV Light Sheet on stream beach, elev. 1000 m, 18 ° 6 ' S, 63 ° 36 ' W, 20 xi – 12 xii 2004, M. V. L. Barclay & H. Mendel (34 BMNH).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E579336A3F8F92BFE66D87A.taxon	diagnosis	Differential diagnosis. The habitus of this species (Fig. 9) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus (Fig. 13). The median lobe is nearly parallel-sided in dorsal view and quite blunt apically; in lateral aspect it is strongly curved. The aedeagus is similar in general shape to that of E. ecuadorensis (Fig. 21), but differs in shapes of the median lobe and parameres, and curvature of the medial lobe. Externally, E. hintoni is slightly larger than E. ecuadorensis (ca. 1.71 vs. 1.56 mm), but otherwise they are very similar.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E579336A3F8F92BFE66D87A.taxon	distribution	Distribution. Currently known from two localities in Bolivia (Fig. 127). Oliva (1986) reported specimens from Tucuman Province, Argentina; these specimens were not seen during this revision, and therefore could not be confirmed as E. hintoni.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E569336A3F8FD88FBF1DE8C.taxon	description	Figs. 10 (habitus), 14 (aedeagus), 127 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E569336A3F8FD88FBF1DE8C.taxon	materials_examined	Type Material. Holotype (male, greatly disarticulated): Brazil: " Matto Grosso, Corumba " (ISNB). New collection records: Bolivia: Beni, Ilha Flores, Rio Itenez, 13 ° 2 ' S, 62 ° 43 ' W, 7 viii 1964, J. Bouseman & J. Lussenhop (5 MCZ); Paraguay: Dep. Concepcion, Arroyo Toro Paso, 23 ° 51 ' S, 56 ° 46 ' W, 25 x 2002, U. Drechsel (5 NMW); Arroyo Toro Paso, Unterlauf (contrib. Rio Apa), seepage, 23 ° 51 ' S, 56 ° 46 ' W, 24 x 2002, U. Drechsel (102 NMW); Stillwasser des Arroyo Toro Paso, Seitenam (Uferschlamm), 23 ° 51 ' S, 56 ° 46 ' W, 25 x 2002, U. Drechsel (8 NMW).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E569336A3F8FD88FBF1DE8C.taxon	diagnosis	Differential diagnosis. Differentiated from all members of the Costatus group, except E. trilobus, by the trilobed apical margin of the pronotal hood (Figs. 10, 32). The aedeagus of E. lacordairei (Fig. 14), with an apically deeply bifid median lobe, differs markedly from all other members of the genus. The aedeagus of E. trilobus (Fig. 36) probably represents a preliminary stage in the evolution of the extreme condition seen in E. lacordairei. This view is supported by the fact that both species have a tri-lobed pronotal apex — the only species now known to have that condition.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E569336A3F8FD88FBF1DE8C.taxon	distribution	Distribution. Currently known from Bolivia, Brazil, and Paraguay (Fig. 127).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E569336A3F8FAD1FBAEDDB9.taxon	description	Figs. 32 (habitus), 36 (aedeagus), 128 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E569336A3F8FAD1FBAEDDB9.taxon	materials_examined	Type Material. Holotype (male): Venezuela: Bolivar, Los Pijiguaos, trampa interceptacion, elev. 600 m, 6 ° 35 ' N, 66 ° 45 ' W, 3 – 13 vi 1992, C. J. Rosales, V. Savini, J. L. Garcia, J. DeMarmels (MIZA). Paratype: Same data as holotype (1 MIZA).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E569336A3F8FAD1FBAEDDB9.taxon	diagnosis	Differential Diagnosis. Differentiated from all members of the Costatus group, except E. lacordairei, by the trilobed apical margin of the pronotal hood (Figs. 10, 32). The aedeagus is distinctive by virtue of the apically bifid apex of the median lobe (Fig 36). Although the aedeagus differs markedly from that of E. lacordairei (Fig. 14), the bifid apex of both, the similarity in elytral structure, and especially the tri-lobed pronotal apex, suggest a relationship.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E569336A3F8FAD1FBAEDDB9.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 1.28 / 0.64; head (width) 0.43; pronotum 0.50 / 0.51; elytra 0.85 / 0.64. Habitus and sculpture as illustrated (Fig. 32). Head bicolored, frons red, clypeus black, venter and coxae reddish brown, remainder of legs ochraceous. Eye with ca. 3 – 4 facets between canthus and posterior margin. Pronotal hood minutely trilobed. Elytra with elongate granules linking punctures; 3 rd carina absent from basal ½ of elytron, otherwise carinae quite high. Protibiae arcuate, anterior surface flat. Metaventral depression shallow.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E569336A3F8FAD1FBAEDDB9.taxon	etymology	Etymology. Named in reference to the tri-lobed pronotal apex.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E569336A3F8FAD1FBAEDDB9.taxon	distribution	Distribution. Currently known only from the type locality in Venezuela (Fig. 128).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E499328A3F8F8C3FCD9D87A.taxon	description	Figs. 16 (habitus), 20 (aedeagus), 130 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E499328A3F8F8C3FCD9D87A.taxon	materials_examined	Type Material. Holotype (male): Peru: Pantiacolla Lodge, Monk Saki Trail, Alto Madre de Dios River, at black light, elev. 400 m, 12 ° 39 ' S, 71 ° 13 ' W, 25 x 2000, R. Brooks (PERU 1 B 00 098 A) (SEMC). Paratypes: Peru: Madre de Dios, Hostel Erica (near Salvacion), elev. 550 m, 12 ° 53 ' S, 71 ° 14 ' W, 3 – 5 ix 1989, R. A. Faitoute et al. (2 USNM).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E499328A3F8F8C3FCD9D87A.taxon	diagnosis	Differential Diagnosis. The habitus of this species (Fig. 16) is very similar to that of other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus is very distinctive by virtue of the spatula-like shape of the median lobe, and its comparatively long length (Fig 20). Also, the parameres are distinctively shaped.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E499328A3F8F8C3FCD9D87A.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 1.68 / 0.85; head (width) 0.43; pronotum 0.56 / 0.57; elytra 1.14 / 0.85. Habitus and sculpture as illustrated (Fig. 16). Head black, dorsum red, venter and coxae dark brown. Eye with ca. 3 – 4 facets between canthus and posterior margin. Elytra with elongate granules linking punctures, non-carinate intervals with uneven surfaces. Metaventral depression large and moderately deep, with a very low midlongitudinal ridge anteriorly, ca. 11 granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E499328A3F8F8C3FCD9D87A.taxon	etymology	Etymology. Named in reference to the spatula-like shape of the median lobe of the aedeagus.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E499328A3F8F8C3FCD9D87A.taxon	distribution	Distribution. Currently known from Peru (Fig. 130).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E489328A3F8FD88FC87DE8B.taxon	description	Figs. 17 (habitus), 21 (aedeagus), 127 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E489328A3F8FD88FC87DE8B.taxon	materials_examined	Type Material. Holotype (male): Ecuador: Napo, 17 km SW Tena, 1 ° 4 ' S, 77 ° 49 ' W, 28 v 1977, W. E. Steiner (USNM). Paratypes (3): Misahualli, 1 ° 2 ' S, 77 ° 40 ' W, 9 xi 1987, M. Huybensz (1 MCZ); Sta. Cecilia, at whitelight, 0 ° 2 ' N, 76 ° 58 ' W, 16 v 1975, Ashley B. Gurney (1 USNM); Tena, 0 ° 59 ' N, 77 ° 49 ' W, 27 v 1977, W. E. Steiner (1 USNM).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E489328A3F8FD88FC87DE8B.taxon	diagnosis	Differential Diagnosis. The habitus of this species (Fig. 17) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus is somewhat similar to that of E. hintoni (Figs. 13, 21). However, the median lobe is slightly wider than that of E. hintoni, whereas the parameres, in both views, are narrower. The median lobe internal structures also differ in the two species, as illustrated.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E489328A3F8FD88FC87DE8B.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 1.56 / 0.75; head (width) 0.40; pronotum 0.56 / 0.57; elytra 1.01 / 0.75. Habitus and sculpture as illustrated (Fig. 17). Head black, dorsum red, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 4 – 5 facets between canthus and posterior margin. Elytra with distinct granules linking punctures. Protibiae slightly arcuate. Metaventral depression moderately deep, ca. eight granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E489328A3F8FD88FC87DE8B.taxon	etymology	Etymology. Named in reference to the geographical distribution.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E489328A3F8FD88FC87DE8B.taxon	distribution	Distribution. Currently known from Ecuador (Fig. 127).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E489328A3F8FADEFCBCDDB4.taxon	description	Figs. 18 (habitus), 22 (aedeagus), 128 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E489328A3F8FADEFCBCDDB4.taxon	materials_examined	Type Material. Holotype (male): Ecuador, Napo, Tena, 0 ° 59 ' N, 77 ° 49 ' W, 27 v 1977, W. E. Steiner (USNM). Paratypes (18): Ecuador: Same data as holotype (5 USNM); Napo, Tena, 0 ° 59 ' N, 77 ° 49 ' W, 26 v 1977, W. E. Steiner (11 USNM); Tena (4 km S), 1 ° 3 ' S, 77 ° 48 ' W, 23 v 1977, P. J. Spangler & D. R. Givens (# 70) (2 USNM).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E489328A3F8FADEFCBCDDB4.taxon	diagnosis	Differential Diagnosis. The habitus of this species (Fig. 18) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus (Fig. 22) has a large and distinctively shaped median lobe, widest at about midlength, tapering toward the apex, then widening slightly apically. The internal sac has many fine ridges and spicules. It might be compared with E. transversoides (Fig. 15), but the median lobe is shaped differently and significantly larger.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E489328A3F8FADEFCBCDDB4.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 2.22 / 1.69; head (width) 0.44; pronotum 0.61 / 0.68; elytra 1.11 / 0.85. Habitus and sculpture as illustrated (Fig. 18). Head black, dorsum red, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 3 – 4 facets between canthus and posterior margin. Elytral punctures round, each with distinct granule at anterior margin, most granules not linking punctures. Protibiae slightly arcuate. Metaventral depression moderately deep, ca. six granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E489328A3F8FADEFCBCDDB4.taxon	etymology	Etymology. Named in honor of the expert collector, long-time friend and colleague Warren E. Steiner.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E489328A3F8FADEFCBCDDB4.taxon	distribution	Distribution. Currently only known from Ecuador (Fig. 128).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4B932AA3F8F917FC2AD8CA.taxon	description	Figs. 19 (habitus), 23 (aedeagus), 129 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4B932AA3F8F917FC2AD8CA.taxon	materials_examined	Type Material. Holotype (male): Ecuador: Napo, Tena, 0 ° 59 ' N, 77 ° 49 ' W, 26 v 1977, W. E. Steiner (USNM). Paratypes (14): Ecuador: Napo, Tena, 0 ° 59 ' N, 77 ° 49 ' W, 26 v 1977, W. E. Steiner (5 USNM); Peru: Madre de Dios, Hostel Erica (near Salvacion), elev. 550 m, 12 ° 53 ' S, 71 ° 14 ' W, 3 – 5 ix 1989, R. A. Faitoute et al. (3 USNM); Pantiacolla Lodge, Monk Saki Trail, Alto Madre de Dios River, at black light, elev. 400 m, 12 ° 39 ' S, 71 ° 13 ' W, 25 x 2000, R. Brooks (PERU 1 B 00 098 A) (5 SEMC); Rio Tambopata Res; 30 air km SW Pto. Maldonado, subtropical moist forest, elev. 290 m, 13 ° 0 ' S, 69 ° 33 ' W, 16 – 20 xi 1979, J. B. Heppner (1 USNM).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4B932AA3F8F917FC2AD8CA.taxon	diagnosis	Differential Diagnosis. A rather coarsely granulate species (Fig. 19), with prominent elongate granules linking the rather large elytral punctures; however, reliable determinations will be based on examination of the aedeagus. The aedeagus is relatively simple in this species (Fig. 23). The median lobe is pointed apically, like that of E. simplex (Fig. 37, 38). However, the parameres are very different in the two species, being pointed apically in E. inaequalis, and much longer than the median lobe.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4B932AA3F8F917FC2AD8CA.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 1.69 / 0.82; head (width) 0.45; pronotum 0.61 / 0.55; elytra 1.11 / 0.82. Habitus and sculpture as illustrated (Fig. 19). Head black, dorsum brown to reddish brown, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 3 – 4 facets between canthus and posterior margin. Protibiae slightly arcuate. Granules of pronotum and elytral carinae quite coarse. Elytral punctures large, granules linking punctures prominent. Metaventral depression moderately deep, ca. six granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4B932AA3F8F917FC2AD8CA.taxon	etymology	Etymology. Named in reference to the unequal lengths of the median lobe and parameres.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4B932AA3F8F917FC2AD8CA.taxon	distribution	Distribution. Currently known only from Ecuador and Peru (Fig. 129).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4A932AA3F8FD18FB5FDCFB.taxon	description	Figs. 24 (habitus), 28 (aedeagus), 130 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4A932AA3F8FD18FB5FDCFB.taxon	materials_examined	Type Material. Holotype (male): Ecuador: Napo, Tena, 0 ° 59 ' N, 77 ° 49 ' W, 27 v 1977, W. E. Steiner (USNM). Paratypes (1479): Ecuador: Napo, Limoncocha, 1 ° 33 ' S, 75 ° 35 ' W, 11 vi 1977, W. E. Steiner (1 USNM); Tena, 0 ° 59 ' N, 77 ° 49 ' W, 26 v 1977, W. E. Steiner (1 USNM); Panama: Panama, El Real, 8 ° 8 ' N, 77 ° 43 ' W, 6 viii 1952, F. S. Blanton (3 USNM); Peru: Madre de Dios, Pantiacolla Lodge, Monk Saki Trail, Alto Madre de Dios River, at black light, elev. 400 m, 12 ° 39 ' S, 71 ° 13 ' W, 25 x 2000, R. Brooks (PERU 1 B 00 098 A) (23 SEMC); Venezuela: Amazonas, Agua Blanca, Cerro d. l. Neblina, elev. 160 m, 0 ° 49 ' N, 66 ° 18 ' W, 20 – 21 v 1984, O. Flint & J. Louton (3 USNM); Cerro de la Neblina, Basecamp, At black light in rainforest clearing near Rio Baria, elev. 140 m, 0 ° 50 ' N, 66 ° 10 ' W, 10 – 11 ii 1985, P. J. Spangler, P. M. Spangler, R. A. Faitoute, W. E. Steiner (268 USNM); Cerro de la Neblina, Basecamp, At black light on bank of Rio Baria, elev. 140 m, 0 ° 50 ' N, 66 ° 10 ' W, 6 ii 1985, W. E. Steiner (887 USNM); same locality, elev. 140 m, 0 ° 50 ' N, 66 ° 10 ' W, 28 ii 1985, P. J. Spangler, P. M. Spangler, R. A. Faitoute, W. E. Steiner (293 USNM).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4A932AA3F8FD18FB5FDCFB.taxon	diagnosis	Differential Diagnosis. The habitus of this species (Fig. 24) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus (Fig. 28) is somewhat similar in general shape to that of E. burruyacu (Fig. 72); both have slender, slightly arching parameres. E. angustus differs from E. burruyacu in that the parameres are longer compared with the length of the basal piece; also, the median lobe is narrower and longer.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4A932AA3F8FD18FB5FDCFB.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 1.31 / 0.73; head (width) 0.38; pronotum 0.52 / 0.53; elytra 0.85 / 0.73. Habitus and sculpture as illustrated (Fig. 24). Head black, dorsum red, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 3 – 4 facets between canthus and posterior margin. Protibiae slightly arcuate. Elytral granules linking punctures. Metaventral depression moderately deep, ca. six granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4A932AA3F8FD18FB5FDCFB.taxon	etymology	Etymology. Named in reference to the narrow median lobe and narrow parameres.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4A932AA3F8FD18FB5FDCFB.taxon	distribution	Distribution. Currently known from Panama, Venezuela, Ecuador, and Peru (Fig. 130).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4A932CA3F8F90EFBC2DBEA.taxon	description	Figs. 25 (habitus), 31 (aedeagus), 127 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4A932CA3F8F90EFBC2DBEA.taxon	materials_examined	Type Material. Holotype (male): Colombia: Cesar, Pueblo Bello; Sierra de Santa Marta, beaten ex tree debris nr. stream, elev. 1200 m, 10 ° 25 ' N, 73 ° 35 ' W, 14 – 15 iv 1968, B. Malkin (FMNH). Paratype: Venezuela: Miranda, Los Chorros, 10 ° 30 ' N, 66 ° 50 ' W, 29 iv 1922, [collr. unknown] (1 FMNH).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4A932CA3F8F90EFBC2DBEA.taxon	diagnosis	Differential Diagnosis. A moderately large and rather coarsely granulate species (Fig. 25); however, reliable determinations will be based on examination of the aedeagus. The aedeagus is similar to that of E. latisoides in that the parameres are very broad, have the medial margins straight or nearly so, and have two very small setae at the apex of each (Figs. 31, 45). However, the aedeagus of E. latus is larger (length 0.38 mm vs. 0.31 mm), the median lobe is wider and the apex is shaped slightly differently.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4A932CA3F8F90EFBC2DBEA.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 1.79 / 0.93; head (width) 0.50; pronotum 0.64 / 0.61; elytra 1.17 / 0.93. Habitus and sculpture as illustrated (Fig. 25). Granules of dorsum rather coarse. Head black, dorsum brown to reddish brown, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 3 – 4 facets between canthus and posterior margin. Protibiae slightly arcuate. Elytral punctures large; elongate granules linking punctures prominent. Metaventral depression moderately deep, ca. eight granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4A932CA3F8F90EFBC2DBEA.taxon	etymology	Etymology. Named in reference to the wide parameres.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4A932CA3F8F90EFBC2DBEA.taxon	distribution	Distribution. Currently known from Colombia and Venezuela (Fig. 127).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4C932CA3F8FE38FC33DF43.taxon	description	Figs. 26 (habitus), 30 (aedeagus), 129 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4C932CA3F8FE38FC33DF43.taxon	materials_examined	Type Material. Holotype (male): Venezuela: Tachira State, El Tama National Park, HG-vapor light, elev. 472 m, 7 ° 35.038 ' N, 72 ° 10.34 ' W, 16 vii 2009, Short, Sites, Garcia, Inciarte, Gustafson & Camaco (VZ 09 - 0716 - 07 A) (MIZA). Paratypes (30): Trinidad: Lovell Estate, 10 ° 22 ' N, 61 ° 14 ' W, 29 x 1937, H. E. Hinton (1 BMNH); Venezuela: Aragua, Ocumare, 10 ° 7 ' N, 66 ° 46 ' W, 19 – 20 ii 1969, P. & P. Spangler (1 USNM); Tachira State, El Tama National Park, HG-vapor light, elev. 472 m, 7 ° 35.038 ' N, 72 ° 10.34 ' W, 16 vii 2009, Short, Sites, Garcia, Inciarte, Gustafson & Camaco (VZ 09 - 0716 - 07 A) (6 SEMC); Trujillo, Mcp. Rafael Rangel, La Gira, Qda. La Amarilla, Trampa interceptacion, elev. 520 m, 9 ° 20 ' N, 70 ° 44 ' W, 20 – 22 v 1995, J. Camacho, M. Garcia (3 MALUZ); Same locality, 23 x 1997, J. Camacho, M. Garcia, E. Gomez (19 MALUZ).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4C932CA3F8FE38FC33DF43.taxon	diagnosis	Differential Diagnosis. The habitus of this species (Fig. 26) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus has arcuate parameres that arch over the median lobe; the extent of the arching is much more than in E. apocinus (Figs. 30, 48). The median lobe, which tapers distinctively in the distal one-half also clearly differs from that of E. apocinus.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4C932CA3F8FE38FC33DF43.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 1.48 / 0.80; head (width) 0.43; pronotum 0.56 / 0.60; elytra 0.95 / 0.80. Habitus and sculpture as illustrated (Fig. 26). Head black, dorsum red, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 3 – 4 facets between canthus and posterior margin. Elytra with elongate granules linking punctures, non-carinate intervals smooth, without granules. Metaventral depression small.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4C932CA3F8FE38FC33DF43.taxon	etymology	Etymology. Named in reference to the shape of the aedeagus, which has the parameres arching over the median lobe, reminiscent of a ballet dancer.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4C932CA3F8FE38FC33DF43.taxon	distribution	Distribution. Currently known from Venezuela and Trinidad (Fig. 129).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4C932FA3F8FA66FBAEDAB9.taxon	description	Figs. 27 (habitus), 29 (aedeagus), 128 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4C932FA3F8FA66FBAEDAB9.taxon	materials_examined	Type Material. Holotype (male): Venezuela: Tachira State, El Tama National Park, HG-vapor light, elev. 472 m, 7 ° 35.038 ' N, 72 ° 10.34 ' W, 16 vii 2009, Short, Sites, Garcia, Inciarte, Gustafson & Camaco (VZ 09 - 0716 - 07 A) (MIZA).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4C932FA3F8FA66FBAEDAB9.taxon	diagnosis	Differential Diagnosis. The habitus of this species (Fig. 27) is very similar to that of other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus of this species has very distinctive parameres (Fig. 29). They are wide, " wrap around " the median lobe, and the ventral median area has an unusual wrinkled appearance. The median lobe is comparatively short, whereas the basal piece is long. The aedeagus cannot be meaningfully compared with that of other species.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4C932FA3F8FA66FBAEDAB9.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 1.49 / 0.73; head (width) 0.40; pronotum 0.54 / 0.56; elytra 0.99 / 0.73. Habitus and sculpture as illustrated (Fig. 27). Head black, dorsum red, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 4 – 5 facets between canthus and posterior margin. Each elytral puncture with granule at anterior and posterior margin, granules usually not linking punctures, non-carinate intervals wide and smooth. Metaventral depression relatively small, only ca. ½ as wide as labrum.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4C932FA3F8FA66FBAEDAB9.taxon	etymology	Etymology. Named in reference to the " wrinkles " of the parameres.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4C932FA3F8FA66FBAEDAB9.taxon	distribution	Distribution. Currently known only from the type locality in Venezuela (Fig. 128).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4F932EA3F8F96CFC2ED8A1.taxon	description	Figs. 33 (habitus), 37, 38 (aedeagus), 144 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4F932EA3F8F96CFC2ED8A1.taxon	diagnosis	Differential diagnosis. The habitus of E. simplex (Fig. 33) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The median lobe is similar to that of E. inaequalis (Figs. 23, 37, 38), but the parameres differ markedly in the two species; those of E. simplex being much narrower basally, rounded apically, and each bearing two short, blunt setae. Externally (Figs. 19, 33) the pronotal hood shape and pronotal hood carinae differ slightly in the two species.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4F932EA3F8F96CFC2ED8A1.taxon	distribution	Distribution. Currently known from Venezuela to Honduras (Fig. 144).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4E932EA3F8FCFBFCBBDF44.taxon	description	Figs. 34 (habitus), 39 (aedeagus), 143 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4E932EA3F8FCFBFCBBDF44.taxon	diagnosis	Differential diagnosis. The habitus of E. panamensis (Fig. 34) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus is somewhat similar to that of E. punctipennis (Figs. 39, 62); differing therefrom in having a narrower and longer median lobe, a wider and longer basal piece, and wider and differently shaped parameres. The aedeagus of E. panamensis also has a larger striate collar. Externally (Figs. 34, 58) the two species differ in color (E. panamensis is brown, E. punctipennis is reddish), and E. panamensis has much smaller elytral punctures and larger elytral granules linking the punctures.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4E932EA3F8FCFBFCBBDF44.taxon	distribution	Distribution. Currently known only from Panama (Fig. 143).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4E932EA3F8FA99FB90DDDD.taxon	description	Figs. 35 (habitus), 40 (aedeagus), 128 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4E932EA3F8FA99FB90DDDD.taxon	materials_examined	Type Material. Holotype (male): Panama: Canal Zone, 4.1 mi NW Gamboa, Rio Frijoles, flood debris and wet leaves along river, 9 ° 9.1 ' N, 79 ° 44.2 ' W, 19 ii 1976, A. Newton (MCZ). Paratype: Same data as holotype (1 MCZ).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4E932EA3F8FA99FB90DDDD.taxon	diagnosis	Differential Diagnosis. The habitus of this species (Fig. 35) is very similar to that of other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus (Fig. 40) is very broad and stout; it cannot be confused with any others.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4E932EA3F8FA99FB90DDDD.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 1.55 / 0.78; head (width) 0.45; pronotum 0.58 / 0.59; elytra 1.01 / 0.78. Habitus and sculpture as illustrated (Fig. 35). Head black, dorsum red, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 4 – 5 facets between canthus and posterior margin. Protibiae slightly arcuate. Many elytral punctures not linked by granules. Metaventral depression moderately deep, ca. eight granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4E932EA3F8FA99FB90DDDD.taxon	etymology	Etymology. Named in reference to the very wide and stout aedeagus.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4E932EA3F8FA99FB90DDDD.taxon	distribution	Distribution. Currently known only from the type locality in Panama (Fig. 128).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E419320A3F8F8E6FB90DB99.taxon	description	Figs. 41 (habitus), 45 (aedeagus), 129 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E419320A3F8F8E6FB90DB99.taxon	materials_examined	Type Material. Holotype (male): Panama: Canal Zone, 12.0 mi NW Gamboa, Rio Agua Salud, 9 ° 9.1 ' N, 79 ° 44.2 ' W, 19 ii 1976, A. Newton (MCZ).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E419320A3F8F8E6FB90DB99.taxon	diagnosis	Differential Diagnosis. The habitus of this species (Fig. 41) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus is similar to E. latus in that the parameres are very broad, have the medial margins straight or nearly so, and have two very small setae at the apex of each (Figs. 31, 45). However, the aedeagus of E. latisoides is much smaller, the median lobe is narrower and the apex is shaped slightly differently.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E419320A3F8F8E6FB90DB99.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 1.67 / 0.88; head (width) 0.45; pronotum 0.58 / 0.61; elytra 1.14 / 0.88. Habitus and sculpture as illustrated (Fig. 41). Head black, dorsum red, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 3 – 4 facets between canthus and posterior margin. Protibiae slightly arcuate. Elongate granules linking elytral punctures. Metaventral depression moderately deep, ca. eight granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E419320A3F8F8E6FB90DB99.taxon	etymology	Etymology. Named in reference to the aedeagal similarity to E. latus.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E419320A3F8F8E6FB90DB99.taxon	distribution	Distribution. Currently known only from the type locality in Panama (Fig. 129).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E439322A3F8F8E6FC26D851.taxon	description	Figs. 42 (habitus), 46 (aedeagus), 134 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E439322A3F8F8E6FC26D851.taxon	materials_examined	Type Material. Holotype (male): Panama: Panama Province, Rio Mamoni at PanAm Hwy, black light, 9 ° 8 ' N, 79 ° 4 ' W, 1 ix 2006, W. D. Shepard and D. Post (SEMC). Paratypes (2): Honduras: Rio Humuya NW Comayagua, 14 ° 29 ' N, 87 ° 39 ' W, 3 viii 1967, O. S. Flint, Jr. (1 USNM); Panama: Chiriqui, Rio Vigui (Pan-Amer. Hwy), 8 ° 12 ' N, 81 ° 34 ' W, 28 ix 1976, Edward S. Ross (1 CAS).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E439322A3F8F8E6FC26D851.taxon	diagnosis	Differential Diagnosis. The habitus of this species (Fig. 42) is very similar to that of other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus is similar to that of E. costaricensis in general proportions and size (Figs. 46, 47); differing therefrom in the shape and degree of sclerotization of the median lobe, which is slightly widened in the distal one-third, and has the margin indented slightly at each pore.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E439322A3F8F8E6FC26D851.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 1.34 / 0.69; head (width) 0.38; pronotum 0.52 / 0.52; elytra 0.83 / 0.69. Habitus and sculpture as illustrated (Fig. 42). Head black, dorsum red, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 3 – 4 facets between canthus and posterior margin. Elytra with elongate granules linking punctures. Metaventral depression small.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E439322A3F8F8E6FC26D851.taxon	etymology	Etymology. Named in reference to the conspicuous micropores of the median lobe of the aedeagus.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E439322A3F8F8E6FC26D851.taxon	distribution	Distribution. Currently known from Panama and Honduras (Fig. 134).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E459325A3F8FB2DFC2CDDC7.taxon	description	Figs. 44 (habitus), 48 (aedeagus), 139 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E459325A3F8FB2DFC2CDDC7.taxon	diagnosis	Differential diagnosis. The habitus of E. apocinus (Fig. 44) is quite similar to other members of the Costatus group; reliable determinations will require examination of the male genitalia. The aedeagus has arching parameres somewhat similar to those of E. mucronatus, but otherwise the aedeagi of the two species are quite dissimilar (Figs. 48, 56). The aedeagus of E. apocinus is larger and wider, the median lobe is less acutely pointed apically, and the parameres are arched differently and, in lateral view, are sinuate. Externally, E. apocinus has elongate granules linking the elytral punctures, whereas in E. mucronatus the granules do not link the punctures. E. ballatoris (Fig. 30) also has arcuate parameres that arch over the median lobe; however, the extent of the arching is much more than in E. apocinus. The median lobe of E. ballatoris, which tapers distinctively in the distal one-half also clearly differs from that of E. apocinus. The pronotal lateral lobes are slightly less angulate in E. apocinus than in E. ballatoris (Figs. 26, 44).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E459325A3F8FB2DFC2CDDC7.taxon	distribution	Distribution. Currently known from Costa Rica and Mexico (Fig. 139).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E459325A3F8FF73FBB5DE18.taxon	description	Figs. 43 (habitus), 47 (aedeagus), 141 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E459325A3F8FF73FBB5DE18.taxon	diagnosis	Differential diagnosis. The habitus of E. costaricensis (Fig. 43) is very similar to that of other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus is similar to that of E. microporus in that both have the parameres lobe-like apically; however, the shapes of the lobes differ in the two species (Figs. 46, 47). The median lobe is very simple and parallel-sided in E. costaricensis, whereas in E. microporus it is widened and minutely indented in the distal one-third, at the locations of micropores. Externally, the pronotal hood carinae are shaped slightly differently in the two species (Figs. 42, 43).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E459325A3F8FF73FBB5DE18.taxon	distribution	Distribution. Currently known to be distributed from Panama to Mexico (Fig. 141).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E449327A3F8F8C3FBB1D8E9.taxon	description	Figs. 49 (habitus), 53 (aedeagus), 146 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E449327A3F8F8C3FBB1D8E9.taxon	materials_examined	Type Material. Holotype (male): Costa Rica: Limon, Reventazon, Hamburg Farm, at light, 10 ° 15 ' N, 83 ° 28 ' W, 25 v 1931, F. Nevermann (FMNH). Paratypes: Same data as holotype, except 28 ix 1934, F. Nevermann (2 FMNH).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E449327A3F8F8C3FBB1D8E9.taxon	diagnosis	Differential Diagnosis. The habitus of this species (Fig. 49) is very similar to that of other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus has parameres that are lobate apically, as does E. costaricensis; however, the paramere lobes are shaped differently in the two species (Figs. 47, 53). Also, the median lobe of E. lobilatus tapers in the distal one-half, whereas in E. costaricensis the median lobe is virtually parallel-sided. The median lobe of E. lobilatus is more similar in shape to that of E. apocinus, but in that species the parameres are narrowed apically, not lobate.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E449327A3F8F8C3FBB1D8E9.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 1.32 / ~ 1.00; head (width) 0.55; pronotum 0.75 / 0.77; elytra 1.33 / ~ 1.00. Habitus and sculpture as illustrated (Fig. 49). Head black, dorsum red, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 3 – 4 facets between canthus and posterior margin. Protibiae slightly arcuate. Elytra with elongate granules linking punctures. Metaventral depression moderately deep, ca. six granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E449327A3F8F8C3FBB1D8E9.taxon	etymology	Etymology. Named in reference to the wide apical lobes of the aedeagal parameres.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E449327A3F8F8C3FBB1D8E9.taxon	distribution	Distribution. Currently known only from the type locality in Costa Rica (Fig. 146).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E479327A3F8FD33FC59DC23.taxon	description	Figs. 50 (habitus), 54 (aedeagus), 147 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E479327A3F8FD33FC59DC23.taxon	materials_examined	Type Material. Holotype (male): Costa Rica: Heredia, OTS La Selva Field Sta., Puerto Viejo de Sarapique, Rio Puerto Viejo, at light, 10 ° 26 ' N, 18 ° 59 ' W, 5 – 11 iii 1973, J. Wagner & J. Kethley (FMNH). Paratypes (10): Costa Rica: Cartago, Cimarrones at Rio Cimarrones, black lights, 10 ° 6 ' N, 83 ° 25 ' W, 23 xi 2002, W. Shepard (3 SEMC); Heredia, La Selva Biol. Res. Stn., 3 km S Puerto Viejo, ex. blacklight, elev. 80 m, 10 ° 26 ' N, 84 ° 1 ' W, 4 vi 2001, Chatzimanolis (CR 1 C 01019) (3 SEMC); La Selva Biological Station, HG-vapor lights, 10 ° 25 ' N, 84 ° 1 ' W, 14 – 15 i 2005, A. E. Z. Short (3 SEMC); OTS La Selva Field Sta., Puerto Viejo de Sarapique, Rio Puerto Viejo, at light, 10 ° 26 ' N, 18 ° 59 ' W, 5 – 11 iii 1973, J. Wagner & J. Kethley (1 FMNH).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E479327A3F8FD33FC59DC23.taxon	diagnosis	Differential Diagnosis. This species has a slightly sinuate anterior margin of the pronotal hood, and the submarginal groove of the hood is relatively wide, otherwise the habitus (Fig. 50) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus is small, very narrow, and has narrow, straight parameres (Fig. 54). The median lobe shape could perhaps be compared with that of E. panamensis (Fig. 39), but internally the median lobes are very different, the internal sac being narrower and much more densely spiculate in E. rectus, and the general proportions of the aedeagi are markedly different.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E479327A3F8FD33FC59DC23.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 1.51 / 0.79; head (width) 0.45; pronotum 0.59 / 0.58; elytra 0.97 / 0.79. Habitus and sculpture as illustrated (Fig. 50). Head black, dorsum brown to reddish brown, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 3 – 4 facets between canthus and posterior margin. Protibiae slightly arcuate. Elytra with elongate granules linking punctures. Metaventral depression moderately deep, ca. six granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E479327A3F8FD33FC59DC23.taxon	etymology	Etymology. Named in reference to the straight parameres of the aedeagus.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E479327A3F8FD33FC59DC23.taxon	distribution	Distribution. Currently known only from Costa Rica (Fig. 147).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E479326A3F8F946FBB1DBEA.taxon	description	Figs. 51 (habitus), 55 (aedeagus), 136 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E479326A3F8F946FBB1DBEA.taxon	materials_examined	Type Material. Holotype (male): Costa Rica: Puntarenas, Monteverde area, 10 ° 18 ' N, 84 ° 50 ' W, 6 – 14 vi 1973, Erwin & Hevel (USNM). Paratype: Same data as holotype (1 USNM).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E479326A3F8F946FBB1DBEA.taxon	diagnosis	Differential Diagnosis. The habitus of this species (Fig. 51) is very similar to that of other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus (Fig. 55) has a large, wide median lobe — unlike any other species. The parameres gradually increase in width from base to apex, and slightly arch; each has two small setae apically.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E479326A3F8F946FBB1DBEA.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 1.79 / 1.00; head (width) 0.53; pronotum 0.67 / 0.71; elytra 1.20 / 1.00. Habitus and sculpture as illustrated (Fig. 51). Head black, dorsum brown to reddish brown, venter dark brown to piceous, legs dark brown. Eye with ca. 3 – 4 facets between canthus and posterior margin. Elytra with elongate granules linking punctures. Metaventral depression moderately deep and wide, ca. nine granules along base, shallow.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E479326A3F8F946FBB1DBEA.taxon	etymology	Etymology. Named in reference to the wide median lobe of the aedeagus.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E479326A3F8F946FBB1DBEA.taxon	distribution	Distribution. Currently known only from the type locality in Costa Rica (Fig. 136).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E799319A3F8FC36FBB0DC44.taxon	description	Figs. 57 (habitus), 61 (aedeagus), 145 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E799319A3F8FC36FBB0DC44.taxon	materials_examined	Type Material. Holotype (male): Guatemala: El Progresso, km. 69 on C. A. 9, 14 ° 51 ' N, 90 ° 4 ' W, 2 vii 1974, W. E. Steiner (USNM).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E799319A3F8FC36FBB0DC44.taxon	diagnosis	Differential Diagnosis. The habitus of this species (Fig. 57) is very similar to that of other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus (Fig. 61) has very distinct parameres, wide basally, and in the distal one-fifth markedly narrowed and abruptly arcuate. The median lobe is somewhat similar in shape to that of E. lobilatus (Fig. 53), but in E. acuminatus it is longer, more nearly parallel-sided; also, the two differ internally, as illustrated. The parameres markedly differ in the two species.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E799319A3F8FC36FBB0DC44.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 1.69 / 0.84; head (width) 0.47; pronotum 0.64 / 0.64; elytra 1.08 / 0.84. Habitus and sculpture as illustrated (Fig. 57). Head black, dorsum red, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 3 – 4 facets between canthus and posterior margin. Protibiae not arcuate. Elongate granules linking most elytral punctures. Metaventral depression moderately deep, ca. eight granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E799319A3F8FC36FBB0DC44.taxon	etymology	Etymology. Named in reference to the acuminate apices of the parameres.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E799319A3F8FC36FBB0DC44.taxon	distribution	Distribution. Currently known only from the type locality in Guatemala (Fig. 145).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E799319A3F8FF73FC23D913.taxon	description	Figs. 52 (habitus), 56 (aedeagus), 148 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E799319A3F8FF73FC23D913.taxon	materials_examined	Type Material. Holotype (male): Honduras: Rio Humuya NW Comayagua, 14 ° 29 ' N, 87 ° 39 ' W, 3 viii 1967, O. S. Flint, Jr. (USNM). Paratypes (9): Mexico: Chiapas, Nr. Pijijiapan, 15 ° 41 ' N, 93 ° 13 ' W, 5 vii 1965, Paul J. Spangler (1 USNM); Nayarit, Tepic, light trap, elev. 914 m, 21 ° 30 ' N, 104 ° 54 ' W, 1 viii 1964, Blanton & Broce (1 USNM); Oaxaca, Rio Jaltepec [no date], 17 ° 21 ' N, 95 ° 25 ' W, R. E. Woodruff (1 BMNH); San Luis Potosi, Tamazunchale, 21 ° 16 ' N, 98 ° 48 ' W, 30 v 1948, Werner, Nutting (5 USNM); Veracruz, Puente Nacional, 19 ° 20 ' N, 96 ° 29 ' W, 31 vii 1966, Flint & Ortiz (1 USNM).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E799319A3F8FF73FC23D913.taxon	diagnosis	Differential Diagnosis. The habitus of this species (Fig. 52) is very similar to that of other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus has arching parameres somewhat similar to those of E. apocinus, but otherwise the aedeagi of the two species are quite dissimilar (Figs. 48, 56). The aedeagus of E. mucronatus is smaller and narrower, the median lobe is more acutely pointed apically, and the parameres are arched differently and, in lateral view, lack the sinuations present in E. apocinus.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E799319A3F8FF73FC23D913.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 1.25 / 0.78; head (width) 0.39; pronotum 0.50 / 0.52; elytra 0.88 / 0.78. Habitus and sculpture as illustrated (Fig. 52). Head black, dorsum red, venter light brown, maxillary palpi testaceous. Eye with ca. 3 – 4 facets between canthus and posterior margin. Protibiae slightly arcuate. Elytral punctures each with a granule at anterior and posterior margins, granules usually not linking punctures. Metaventral depression moderately deep, ca. eight granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E799319A3F8FF73FC23D913.taxon	etymology	Etymology. Named in reference to the pointed apex of the median lobe of the aedeagus.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E799319A3F8FF73FC23D913.taxon	distribution	Distribution. Currently known from Mexico and Honduras (Fig. 148).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E799318A3F8F98BFACFDBEA.taxon	description	Figs. 58 (habitus), 62 (aedeagus), 137 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E799318A3F8F98BFACFDBEA.taxon	diagnosis	Differential diagnosis. The habitus of E. punctipennis (Fig. 58) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus is somewhat similar to that of E. panamensis (Figs. 39, 62; differing therefrom in having a wider and shorter median lobe, a narrower and shorter basal piece, and narrower and differently shaped parameres. Externally (Figs. 34, 58) the two species differ in color (E. punctipennis is reddish, E. panamensis is brown), and E. punctipennis has much larger elytral punctures and smaller elytral granules, which do not link the punctures.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E799318A3F8F98BFACFDBEA.taxon	distribution	Distribution. Currently known from northern Mexico, Arizona, Texas, and Oklahoma (Fig. 137).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7B931AA3F8F8E0FB8CDBC1.taxon	description	Figs. 59 (habitus), 63 (aedeagus), 135 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7B931AA3F8F8E0FB8CDBC1.taxon	materials_examined	Type Material. Holotype (male): Mexico: Oaxaca, Tapanatepec, 8 mi. W Oaxaca, elev. 122 m, 16 ° 22 ' N, 94 ° 12 ' W, 10 vii 1953, Univ. Kans. Mex. Expedition (SEMC). Paratypes: Same data as holotype (10 SEMC).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7B931AA3F8F8E0FB8CDBC1.taxon	diagnosis	Differential Diagnosis. The habitus of this species (Fig. 59) is very similar to that of other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus is similar in general proportions to that of E. mucronatus (Figs. 56, 63). However, the median lobe in E. bifidus is apically divided into two lobes; also, the internal sac is different in the two species, as illustrated.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7B931AA3F8F8E0FB8CDBC1.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 1.40 / 0.73; head (width) 0.42; pronotum 0.55 / 0.58; elytra 0.92 / 0.73. Habitus and sculpture as illustrated (Fig. 59). Head black, frons red, clypeus black, dorsum red, venter rufous. Eye with ca. 3 – 4 facets between canthus and posterior margin. Elytra with elongate granules linking punctures. Metaventral depression small.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7B931AA3F8F8E0FB8CDBC1.taxon	etymology	Etymology. Named in reference to the bifid apex of the median lobe of the aedeagus.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7B931AA3F8F8E0FB8CDBC1.taxon	distribution	Distribution. Currently known only from the type locality in Mexico (Fig. 135).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7A931AA3F8FB86FC23DCBA.taxon	description	Figs. 65 (habitus), 69 (aedeagus), 142 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7A931AA3F8FB86FC23DCBA.taxon	diagnosis	Differential diagnosis. The habitus of E. fisheri (Fig. 65) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus is somewhat similar to that of E. costatus in general proportions (Figs. 69, 71), including being broad in lateral view. However, the median lobe shape and the shape of the parameres differ markedly in the two species.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7A931AA3F8FB86FC23DCBA.taxon	distribution	Distribution. Currently known from Honduras and Mexico (Fig. 142).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7A931AA3F8FE1BFB8CDE63.taxon	description	Figs. 60 (habitus), 64 (aedeagus), 135 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7A931AA3F8FE1BFB8CDE63.taxon	materials_examined	Type Material. Holotype (male) :: Mexico: Oaxaca, Tapanatepec, 8 mi. W Oaxaca, elev. 122 m, 16 ° 22 ' N, 94 ° 12 ' W, 10 vii 1953, Univ. Kans. Mex. Expedition (SEMC).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7A931AA3F8FE1BFB8CDE63.taxon	diagnosis	Differential Diagnosis. The habitus of this species (Fig. 60) is very similar to that of other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus has a median lobe shaped somewhat like that of E. spatulus (Figs. 20, 64), but the basal part, between the striate collar and the base, is wider and much shorter. The parameres and basal piece differ markedly in the two species.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7A931AA3F8FE1BFB8CDE63.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 1.63 / 0.86; head (width) 0.47; pronotum 0.66 / 0.67; elytra 1.09 / 0.86. Habitus and sculpture as illustrated (Fig. 60). Head black, dorsum brown to reddish brown, venter reddish brown. Eye with ca. 4 – 5 facets between canthus and posterior margin. Elytra with elongate granules linking punctures. Metaventral depression very small.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7A931AA3F8FE1BFB8CDE63.taxon	etymology	Etymology. Named in reference to the geographical distribution.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7A931AA3F8FE1BFB8CDE63.taxon	distribution	Distribution. Currently known only from the type locality in Mexico (Fig. 135).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7A931CA3F8F8C8FB91DB09.taxon	description	Figs. 66 (habitus), 70 (aedeagus), 138 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7A931CA3F8F8C8FB91DB09.taxon	materials_examined	Type Material. Holotype (male): USA: Arizona, Pajarito Mts., Sycamore Cyn., 31 ° 56 ' N, 110 ° 48 ' W, 24 viii 1968, [no collector on label] (MCZ). Paratypes: Same data as holotype (4 MCZ).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7A931CA3F8F8C8FB91DB09.taxon	diagnosis	Differential Diagnosis. This is a markedly granulate species, with dark brown dorsum; otherwise the habitus (Fig. 66) is very similar to that of other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus has a long median lobe, which is slightly widened at about the distal one-third; it is probably related to E. costatus, which see for comparisons (Figs. 70, 71).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7A931CA3F8F8C8FB91DB09.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 1.79 / 0.94; head (width) 0.50; pronotum 0.64 / 0.69; elytra 1.20 / 0.94. Habitus and sculpture as illustrated (Fig. 66). Head black, dorsum brown to reddish brown, venter and legs dark brown, maxillary palpi brown. Pronotal and elytral granules prominent, including elongate granules linking punctures. Eye with ca. 3 – 4 facets between canthus and posterior margin. Protibiae stout, flattened in distal 1 / 5, outer margin arcuate, inner margin straight. Metaventral depression moderately deep and wide, ca. eight granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7A931CA3F8F8C8FB91DB09.taxon	etymology	Etymology. Named in reference to the geographical distribution.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7A931CA3F8F8C8FB91DB09.taxon	distribution	Distribution. Currently known only from the type locality in Arizona (Fig. 138).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7C931FA3F8FE53FC05DBC1.taxon	description	Figs. 67 (habitus), 71 (aedeagus), 140 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7C931FA3F8FE53FC05DBC1.taxon	diagnosis	Differential diagnosis. The habitus of E. costatus (Fig. 67) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus (Fig. 71) has a very distinctive median lobe, which is markedly widened at the distal one-third. It is nearest in form to the aedeagus of E. arizonicus (Fig. 70), but that species has a much narrower and longer median lobe, which extends further beyond the slightly differently shaped parameres. Externally (Figs. 66, 67), the two species differ in color, E. costatus is reddish and E. arizonicus is brown, and E. costatus has the non-carinate elytral intervals smoother.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7C931FA3F8FE53FC05DBC1.taxon	distribution	Distribution. Currently known from Texas and Arkansas (Fig. 140).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7F931FA3F8FE1BFB2CDED0.taxon	description	Figs. 68 (habitus), 72 (aedeagus), 126 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7F931FA3F8FE1BFB2CDED0.taxon	materials_examined	Type material: Holotype (male): " ARGENTINA, Prov. Tucuman, Dpto. Burruyacú rio Salas 12. II. 1982 Leg. R. Golbach; Epimetopus burruyacu A. Oliva 1984; HOLOTYPO; aedeagus drawn by P. D. Perkins; DIGITAL IMAGE captured 2012, P. D. Perkins " (MACN).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7F931FA3F8FE1BFB2CDED0.taxon	diagnosis	Differential diagnosis. The habitus (Fig. 68) is similar to that of several species in the Costatus group; reliable determinations will be based on examination of the male genitalia. The aedeagus is somewhat similar in general to that of E. angustus (Figs. 28, 72); both have slender, slightly arching parameres. E. burruyacu differs from E. angustus in that the parameres are shorter compared with the length of the basal piece; also, the median lobe is wider and shorter. Externally, E. burruyacu is larger that E. angustus (ca. 1.55 vs. 1.31 mm), has the elytral carinae and pronotal sculpture slightly less developed, and has smaller elytral punctures (Figs. 24, 68).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7F931FA3F8FE1BFB2CDED0.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 1.55 / 0.78; head (width) 0.41; pronotum 0.55 / 0.56; elytra 1.00 / 0.78. Habitus and sculpture as illustrated (Fig. 68). Head piceous, dorsum red, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 3 – 4 facets between canthus and posterior margin. Protibiae slightly arcuate. Elytral granules linking punctures. Metaventral depression moderately deep, ca. six granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7F931FA3F8FE1BFB2CDED0.taxon	distribution	Distribution. Currently known from two narrowly separated localities in Argentina (Fig. 126).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7F9310A3F8F978FB93DB8A.taxon	description	Figs. 73 – 76 (habitus), 77 (aedeagus), 126 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7F9310A3F8F978FB93DB8A.taxon	materials_examined	Type material: Holotype (male; examined): " Colombia " (MNHN). New collection record: Colombia: C. Amara, La Aguadita, 4 ° 36 ' N, 74 ° 5 ' W, 7 vii 1970, J. M. Campbell (1 CNC).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7F9310A3F8F978FB93DB8A.taxon	diagnosis	Differential diagnosis. Distinguished from other members of the Mendeli group by the very ovate elytra (l / w ca. 1.71 / 1.55), the relatively small elytral punctures, and the multiple grooves of the pronotal hood carinae (Figs. 73 – 76). The aedeagus (Fig. 77) is not closely similar to other members of the Mendeli group. The distal part of the median lobe of the aedeagus differs from that of other members: the two apical processes are long, slender, and pointed; the hooks are large and claw-like.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7F9310A3F8F978FB93DB8A.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 2.60 / 1.55; head (width) 0.73; pronotum 0.98 / 1.05; elytra 1.71 / 1.55. Habitus as illustrated (fig. 73). Dorsal surface piceous brown; elytra without hint of fascia; ventral surface piceous black, epipleura reddish brown; antennae reddish brown, maxillary palpi reddish with palpomere 4 brown; trochanters and bases of femora dark brown to brown, distal portions of femora and entire tibiae and tarsi reddish. Body widely elongate, widest at about midlength of elytra, body moderately convex in lateral view, elytra very transversely convex. Eye canthus incomplete, ca. 3 facets between canthus and posterior margin. Pronotum with hood carinae slightly arcuate, confluent anteriorly, grooved throughout, grooves multiple in posterior 1 / 2; primary lateral lobe with anterior angle roundly produced; secondary lateral lobe extremely small or absent. Elytron with four granulate carinae, third interrupted anteriorly for distance of ca. 3 – 4 punctures; first three carinae (from suture) very low and very weakly granulate; punctures round, small, each with moderately large granule at anterior margin, non-carinate intervals with irregular row of small granules. Protibiae straight or very slightly emarginate on inner margin, outer margin weakly arcuate. Metaventral depression with less dense granules than remainder of mesoventrite.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7F9310A3F8F978FB93DB8A.taxon	distribution	Distribution. Currently known from one unambiguous site in Colombia (Fig. 126); the holotype locality data is only " Colombia. " Notes. The holotype (Figs. 73 – 76), which had not previously been dissected, luckily proved to be a male. Two other specimens, one of each gender, were received with the holotype. Neither of these two specimens had any labels, but both were pinned with the same kind of very fine insect pin as the holotype. The second male, which I dissected, is conspecific with the holotype. The female probably is also conspecific. The female differs from the two males in the following: the body size is slightly smaller, the pronotal lateral lobes have the anterior part slightly more indented, the elytral granules are slightly less developed, and the circular transparent serial punctures of the elytra are smaller. The lateral margins of the elytra of the female, in the area slightly anterior to midlength, are straight or slightly emarginate, whereas in males the margins are clearly arcuate; this is probably a sexual dimorphism.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E739315A3F8FF73FB94DBB2.taxon	description	Figs. 78 (habitus), 79, 80 (aedeagus), 126 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E739315A3F8FF73FB94DBB2.taxon	materials_examined	Type material: Holotype (male): " Type [red disc]; Yungas Valley. Bolivia. 12 – 13. vi. 1937; H. E. Hinton Collector; Epimetopus angulatus Type! J. Balfour-Browne det.; Hinton Coll. B. M. 1939 - 583 " (BMNH).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E739315A3F8FF73FB94DBB2.taxon	diagnosis	Differential Diagnosis. The dorsal habitus of E. angulatus is very similar to that of E. peruvianus; E. angulatus is slightly larger (ca. 2.73 vs. 2.62 mm) but otherwise very similar (Figs. 78, 82). Reliable determinations will be based on examination of the aedeagi of the two species, which are markedly dissimilar (Figs. 78, 80, 82). The aedeagus could be compared with that of E. coleuncus (Fig. 85), but differs from that in several respects, including the following: 1) the aedeagus is wider, the lateral processes of the median lobe (the part contacting the parameres) being notably larger; 2) the median lobe apical part is narrower and is emarginate laterally, whereas in E. coleuncus the apical part is wider and arcuate laterally, 3) the hooks rest in a slot on each side of the median lobe, whereas in E. coleuncus the hooks each rest in a groove on the dorsal surface of the median lobe; 4) the parameres are shaped differently; and 5) the basal piece is longer in E. coleuncus. The aedeagus of E. angulatus differs markedly from E. mendeli in overall shape, and many details.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E739315A3F8FF73FB94DBB2.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 2.73 / 1.43; head (width) 0.68; pronotum 0.95 / 1.00; elytra 1.82 / 1.43. Habitus as illustrated (Fig. 78). Head and pronotum dark brown to piceous; elytra brown to dark brown, without fascia; maxillary palpi light brown; venter and coxae piceous, remainder of legs reddish brown. Eye canthus incomplete, ca. 3 facets between canthus and posterior margin. Pronotum with hood carinae slightly arcuate, confluent at anterior tip, grooved throughout; primary lateral lobe with anterior angle produced and acute; secondary lateral lobe extremely small or absent. Elytron with four granulate carinae, third interrupted anteriorly for distance of ca. 3 – 4 punctures; punctures round, small, each with moderately large granule at anterior margin, non-carinate intervals with irregular row of granules. Protibiae straight or very slightly emarginate on inner margin, outer margin weakly arcuate. Metaventral depression small, ca. 8 granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E739315A3F8FF73FB94DBB2.taxon	distribution	Distribution. Currently known only from the type locality in Bolivia (Fig. 126). Notes. Oliva (1986; fig. 4), illustrated an aedeagus supposedly of E. angulatus. However, it is not, and appears instead to perhaps be the aedeagus of E. coleuncus. The aedeagus of the holotype of E. angulatus was illustrated by Fikáček et al. (2011), and a detail of the apex of the same aedeagus is illustrated herein (Fig. 79). Because of the error, and the fact that other species have a habitus similar to that E. angulatus, the distribution records given by Oliva (1986) must be considered invalid until being confirmed by the dissection of males. Epimetopus mendeli Fiká č ek, Barclay & Perkins, 2011 Fig. 126 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E739315A3F8FF73FB94DBB2.taxon	diagnosis	Differential diagnosis. The dorsal habitus is similar to that of E. angulatus, but E. mendeli is considerably larger (ca. 3.10 vs. 2.73 mm), and has more elongate elytra (l / w ratio ca. 1.40 vs. 1.26), and a distinctive elytral fascia (see Figs. in Fikáček et al., 2011). The aedeagus of E. mendeli differs markedly from E. angulatus in overall shape, and many details (Figs. 79, 80). The aedeagus of E. mendeli is somewhat similar to that of E. coleuncus. It differs from E. coleuncus in the following: 1) the shape and length of the parameres, which are shorter and lack the subapical lateral sinuations; 2) the shape of the apical part of the median lobe (more sharply rounded at the tips); and 3) in having narrower lateral processes of the median lobe.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E739315A3F8FF73FB94DBB2.taxon	distribution	Distribution. Currently known only from Peru (Fig. 126). Notes. For a detailed description see Fikáček et al. 2011. Epimetopus flavicaptus Fiká č ek, Barclay & Perkins, 2011 Figs. 81 (habitus), 126 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E739315A3F8FF73FB94DBB2.taxon	materials_examined	Type material: Holotype (male): Ecuador, Napo Province [see data in Fikáček, et al. 2011].	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E739315A3F8FF73FB94DBB2.taxon	diagnosis	Differential diagnosis. Distinguished from other members of the Mendeli group by the transverse or slightly emarginate anterior margin of the pronotal hood (Fig. 81). The aedeagus is illustrated by Fikáček et al. (2011). The distal part of the median lobe is clearly diagnostically different than that of the other members of the Mendeli group.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E739315A3F8FF73FB94DBB2.taxon	distribution	Distribution. Currently known only from the type locality in Ecuador (Fig. 126).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E749309A3F8F8C6FC5CDAB9.taxon	description	Figs. 82 (habitus), 83 (aedeagus), 128 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E749309A3F8F8C6FC5CDAB9.taxon	materials_examined	Type Material. Holotype (male): Peru: Ayacucho, La Mar, Santa Rosa, elev. 640 m, 13 ° 0 ' S, 73 ° 41 ' W, 19 – 25 ix 1976, Robert Gordon (USNM). Paratypes: Same data as holotype (2 USNM); same data, except 8 – 15 ix 1976 (10 USNM).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E749309A3F8F8C6FC5CDAB9.taxon	diagnosis	Differential Diagnosis. The dorsal habitus of E. peruvianus is very similar to that of E. angulatus (Figs. 78, 82); E. peruvianus is slightly smaller (ca. 2.62 vs. 2.73 mm). Reliable determinations will be based on examination of the aedeagi of the two species (Figs. 79, 80, 82). The aedeagus clearly differs from that of other members of the Mendeli group: the distal part of the median lobe is deeply divided into two processes that diverge from one another; the membranous part of the median lobe is very wide; and the parameres are wide at the base, but abruptly narrowed at about midlength, and from there to the apices are parallel.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E749309A3F8F8C6FC5CDAB9.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 2.62 / 1.34; head (width) 0.64; pronotum 0.90 / 0.94; elytra 1.76 / 1.34. Habitus as illustrated (Fig. 82). Head and pronotum piceous, maxillary palpi light brown; elytra brown to dark brown except yellowish fascia slightly past midlength (very faint in some specimens); venter and coxae piceous, remainder of legs dark brown. Eye canthus incomplete, ca. 3 facets between canthus and posterior margin. Pronotum with hood carinae arcuate, confluent at anterior tip, grooved throughout; primary lateral lobe with anterior angle produced; secondary lateral lobe very small, almost absent. Elytron with four granulate carinae, third interrupted anteriorly for distance of ca. 2 – 3 punctures; 4 th carina slightly higher than others; punctures round, moderately large, each with small granule at anterior margin, non-carinate intervals with irregular row of small granules. Protibiae rather long and slender, emarginate on inner margin in distal ½. Metaventral depression moderately deep and wide, granules sparse, rather widely separated, ca. seven granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E749309A3F8F8C6FC5CDAB9.taxon	distribution	Distribution. Currently known only from the type locality in Peru (Fig. 128).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E749309A3F8F8C6FC5CDAB9.taxon	etymology	Etymology. Named in reference to the geographical distribution.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E69930BA3F8F8C9FC22DBC1.taxon	description	Figs. 84 (habitus), 85 (aedeagus), 127 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E69930BA3F8F8C9FC22DBC1.taxon	materials_examined	Type Material. Holotype (male): Argentina: Tucuman, Rio Salas, departamento de Burruyacu, 26 ° 30 ' S, 64 ° 45 ' W, 12 ii 1982, Golbach (MACN). Paratypes (7): Same data as holotype (6 MACN); Bolivia: Tarija, Ing. Bermejo, 21 ° 32 ' S, 64 ° 44 ' W, 14 – 28 ii 1969, R. Golbach (1 MACN).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E69930BA3F8F8C9FC22DBC1.taxon	diagnosis	Differential Diagnosis. Differing externally from other members of the Mendeli group by the pit-like depression at the base of the fused pronotal carinae (Fig. 84). The aedeagus differs from that of E. angulatus (Figs. 79, 80, 85) in several respects, including the following: 1) it is not as wide, the lateral processes of the median lobe being notably smaller; 2) the median lobe apical part is wider, arcuate laterally, whereas in E. angulatus the apical part is emarginate laterally; 3) the hooks each rest in a groove on the dorsal surface of the median lobe, whereas in E. angulatus the hooks rest in a slot on each side; 4) the parameres are shaped differently; and 5) the basal piece is longer in E. coleuncus. It differs from E. mendeli in 1) the shape and length of the parameres, 2) the shape of the apical part of the median lobe (more rounded at the tips), and 3) in having wider lateral processes of the median lobe.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E69930BA3F8F8C9FC22DBC1.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 2.74 / 1.45; head (width) 0.69; pronotum 0.95 / 1.05; elytra 1.90 / 1.45. Habitus as illustrated (Fig. 84). Head and pronotum piceous, maxillary palpi brown; elytra dark brown without hint of fascia; venter black, coxae and basal ½ of femora piceus, remainder of legs dark brown. Eye canthus incomplete, ca. 3 facets between canthus and posterior margin. Pronotum with hood carinae arcuate, completely confluent in anterior ½, grooved throughout; primary lateral lobe with anterior angle produced and acute; secondary lateral lobe very small. Elytron with four carinae, third interrupted anteriorly for distance of ca. 3 – 4 punctures; first three carinae (from suture) very low and very weakly granulate; 4 th carina higher and more strongly granulate; punctures round, large, each with minute granule at anterior margin, non-carinate intervals with unilinear row of minute, well separated granules. Protibiae straight or very slightly emarginate on inner margin, outer margin weakly arcuate. Metaventral depression shallow, ca. six granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E69930BA3F8F8C9FC22DBC1.taxon	etymology	Etymology. Named in reference to the sheathed hooks of the median lobe of the aedeagus.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E69930BA3F8F8C9FC22DBC1.taxon	distribution	Distribution. Currently known from Argentina and Bolivia (Fig. 127).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E6D930FA3F8FF73FE4AD979.taxon	description	Figs. 86 (habitus), 87, 91 (aedeagus), 133 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E6D930FA3F8FF73FE4AD979.taxon	materials_examined	Type material: Holotype (female; examined): “ Hot Sprgs Ar 26.6.01; Schwarz & Barber Coll; Epimetopus thermarum S & B 1917; Type No. 21052 U. S. N. M ” (USNM). Many paratypes with same locality data (USNM). New collection records (219 specimens): Belize: Cayo District, nr Teakettle Bank, Pook’s Hill, Sweep, 17 ° 13 ' N, 88 ° 51 ' W, 6 i 2003, C. R. Bartlett (1 SEMC); Stann Creek District, Nr Armenia at river, 16 ° 48 ' N, 88 ° 29 ' W, 7 i 2003, C. R. Bartlett (2 SEMC); Costa Rica: Guanacaste, 18.7 km NW Bagaces, Hwy 1, Rio Carana, elev. 91 m, 10 ° 39 ' N, 85 ° 22 ' W, 4 vi 1973, Erwin & Hevel (28 USNM); Est. Manglar Ju Villa Nandayure, at light, 10 ° 2 ' N, 85 ° 12 ' W, 11 xii 2002, W. Porras (1 SEMC); Puntarenas, Corcovado National Park, Sirena Stn., ex UV light, elev. 5 m, 8 ° 24 ' N, 83 ° 35 ' W, 28 vi 2000, Z. H. Falin (CR 1 ABF 00 038) (3 SEMC); Guatemala: Baja Verapaz, 4 mi. S. Rabinal, stream, transition xeric-tropical, 15 ° 3 ' N, 90 ° 27 ' W, 10 vi 1974, M. E. & P. D. Perkins (1 MCZ); El Progresso, El Progresso, interm. desert stream, 14 ° 51 ' N, 90 ° 4 ' W, 2 vi 1974, M. E. & P. D. Perkins (1 MCZ); El Progresso, Km. 69 on C. A. 9, 14 ° 51 ' N, 90 ° 4 ' W, 2 vii 1974, W. E. Steiner (2 MCZ); Mexico: Baja California Sur, 1.5 mi. NW Miraflores, collected at blacklight, elev. 213 m, 23 ° 22 ' N, 109 ° 46 ' W, 28 – 29 x 1968, E. L. Sleeper & F. J. Moore (2 CAS); 13 mi. up Canyon San Bernardo fr. Boca de la Sierra — San Bernardo, elev. 609 m, 23 ° 23 ' N, 110 ° 1 ' W, 6 viii 1971, H. G. Real & R. E. Main (1 CAS); 3 mi N Miraflores, Boca de la Sierra, 28 ° 5 ' N, 113 ° 14 ' W, 22 vi 1997, W. D. Shepard (1 NMW); 5 mi. S. Miraflores, 28 ° 1 ' N, 113 ° 14 ' W, 10 vii 1938, Michelbacher & Ross (1 FMNH); 5 mi W San Bartolo, 23 ° 44 ' N, 109 ° 57 ' W, 13 vii 1938, Michelbacher & Ross (1 FMNH, 9 CAS); 8 mi. SE La Paz, collected at blacklight, elev. 304 m, 24 ° 5 ' N, 110 ° 13 ' W, 13 x 1968, E. L. Sleeper & F. J. Moore (1 CAS); 9 mi. N San Lucas, collected at blacklight, elev. 304 m, 27 ° 17 ' N, 112 ° 17 ' W, 24 – 25 x 1968, E. L. Sleeper & F. J. Moore (1 CAS); 9 mi. up Canyon San Bernardo fr. Boca de la Sierra (Potrero), elev. 457 m, 23 ° 23 ' N, 109 ° 57 ' W, 2 viii 1971, H. G. Real & R. E. Main (2 FMNH, 98 CAS); Sa. Victoria, 7 mi. W Santiago, collected at blacklight, 23 ° 28 ' N, 109 ° 50 ' W, 30 – 31 x 1968, E. L. Sleeper & F. J. Moore (10 CAS); Sa. Victoria, El Chorro, collected at blacklight, elev. 304 m, 23 ° 28 ' N, 109 ° 50 ' W, 29 – 30 x 1968, E. L. Sleeper & F. J. Moore (4 CAS); Chiapas, Chintul, Rio Chintul (trib. R. Grijalva), Rt. 187, 17 ° 14.55 ' N, 93 ° 35.11 ' W, 20 ix 1966, Bueno, Barba, Contreras (1 USNM); Municipio de Cintalapa La Mina, elev. 914 m, 16 ° 39 ' N, 93 ° 44 ' W, 14 ix 1981, D. E. & P. M. Breedlove and C. G. Whitefield (3 CAS); Palenque, 17 ° 31 ' N, 91 ° 59 ' W, 22 – 23 vi 1969, D. Bright & J. M. Campbell (6 CNC); Jalisco, 25 mi NW of Barra de Navidad, intermittent stream, scrub tropics, 19 ° 28 ' N, 104 ° 56 ' W, 22 vii 1974, M. E. & P. D. Perkins (1 MCZ); SE of Chamela, Rio Cuitzmala, elev. 55 m, 19 ° 32 ' N, 105 ° 4 ' W, 6 i 2005, W. D. Shepard (1 SEMC); Nayarit, 32 miles S. Acaponeta, 22 ° 1 ' N, 105 ° 18 ' W, 24 xi 1948, H. B. Leech (1 CAS); Sinaloa, 21 mi. E. Villa Union, 23 ° 12 ' N, 105 ° 55 ' W, 25 vii 1964, H. F. Howden (2 CNC); Concordia, 23 ° 17 ' N, 106 ° 4 ' W, 1 – 30 vii 1974, D. Chandler (1 USNM); Culiacan, 24 ° 48 ' N, 107 ° 23 ' W, 16 vii 1963, P. J. Spangler (1 USNM); Mazatlan, blk. lite trap, 23 ° 14 ' N, 106 ° 24 ' W, 17 – 23 vii 1963, P. J. Spangler (2 USNM); Sonora, 6 mi. W Alamos, 27 ° 1 ' N, 109 ° 2 ' W, 14 viii 1963, Werner & Nutting (1 USNM); Alamos, 27 ° 1 ' N, 108 ° 56 ' W, 12 viii 1960, P. H. Arnaud, Jr., E. S. Ross, D. C. Rentz (1 CAS); same locality, 27 ° 1 ' N, 108 ° 56 ' W, 17 ix 1980, Robert Gordon (1 USNM); Rio Cuchujaqui, 7 mi. SE Alamos, black lite, 26 ° 57 ' N, 108 ° 51 ' W, 25 x 1972, E. M. Fisher (2 MCZ); Panama: Panama Province, Parque Nac. Soberania, Pipeline Rd., ex. light, 9 ° 4 ' N, 79 ° 40 ' W, 9 vi 1993, M. L. Jameson (2 SEMC); U. S. A.: Texas, Cypress Mills [no date or collector], 29 ° 59 ' N, 95 ° 40 ' W, (1 MCZ); Venezuela: Aragua, Ocumare, 10 ° 7 ' N, 66 ° 46 ' W, 19 – 20 ii 1969, P. & P. Spangler (5 USNM); Barinas, Pte. Parangula, 8 km S. Barinitas, 8 ° 41 ' N, 70 ° 25 ' W, 18 ii 1976, C. M. & O. S. Flint, Jr. (1 USNM); Guarico, ~ 15 km S. San Juan, stream margin & rock pools, elev. 255 m, 9 ° 46.321 ' N, 67 ° 21.201 ' W, 3 i 2006, A. E. Z. Short (AS- 06 - 005) (3 AEZS); Trujillo, Mcp. Rafael Rangel, La Gira, Qda. La Amarilla, elev. 520 m, 9 ° 20 ' N, 70 ° 44 ' W, 10 – 12 vi 1994, J. Camacho, M. Garcia (1 MALUZ); Mcp. Rafael Rangel, La Gira, Qda. La Amarilla, Trampa interceptacion, elev. 520 m, 9 ° 20 ' N, 70 ° 44 ' W, 20 – 22 v 1995, J. Camacho, M. Garcia (3 MALUZ); Mcpio. Rafael Rangel, La Gira, Trampa Interceptacion, elev. 520 m, 9 ° 20 ' N, 70 ° 44 ' W, 16 – 17 ix 1995, M. Garcia, D. Contreras (2 MALUZ); Same locality, 23 x 1997, J. Camacho, M. Garcia, E. Gomez (3 MALUZ); Zulia, Mcpio. Lagunillas, Zona de Reserva de Burro Negro, elev. 250 m, 10 ° 7 ' N, 71 ° 14 ' W, 29 – 31 vii 1995, J. Comacho, M. Garcia (3 MALUZ).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E6D930FA3F8FF73FE4AD979.taxon	diagnosis	Differential diagnosis. The dorsal habitus of E. thermarum is quite similar to that of E. balfourbrownei (Figs. 86, 92); differing therefrom in the larger size (ca. 2.73 vs. 2.16 mm) and the slightly more elongate elytra which are slightly more convex and have a more abrupt posterior declivity. However, reliable determinations will require examination of the aedeagus. The aedeagus of E. thermarum is recognized by the following combination of characters: 1) the very long and very slender dorsal processes; 2) the slender median lobe, which apically is comparatively long and with narrow, pointed tips; and 3) the parameres bend outward slightly and have truncate tips. There is some variation in aedeagal form (Figs. 87, 91) across the rather large geographical distribution of E. thermarum, which ranges from Arizona to Venezuela (Fig. 133). This variation mostly involves the degree to which the median lobe distal part is narrowed, elongated, and produced in sharp tips. The most extreme variant in this respect is from Venezuela, but other specimens from Venezuela, which have a shape more like those from Panama (Fig. 91), have been seen.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E6D930FA3F8FF73FE4AD979.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 2.73 / 1.45; head (width) 0.68; pronotum 0.91 / 1.05; elytra 1.86 / 1.45. Habitus as illustrated (Fig. 86). Head piceous, maxillary palpi light brown; pronotum dark brown to piceous; elytra dark brown except yellowish large Vshaped fascia near midlength, and at apical 1 / 3 of elytra, which joins fascia laterally; venter piceous; legs rufous. Pronotum with hood carinae very slightly arcuate, almost straight, converging and confluent anteriorly, grooved throughout; secondary lateral lobe very small. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 3 – 4 punctures; punctures round or oval, large, each with small granule at anterior margin, non-carinate intervals usually with irregular row of granules. Protibiae slightly emarginate on inner margin, almost straight. Metaventral depression moderately deep and wide, ca. seven granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E6D930FA3F8FF73FE4AD979.taxon	distribution	Distribution. The most widely distributed species of Epimetopus, currently known from Venezuela north to Texas and southern Arizona (Fig. 133). Notes. Specimens generally have an irregular row of granules on the non-carinate elytral intervals. These granules are usually somewhat larger in specimens from Arizona than in specimens from areas south. A small percentage of specimens from Costa Rica and Venezuela lack these granules. There is little variation in size throughout the range.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E6F930FA3F8FC83FAC9DD83.taxon	description	Figs. 92, 93 (habitus), 89, 90 (aedeagus), 127 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E6F930FA3F8FC83FAC9DD83.taxon	materials_examined	Type material: Holotype (male): Brazil, Mato Grosso, " Jacaré P. N. Xingu M. T. XI- 1965, M. Alvarenga; [male symbol]; 3; [label with measurements]; Epimetopus balfourbrownei A. A. Rocha det.; HOLOTYPO [red label] " (MSP). Epimetopus leechi (new synonymy): Holotype (male): " BRAZIL: Matto Grosso side of Rio Araguaia, Santa Isabel; VIII- 10 to 20 – 57 Borys Malkin; 2; [male symbol]; [label with measurements]; HOLOTIPO [red]; Epimetopus leechi A. A. Rocha det.; California Academy of Sciences Type No. 10192 / " (CAS). New collection records: Four specimens with same locality data as holotype of E. balfourbrownei (3 FMNH, 1 USNM).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E6F930FA3F8FC83FAC9DD83.taxon	diagnosis	Differential diagnosis. The dorsal habitus of E. balfourbrownei is quite similar to that of E. thermarum (Figs. 86, 92); differing therefrom in the smaller size (ca. 2.16 vs. 2.73 mm) and the slightly less elongate elytra which are slightly less convex and have a less abrupt posterior declivity. However, reliable determinations will require examination of the aedeagus. The aedeagus is similar to that of E. thermarum, but notably differs in the following: 1) the aedeagus is considerably shorter (ca. 0.65 vs. 0.80 mm); 2) the dorsal processes are wider, stouter; 3) the median lobe distal part that extends beyond the tips of the dorsal processes is shorter, wider, and less acutely pointed apically; and 4) the basal piece is shorter. The aedeagus of the holotype of E. leechi (Fig. 89) is virtually identical to the aedeagus of the holotype of E. balfourbrownei (Fig. 90); E. leechi is therefore herein synonomized with E. balfourbrownei.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E6F930FA3F8FC83FAC9DD83.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 2.16 / 1.17; head (width) 0.62; pronotum 0.78 / 0.90; elytra 1.38 / 1.17. Habitus as illustrated (Figs. 92, 93). Head piceous, maxillary palpi light brown; pronotum dark brown to piceous; elytra dark brown except yellowish large V-shaped fascia near midlength, and very faintly yellowish apically; venter piceous; legs rufous. Pronotum with hood carinae straight, converging and confluent anteriorly, grooved throughout; secondary lateral lobes very small, almost absent. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 3 – 4 punctures; punctures round or oval, large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae slightly emarginate on inner margin, almost straight. Metaventral depression moderately deep and wide, ca. nine granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E6F930FA3F8FC83FAC9DD83.taxon	distribution	Distribution. Currently known from two rather narrowly separated localities in Brazil (Fig. 127).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E6E9300A3F8F911FBA3DBC1.taxon	description	Figs. 94 (habitus), 88 (aedeagus), 128 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E6E9300A3F8F911FBA3DBC1.taxon	materials_examined	Type Material. Holotype (male): Paraguay: Dep. Concepcion, Arroyo Toro Paso, Unterlauf (contrib. Rio Apa), seepage, 23 ° 51 ' S, 56 ° 46 ' W, 24 x 2002, U. Drechsel (NMW). Paratype: Same data as holotype (1 NMW).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E6E9300A3F8F911FBA3DBC1.taxon	diagnosis	Differential Diagnosis. The dorsal habitus (Fig. 94) differs from that of other members of the Thermarum group by the elytral carinae which, in dorsal view, appear arcuate and, in lateral view, appear sinuate posteriorly. The elytral punctures are quite large and round. The pronotal lateral lobes are slightly more angulate that those of the other members of the group. The aedeagus is similar to that of E. thermarum (Figs. 87, 88), but notably differs in the following: 1) the median lobe is wider, and less acutely pointed apically; 2) the lateral margins of the parameres are virtually parallel, not bending outward; and 3) the median lobe distal part that extends beyond the tips of the dorsal processes is shorter (best seen in lateral view).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E6E9300A3F8F911FBA3DBC1.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 2.11 / 1.11; head (width) 0.55; pronotum 0.75 / 0.86; elytra 1.34 / 1.11. Habitus as illustrated (Fig. 94). Head piceous, maxillary palpi light brown; pronotum brown; elytra brown to dark brown except very faint lighter fascia in front of middle, and one behind middle (specimen may be slightly teneral); venter and coxae piceous, remainder of legs brown to reddish brown. Head (male) with sharply upturned clypeus. Pronotum with hood carinae slightly arcuate, converging and confluent anteriorly, grooved throughout; secondary lateral lobe small. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 3 punctures; punctures round or oval, large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae with inner margin straight, outer margin arcuate. Metaventral depression shallow, moderately wide, ca. seven granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E6E9300A3F8F911FBA3DBC1.taxon	etymology	Etymology. Named in reference to the arcuate elytral carinae.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E6E9300A3F8F911FBA3DBC1.taxon	distribution	Distribution. Currently known only from the type locality in Paraguay (Fig. 128).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E639305A3F8FF73FB92DBC1.taxon	description	Figs. 95 (habitus), 97 (aedeagus), 126 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E639305A3F8FF73FB92DBC1.taxon	materials_examined	Type Material. Holotype (male): Guyana: Region 8, Iwokrama Forest, 1 km W Kurupukan, Iwokrama Field Stn., ex. mercury vapor / UV light, elev. 60 m, 4 ° 40 ' N, 58 ° 41 ' W, 20 v 2001, R. Brooks, Z. Falin (GUY 1 BF 01 001) (SEMC).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E639305A3F8FF73FB92DBC1.taxon	diagnosis	Differential Diagnosis. The dorsal habitus (Fig. 95) is very similar to that of other members of the Thermarum group; the pronotal lateral lobes are slightly smaller than is usual in the group. Males are distinguished from other members of the group by the distinctively upturned and medially pointed clypeus. The aedeagus (Fig. 97) differs from that of other members of the group in several respects: 1) the dorsal processes converge slightly, and at the tips are produced in a sharp point directed medially; 2) the median lobe distal part is widened and is deeply cleft in the midline; and 3) each paramere is produced in a point apically, and bear two microsetae on the point.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E639305A3F8FF73FB92DBC1.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 1.92 / 1.05; head (width) 0.53; pronotum 0.68 / 0.73; elytra 1.30 / 1.05. Head piceous, maxillary palpi light brown; pronotum brown; elytra brown except very faint lighter fascia near middle, (specimen may be slightly teneral); venter and coxae piceous, remainder of legs brown to reddish brown. Head (male) with sharply upturned clypeus. Pronotum with hood carinae very slightly arcuate, converging and confluent anteriorly, grooved throughout; secondary lateral lobe small. Elytron with four strong carinae, weakly granulate, third interrupted anteriorly for distance of ca. 3 punctures; punctures round or oval, small, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae with inner margin straight, outer margin arcuate. Metaventral depression moderately wide, ca. seven granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E639305A3F8FF73FB92DBC1.taxon	etymology	Etymology. Named in reference to the modified male clypeus.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E639305A3F8FF73FB92DBC1.taxon	distribution	Distribution. Currently known only from the type locality in Guyana (Fig. 126).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E649304A3F8FF73FBA6D9AB.taxon	description	Figs. 96 (habitus), 98 (aedeagus), 126 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E649304A3F8FF73FBA6D9AB.taxon	materials_examined	Type Material. Holotype (male): Suriname: Sipaliwini District, Camp 3, Wehepai; 2010 CI-RAP Survey, sandy forest creek, elev. 237 m, 2 ° 21.776 ' N, 56 ° 41.861 ' W, 4 – 6 ix 2010, Short & Kadosoe (SR 10 - 0904 - 01 A) (SEMC). Paratypes: Same data as holotype (2 SEMC). 77	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E649304A3F8FF73FBA6D9AB.taxon	diagnosis	Differential Diagnosis. The dorsal habitus of E. surinamensis is very similar to that of E. clypeatus (Figs. 95, 96); differing therefrom in the larger size (ca. 2.44 vs. 1.92 mm), and the unmodified male clypeus. The aedeagi of the two species differ markedly (Figs. 97, 98). The aedeagus differs from that of other members of the Thermarum group in several respects: 1) the dorsal processes are relatively large, arch away from one another, and do not have ridges on the tips; 2) the median lobe distal part is notably widened, and strongly sclerotized on each side of the median slit; and 3) the parameres gradually decrease in width, to a narrow tip.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E649304A3F8FF73FBA6D9AB.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 2.44 / 1.32; head (width) 0.64; pronotum 0.88 / 0.96; elytra 1.60 / 1.32. Habitus as illustrated (Fig. 96). Head piceous, maxillary palpi light brown; pronotum dark brown to piceous; elytra dark brown except yellowish at Vshaped fascia near midlength, and second fascia at posterior declivity; venter and coxae black, remainder of legs brown to reddish brown. Pronotum with hood carinae slightly arcuate, converging and confluent anteriorly, grooved throughout; secondary lateral lobe small. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 3 – 4 punctures; punctures round or oval, large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae slightly emarginate on inner margin, outer margin arcuate. Metaventral depression moderately wide, ca. seven granules along base, shallow.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E649304A3F8FF73FBA6D9AB.taxon	etymology	Etymology. Named in reference to the geographical distribution.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E649304A3F8FF73FBA6D9AB.taxon	distribution	Distribution. Currently known only from the type locality in Suriname (Fig. 126).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E649306A3F8FA09FBFFDBEA.taxon	description	Figs. 99 (habitus), 103 (aedeagus), 126 (map) Georyssus plaumanni Costa Lima, 1954: 177.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E649306A3F8FA09FBFFDBEA.taxon	materials_examined	Type material: Holotype (male): “ Brasilien, Nova Teutonia, 27 ° 11 ’ B – 52 ° 23 ’ l, 300 – 500 m, IX 1952, Fritz Plaumann; Georyssus plaumanni n. sp. Costa Lima det.; TIPO [red]; Epimetopus plaumanni n. comb. Costa Lima 1954, A. A. Rocha det.; 10.989; HOLOTIPO [red] ” (MSP). New collection record: Brazil: Santa Catarina, Nova Teutonia, 27 ° 3 ' S, 52 ° 24 ' W, 1 – 30 i 1971, F. Plaumann (1 MCZ).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E649306A3F8FA09FBFFDBEA.taxon	diagnosis	Differential diagnosis. The dorsal habitus of E. plaumanni (Fig. 99) is very similar to that of other members of the Plaumanni group. It differs from the others in the larger body size (ca. 2.50 mm) and the fact that males do not have the clypeus upturned. Dissection of males will be necessary for reliable identifications. The aedeagus is similar in size and general proportions to that of E. vianai (Figs. 103, 104); it differs therefrom in the following: 1) the dorsal processes are shaped differently, being smaller and not having the apices turning outward; 2) the length of the median lobe above the dorsal processes is greater; 3) the apex of the median lobe is shaped differently; and 4) the apices of the parameres are shaped differently, in E. plaumanni being produced in a point on the medial side, a character not seen in the other species. There are also differences in the internal structure of the median lobe in the two species, E. plaumanni having the internal sac narrower and much more densely spiculate.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E649306A3F8FA09FBFFDBEA.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 2.50 / 1.25; head (width) 0.65; pronotum 0.88 / 0.96; elytra 1.69 / 1.25. Habitus as illustrated (Fig. 99). Head piceous, maxillary palpi light brown; pronotum dark brown to piceous; elytra dark brown except yellowish fascia near midlength; venter piceous; legs rufous. Pronotum with hood carinae very slightly arcuate, converging and confluent anteriorly, grooved throughout; secondary lateral very small, almost absent. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 3 punctures; punctures round or oval, large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae slightly emarginate on inner margin, almost straight. Metaventral depression moderately deep and wide, ca. nine granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E649306A3F8FA09FBFFDBEA.taxon	distribution	Distribution. Currently known only from the type locality in Brazil (Fig. 126).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E199379A3F8FF73FEE3DE92.taxon	description	Figs. 100 (habitus), 104 (aedeagus), 126 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E199379A3F8FF73FEE3DE92.taxon	materials_examined	Type material: Holotype (male; examined): " CORDOBA — ARGENTINA Dep. de Calamuchita " El Sauce " XII - 1938, Manuel J. Viana; 17 - [green label]; Type [red disc]; Brit. Mus. 1966 - 473; Epimetopus vianai Type J. Balfour- Browne det. " (BMNH).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E199379A3F8FF73FEE3DE92.taxon	diagnosis	Differential diagnosis. The dorsal habitus of E. vianai (Fig. 100) is very similar to that of other members of the Plaumanni group. It differs from the others in having a slightly more elongate pronotum. Males have the clypeus upturned, as do E. vulpinus and E. multiportus. Examination of the male genitalia will be essential for reliable determinations. The aedeagus is similar in size and general proportions to that of E. plaumanni (Figs. 103, 104); it differs therefrom in the following: 1) the dorsal processes are shaped differently, being larger and having the apices turning outward and sharply pointed at the tip, a character not seen in the other species; 2) the length of the median lobe above the dorsal processes is less; 3) the apex of the median lobe is shaped differently, being more deeply bifurcate in E. vianai; and 4) the apices of the parameres are shaped differently, in E. vianai being more truncate, and in E. plaumanni being produced in a point on the medial side. There are also differences in the internal structure of the median lobe in the two species, E. vianai having the internal sac wider and much less densely spiculate.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E199379A3F8FF73FEE3DE92.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 2.44 / 1.20; head (width) 0.62; pronotum 0.88 / 0.90; elytra 1.60 / 1.20. Habitus as illustrated (Fig. 100). Head piceous, maxillary palpi light brown; pronotum dark brown to piceous; elytra dark brown except yellowish, large V-shaped fascia near midlength, and faintly yellowish at apical 1 / 3 of elytra; venter and coxae piceous, remainder of legs reddish brown; legs rufous. Pronotum with hood carinae very slightly arcuate, almost straight, converging and confluent anteriorly, grooved throughout; secondary lateral very small, almost absent. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 3 punctures; punctures round or oval, large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae straight on inner margin, outer margin weakly arcuate. Metaventral depression moderately large, ca. 9 granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E199379A3F8FF73FEE3DE92.taxon	distribution	Distribution. Currently known from Argentina (Fig. 126). Oliva (1986) reported specimens from several Provinces in Argentina; these specimens were not seen during this revision, and therefore could not be confirmed as E. vianai.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E199378A3F8FAB0FB9ADB09.taxon	description	Figs. 101 (habitus), 105 (aedeagus), 126 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E199378A3F8FAB0FB9ADB09.taxon	materials_examined	Type Material. Holotype (male): Brazil: Rio Grande do Sul, Pelotas, 31 ° 46 ' S, 52 ° 20 ' W, 28 xi 1952, C. Biezanko (USNM). Paratype: Uruguay: Lavalleja, Rio Cebollati, Picada de Rodriguez, 33 ° 32 ' S, 54 ° 23 ' W, 5 i 1957, C. S. Carbonell (1 USNM).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E199378A3F8FAB0FB9ADB09.taxon	diagnosis	Differential Diagnosis. The dorsal habitus of E. vulpinus (Fig. 101) is very similar to that of other members of the Plaumanni group. It differs from the others in having slightly more ovate elytra, and usually a larger V-shaped fascia on the elytra. It is the smallest member of the group (ca. 2.17 mm). Males have the clypeus upturned, as do E. vianai and E. multiportus. Examination of the male genitalia will be essential for reliable determinations. The aedeagus (Fig. 105) is much broader (in both views) than others in the species group; the dorsal processes are shaped differently and are broader basally than those of other species; and the median lobe above the bases of the dorsal processes is not parallel-sided.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E199378A3F8FAB0FB9ADB09.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 2.17 / 1.19; head (width) 0.59; pronotum 0.81 / 0.87; elytra 1.44 / 1.19. Habitus as illustrated (Fig. 101). Head piceous, maxillary palpi light brown; pronotum dark brown to piceous; elytra dark brown except yellowish on carinae in basal ¼, large V-shaped fascia near midlength, and at apical ¼ of elytra; venter piceous; legs rufous. Pronotum with hood carinae very slightly arcuate, almost straight, converging and confluent anteriorly, grooved throughout; secondary lateral very small, almost absent. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 3 punctures; punctures round or oval, large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae slightly emarginate on inner margin, almost straight. Metaventral depression moderately deep and wide, ca. nine granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E199378A3F8FAB0FB9ADB09.taxon	etymology	Etymology. Named in reference to the dorsal processes of the median lobe of the aedeagus, which are shaped like fox ears.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E199378A3F8FAB0FB9ADB09.taxon	distribution	Distribution. Currently known from southeastern Brazil and Uruguay (Fig. 126).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E1B937DA3F8F8ECFBA1D856.taxon	description	Figs. 102 (habitus), 106 (aedeagus), 127 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E1B937DA3F8F8ECFBA1D856.taxon	materials_examined	Type Material. Holotype (male): Uruguay: 90 km SW Artigas, 31 ° 1 ' S, 57 ° 52 ' W, 27 – 30 ix 2001, R. Linek (NMW). Paratypes (32): Paraguay: Dep. Concepcion, Arroyo Toro Paso, Unterlauf (contrib. Rio Apa), seepage, 23 ° 51 ' S, 56 ° 46 ' W, 24 x 2002, U. Drechsel (19 NMW); Uruguay: 90 km SW Artigas, 31 ° 1 ' S, 57 ° 52 ' W, 27 – 30 ix 2001, R. Linek (13 NMW).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E1B937DA3F8F8ECFBA1D856.taxon	diagnosis	Differential Diagnosis. The dorsal habitus of E. multiportus (Fig. 102) is very similar to that of other members of the Plaumanni group. Males have the clypeus upturned, as do E. vianai and E. vulpinus. Examination of the male genitalia will be essential for reliable determinations. The aedeagus (Fig. 106) is smaller than that of other members of the species group (ca. 0.66 vs. 0.76 mm), and the apices of the dorsal processes fit into indentations in the median lobe, a character not seen in the other species.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E1B937DA3F8F8ECFBA1D856.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 2.30 / 1.13; head (width) 0.59; pronotum 0.80 / 0.85; elytra 1.48 / 1.13. Habitus as illustrated (Fig. 102). Head piceous, maxillary palpi light brown; pronotum dark brown to piceous; elytra dark brown except yellowish, large V-shaped fascia near midlength, and at apical ¼ of elytra; venter and coxae piceous, remainder of legs dark brown. Pronotum with hood carinae very slightly arcuate, converging and confluent anteriorly, grooved throughout; secondary lateral very small, almost absent. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 3 punctures; punctures round or oval, large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae with inner margin straight, outer arcuate. Metaventral depression moderately deep, ca. seven granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E1B937DA3F8F8ECFBA1D856.taxon	distribution	Distribution. Currently known from Paraguay and Uruguay (Fig. 127).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E1B937DA3F8F8ECFBA1D856.taxon	etymology	Etymology. Named in reference to the transparent, porthole-like elytral punctures.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E1D937DA3F8FBDAFC17DDB4.taxon	materials_examined	Type Material. Holotype (male): Brazil: Mato Grosso, Caceres, MT., 16 ° 4 ' S, 57 ° 41 ' W, 1 – 30 xii 1955, M. Alvarenga (MSP). Paratype: Brazil: Mato Grosso, Caceres, MT., 16 ° 4 ' S, 57 ° 41 ' W, 2 – 4 xii 1955, M. Alvarenga (1 CAS).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E1D937DA3F8FBDAFC17DDB4.taxon	diagnosis	Differential Diagnosis. Differentiated from E. lanceolatulus (Figs. 107, 108) by the much larger size (ca. 3.63 vs. 2.61 mm), the longer and more sharply arcuate pronotal hood, the different spearhead-like shape formed by the pronotal carinae, the presence of a pronotal ridge leading posteriorly from the posterior margin of the pronotal hood, and more robust elytra. In addition, the pronotal hood carinae are anteriorly confluent in E. lanceolatus, and separated in E. lanceolatulus. The aedeagus is notably broader than that of E. lanceolatulus (Figs. 109, 110), but despite the great difference in size of adults of the two species the aedeagi are of approximately the same length. Also, the parameres are shaped differently in the two species.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E1D937DA3F8FBDAFC17DDB4.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 3.63 / 2.03; head (width) 0.94; pronotum 1.34 / 1.44; elytra 2.44 / 2.03. Habitus as illustrated (Fig. 107). Head dark brown to piceous, maxillary palpi light brown, dorsum dark brown except light brown narrow transverse fascia on elytra, venter and coxae dark brown to piceous, remainder of legs brown, abdominal ventrites brown, margin slightly lighter. Pronotum with primary lateral lobes large, angulate, secondary lobes small; hood anterior margin rather sharply arcuate; hood carinae anteriorly converging, confluent at anterior tip, together forming very sharp angle; pronotal ridges strong. Elytra with carinae strong, except third carina absent in short area subbasally; non-carinate intervals each with an irregular row of small granules. Metaventral depression moderately deep and wide, ca. eight large granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E1D937DA3F8FBDAFC17DDB4.taxon	etymology	Etymology. Named in reference to the shape of the aedeagus.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E1D937DA3F8FBDAFC17DDB4.taxon	distribution	Distribution. Currently known only from the type locality (Fig. 128).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E1C937EA3F8F8C3FC09DBC1.taxon	description	Figs. 108 (habitus), 110 (aedeagus), 128 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E1C937EA3F8F8C3FC09DBC1.taxon	materials_examined	Type Material. Holotype (male): Brazil: Mato Grosso, Caceres, MT., 16 ° 4 ' S, 57 ° 41 ' W, 1 – 30 xii 1955, M. Alvarenga (MSP). Paratypes (11): Brazil: Mato Grosso, Corumba, 17 ° 32 ' S, 56 ° 27 ' W, 14 – 19 xii 1919 (3 MCZ); Paraguay: Ita, 25 ° 29 ' S, 57 ° 21 ' W, 12 iv 1980, Spangler et al. (6 USNM); Chaco, P. N. Defensores del Chaco, Madrejon, at black light, 20 ° 38 ' S, 59 ° 52 ' W, 10 xii 1981, J. A. Kochalka (1 MCZ); Dep. San Pedro, [Estancia] Carumbé, 25 ° 33 ' S, 56 ° 40 ' W, 1 – 30 i 1971, R. Goldbach (1 MACN).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E1C937EA3F8F8C3FC09DBC1.taxon	diagnosis	Differential Diagnosis. Differentiated from E. lanceolatus (which see) by the much smaller body size (ca. 2.61 vs. 3.63 mm), several characters of the pronotum and elytra, and form of the aedeagus.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E1C937EA3F8F8C3FC09DBC1.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 2.61 / 1.33; head (width) 0.72; pronotum 0.85 / 1.04; elytra 1.74 / 1.33. Habitus as illustrated (Fig. 108). Head dark brown to piceous, maxillary palpi light brown, dorsum dark brown except light brown narrow transverse fascia on elytra, venter dark brown to piceous, legs dark brown. Pronotum with primary lateral lobes large, more angulate anteriorly than laterally, secondary lobes small; hood anterior margin not sharply arcuate; hood carinae very straight, not confluent at anterior tip; pronotal ridges only moderately strong. Elytra with carinae strong, except third carina absent in short area subbasally; non-carinate intervals without granules. Metaventral depression moderately deep, wide, ca. 11 granules along base; granules lateral to depression very coarse.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E1C937EA3F8F8C3FC09DBC1.taxon	etymology	Etymology. Named in reference to the shape of the aedeagus, and relatively small size, compared to E. lanceolatus.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E1C937EA3F8F8C3FC09DBC1.taxon	distribution	Distribution. Currently known from Brazil and Paraguay (Fig. 128).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E119370A3F8FD56FCC3DB09.taxon	description	Figs. 111 (habitus), 112 (aedeagus), 126 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E119370A3F8FD56FCC3DB09.taxon	materials_examined	Type material: Holotype (male): " Type H. T. [red disc]; Amer. mer.; Sharp Coll. 1905 - 313; Aedeagus drawn by P. D. Perkins 1993 " (BMNH). New collection records: Brazil: Mato Grosso, Caceres, MT., 16 ° 4 ' S, 57 ° 41 ' W, 2 – 4 xii 1955, M. Alvarenga (1 CAS); Jacaré, Parque Nacional do Xingu, 15 ° 16 ' S, 57 ° 42 ' W, 1 – 30 xi 1965, M. Alvarenga & Werner (1 FMNH). Sao Paulo, Piracicaba, collected at black light, 22 ° 44 ' S, 47 ° 39 ' W, 7 x 1965, C. A. Triplehorn (2 USNM).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E119370A3F8FD56FCC3DB09.taxon	diagnosis	Differential diagnosis. This species is very similar externally to E. clandestinus (Figs. 111, 115); it is larger (ca. 2.88 vs. 2.50 mm) and the pronotal lateral lobes are shaped slightly differently. Reliable determinations will require dissection of males. The aedeagus is superficially similar to that of E. deceptus (Figs. 112, 114). It differs in several respects, including the following: 1) the parameres apically are narrowed and pointed, have the lateral margin curved subapically, and have very thin walls; 2) the dorsal processes are longer; 3) the distance from the apex of the dorsal processes to the tip of the median lobe is shorter; 4) the tip of the median lobe is more acute; 5) the distance that the apex of the paramere extends beyond the tip of the dorsal process is less; 6) the basal piece is less arcuate; and 7) the oriface of the basal piece is larger.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E119370A3F8FD56FCC3DB09.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 2.88 / 1.53; head (width) 0.80; pronotum 0.95 / 1.20; elytra 2.53 / 1.53. Habitus as illustrated (Fig. 111). Head dark brown to piceous, maxillary palpi light brown; dorsum dark brown with lighter fascia at about midlength of elytra; venter and coxae piceous, remainder of legs reddish brown. Pronotum with hood carinae straight, converging and very narrowly separated anteriorly; secondary lateral lobe moderately large. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 4 – 5 punctures; punctures round or oval, large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae slightly emarginate on inner margin, outer margin arcuate. Metaventral depression small, ca. 7 granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E119370A3F8FD56FCC3DB09.taxon	distribution	Distribution. Currently known (reliable determinations) from three localities in Brazil (Fig. 126). According to Sharp (1875), in the original description, the holotype is from “ South America, or possibly Mexico. " Notes. This species has a long history of misidentifications. These errors are the result of the external similarity of several species related to E. trogoides, combined with the failure to examine the holotype aedeagus, or uncritical examination of the aedeagus of the holotype. Balfour-Browne (1949; fig. 1) illustrated an aedeagus supposedly of E. trogoides. However, it is not, and may represent the aedeagus of E. lanceolatulus n. sp., but the detail is inadequate to be sure. Rocha (1969; fig. 13) illustrated an aedeagus supposedly of E. trogoides. However, it is not, nor is it the same species as illustrated by Balfour-Browne (1949); the detail of the illustration is inadequate to positively determine what other species it represents. Likewise, Oliva (1986; fig. 5), illustrated an aedeagus supposedly of E. trogoides. However, it also is not, and appears instead to be the aedeagus of E. lanceolatulus. The distribution records given by Rocha (1969) and Oliva (1986) for E. trogoides are therefore invalid until proven, by careful examination of the aedeagi, to actually be E. trogoides. The aedeagus of the holotype of E. trogoides is illustrated herein (Fig. 112).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E139374A3F8F917FA9CDBEA.taxon	description	Figs. 113 (habitus), 114 (aedeagus), 129 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E139374A3F8F917FA9CDBEA.taxon	materials_examined	Type Material. Holotype (male): Brazil: Mato Grosso, Caceres, MT., 16 ° 4 ' S, 57 ° 41 ' W, d 16 ° 4 ' S, 57 ° 41 ' W, 2 – 4 xii 1955, M. Alvarenga (MSP). Paratypes (24): Brazil: Same data as holotype (1 MSP, 6 CAS); Mato Grosso, Caceres, MT., 16 ° 4 ' S, 57 ° 41 ' W, 1 – 30 xii 1955, M. Alvarenga (12 MSP); Pantanal, 17 km POCONE Pousada (farm), PORTAL PARADISO (B 10), 16 ° 16 ' S, 56 ° 38 ' W, 1 viii 2000, Jan Ruzicka (5 NMPC).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E139374A3F8F917FA9CDBEA.taxon	diagnosis	Differential Diagnosis. In dorsal habitus (Fig. 113), this species is very similar to E. trogoides and E. clandestinus, including having the carinae of the pronotal hood not confluent with one another anteriorly. It differs from E. fimbriatus in having fewer and larger tubercles on the pronotum, smaller body size (ca. 2.73 vs. 3.59 mm), and a very different aedeagus (Figs. 114, 122). The aedeagus is superficially similar to that of E. trogoides (Fig. 112). It differs in several respects, including the following: 1) the parameres apically are wider and rounded, having the lateral margin slightly emarginate subapically, and having the walls thicker, especially the medial walls; 2) the dorsal processes are shorter, more distinctly knobbed and arcuate apically, and in repose fitting into grooves in the median lobe; 3) the distance from the apex of the dorsal processes to the tip of the median lobe is greater; 4) the tip of the median lobe is more rounded; 5) the distance that the apex of the paramere extends beyond the tip of the dorsal process is greater; 6) the basal piece is more arcuate; and 7) the oriface of the basal piece is smaller.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E139374A3F8F917FA9CDBEA.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 2.73 / 1.39; head (width) 0.75; pronotum 0.89 / 1.14; elytra 1.91 / 1.39. Habitus as illustrated (Fig. 113). Head dark brown to piceous, maxillary palpi light brown; dorsum dark brown with lighter fascia at about midlength of elytra; venter and coxae piceous, remainder of legs brown to reddish brown. Pronotum with hood carinae straight, converging and narrowly separated anteriorly; secondary lateral lobe moderately large; sometimes with very small third lateral lobe near base. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 3 – 4 punctures; punctures round or oval, large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae slightly emarginate on inner margin, outer margin arcuate. Metaventral depression moderately wide, shallow, ca. nine granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E139374A3F8F917FA9CDBEA.taxon	etymology	Etymology. Named in reference to the similarity to E. trogoides and E. clandestinus.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E139374A3F8F917FA9CDBEA.taxon	distribution	Distribution. Currently known from two narrowly separated localities in southwestern Brazil (Fig. 129).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E179376A3F8FF73FF24D9AA.taxon	description	Figs. 115 (habitus), 116 (aedeagus), 128 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E179376A3F8FF73FF24D9AA.taxon	materials_examined	Type Material. Holotype (male): Brazil: Mato Grosso, Caceres, MT., 16 ° 4 ' S, 57 ° 41 ' W, 16 ° 4 ' S, 57 ° 41 ' W, 2 – 4 xii 1955, M. Alvarenga (MSP). Paratypes (6): Same data as holotype (1 CAS, 1 MSP); Brazil: Mato Grosso, Caceres, MT., 16 ° 4 ' S, 57 ° 41 ' W, 1 – 30 xii 1955, M. Alvarenga (3 MSP); Venezuela: Barinas, Rio Caparo Res. Station, 32 km E El Canton, seasonal forest, black light, 7 ° 32 ' N, 70 ° 59 ' W, 3 – 5 ii 1978, J. B. Heppner (1 USNM).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E179376A3F8FF73FF24D9AA.taxon	diagnosis	Differential Diagnosis. This species is very similar externally to E. trogoides (Figs. 111, 115); it is smaller (ca. 2.50 vs. 2.88 mm) and the pronotal lateral lobes are shaped slightly differently. Reliable determinations will require dissection of males. The aedeagus is superficially similar to that of E. trogoides (Figs. 112, 116). It differs in several respects, including the following: 1) the parameres apically are less pointed, subapically are narrower, and the medial margin is distinctly bisinuate; 2) the dorsal processes are shorter and more arcuate; 3) the distance from the apex of the dorsal processes to the tip of the median lobe is much shorter; 4) the apex of the paramere does not attain the level of the tip of the dorsal process; 6) the basal piece is more arcuate; and 7) the oriface of the basal piece is smaller.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E179376A3F8FF73FF24D9AA.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 2.50 / 1.26; head (width) 0.72; pronotum 0.80 / 1.06; elytra 1.70 / 1.26. Habitus as illustrated (Fig. 115). Head dark brown to piceous, maxillary palpi light brown; dorsum dark brown with lighter fascia at about midlength of elytra; venter and legs dark brown. Pronotum with hood carinae straight, converging and very narrowly separated anteriorly; secondary lateral lobe moderately large. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 3 – 4 punctures; punctures round or oval, large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae slightly emarginate on inner margin, outer margin arcuate. Metaventral depression moderately wide, ca. 10 granules along base, shallow.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E179376A3F8FF73FF24D9AA.taxon	distribution	Distribution. Currently known from the type locality in Brazil, and one questionable locality in Venezuela (Fig. 128). Notes. The one record from Venezuela is based on a male. The aedeagus of this specimen does not significantly differ from that of the specimens from Brazil. However, the distance separating these two populations is so great that one must question whether the specimen was mislabeled. More material is needed to verify this record.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E169376A3F8FBF8FC5CDDDB.taxon	description	Figs. 117 (habitus), 121 (aedeagus), 126 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E169376A3F8FBF8FC5CDDDB.taxon	materials_examined	Type Material. Holotype (male): Venezuela: Apure State, ca. 1 km N. Rio Claro, roadside swale, elev. 52 m, 7 ° 10.162 ' N, 67 ° 38.69 ' W, 4 i 2006, A. E. Z. Short (AS- 06 - 009) (MIZA). Paratypes (33): Venezuela: Same data as holotype (29 SEMC); Apure, Hato El Frio, Fundo Ceibote, elev. 100 m, 7 ° 49 ' N, 68 ° 54 ' W, 20 v 1975, C. J. Rosales (1 MIZA); Cojedes, Hato Mata Clara, cr. El Baul, 9 ° 17 ' N, 68 ° 21 ' W, 1 – 3 iii 1981, F. Fernandes (2 MIZA); Guarico, Calabozo, 8 ° 56 ' N, 67 ° 26 ' W, 7 ii 1969, P. & P. Spangler (1 USNM).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E169376A3F8FBF8FC5CDDDB.taxon	diagnosis	Differential Diagnosis. This species differs from all others in the Trogoides group by the entirely dull, shagreened dorsal surface. It is otherwise similar in dorsal habitus to E. fimbriatus (Figs. 117, 119). It differs therefrom in some subtle characters, such as the less denticulate pronotal lateral lobes, and a slightly narrower body form. The aedeagi of the two species differ markedly, especially the shape of the apex of the median lobe, and the fact that the dorsal processes of E. venezuelensis fit into grooves in the median lobe (Figs. 121, 122). The aedeagus shows some very weak superficial similarity to that of E. deceptus (Fig. 114); however, the parameres and dorsal processes are shaped very differently in the two species.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E169376A3F8FBF8FC5CDDDB.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 3.50 / 1.79; head (width) 0.94; pronotum 1.18 / 1.44; elytra 2.41 / 1.79. Habitus as illustrated (Fig. 117). Head dark brown to piceous, maxillary palpi light brown; dorsum dark brown with lighter fascia at about midlength of elytra and another, less distinct, at posterior declivity; venter and coxae piceous, remainder of legs reddish brown. Entire dorsal surface dull, matt. Pronotum with hood carinae straight, converging and narrowly separated anteriorly; secondary lateral lobe moderately large; smaller, third lateral lobe near base. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 3 punctures; punctures round or oval, large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae slightly emarginate on inner margin, outer margin arcuate. Metaventral depression small, ca. 7 granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E169376A3F8FBF8FC5CDDDB.taxon	distribution	Distribution. Currently known only from Venezuela (Fig. 126).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E169376A3F8FBF8FC5CDDDB.taxon	etymology	Etymology. Named in reference to the geographical distribution.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E09936BA3F8F8E4FBFFDBA2.taxon	description	Figs. 118 (habitus), 123 (aedeagus), 126 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E09936BA3F8F8E4FBFFDBA2.taxon	materials_examined	Type Material. Holotype (male): Brazil: Sao Paulo, Campos do Jordão, Parque do Estado Rio Galharada, 22 ° 44 ' S, 45 ° 36 ' W, 17 x 1992, Exp. MZSP (MSP). Paratypes: Same data as holotype (3 MSP).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E09936BA3F8F8E4FBFFDBA2.taxon	diagnosis	Differential Diagnosis. This distinctive species (Fig. 118) is differentiated from other members of the Trogoides group by several characters: 1) the longer and more sharply rounded pronotal hood; 2) the anteriorly confluent pronotal hood carinae; 3) the very small, almost absent, secondary pronotal lateral lobes, 4) the relatively deep pronotal depressions; and 5) the comparatively small elytral punctures. All carinae are well developed, and the carinae of the pronotal hood are confluent and raised anteriorly, thereby differing from all other members of the Trogoides group. Also, males have an upturned clypeus, a character not present in the others species in the group. The aedeagus (Fig. 123), which has a very long and slender median lobe, and slender parameres, cannot be confused with any other known Epimetopus. It is similar in general plan to that of E. fimbriatus (Fig. 122), a species with well developed secondary lateral lobes of the pronotum.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E09936BA3F8F8E4FBFFDBA2.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 3.16 / 1.63; head (width) 0.82; pronotum 1.26 / 1.21; elytra 2.08 / 1.63. Habitus as illustrated (Fig. 118). Head dark brown to piceous, maxillary palpi light brown; dorsum dark brown to reddish brown with lighter V-shaped fascia at about midlength of elytra; venter and coxae reddish brown, remainder of legs ochraceous. Head (male) with clypeus upturned. Pronotum with hood carinae straight, confluent and raised anteriorly; secondary lateral lobe very small, almost absent; all ridges well raised, depressions deep. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 4 – 5 punctures; punctures round or oval, not large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae slightly emarginate on inner margin in distal 1 / 2, outer margin arcuate. Metaventral depression moderately wide, ca. 14 granules along base, shallow.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E09936BA3F8F8E4FBFFDBA2.taxon	etymology	Etymology. Named in reference to the trident-like median lobe of the aedeagus.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E09936BA3F8F8E4FBFFDBA2.taxon	distribution	Distribution. Currently known only from the type locality in Brazil (Fig. 126).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E0A936FA3F8F8C1FBC7DDA0.taxon	description	Figs. 119 (habitus), 122 (aedeagus), 126 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E0A936FA3F8F8C1FBC7DDA0.taxon	materials_examined	Type Material. Holotype (male): Brazil: Mato Grosso, São Felix, 11 ° 37 ' S, 50 ° 40 ' W, 1 – 30 vi 1961, M. Alvarenga (MSP). Paratype: Brazil: Mato Grosso, Barra do Tapirapé, 10 ° 50 ' S, 50 ° 40 ' W, 1 – 30 xi 1964, B. Malkin (1 MSP).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E0A936FA3F8F8C1FBC7DDA0.taxon	diagnosis	Differential Diagnosis. Habitus similar to that of E. deceptus (Figs. 113, 119); differing therefrom in the smaller and denser pronotal tubercles, the larger denticles on the anterior angles of the elytra, and the larger size (ca. 3.59 v. 2.73 mm). The aedeagi of the two species are marked dissimilar (Figs. 114, 122). The aedeagus can perhaps be compared with that of E. tridens (Fig. 123); they have a similar general plan, but differ markedly in proportions. In E. fimbriatus the median lobe is much wider, as are the parameres and the basal piece; also, the dorsal processes of the median lobe are located more proximally, and the basal piece is markedly more arcuate. The habitus of E. fimbriatus and E. tridens are markedly dissimilar (Figs. 118, 119).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E0A936FA3F8F8C1FBC7DDA0.taxon	description	Description. Size: holotype (length / width, mm): body (length from anterior margin of pronotum to elytral apices) 3.59 / 1.79; head (width) 0.94; pronotum 1.18 / 1.50; elytra 2.50 / 1.79. Habitus as illustrated (Fig. 119). Head dark brown to piceous, maxillary palpi light brown; dorsum dark brown with lighter fascia at about midlength of elytra; venter and coxae dark reddish brown, remainder of legs ochraceous. Pronotum with hood carinae straight, converging and well separated anteriorly; secondary lateral lobe moderately large; sometimes with smaller, third lobe near base. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 3 – 4 punctures; punctures round or oval, large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae slightly emarginate on inner margin, outer margin arcuate. Metaventral depression moderately deep and wide, ca. eight large granules along base.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E0A936FA3F8F8C1FBC7DDA0.taxon	etymology	Etymology. Named in reference to the fimbriate lateral margins of the pronotum and anterior angles of the elytra. Distribution. Currently known only from Mato Grosso, Brazil (Fig. 126).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E019360A3F8FF73FBFFDB52.taxon	description	Figs. 124 (habitus), 120 (aedeagus), 126 (map)	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E019360A3F8FF73FBFFDB52.taxon	materials_examined	Type material: Holotype (male): “ Arinos M. Gerais — Brasil, 6 – 8. XI. 1964, Exp. Dep. Zool.; 1; [label with measurements]; Epimetopus tuberculatus A. A. Rocha det.; HOLOTIPO [red] ” (MSP).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E019360A3F8FF73FBFFDB52.taxon	diagnosis	Differential diagnosis. Immediately differentiated from all other members of the genus by the markedly tuberculate dorsum and pronotal sculpture (Fig. 124). Although the outline of the pronotum somewhat resembles that of members of the Trogoides group, the aedeagus is markedly different, much less complex and lacking dorsal processes.	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E019360A3F8FF73FBFFDB52.taxon	distribution	Distribution. Currently known only from the type locality in Brazil (Fig. 126).	en	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
