identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
3601F35E9E529332A3F8FB5EFA59DD9E.text	3601F35E9E529332A3F8FB5EFA59DD9E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus : Orchymont 1933	<div><p>Key to the groups of Epimetopus</p> <p>1 Small to very small species (ca. 1.25–2.25 mm); pronotum and elytra usually reddish, head dark brown to piceous, elytra without fascia (e.g., Figs. 8–11); pronotum with, at most, minute secondary lobes; eye with incomplete canthus; aedeagus simple, without dorsal processes (e.g., Figs. 12–15)..................................................... Costatus Group</p> <p>1' Usually larger species (ca. 2.00– 3.70 mm, but seldom less than 2.25 mm); head, pronotum and elytra usually dark brown to piceous, elytra often with lighter V-shaped fascia (e.g., Figs. 82, 107); pronotum with small to large secondary lobes; eye with canthus incomplete or complete; aedeagus with or without dorsal processes....................................... 2</p> <p>2 Eye with canthus incomplete; size ca. 2.60–3.20 mm; pronotum with secondary lobes very small; pronotal hood carinae grooved (e.g., Figs. 73, 78); aedeagus with extendable hooks near apex of median lobe (e.g., Figs. 79–80).... Mendeli Group</p> <p>2' Eye with canthus complete; pronotum with secondary lobes small to large; pronotal hood carinae grooved or non-grooved; aedeagus without extendable hooks near apex of median lobe................................................... 3</p> <p>3 Aedeagus simple, without dorsal processes (e.g., Figs. 109–110); pronotum with large secondary lobes; pronotal hood carinae not grooved........................................................................................... 4</p> <p>3' Aedeagus with dorsal processes on median lobe (e.g., Figs. 90, 98); pronotum with small to large secondary lobes; pronotal hood carinae grooved................................................................................... 5</p> <p>4 Dorsum strongly tuberculate (Fig. 124); pronotal hood carinae not confluent anteriorly; pronotal hood emarginate anteriorly...................................................................................... Tuberculatus Group</p> <p>4' Dorsum strongly carinate, but not tuberculate (e.g., Figs. 107–108); pronotal hood carinae confluent anteriorly; pronotal hood sharply rounded anteriorly................................................................ Lanceolatus Group</p> <p>5 Aedeagus with dorsal processes short, wide basally and pointed apically (e.g., Figs. 103–106); pronotum with secondary lobes very small............................................................................. Plaumanni Group</p> <p>5' Aedeagus with dorsal processes long and slender, often with apices with ridges (e.g., Figs. 91, 112); pronotum with secondary lobes small to large.................................................................................... 6</p> <p>6 Pronotum with very small secondary lobes (e.g., Figs. 86, 95); aedeagus with dorsal processes attaching to median lobe proximal to its midlength (e.g., Figs. 97, 98); usually smaller species (ca. 2.10–2.75 mm)................... Thermarum Group</p> <p>6' Pronotum with large (five species) or small (E. tridens) secondary lobes (e.g., Figs. 111, 118); aedeagus with dorsal processes attaching to median lobe distal to its midlength (e.g., Figs. 112, 116); usually larger species (ca. 2.50–3.65 mm).. Trogoides Group</p> </div>	https://treatment.plazi.org/id/3601F35E9E529332A3F8FB5EFA59DD9E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E529332A3F8FC90FDCBDE04.text	3601F35E9E529332A3F8FC90FDCBDE04.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus Lacordaire 1854	<div><p>Epimetopus Lacordaire, 1854</p> <p>Type species: Ceratoderus graniger Mulsant, 1851 (by monotypy).</p> <p>Ceratoderus Mulsant 1851: 1 (nec Ceratoderus Westwood 1841 (Paussidae)</p> <p>Epimetopus Lacordaire 1854: 465, 467.</p> <p>Sepidulum Leconte 1874: 48.</p> <p>Epimetopus: Orchymont, 1933: 287.</p> </div>	https://treatment.plazi.org/id/3601F35E9E529332A3F8FC90FDCBDE04	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E549334A3F8FB16FC96DDFC.text	3601F35E9E549334A3F8FB16FC96DDFC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus transversoides Perkins 2012	<div><p>Epimetopus transversoides, new species</p> <p>Figs. 11 (habitus), 15 (aedeagus), 128 (map)</p> <p>Type Material. Holotype (male): <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.21667&amp;materialsCitation.latitude=-12.65" title="Search Plazi for locations around (long -71.21667/lat -12.65)">Peru</a>: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.21667&amp;materialsCitation.latitude=-12.65" title="Search Plazi for locations around (long -71.21667/lat -12.65)">Pantiacolla Lodge</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.21667&amp;materialsCitation.latitude=-12.65" title="Search Plazi for locations around (long -71.21667/lat -12.65)">Monk Saki Trail</a>, Alto Madre de Dios River, at black light, elev. 400 m, 12° 39' S, 71° 13' W, 25 x 2000, R. Brooks (PERU 1B00 098A) (SEMC). Paratypes (42): Peru: Same data as holotype (32 SEMC); Madre de Dios, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.23333&amp;materialsCitation.latitude=-12.883333" title="Search Plazi for locations around (long -71.23333/lat -12.883333)">Hostel Erica</a> (near Salvacion), elev. 550 m, 12° 53' S, 71° 14' W, 3–5 ix 1989, R. A. Faitoute et al. (10 USNM).</p> <p>Differential Diagnosis. The habitus of this species (Fig. 11) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus could be compared with that of E. transversus (Figs. 12, 15); however, the median lobe is shaped differently, considerably smaller in E. transversoides, and the internal sac is different in extent and sculpture in the two species, being narrower and with denser spicules in E. transversoides.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 1.36/0.69; head (width) 0.41; pronotum 0.52/0.51; elytra 0.89/0.69. Habitus and sculpture as illustrated (Fig. 11). Head black, dorsum brown to reddish brown, venter and coxae dark brown. Eye with ca. 4–5 facets between canthus and posterior margin. Elytra with elongate granules linking punctures. Metaventral depression moderately deep, ca. eight granules along base.</p> <p>Etymology. Named in reference to the similarity of the aedeagus to that of E. transversus.</p> <p>Distribution. Currently known only from Peru (Fig. 128).</p></div> 	https://treatment.plazi.org/id/3601F35E9E549334A3F8FB16FC96DDFC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E549334A3F8FDC0FB89DEF3.text	3601F35E9E549334A3F8FDC0FB89DEF3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus transversus Perkins 2012	<div><p>Epimetopus transversus, new species</p> <p>Figs. 8 (habitus), 12 (aedeagus), 127 (map)</p> <p>Type Material. Holotype (male): Bolivia: La Paz, Alcoche, lamping, elev. 600 m, 15° 45' S, 67° 40' W, 19 xii 1966, Balough, Mahunka, Zicsi (493 Soil Zoological Exp.) (HNHM). Paratypes: Same data as holotype (12 HNHM).</p> <p>Differential Diagnosis. The habitus of this species (Fig. 8) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus (Fig. 12) has a median lobe somewhat similar in general shape to that of E. transversoides (Fig. 15), but it is much longer and apically is shaped differently. The parameres are similar in shape in the two species, but in E. transversus they are longer compared to the length of the basal piece.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 1.65/0.81; head (width) 0.44; pronotum 0.59/0.60; elytra 1.10/0.81. Habitus and sculpture as illustrated (Fig. 8). Head black, dorsum red, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 3–4 facets between canthus and posterior margin. Protibiae slightly arcuate. Elytral punctures linked by elongate granules. Metaventral depression moderately deep, ca. eight granules along base.</p> <p>Etymology. Named in reference to the transverse apex of the median lobe of the aedeagus.</p> <p>Distribution. Currently known only from the type locality in Bolivia (Fig. 127).</p></div> 	https://treatment.plazi.org/id/3601F35E9E549334A3F8FDC0FB89DEF3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E579336A3F8F92BFE66D87A.text	3601F35E9E579336A3F8F92BFE66D87A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus hintoni Balfour-Browne 1949	<div><p>Epimetopus hintoni Balfour-Browne, 1949</p> <p>Figs. 9 (habitus), 13 (aedeagus), 127 (map)</p> <p>Epimetopus hintoni Balfour-Browne, 1949: 17.</p> <p>Epimetopus hintoni: Oliva, 1986: 8.</p> <p>Type material: Holotype (male): " Type [red disc]; Yungas Valley Bolivia. 12–13.vi.1937; H. E. Hinton Collector; Epimetopus hintoni Type! J. Balfour-Browne det.; Hinton Coll. B.M. 1939-583" (BMNH).</p> <p>New collection record: Bolivia: Santa Cruz, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-63.6&amp;materialsCitation.latitude=-18.1" title="Search Plazi for locations around (long -63.6/lat -18.1)">Amboro National Park</a>, Los Volcanes, MV Light Sheet on stream beach, elev. 1000 m, 18° 6' S, 63° 36' W, 20 xi–12 xii 2004, M.V.L. Barclay &amp; H. Mendel (34 BMNH).</p> <p>Differential diagnosis. The habitus of this species (Fig. 9) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus (Fig. 13). The median lobe is nearly parallel-sided in dorsal view and quite blunt apically; in lateral aspect it is strongly curved. The aedeagus is similar in general shape to that of E. ecuadorensis (Fig. 21), but differs in shapes of the median lobe and parameres, and curvature of the medial lobe. Externally, E. hintoni is slightly larger than E. ecuadorensis (ca. 1.71 vs. 1.56 mm), but otherwise they are very similar.</p> <p>Distribution. Currently known from two localities in Bolivia (Fig. 127). Oliva (1986) reported specimens from Tucuman Province, Argentina; these specimens were not seen during this revision, and therefore could not be confirmed as E. hintoni.</p> </div>	https://treatment.plazi.org/id/3601F35E9E579336A3F8F92BFE66D87A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E569336A3F8FD88FBF1DE8C.text	3601F35E9E569336A3F8FD88FBF1DE8C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus lacordairei Orchymont 1933	<div><p>Epimetopus lacordairei Orchymont, 1933</p> <p>Figs. 10 (habitus), 14 (aedeagus), 127 (map)</p> <p>Epimetopus lacordairei Orchymont, 1933: 288.</p> <p>Type Material. Holotype (male, greatly disarticulated): Brazil: "Matto Grosso, Corumba" (ISNB).</p> <p>New collection records: Bolivia: Beni, Ilha Flores, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-62.716667&amp;materialsCitation.latitude=-13.033334" title="Search Plazi for locations around (long -62.716667/lat -13.033334)">Rio Itenez</a>, 13° 2' S, 62° 43' W, 7 viii 1964, J. Bouseman &amp; J. Lussenhop (5 MCZ); Paraguay: Dep. Concepcion, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-56.766666&amp;materialsCitation.latitude=-23.85" title="Search Plazi for locations around (long -56.766666/lat -23.85)">Arroyo Toro Paso</a>, 23° 51' S, 56° 46' W, 25 x 2002, U. Drechsel (5 NMW); Arroyo Toro Paso, Unterlauf (contrib. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-56.766666&amp;materialsCitation.latitude=-23.85" title="Search Plazi for locations around (long -56.766666/lat -23.85)">Rio Apa</a>), seepage, 23° 51' S, 56° 46' W, 24 x 2002, U. Drechsel (102 NMW); Stillwasser des Arroyo Toro Paso, Seitenam (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-56.766666&amp;materialsCitation.latitude=-23.85" title="Search Plazi for locations around (long -56.766666/lat -23.85)">Uferschlamm</a>), 23° 51' S, 56° 46' W, 25 x 2002, U. Drechsel (8 NMW).</p> <p>Differential diagnosis. Differentiated from all members of the Costatus group, except E. trilobus, by the trilobed apical margin of the pronotal hood (Figs. 10, 32). The aedeagus of E. lacordairei (Fig. 14), with an apically deeply bifid median lobe, differs markedly from all other members of the genus. The aedeagus of E. trilobus (Fig. 36) probably represents a preliminary stage in the evolution of the extreme condition seen in E. lacordairei. This view is supported by the fact that both species have a tri-lobed pronotal apex—the only species now known to have that condition.</p> <p>Distribution. Currently known from Bolivia, Brazil, and Paraguay (Fig. 127).</p></div> 	https://treatment.plazi.org/id/3601F35E9E569336A3F8FD88FBF1DE8C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E569336A3F8FAD1FBAEDDB9.text	3601F35E9E569336A3F8FAD1FBAEDDB9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus trilobus Perkins 2012	<div><p>Epimetopus trilobus, new species</p> <p>Figs. 32 (habitus), 36 (aedeagus), 128 (map)</p> <p>Type Material. Holotype (male): Venezuela: Bolivar, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-66.75&amp;materialsCitation.latitude=6.5833335" title="Search Plazi for locations around (long -66.75/lat 6.5833335)">Los Pijiguaos</a>, trampa interceptacion, elev. 600 m, 6° 35' N, 66° 45' W, 3–13 vi 1992, C. J. Rosales, V. Savini, J. L. Garcia, J. DeMarmels (MIZA). Paratype: Same data as holotype (1 MIZA).</p> <p>Differential Diagnosis. Differentiated from all members of the Costatus group, except E. lacordairei, by the trilobed apical margin of the pronotal hood (Figs. 10, 32). The aedeagus is distinctive by virtue of the apically bifid apex of the median lobe (Fig 36). Although the aedeagus differs markedly from that of E. lacordairei (Fig. 14), the bifid apex of both, the similarity in elytral structure, and especially the tri-lobed pronotal apex, suggest a relationship.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 1.28/0.64; head (width) 0.43; pronotum 0.50/0.51; elytra 0.85/0.64. Habitus and sculpture as illustrated (Fig. 32). Head bicolored, frons red, clypeus black, venter and coxae reddish brown, remainder of legs ochraceous. Eye with ca. 3–4 facets between canthus and posterior margin. Pronotal hood minutely trilobed. Elytra with elongate granules linking punctures; 3rd carina absent from basal ½ of elytron, otherwise carinae quite high. Protibiae arcuate, anterior surface flat. Metaventral depression shallow.</p> <p>Etymology. Named in reference to the tri-lobed pronotal apex.</p> <p>Distribution. Currently known only from the type locality in Venezuela (Fig. 128).</p></div> 	https://treatment.plazi.org/id/3601F35E9E569336A3F8FAD1FBAEDDB9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E499328A3F8F8C3FCD9D87A.text	3601F35E9E499328A3F8F8C3FCD9D87A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus spatulus Perkins 2012	<div><p>Epimetopus spatulus, new species</p> <p>Figs. 16 (habitus), 20 (aedeagus), 130 (map)</p> <p>Type Material. Holotype (male): <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.21667&amp;materialsCitation.latitude=-12.65" title="Search Plazi for locations around (long -71.21667/lat -12.65)">Peru</a>: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.21667&amp;materialsCitation.latitude=-12.65" title="Search Plazi for locations around (long -71.21667/lat -12.65)">Pantiacolla Lodge</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.21667&amp;materialsCitation.latitude=-12.65" title="Search Plazi for locations around (long -71.21667/lat -12.65)">Monk Saki Trail</a>, Alto Madre de Dios River, at black light, elev. 400 m, 12° 39' S, 71° 13' W, 25 x 2000, R. Brooks (PERU 1B00 098A) (SEMC). Paratypes: Peru: Madre de Dios, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.23333&amp;materialsCitation.latitude=-12.883333" title="Search Plazi for locations around (long -71.23333/lat -12.883333)">Hostel Erica</a> (near Salvacion), elev. 550 m, 12° 53' S, 71° 14' W, 3–5 ix 1989, R. A. Faitoute et al. (2 USNM).</p> <p>Differential Diagnosis. The habitus of this species (Fig. 16) is very similar to that of other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus is very distinctive by virtue of the spatula-like shape of the median lobe, and its comparatively long length (Fig 20). Also, the parameres are distinctively shaped.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 1.68/0.85; head (width) 0.43; pronotum 0.56/0.57; elytra 1.14/0.85. Habitus and sculpture as illustrated (Fig. 16). Head black, dorsum red, venter and coxae dark brown. Eye with ca. 3–4 facets between canthus and posterior margin. Elytra with elongate granules linking punctures, non-carinate intervals with uneven surfaces. Metaventral depression large and moderately deep, with a very low midlongitudinal ridge anteriorly, ca. 11 granules along base.</p> <p>Etymology. Named in reference to the spatula-like shape of the median lobe of the aedeagus.</p> <p>Distribution. Currently known from Peru (Fig. 130).</p></div> 	https://treatment.plazi.org/id/3601F35E9E499328A3F8F8C3FCD9D87A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E489328A3F8FD88FC87DE8B.text	3601F35E9E489328A3F8FD88FC87DE8B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus ecuadorensis Perkins 2012	<div><p>Epimetopus ecuadorensis, new species</p> <p>Figs. 17 (habitus), 21 (aedeagus), 127 (map)</p> <p>Type Material. Holotype (male): Ecuador: Napo, 17 km SW <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.816666&amp;materialsCitation.latitude=-1.0666667" title="Search Plazi for locations around (long -77.816666/lat -1.0666667)">Tena</a>, 1° 4' S, 77° 49' W, 28 v 1977, W. E. Steiner (USNM). Paratypes (3): <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.666664&amp;materialsCitation.latitude=-1.0333333" title="Search Plazi for locations around (long -77.666664/lat -1.0333333)">Misahualli</a>, 1° 2' S, 77° 40' W, 9 xi 1987, M. Huybensz (1 MCZ); Sta. Cecilia, at whitelight, 0° 2' N, 76° 58' W, 16 v 1975, Ashley B. Gurney (1 USNM); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.816666&amp;materialsCitation.latitude=0.98333335" title="Search Plazi for locations around (long -77.816666/lat 0.98333335)">Tena</a>, 0° 59' N, 77° 49' W, 27 v 1977, W. E. Steiner (1 USNM).</p> <p>Differential Diagnosis. The habitus of this species (Fig. 17) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus is somewhat similar to that of E. hintoni (Figs. 13, 21). However, the median lobe is slightly wider than that of E. hintoni, whereas the parameres, in both views, are narrower. The median lobe internal structures also differ in the two species, as illustrated.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 1.56/0.75; head (width) 0.40; pronotum 0.56/0.57; elytra 1.01/0.75. Habitus and sculpture as illustrated (Fig. 17). Head black, dorsum red, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 4–5 facets between canthus and posterior margin. Elytra with distinct granules linking punctures. Protibiae slightly arcuate. Metaventral depression moderately deep, ca. eight granules along base.</p> <p>Etymology. Named in reference to the geographical distribution.</p> <p>Distribution. Currently known from Ecuador (Fig. 127).</p></div> 	https://treatment.plazi.org/id/3601F35E9E489328A3F8FD88FC87DE8B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E489328A3F8FADEFCBCDDB4.text	3601F35E9E489328A3F8FADEFCBCDDB4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus steineri Perkins 2012	<div><p>Epimetopus steineri, new species</p> <p>Figs. 18 (habitus), 22 (aedeagus), 128 (map)</p> <p>Type Material. Holotype (male): Ecuador, Napo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.816666&amp;materialsCitation.latitude=0.98333335" title="Search Plazi for locations around (long -77.816666/lat 0.98333335)">Tena</a>, 0° 59' N, 77° 49' W, 27 v 1977, W. E. Steiner (USNM). Paratypes (18): Ecuador: Same data as holotype (5 USNM); Napo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.816666&amp;materialsCitation.latitude=0.98333335" title="Search Plazi for locations around (long -77.816666/lat 0.98333335)">Tena</a>, 0° 59' N, 77° 49' W, 26 v 1977, W. E. Steiner (11 USNM); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.8&amp;materialsCitation.latitude=-1.05" title="Search Plazi for locations around (long -77.8/lat -1.05)">Tena</a> (4 km S), 1° 3' S, 77° 48' W, 23 v 1977, P. J. Spangler &amp; D. R. Givens (#70) (2 USNM).</p> <p>Differential Diagnosis. The habitus of this species (Fig. 18) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus (Fig. 22) has a large and distinctively shaped median lobe, widest at about midlength, tapering toward the apex, then widening slightly apically. The internal sac has many fine ridges and spicules. It might be compared with E. transversoides (Fig. 15), but the median lobe is shaped differently and significantly larger.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 2.22/1.69; head (width) 0.44; pronotum 0.61/0.68; elytra 1.11/0.85. Habitus and sculpture as illustrated (Fig. 18). Head black, dorsum red, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 3–4 facets between canthus and posterior margin. Elytral punctures round, each with distinct granule at anterior margin, most granules not linking punctures. Protibiae slightly arcuate. Metaventral depression moderately deep, ca. six granules along base.</p> <p>Etymology. Named in honor of the expert collector, long-time friend and colleague Warren E. Steiner.</p> <p>Distribution. Currently only known from Ecuador (Fig. 128).</p></div> 	https://treatment.plazi.org/id/3601F35E9E489328A3F8FADEFCBCDDB4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4B932AA3F8F917FC2AD8CA.text	3601F35E9E4B932AA3F8F917FC2AD8CA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus inaequalis Perkins 2012	<div><p>Epimetopus inaequalis, new species</p> <p>Figs. 19 (habitus), 23 (aedeagus), 129 (map)</p> <p>Type Material. Holotype (male): Ecuador: Napo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.816666&amp;materialsCitation.latitude=0.98333335" title="Search Plazi for locations around (long -77.816666/lat 0.98333335)">Tena</a>, 0° 59' N, 77° 49' W, 26 v 1977, W. E. Steiner (USNM). Paratypes (14): Ecuador: Napo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.816666&amp;materialsCitation.latitude=0.98333335" title="Search Plazi for locations around (long -77.816666/lat 0.98333335)">Tena</a>, 0° 59' N, 77° 49' W, 26 v 1977, W. E. Steiner (5 USNM); Peru: Madre de Dios, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.23333&amp;materialsCitation.latitude=-12.883333" title="Search Plazi for locations around (long -71.23333/lat -12.883333)">Hostel Erica</a> (near Salvacion), elev. 550 m, 12° 53' S, 71° 14' W, 3–5 ix 1989, R. A. Faitoute et al. (3 USNM); Pantiacolla Lodge, Monk Saki Trail, Alto Madre de Dios <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.21667&amp;materialsCitation.latitude=-12.65" title="Search Plazi for locations around (long -71.21667/lat -12.65)">River</a>, at black light, elev. 400 m, 12° 39' S, 71° 13' W, 25</p> <p>x 2000, R. Brooks (PERU 1B00 098A) (5 SEMC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-69.55&amp;materialsCitation.latitude=-13.0" title="Search Plazi for locations around (long -69.55/lat -13.0)">Rio Tambopata Res</a>; 30 air km SW Pto. Maldonado, subtropical moist forest, elev. 290 m, 13° 0' S, 69° 33' W, 16–20 xi 1979, J. B. Heppner (1 USNM).</p> <p>Differential Diagnosis. A rather coarsely granulate species (Fig. 19), with prominent elongate granules linking the rather large elytral punctures; however, reliable determinations will be based on examination of the aedeagus. The aedeagus is relatively simple in this species (Fig. 23). The median lobe is pointed apically, like that of E. simplex (Fig. 37, 38). However, the parameres are very different in the two species, being pointed apically in E. inaequalis, and much longer than the median lobe.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 1.69/0.82; head (width) 0.45; pronotum 0.61/0.55; elytra 1.11/0.82. Habitus and sculpture as illustrated (Fig. 19). Head black, dorsum brown to reddish brown, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 3–4 facets between canthus and posterior margin. Protibiae slightly arcuate. Granules of pronotum and elytral carinae quite coarse. Elytral punctures large, granules linking punctures prominent. Metaventral depression moderately deep, ca. six granules along base.</p> <p>Etymology. Named in reference to the unequal lengths of the median lobe and parameres.</p> <p>Distribution. Currently known only from Ecuador and Peru (Fig. 129).</p></div> 	https://treatment.plazi.org/id/3601F35E9E4B932AA3F8F917FC2AD8CA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4A932AA3F8FD18FB5FDCFB.text	3601F35E9E4A932AA3F8FD18FB5FDCFB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus angustus Perkins 2012	<div><p>Epimetopus angustus, new species</p> <p>Figs. 24 (habitus), 28 (aedeagus), 130 (map)</p> <p>Type Material. Holotype (male): Ecuador: Napo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.816666&amp;materialsCitation.latitude=0.98333335" title="Search Plazi for locations around (long -77.816666/lat 0.98333335)">Tena</a>, 0° 59' N, 77° 49' W, 27 v 1977, W. E. Steiner (USNM). Paratypes (1479): Ecuador: Napo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-75.583336&amp;materialsCitation.latitude=-1.55" title="Search Plazi for locations around (long -75.583336/lat -1.55)">Limoncocha</a>, 1° 33' S, 75° 35' W, 11 vi 1977, W. E. Steiner (1 USNM); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.816666&amp;materialsCitation.latitude=0.98333335" title="Search Plazi for locations around (long -77.816666/lat 0.98333335)">Tena</a>, 0° 59' N, 77° 49' W, 26 v 1977, W. E. Steiner (1 USNM); Panama: Panama, El Real, 8° 8' N, 77° 43' W, 6 viii 1952, F. S. Blanton (3 USNM); Peru: Madre de Dios, Pantiacolla Lodge, Monk Saki Trail, Alto Madre de Dios <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.21667&amp;materialsCitation.latitude=-12.65" title="Search Plazi for locations around (long -71.21667/lat -12.65)">River</a>, at black light, elev. 400 m, 12° 39' S, 71° 13' W, 25 x 2000, R. Brooks (PERU1B00 098A) (23 SEMC); Venezuela: Amazonas, Agua Blanca, Cerro d. l. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-66.3&amp;materialsCitation.latitude=0.81666666" title="Search Plazi for locations around (long -66.3/lat 0.81666666)">Neblina</a>, elev. 160 m, 0° 49' N, 66° 18' W, 20–21 v 1984, O. Flint &amp; J. Louton (3 USNM); Cerro de la Neblina, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-66.166664&amp;materialsCitation.latitude=0.8333333" title="Search Plazi for locations around (long -66.166664/lat 0.8333333)">Basecamp</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-66.166664&amp;materialsCitation.latitude=0.8333333" title="Search Plazi for locations around (long -66.166664/lat 0.8333333)">At</a> black light in rainforest clearing near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-66.166664&amp;materialsCitation.latitude=0.8333333" title="Search Plazi for locations around (long -66.166664/lat 0.8333333)">Rio Baria</a>, elev. 140 m, 0° 50' N, 66° 10' W, 10–11 ii 1985, P. J. Spangler, P. M. Spangler, R. A. Faitoute, W. E. Steiner (268 USNM); Cerro de la Neblina, Basecamp, At black light on bank of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-66.166664&amp;materialsCitation.latitude=0.8333333" title="Search Plazi for locations around (long -66.166664/lat 0.8333333)">Rio Baria</a>, elev. 140 m, 0° 50' N, 66° 10' W, 6 ii 1985, W. E. Steiner (887 USNM); same locality, elev. 140 m, 0° 50' N, 66° 10' W, 28 ii 1985, P. J. Spangler, P. M. Spangler, R. A. Faitoute, W. E. Steiner (293 USNM).</p> <p>Differential Diagnosis. The habitus of this species (Fig. 24) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus (Fig. 28) is somewhat similar in general shape to that of E. burruyacu (Fig. 72); both have slender, slightly arching parameres. E. angustus differs from E. burruyacu in that the parameres are longer compared with the length of the basal piece; also, the median lobe is narrower and longer.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 1.31/0.73; head (width) 0.38; pronotum 0.52/0.53; elytra 0.85/0.73. Habitus and sculpture as illustrated (Fig. 24). Head black, dorsum red, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 3–4 facets between canthus and posterior margin. Protibiae slightly arcuate. Elytral granules linking punctures. Metaventral depression moderately deep, ca. six granules along base.</p> <p>Etymology. Named in reference to the narrow median lobe and narrow parameres.</p> <p>Distribution. Currently known from Panama, Venezuela, Ecuador, and Peru (Fig. 130).</p></div> 	https://treatment.plazi.org/id/3601F35E9E4A932AA3F8FD18FB5FDCFB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4A932CA3F8F90EFBC2DBEA.text	3601F35E9E4A932CA3F8F90EFBC2DBEA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus latus Perkins 2012	<div><p>Epimetopus latus, new species</p> <p>Figs. 25 (habitus), 31 (aedeagus), 127 (map)</p> <p>Type Material. Holotype (male): Colombia: Cesar, Pueblo Bello; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.583336&amp;materialsCitation.latitude=10.416667" title="Search Plazi for locations around (long -73.583336/lat 10.416667)">Sierra de Santa Marta</a>, beaten ex tree debris nr. stream, elev. 1200 m, 10° 25' N, 73° 35' W, 14–15 iv 1968, B. Malkin (FMNH). Paratype: Venezuela: Miranda, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-66.833336&amp;materialsCitation.latitude=10.5" title="Search Plazi for locations around (long -66.833336/lat 10.5)">Los Chorros</a>, 10° 30' N, 66° 50' W, 29 iv 1922, [collr. unknown] (1 FMNH).</p> <p>Differential Diagnosis. A moderately large and rather coarsely granulate species (Fig. 25); however, reliable determinations will be based on examination of the aedeagus. The aedeagus is similar to that of E. latisoides in that the parameres are very broad, have the medial margins straight or nearly so, and have two very small setae at the apex of each (Figs. 31, 45). However, the aedeagus of E. latus is larger (length 0.38 mm vs. 0.31 mm), the median lobe is wider and the apex is shaped slightly differently.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 1.79/0.93; head (width) 0.50; pronotum 0.64/0.61; elytra 1.17/0.93. Habitus and sculpture as illustrated (Fig. 25). Granules of dorsum rather coarse. Head black, dorsum brown to reddish brown, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 3–4 facets between canthus and posterior margin. Protibiae slightly arcuate.</p> <p>Elytral punctures large; elongate granules linking punctures prominent. Metaventral depression moderately deep, ca. eight granules along base.</p> <p>Etymology. Named in reference to the wide parameres.</p> <p>Distribution. Currently known from Colombia and Venezuela (Fig. 127).</p></div> 	https://treatment.plazi.org/id/3601F35E9E4A932CA3F8F90EFBC2DBEA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4C932CA3F8FE38FC33DF43.text	3601F35E9E4C932CA3F8FE38FC33DF43.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus ballatoris Perkins 2012	<div><p>Epimetopus ballatoris, new species</p> <p>Figs. 26 (habitus), 30 (aedeagus), 129 (map)</p> <p>Type Material. Holotype (male): Venezuela: Tachira State, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.17233&amp;materialsCitation.latitude=7.5839667" title="Search Plazi for locations around (long -72.17233/lat 7.5839667)">El Tama National Park</a>, HG-vapor light, elev. 472 m, 7° 35.038' N, 72° 10.34' W, 16 vii 2009, Short, Sites, Garcia, Inciarte, Gustafson &amp; Camaco (VZ09-0716-07A) (MIZA). Paratypes (30): Trinidad: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-61.233334&amp;materialsCitation.latitude=10.366667" title="Search Plazi for locations around (long -61.233334/lat 10.366667)">Lovell Estate</a>, 10° 22' N, 61° 14' W, 29 x 1937, H. E. Hinton (1 BMNH); Venezuela: Aragua, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-66.76667&amp;materialsCitation.latitude=10.116667" title="Search Plazi for locations around (long -66.76667/lat 10.116667)">Ocumare</a>, 10° 7' N, 66° 46' W, 19–20 ii 1969, P. &amp; P. Spangler (1 USNM); Tachira State, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.17233&amp;materialsCitation.latitude=7.5839667" title="Search Plazi for locations around (long -72.17233/lat 7.5839667)">El Tama National Park</a>, HG-vapor light, elev. 472 m, 7° 35.038' N, 72° 10.34' W, 16 vii 2009, Short, Sites, Garcia, Inciarte, Gustafson &amp; Camaco (VZ09-0716-07A) (6 SEMC); Trujillo, Mcp. Rafael Rangel, La Gira, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-70.73333&amp;materialsCitation.latitude=9.333333" title="Search Plazi for locations around (long -70.73333/lat 9.333333)">Qda. La Amarilla</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-70.73333&amp;materialsCitation.latitude=9.333333" title="Search Plazi for locations around (long -70.73333/lat 9.333333)">Trampa</a> interceptacion, elev. 520 m, 9° 20' N, 70° 44' W, 20–22 v 1995, J. Camacho, M. Garcia (3 MALUZ); Same locality, 23 x 1997, J. Camacho, M. Garcia, E. Gomez (19 MALUZ).</p> <p>Differential Diagnosis. The habitus of this species (Fig. 26) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus has arcuate parameres that arch over the median lobe; the extent of the arching is much more than in E. apocinus (Figs. 30, 48). The median lobe, which tapers distinctively in the distal one-half also clearly differs from that of E. apocinus.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 1.48/0.80; head (width) 0.43; pronotum 0.56/0.60; elytra 0.95/0.80. Habitus and sculpture as illustrated (Fig. 26). Head black, dorsum red, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 3–4 facets between canthus and posterior margin. Elytra with elongate granules linking punctures, non-carinate intervals smooth, without granules. Metaventral depression small.</p> <p>Etymology. Named in reference to the shape of the aedeagus, which has the parameres arching over the median lobe, reminiscent of a ballet dancer.</p> <p>Distribution. Currently known from Venezuela and Trinidad (Fig. 129).</p></div> 	https://treatment.plazi.org/id/3601F35E9E4C932CA3F8FE38FC33DF43	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4C932FA3F8FA66FBAEDAB9.text	3601F35E9E4C932FA3F8FA66FBAEDAB9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus plicatus Perkins 2012	<div><p>Epimetopus plicatus, new species</p> <p>Figs. 27 (habitus), 29 (aedeagus), 128 (map)</p> <p>Type Material. Holotype (male): Venezuela: Tachira State, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.17233&amp;materialsCitation.latitude=7.5839667" title="Search Plazi for locations around (long -72.17233/lat 7.5839667)">El Tama National Park</a>, HG-vapor light, elev. 472 m, 7° 35.038' N, 72° 10.34' W, 16 vii 2009, Short, Sites, Garcia, Inciarte, Gustafson &amp; Camaco (VZ09-0716-07A) (MIZA).</p> <p>Differential Diagnosis. The habitus of this species (Fig. 27) is very similar to that of other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus of this species has very distinctive parameres (Fig. 29). They are wide, "wrap around" the median lobe, and the ventral median area has an unusual wrinkled appearance. The median lobe is comparatively short, whereas the basal piece is long. The aedeagus cannot be meaningfully compared with that of other species.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 1.49/0.73; head (width) 0.40; pronotum 0.54/0.56; elytra 0.99/0.73. Habitus and sculpture as illustrated (Fig. 27). Head black, dorsum red, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 4–5 facets between canthus and posterior margin. Each elytral puncture with granule at anterior and posterior margin, granules usually not linking punctures, non-carinate intervals wide and smooth. Metaventral depression relatively small, only ca. ½ as wide as labrum.</p> <p>Etymology. Named in reference to the "wrinkles" of the parameres.</p> <p>Distribution. Currently known only from the type locality in Venezuela (Fig. 128).</p></div> 	https://treatment.plazi.org/id/3601F35E9E4C932FA3F8FA66FBAEDAB9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4F932EA3F8F96CFC2ED8A1.text	3601F35E9E4F932EA3F8F96CFC2ED8A1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus simplex Perkins 1979	<div><p>Epimetopus simplex Perkins, 1979</p> <p>Figs. 33 (habitus), 37, 38 (aedeagus), 144 (map)</p> <p>Epimetopus simplex Perkins, 1979: 323.</p> <p>New collection records: Costa Rica: Guanacaste, 2 km SE Santa Rita, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.25&amp;materialsCitation.latitude=10.016666" title="Search Plazi for locations around (long -85.25/lat 10.016666)">Rio Carmona</a>, elev. 64 m, 10° 1' N, 85° 15' W, 26 xi 2002, William D. Shepard (4 SEMC); Finca Agua Fria, N. of Carmona, UV light by <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.23333&amp;materialsCitation.latitude=10.016666" title="Search Plazi for locations around (long -85.23333/lat 10.016666)">Rio Carmona</a>, elev. 64 m, 10° 1' N, 85° 14' W, 26 xi 2002, W. Shepard (4 SEMC); nr. Carmona—Finca Agua Frio, at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.23333&amp;materialsCitation.latitude=10.016666" title="Search Plazi for locations around (long -85.23333/lat 10.016666)">Rio Carmona</a>, HG light, elev. 30 m, 10° 1' N, 85° 14' W, 15 i 2003, A. Short, R. Roughley, W. Porras (35 SEMC); Puntarenas, Corcovado National Park, Sirena Stn., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.583336&amp;materialsCitation.latitude=8.4" title="Search Plazi for locations around (long -83.583336/lat 8.4)">lower Ollas Trail</a>, ex. UV light, elev. 5 m, 8° 24' N, 83° 35' W, 25 v 2000, Z. H. Falin (CR1ABF00) (4 SEMC); Rio Bonito, ca. 9.2 km NW Golfito, ca. 1.6 km WNW <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.21667&amp;materialsCitation.latitude=8.683333" title="Search Plazi for locations around (long -83.21667/lat 8.683333)">Field Station La Gamba</a>, elev. 109 m, 8° 41' N, 83° 13' W, 4 iii 2009, M. Bojer (9 NMW); Nicaragua: Rio San Juan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-84.333336&amp;materialsCitation.latitude=10.983334" title="Search Plazi for locations around (long -84.333336/lat 10.983334)">Refugio Bartola</a>, light sheets, 10° 59' N, 84° 20' W, 5–10 viii 2002, W. D. Shepard (3 SEMC); Panama: Chiriqui, Las Lajas (2 km N) at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.86667&amp;materialsCitation.latitude=8.25" title="Search Plazi for locations around (long -81.86667/lat 8.25)">Rio San Felix</a>, blacklight, 8° 15' N, 81° 52' W, 28 v 1983, P. J. Spangler, R. A. Faitoute, W. E. Steiner (2 USNM); Panama Province, Rio Mamoni at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.066666&amp;materialsCitation.latitude=9.133333" title="Search Plazi for locations around (long -79.066666/lat 9.133333)">PanAm Hwy</a>, black light, 9° 8' N, 79° 4' W, 1 ix 2006, W. D. Shepard and D. Post (2 SEMC); Venezuela: Aragua, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-66.76667&amp;materialsCitation.latitude=10.116667" title="Search Plazi for locations around (long -66.76667/lat 10.116667)">Ocumare</a>, 10° 7' N, 66° 46' W, 19–20 ii 1969, P. &amp; P. Spangler (2 USNM); Guarico, Rio San Antonio, N. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.35295&amp;materialsCitation.latitude=9.772" title="Search Plazi for locations around (long -67.35295/lat 9.772)">Dos Caminos</a>, river margins, elev. 280 m, 9° 46.32' N, 67° 21.177' W, 8 i 2009, Short, Miller, Garcia, Camacho &amp; Joly (VZ09-0108-02A) (3 SEMC); Zulia, El Tucuco (51 Km. S. O. de <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.55&amp;materialsCitation.latitude=10.066667" title="Search Plazi for locations around (long -72.55/lat 10.066667)">Machiques</a>), 10° 4' N, 72° 33' W, 17–18 ix 1982, T. Borrego, E. Rubio, E. Inciarte (3 MALUZ); same locality, 10° 4' N, 72° 33' W, 1 ii 1985, T. Borrego, E. Rubio, E. Inciarte (1 MALUZ).</p> <p>Differential diagnosis. The habitus of E. simplex (Fig. 33) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The median lobe is similar to that of E. inaequalis (Figs. 23, 37, 38), but the parameres differ markedly in the two species; those of E. simplex being much narrower basally, rounded apically, and each bearing two short, blunt setae. Externally (Figs. 19, 33) the pronotal hood shape and pronotal hood carinae differ slightly in the two species.</p> <p>Distribution. Currently known from Venezuela to Honduras (Fig. 144).</p></div> 	https://treatment.plazi.org/id/3601F35E9E4F932EA3F8F96CFC2ED8A1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4E932EA3F8FCFBFCBBDF44.text	3601F35E9E4E932EA3F8FCFBFCBBDF44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus panamensis Perkins 1979	<div><p>Epimetopus panamensis Perkins, 1979</p> <p>Figs. 34 (habitus), 39 (aedeagus), 143 (map)</p> <p>Epimetopus panamensis Perkins, 1979: 324.</p> <p>New collection record: Panama: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.833336&amp;materialsCitation.latitude=9.166667" title="Search Plazi for locations around (long -79.833336/lat 9.166667)">Canal Zone</a>, Barro Colorado, leaf litter, 9° 10' N, 79° 50' W, 19 ii – 9 iii 1975, Lawrence, Erwin (1 MCZ).</p> <p>Differential diagnosis. The habitus of E. panamensis (Fig. 34) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus is somewhat similar to that of E. punctipennis (Figs. 39, 62); differing therefrom in having a narrower and longer median lobe, a wider and longer basal piece, and wider and differently shaped parameres. The aedeagus of E. panamensis also has a larger striate collar. Externally (Figs. 34, 58) the two species differ in color (E. panamensis is brown, E. punctipennis is reddish), and E. panamensis has much smaller elytral punctures and larger elytral granules linking the punctures.</p> <p>Distribution. Currently known only from Panama (Fig. 143).</p></div> 	https://treatment.plazi.org/id/3601F35E9E4E932EA3F8FCFBFCBBDF44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E4E932EA3F8FA99FB90DDDD.text	3601F35E9E4E932EA3F8FA99FB90DDDD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus robustus Perkins 2012	<div><p>Epimetopus robustus, new species</p> <p>Figs. 35 (habitus), 40 (aedeagus), 128 (map)</p> <p>Type Material. Holotype (male): Panama: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.736664&amp;materialsCitation.latitude=9.151667" title="Search Plazi for locations around (long -79.736664/lat 9.151667)">Canal Zone</a>, 4.1 mi NW Gamboa, Rio Frijoles, flood debris and wet leaves along river, 9° 9.1' N, 79° 44.2' W, 19 ii 1976, A. Newton (MCZ). Paratype: Same data as holotype (1 MCZ).</p> <p>Differential Diagnosis. The habitus of this species (Fig. 35) is very similar to that of other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus (Fig. 40) is very broad and stout; it cannot be confused with any others.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 1.55/0.78; head (width) 0.45; pronotum 0.58/0.59; elytra 1.01/0.78. Habitus and sculpture as illustrated (Fig. 35). Head black, dorsum red, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 4–5 facets between canthus and posterior margin. Protibiae slightly arcuate. Many elytral punctures not linked by granules. Metaventral depression moderately deep, ca. eight granules along base.</p> <p>Etymology. Named in reference to the very wide and stout aedeagus.</p> <p>Distribution. Currently known only from the type locality in Panama (Fig. 128).</p></div> 	https://treatment.plazi.org/id/3601F35E9E4E932EA3F8FA99FB90DDDD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E419320A3F8F8E6FB90DB99.text	3601F35E9E419320A3F8F8E6FB90DB99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus latisoides Perkins 2012	<div><p>Epimetopus latisoides, new species</p> <p>Figs. 41 (habitus), 45 (aedeagus), 129 (map)</p> <p>Type Material. Holotype (male): Panama: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.736664&amp;materialsCitation.latitude=9.151667" title="Search Plazi for locations around (long -79.736664/lat 9.151667)">Canal Zone</a>, 12.0 mi NW Gamboa, Rio Agua Salud, 9° 9.1' N, 79° 44.2' W, 19 ii 1976, A. Newton (MCZ).</p> <p>Differential Diagnosis. The habitus of this species (Fig. 41) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus is similar to E. latus in that the parameres are very broad, have the medial margins straight or nearly so, and have two very small setae at the apex of each (Figs. 31, 45). However, the aedeagus of E. latisoides is much smaller, the median lobe is narrower and the apex is shaped slightly differently.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 1.67/0.88; head (width) 0.45; pronotum 0.58/0.61; elytra 1.14/0.88. Habitus and sculpture as illustrated (Fig. 41). Head black, dorsum red, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 3–4 facets between canthus and posterior margin. Protibiae slightly arcuate. Elongate granules linking elytral punctures. Metaventral depression moderately deep, ca. eight granules along base.</p> <p>Etymology. Named in reference to the aedeagal similarity to E. latus.</p> <p>Distribution. Currently known only from the type locality in Panama (Fig. 129).</p></div> 	https://treatment.plazi.org/id/3601F35E9E419320A3F8F8E6FB90DB99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E439322A3F8F8E6FC26D851.text	3601F35E9E439322A3F8F8E6FC26D851.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus microporus Perkins 2012	<div><p>Epimetopus microporus, new species</p> <p>Figs. 42 (habitus), 46 (aedeagus), 134 (map)</p> <p>Type Material. Holotype (male): Panama: Panama Province, Rio Mamoni at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.066666&amp;materialsCitation.latitude=9.133333" title="Search Plazi for locations around (long -79.066666/lat 9.133333)">PanAm Hwy</a>, black light, 9° 8' N, 79° 4' W, 1 ix 2006, W. D. Shepard and D. Post (SEMC). Paratypes (2): Honduras: Rio Humuya NW Comayagua, 14° 29' N, 87° 39' W, 3 viii 1967, O. S. Flint, Jr. (1 USNM); Panama: Chiriqui, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.566666&amp;materialsCitation.latitude=8.2" title="Search Plazi for locations around (long -81.566666/lat 8.2)">Rio Vigui</a> (Pan-Amer. Hwy), 8° 12' N, 81° 34' W, 28 ix 1976, Edward S. Ross (1 CAS).</p> <p>Differential Diagnosis. The habitus of this species (Fig. 42) is very similar to that of other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus is similar to that of E. costaricensis in general proportions and size (Figs. 46, 47); differing therefrom in the shape and degree of sclerotization of the median lobe, which is slightly widened in the distal one-third, and has the margin indented slightly at each pore.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 1.34/0.69; head (width) 0.38; pronotum 0.52/0.52; elytra 0.83/0.69. Habitus and sculpture as illustrated (Fig. 42). Head black, dorsum red, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 3–4 facets between canthus and posterior margin. Elytra with elongate granules linking punctures. Metaventral depression small.</p> <p>Etymology. Named in reference to the conspicuous micropores of the median lobe of the aedeagus.</p> <p>Distribution. Currently known from Panama and Honduras (Fig. 134).</p></div> 	https://treatment.plazi.org/id/3601F35E9E439322A3F8F8E6FC26D851	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E459325A3F8FB2DFC2CDDC7.text	3601F35E9E459325A3F8FB2DFC2CDDC7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus apocinus Perkins 1979	<div><p>Epimetopus apocinus Perkins, 1979</p> <p>Figs. 44 (habitus), 48 (aedeagus), 139 (map)</p> <p>Epimetopus apocinus Perkins, 1979: 323.</p> <p>New collection records: Costa Rica: Guanacaste, 18.7 km NW Bagaces, Hwy 1, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.36667&amp;materialsCitation.latitude=10.65" title="Search Plazi for locations around (long -85.36667/lat 10.65)">Rio Carana</a>, elev. 91 m, 10° 39' N, 85° 22' W, 4 vi 1973, Erwin &amp; Hevel (5 USNM); Limon, Reventazon, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.46667&amp;materialsCitation.latitude=10.25" title="Search Plazi for locations around (long -83.46667/lat 10.25)">Hamburg Farm</a>, at light, 10° 15' N, 83° 28' W, 27 v 1935, F. Nevermann (2 FMNH); Puntarenas, Corcovado National Park, Sirena Stn., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.583336&amp;materialsCitation.latitude=8.4" title="Search Plazi for locations around (long -83.583336/lat 8.4)">lower Ollas Trail</a>, ex. UV light, elev. 5 m, 8° 24' N, 83° 35' W, 25 v 2000, Z. H. Falin (CR1ABF00) (5 SEMC); Mexico: Tabasco, Rio Puyacatengo, E. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-92.95&amp;materialsCitation.latitude=17.55" title="Search Plazi for locations around (long -92.95/lat 17.55)">Teapa</a>, 17° 33' N, 92° 57' W, 28–29 vii 1966, Flint &amp; Ortiz (1 USNM).</p> <p>Differential diagnosis. The habitus of E. apocinus (Fig. 44) is quite similar to other members of the Costatus group; reliable determinations will require examination of the male genitalia. The aedeagus has arching parameres somewhat similar to those of E. mucronatus, but otherwise the aedeagi of the two species are quite dissimilar (Figs. 48, 56). The aedeagus of E. apocinus is larger and wider, the median lobe is less acutely pointed apically, and the parameres are arched differently and, in lateral view, are sinuate. Externally, E. apocinus has elongate granules linking the elytral punctures, whereas in E. mucronatus the granules do not link the punctures. E. ballatoris (Fig. 30) also has arcuate parameres that arch over the median lobe; however, the extent of the arching is much more than in E. apocinus. The median lobe of E. ballatoris, which tapers distinctively in the distal one-half also clearly differs from that of E. apocinus. The pronotal lateral lobes are slightly less angulate in E. apocinus than in E. ballatoris (Figs. 26, 44).</p> <p>Distribution. Currently known from Costa Rica and Mexico (Fig. 139).</p></div> 	https://treatment.plazi.org/id/3601F35E9E459325A3F8FB2DFC2CDDC7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E459325A3F8FF73FBB5DE18.text	3601F35E9E459325A3F8FF73FBB5DE18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus costaricensis Perkins 1979	<div><p>Epimetopus costaricensis Perkins, 1979</p> <p>Figs. 43 (habitus), 47 (aedeagus), 141 (map)</p> <p>Epimetopus costaricensis Perkins, 1979: 324.</p> <p>New collection records: Belize: Cayo District, nr Teakettle Bank, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-88.85&amp;materialsCitation.latitude=17.216667" title="Search Plazi for locations around (long -88.85/lat 17.216667)">Pook’s Hill</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-88.85&amp;materialsCitation.latitude=17.216667" title="Search Plazi for locations around (long -88.85/lat 17.216667)">Lt.</a> trap at river, 17° 13' N, 88° 51' W, 6 i 2003, C. R. Bartlett (12 SEMC); Costa Rica: Cartago, Cimarrones at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.416664&amp;materialsCitation.latitude=10.1" title="Search Plazi for locations around (long -83.416664/lat 10.1)">Rio Cimarrones</a>, black lights, 10° 6' N, 83° 25' W, 23 xi 2002, W. Shepard (1 SEMC); Guanacaste, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.2&amp;materialsCitation.latitude=10.033334" title="Search Plazi for locations around (long -85.2/lat 10.033334)">Est. Manglar Ju Villa Nandayure</a>, at light, 10° 2' N, 85° 12' W, 11 xii 2002, W. Porras (2 SEMC); Heredia, OTS La Selva Field Sta., Puerto Viejo de Sarapique, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-18.983334&amp;materialsCitation.latitude=10.433333" title="Search Plazi for locations around (long -18.983334/lat 10.433333)">Rio Puerto Viejo</a>, at light, 10° 26' N, 18° 59' W, 5–11 iii 1973, J. Wagner &amp; J. Kethley (1 FMNH); Limon, Reventazon, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.46667&amp;materialsCitation.latitude=10.25" title="Search Plazi for locations around (long -83.46667/lat 10.25)">Hamburg Farm</a>, at light, 10° 15' N, 83° 28' W, 25 v 1931, F. Nevermann (1 FMNH); Puntarenas, 4 km N. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.333336&amp;materialsCitation.latitude=8.566667" title="Search Plazi for locations around (long -83.333336/lat 8.566667)">Puerto Jimenez</a>, blacklights, 8° 34' N, 83° 20' W, 18 vi 2003, W. Shepard (11 SEMC); Corcovado National Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.583336&amp;materialsCitation.latitude=8.4" title="Search Plazi for locations around (long -83.583336/lat 8.4)">Sirena Stn.</a>, ex UV light, elev. 5 m, 8° 24' N, 83° 35' W, 28 vi 2000, Z. H. Falin (CR1ABF00 038) (15 SEMC); Corcovado National Park, Sirena Stn., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.583336&amp;materialsCitation.latitude=8.4" title="Search Plazi for locations around (long -83.583336/lat 8.4)">lower Ollas Trail</a>, ex. UV light, elev. 5 m, 8° 24' N, 83° 35' W, 25 v 2000, Z. H. Falin (CR1ABF00) (6 SEMC); Guatemala: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-90.066666&amp;materialsCitation.latitude=14.85" title="Search Plazi for locations around (long -90.066666/lat 14.85)">El Progresso</a>, km. 69 on C. A. 9, 14° 51' N, 90° 4' W, 2 vii 1974, W. E. Steiner (3 USNM); Honduras: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-87.46667&amp;materialsCitation.latitude=15.7" title="Search Plazi for locations around (long -87.46667/lat 15.7)">Lancetilla</a> [Aug., no year], 15° 42' N, 87° 28' W,, Stadelmann (1 MCZ); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-87.6&amp;materialsCitation.latitude=13.533334" title="Search Plazi for locations around (long -87.6/lat 13.533334)">Nacaome</a>, 17.4 km W Jicaro Galan Jct. Hwy CA1, elev. 100 m, 13° 32' N, 87° 36' W, 2 vi 1973, Erwin &amp; Hevel (1 USNM); Mexico: Tabasco, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-92.95&amp;materialsCitation.latitude=17.55" title="Search Plazi for locations around (long -92.95/lat 17.55)">Rio Puyacatengo</a>, E. Teapa, 17° 33' N, 92° 57' W, 28–29 vii 1966, Flint &amp; Ortiz (2 USNM); Panama: Panama Province, Rio Mamoni at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.066666&amp;materialsCitation.latitude=9.133333" title="Search Plazi for locations around (long -79.066666/lat 9.133333)">PanAm Hwy</a>, black light, 9° 8' N, 79° 4' W, 1 ix 2006, W. D. Shepard and D. Post (13 SEMC).</p> <p>Differential diagnosis. The habitus of E. costaricensis (Fig. 43) is very similar to that of other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus is similar to that of E. microporus in that both have the parameres lobe-like apically; however, the shapes of the lobes differ in the two species (Figs. 46, 47). The median lobe is very simple and parallel-sided in E. costaricensis, whereas in E. microporus it is widened and minutely indented in the distal one-third, at the locations of micropores. Externally, the pronotal hood carinae are shaped slightly differently in the two species (Figs. 42, 43).</p> <p>Distribution. Currently known to be distributed from Panama to Mexico (Fig. 141).</p></div> 	https://treatment.plazi.org/id/3601F35E9E459325A3F8FF73FBB5DE18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E449327A3F8F8C3FBB1D8E9.text	3601F35E9E449327A3F8F8C3FBB1D8E9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus lobilatus Perkins 2012	<div><p>Epimetopus lobilatus, new species</p> <p>Figs. 49 (habitus), 53 (aedeagus), 146 (map)</p> <p>Type Material. Holotype (male): Costa Rica: Limon, Reventazon, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.46667&amp;materialsCitation.latitude=10.25" title="Search Plazi for locations around (long -83.46667/lat 10.25)">Hamburg Farm</a>, at light, 10° 15' N, 83° 28' W, 25 v 1931, F. Nevermann (FMNH). Paratypes: Same data as holotype, except 28 ix 1934, F. Nevermann (2 FMNH).</p> <p>Differential Diagnosis. The habitus of this species (Fig. 49) is very similar to that of other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus has parameres that are lobate apically, as does E. costaricensis; however, the paramere lobes are shaped differently in the two species (Figs. 47, 53). Also, the median lobe of E. lobilatus tapers in the distal one-half, whereas in E. costaricensis the median lobe is virtually parallel-sided. The median lobe of E. lobilatus is more similar in shape to that of E. apocinus, but in that species the parameres are narrowed apically, not lobate.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 1.32/~1.00; head (width) 0.55; pronotum 0.75/0.77; elytra 1.33/~1.00. Habitus and sculpture as illustrated (Fig. 49). Head black, dorsum red, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 3–4 facets between canthus and posterior margin. Protibiae slightly arcuate. Elytra with elongate granules linking punctures. Metaventral depression moderately deep, ca. six granules along base.</p> <p>Etymology. Named in reference to the wide apical lobes of the aedeagal parameres.</p> <p>Distribution. Currently known only from the type locality in Costa Rica (Fig. 146).</p></div> 	https://treatment.plazi.org/id/3601F35E9E449327A3F8F8C3FBB1D8E9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E479327A3F8FD33FC59DC23.text	3601F35E9E479327A3F8FD33FC59DC23.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus rectus Perkins 2012	<div><p>Epimetopus rectus, new species</p> <p>Figs. 50 (habitus), 54 (aedeagus), 147 (map)</p> <p>Type Material. Holotype (male): Costa Rica: Heredia, OTS La Selva Field Sta., Puerto Viejo de Sarapique, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-18.983334&amp;materialsCitation.latitude=10.433333" title="Search Plazi for locations around (long -18.983334/lat 10.433333)">Rio Puerto Viejo</a>, at light, 10° 26' N, 18° 59' W, 5–11 iii 1973, J. Wagner &amp; J. Kethley (FMNH). Paratypes (10): Costa Rica: Cartago, Cimarrones at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.416664&amp;materialsCitation.latitude=10.1" title="Search Plazi for locations around (long -83.416664/lat 10.1)">Rio Cimarrones</a>, black lights, 10° 6' N, 83° 25' W, 23 xi 2002, W. Shepard (3 SEMC); Heredia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-84.01667&amp;materialsCitation.latitude=10.433333" title="Search Plazi for locations around (long -84.01667/lat 10.433333)">La Selva Biol. Res. Stn.</a>, 3 km S Puerto Viejo, ex. blacklight, elev. 80 m, 10° 26' N, 84° 1' W, 4 vi 2001, Chatzimanolis (CR1C01019) (3 SEMC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-84.01667&amp;materialsCitation.latitude=10.416667" title="Search Plazi for locations around (long -84.01667/lat 10.416667)">La Selva Biological Station</a>, HG-vapor lights, 10° 25' N, 84° 1' W, 14–15 i 2005, A.E.Z. Short (3 SEMC); OTS La Selva Field Sta., Puerto Viejo de Sarapique, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-18.983334&amp;materialsCitation.latitude=10.433333" title="Search Plazi for locations around (long -18.983334/lat 10.433333)">Rio Puerto Viejo</a>, at light, 10° 26' N, 18° 59' W, 5–11 iii 1973, J. Wagner &amp; J. Kethley (1 FMNH).</p> <p>Differential Diagnosis. This species has a slightly sinuate anterior margin of the pronotal hood, and the submarginal groove of the hood is relatively wide, otherwise the habitus (Fig. 50) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus is small, very narrow, and has narrow, straight parameres (Fig. 54). The median lobe shape could perhaps be compared with that of E. panamensis (Fig. 39), but internally the median lobes are very different, the internal sac being narrower and much more densely spiculate in E. rectus, and the general proportions of the aedeagi are markedly different.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 1.51/0.79; head (width) 0.45; pronotum 0.59/0.58; elytra 0.97/0.79. Habitus and sculpture as illustrated (Fig. 50). Head black, dorsum brown to reddish brown, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 3–4 facets between canthus and posterior margin. Protibiae slightly arcuate. Elytra with elongate granules linking punctures. Metaventral depression moderately deep, ca. six granules along base.</p> <p>Etymology. Named in reference to the straight parameres of the aedeagus.</p> <p>Distribution. Currently known only from Costa Rica (Fig. 147).</p></div> 	https://treatment.plazi.org/id/3601F35E9E479327A3F8FD33FC59DC23	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E479326A3F8F946FBB1DBEA.text	3601F35E9E479326A3F8F946FBB1DBEA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus latilobus Perkins 2012	<div><p>Epimetopus latilobus, new species</p> <p>Figs. 51 (habitus), 55 (aedeagus), 136 (map)</p> <p>Type Material. Holotype (male): Costa Rica: Puntarenas, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-84.833336&amp;materialsCitation.latitude=10.3" title="Search Plazi for locations around (long -84.833336/lat 10.3)">Monteverde area</a>, 10° 18' N, 84° 50' W, 6–14 vi 1973, Erwin &amp; Hevel (USNM). Paratype: Same data as holotype (1 USNM).</p> <p>Differential Diagnosis. The habitus of this species (Fig. 51) is very similar to that of other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus (Fig. 55) has a large, wide median lobe—unlike any other species. The parameres gradually increase in width from base to apex, and slightly arch; each has two small setae apically.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 1.79/1.00; head (width) 0.53; pronotum 0.67/0.71; elytra 1.20/1.00. Habitus and sculpture as illustrated (Fig. 51). Head black, dorsum brown to reddish brown, venter dark brown to piceous, legs dark brown. Eye with ca. 3–4 facets between canthus and posterior margin. Elytra with elongate granules linking punctures. Metaventral depression moderately deep and wide, ca. nine granules along base, shallow.</p> <p>Etymology. Named in reference to the wide median lobe of the aedeagus.</p> <p>Distribution. Currently known only from the type locality in Costa Rica (Fig. 136).</p></div> 	https://treatment.plazi.org/id/3601F35E9E479326A3F8F946FBB1DBEA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E799319A3F8FC36FBB0DC44.text	3601F35E9E799319A3F8FC36FBB0DC44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus acuminatus Perkins 2012	<div><p>Epimetopus acuminatus, new species</p> <p>Figs. 57 (habitus), 61 (aedeagus), 145 (map)</p> <p>Type Material. Holotype (male): Guatemala: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-90.066666&amp;materialsCitation.latitude=14.85" title="Search Plazi for locations around (long -90.066666/lat 14.85)">El Progresso</a>, km. 69 on C. A. 9, 14° 51' N, 90° 4' W, 2 vii 1974, W. E. Steiner (USNM).</p> <p>Differential Diagnosis. The habitus of this species (Fig. 57) is very similar to that of other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus (Fig. 61) has very distinct parameres, wide basally, and in the distal one-fifth markedly narrowed and abruptly arcuate. The median lobe is somewhat similar in shape to that of E. lobilatus (Fig. 53), but in E. acuminatus it is longer, more nearly parallel-sided; also, the two differ internally, as illustrated. The parameres markedly differ in the two species.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 1.69/0.84; head (width) 0.47; pronotum 0.64/0.64; elytra 1.08/0.84. Habitus and sculpture as illustrated (Fig. 57). Head black, dorsum red, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 3–4 facets between canthus and posterior margin. Protibiae not arcuate. Elongate granules linking most elytral punctures. Metaventral depression moderately deep, ca. eight granules along base.</p> <p>Etymology. Named in reference to the acuminate apices of the parameres.</p> <p>Distribution. Currently known only from the type locality in Guatemala (Fig. 145).</p></div> 	https://treatment.plazi.org/id/3601F35E9E799319A3F8FC36FBB0DC44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E799319A3F8FF73FC23D913.text	3601F35E9E799319A3F8FF73FC23D913.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus mucronatus Perkins 2012	<div><p>Epimetopus mucronatus, new species</p> <p>Figs. 52 (habitus), 56 (aedeagus), 148 (map)</p> <p>Type Material. Holotype (male): Honduras: Rio Humuya NW Comayagua, 14° 29' N, 87° 39' W, 3 viii 1967, O. S. Flint, Jr. (USNM). Paratypes (9): Mexico: Chiapas, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-93.21667&amp;materialsCitation.latitude=15.683333" title="Search Plazi for locations around (long -93.21667/lat 15.683333)">Nr. Pijijiapan</a>, 15° 41' N, 93° 13' W, 5 vii 1965, Paul J. Spangler (1 USNM); Nayarit, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-104.9&amp;materialsCitation.latitude=21.5" title="Search Plazi for locations around (long -104.9/lat 21.5)">Tepic</a>, light trap, elev. 914 m, 21° 30' N, 104° 54' W, 1 viii 1964, Blanton &amp; Broce (1 USNM); Oaxaca, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.416664&amp;materialsCitation.latitude=17.35" title="Search Plazi for locations around (long -95.416664/lat 17.35)">Rio Jaltepec</a> [no date], 17° 21' N, 95° 25' W, R. E. Woodruff (1 BMNH); San Luis Potosi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-98.8&amp;materialsCitation.latitude=21.266666" title="Search Plazi for locations around (long -98.8/lat 21.266666)">Tamazunchale</a>, 21° 16' N, 98° 48' W, 30 v 1948, Werner, Nutting (5 USNM); Veracruz, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-96.48333&amp;materialsCitation.latitude=19.333334" title="Search Plazi for locations around (long -96.48333/lat 19.333334)">Puente Nacional</a>, 19° 20' N, 96° 29' W, 31 vii 1966, Flint &amp; Ortiz (1 USNM).</p> <p>Differential Diagnosis. The habitus of this species (Fig. 52) is very similar to that of other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus has arching parameres somewhat similar to those of E. apocinus, but otherwise the aedeagi of the two species are quite dissimilar (Figs. 48, 56). The aedeagus of E. mucronatus is smaller and narrower, the median lobe is more acutely pointed apically, and the parameres are arched differently and, in lateral view, lack the sinuations present in E. apocinus.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 1.25/0.78; head (width) 0.39; pronotum 0.50/0.52; elytra 0.88/0.78. Habitus and sculpture as illustrated (Fig. 52). Head black, dorsum red, venter light brown, maxillary palpi testaceous. Eye with ca. 3–4 facets between canthus and posterior margin. Protibiae slightly arcuate. Elytral punctures each with a granule at anterior and posterior margins, granules usually not linking punctures. Metaventral depression moderately deep, ca. eight granules along base.</p> <p>Etymology. Named in reference to the pointed apex of the median lobe of the aedeagus.</p> <p>Distribution. Currently known from Mexico and Honduras (Fig. 148).</p></div> 	https://treatment.plazi.org/id/3601F35E9E799319A3F8FF73FC23D913	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E799318A3F8F98BFACFDBEA.text	3601F35E9E799318A3F8F98BFACFDBEA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus punctipennis Perkins 1979	<div><p>Epimetopus punctipennis Perkins, 1979</p> <p>Figs. 58 (habitus), 62 (aedeagus), 137 (map)</p> <p>Epimetopus punctipennis Perkins, 1979: 321.</p> <p>New collection records: Mexico: Sinaloa, 21 mi. E. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.916664&amp;materialsCitation.latitude=23.2" title="Search Plazi for locations around (long -105.916664/lat 23.2)">Villa Union</a>, 23° 12' N, 105° 55' W, 25 vii 1964, H. F. Howden (2 CNC); 54 mi S <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-107.2&amp;materialsCitation.latitude=24.133333" title="Search Plazi for locations around (long -107.2/lat 24.133333)">Culiacan</a>, elev. 164 m, 24° 8' N, 107° 12' W, 23 iv 1969, M. E. Irwin (2 USNM); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-106.066666&amp;materialsCitation.latitude=23.283333" title="Search Plazi for locations around (long -106.066666/lat 23.283333)">Concordia</a>, 23° 17' N, 106° 4' W, 1–30 vii 1974, D. Chandler (7 USNM); U.S.A.: Oklahoma, 9 mi W Broken Bow, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-94.73333&amp;materialsCitation.latitude=34.033333" title="Search Plazi for locations around (long -94.73333/lat 34.033333)">Glover River</a>, black light trap, 34° 2' N, 94° 44' W, 26 vii 1987, W. D. Shepard (4 CAS); Texas, Gillespie Co., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-98.9&amp;materialsCitation.latitude=30.3" title="Search Plazi for locations around (long -98.9/lat 30.3)">Lange’s Mill</a>, 30° 18' N, 98° 54' W, 5 vii 1969, T. W. Taylor (3 USNM); Llano Co., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-98.816666&amp;materialsCitation.latitude=30.5" title="Search Plazi for locations around (long -98.816666/lat 30.5)">Enchanted Rock</a>, collected at light, 30° 30' N, 98° 49' W, 18 iv 1959, H. F. Howden (1 CNC); USA: Burnett Co. [no other data, or date; L/L very approximate], 30° 44' N, 98° 13' W, (3 MCZ).</p> <p>Differential diagnosis. The habitus of E. punctipennis (Fig. 58) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus is somewhat similar to that of E. panamensis (Figs. 39, 62; differing therefrom in having a wider and shorter median lobe, a narrower and shorter basal piece, and narrower and differently shaped parameres. Externally (Figs. 34, 58) the two species differ in color (E. punctipennis is reddish, E. panamensis is brown), and E. punctipennis has much larger elytral punctures and smaller elytral granules, which do not link the punctures.</p> <p>Distribution. Currently known from northern Mexico, Arizona, Texas, and Oklahoma (Fig. 137).</p></div> 	https://treatment.plazi.org/id/3601F35E9E799318A3F8F98BFACFDBEA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7B931AA3F8F8E0FB8CDBC1.text	3601F35E9E7B931AA3F8F8E0FB8CDBC1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus bifidus Perkins 2012	<div><p>Epimetopus bifidus, new species</p> <p>Figs. 59 (habitus), 63 (aedeagus), 135 (map)</p> <p>Type Material. Holotype (male): Mexico: Oaxaca, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-94.2&amp;materialsCitation.latitude=16.366667" title="Search Plazi for locations around (long -94.2/lat 16.366667)">Tapanatepec</a>, 8 mi. W Oaxaca, elev. 122 m, 16° 22' N, 94° 12' W, 10 vii 1953, Univ. Kans. Mex. Expedition (SEMC). Paratypes: Same data as holotype (10 SEMC).</p> <p>Differential Diagnosis. The habitus of this species (Fig. 59) is very similar to that of other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus is similar in general proportions to that of E. mucronatus (Figs. 56, 63). However, the median lobe in E. bifidus is apically divided into two lobes; also, the internal sac is different in the two species, as illustrated.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 1.40/0.73; head (width) 0.42; pronotum 0.55/0.58; elytra 0.92/0.73. Habitus and sculpture as illustrated (Fig. 59). Head black, frons red, clypeus black, dorsum red, venter rufous. Eye with ca. 3–4 facets between canthus and posterior margin. Elytra with elongate granules linking punctures. Metaventral depression small.</p> <p>Etymology. Named in reference to the bifid apex of the median lobe of the aedeagus.</p> <p>Distribution. Currently known only from the type locality in Mexico (Fig. 135).</p></div> 	https://treatment.plazi.org/id/3601F35E9E7B931AA3F8F8E0FB8CDBC1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7A931AA3F8FB86FC23DCBA.text	3601F35E9E7A931AA3F8FB86FC23DCBA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus fisheri Perkins 1979	<div><p>Epimetopus fisheri Perkins, 1979</p> <p>Figs. 65 (habitus), 69 (aedeagus), 142 (map)</p> <p>Epimetopus fisheri Perkins, 1979: 322.</p> <p>New collection records: Honduras: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-87.6&amp;materialsCitation.latitude=13.533334" title="Search Plazi for locations around (long -87.6/lat 13.533334)">Nacaome</a>, 17.4 km W Jicaro Galan Jct. Hwy CA1, elev. 100 m, 13° 32' N, 87° 36' W, 2 vi 1973, Erwin &amp; Hevel (3 USNM); Mexico: Chiapas, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-93.816666&amp;materialsCitation.latitude=16.266666" title="Search Plazi for locations around (long -93.816666/lat 16.266666)">2 mi. SW Las Cruces</a>, 16° 16' N, 93° 49' W, 3 viii 1954, A. A. Alcorn (1 SEMC); Guerrero, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-99.9&amp;materialsCitation.latitude=16.866667" title="Search Plazi for locations around (long -99.9/lat 16.866667)">Acapulco</a>, 16° 52' N, 99° 54' W, 16–30 viii 1938, L. J. Lipovsky (3 SEMC); same locality, 16° 52' N, 99° 54' W, 18 viii 1938, L. J. Lipovsky (15 SEMC); same locality, 16° 52' N, 99° 54' W, 20 viii 1938, L. J. Lipovsky (2 SEMC); Jalisco, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-104.933334&amp;materialsCitation.latitude=19.466667" title="Search Plazi for locations around (long -104.933334/lat 19.466667)">25 mi NW of Barra de Navidad</a>, intermittent stream, scrub tropics, 19° 28' N, 104° 56' W, 22 vii 1974, M. E. &amp; P. D. Perkins (11 MCZ); Agua Caliente, unnamed stream, elev. 290 m, 20° 23' N, 105° 26', 15 i 2005, W. D. Shepard (WDS-A-1624) (6 AEZS); Oaxaca, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-94.2&amp;materialsCitation.latitude=16.366667" title="Search Plazi for locations around (long -94.2/lat 16.366667)">Tapanatepec</a>, 8 mi. W Oaxaca, elev. 122 m, 16° 22' N, 94° 12' W, 10 vii 1953, Univ. Kans. Mex. Expedition (1 SEMC).</p> <p>Differential diagnosis. The habitus of E. fisheri (Fig. 65) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus is somewhat similar to that of E. costatus in general proportions (Figs. 69, 71), including being broad in lateral view. However, the median lobe shape and the shape of the parameres differ markedly in the two species.</p> <p>Distribution. Currently known from Honduras and Mexico (Fig. 142).</p></div> 	https://treatment.plazi.org/id/3601F35E9E7A931AA3F8FB86FC23DCBA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7A931AA3F8FE1BFB8CDE63.text	3601F35E9E7A931AA3F8FE1BFB8CDE63.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus oaxacus Perkins 2012	<div><p>Epimetopus oaxacus, new species</p> <p>Figs. 60 (habitus), 64 (aedeagus), 135 (map)</p> <p>Type Material. Holotype (male):: Mexico: Oaxaca, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-94.2&amp;materialsCitation.latitude=16.366667" title="Search Plazi for locations around (long -94.2/lat 16.366667)">Tapanatepec</a>, 8 mi. W Oaxaca, elev. 122 m, 16° 22' N, 94° 12' W, 10 vii 1953, Univ. Kans. Mex. Expedition (SEMC).</p> <p>Differential Diagnosis. The habitus of this species (Fig. 60) is very similar to that of other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus has a median lobe shaped somewhat like that of E. spatulus (Figs. 20, 64), but the basal part, between the striate collar and the base, is wider and much shorter. The parameres and basal piece differ markedly in the two species.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 1.63/0.86; head (width) 0.47; pronotum 0.66/0.67; elytra 1.09/0.86. Habitus and sculpture as illustrated (Fig. 60). Head black, dorsum brown to reddish brown, venter reddish brown. Eye with ca. 4–5 facets between canthus and posterior margin. Elytra with elongate granules linking punctures. Metaventral depression very small.</p> <p>Etymology. Named in reference to the geographical distribution.</p> <p>Distribution. Currently known only from the type locality in Mexico (Fig. 135).</p></div> 	https://treatment.plazi.org/id/3601F35E9E7A931AA3F8FE1BFB8CDE63	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7A931CA3F8F8C8FB91DB09.text	3601F35E9E7A931CA3F8F8C8FB91DB09.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus arizonicus Perkins 2012	<div><p>Epimetopus arizonicus, new species</p> <p>Figs. 66 (habitus), 70 (aedeagus), 138 (map)</p> <p>Type Material. Holotype (male): USA: Arizona, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-110.8&amp;materialsCitation.latitude=31.933332" title="Search Plazi for locations around (long -110.8/lat 31.933332)">Pajarito Mts.</a>, Sycamore Cyn., 31° 56' N, 110° 48' W, 24 viii 1968, [no collector on label] (MCZ). Paratypes: Same data as holotype (4 MCZ).</p> <p>Differential Diagnosis. This is a markedly granulate species, with dark brown dorsum; otherwise the habitus (Fig. 66) is very similar to that of other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus has a long median lobe, which is slightly widened at about the distal one-third; it is probably related to E. costatus, which see for comparisons (Figs. 70, 71).</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 1.79/0.94; head (width) 0.50; pronotum 0.64/0.69; elytra 1.20/0.94. Habitus and sculpture as illustrated (Fig. 66). Head black, dorsum brown to reddish brown, venter and legs dark brown, maxillary palpi brown. Pronotal and elytral granules prominent, including elongate granules linking punctures. Eye with ca. 3–4 facets between canthus and posterior margin. Protibiae stout, flattened in distal 1/5, outer margin arcuate, inner margin straight. Metaventral depression moderately deep and wide, ca. eight granules along base.</p> <p>Etymology. Named in reference to the geographical distribution.</p> <p>Distribution. Currently known only from the type locality in Arizona (Fig. 138).</p></div> 	https://treatment.plazi.org/id/3601F35E9E7A931CA3F8F8C8FB91DB09	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7C931FA3F8FE53FC05DBC1.text	3601F35E9E7C931FA3F8FE53FC05DBC1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus costatus (LeConte 1874)  1874	<div><p>Epimetopus costatus (LeConte), 1874</p> <p>Figs. 67 (habitus), 71 (aedeagus), 140 (map)</p> <p>Sepidulum costatum LeConte, 1874: 48.</p> <p>Epimetopus costatus (LeConte): Schwarz &amp; Barber 1917: 132 (key).</p> <p>Epimetopus costatus (LeConte): Perkins 1979: 320 (lectotype design., desc., fig.). New collection record: USA: Texas, [no other locality data] (1 FMNH).</p> <p>Differential diagnosis. The habitus of E. costatus (Fig. 67) is very similar to that of several other members of the Costatus group; reliable determinations will be based on examination of the aedeagus. The aedeagus (Fig. 71) has a very distinctive median lobe, which is markedly widened at the distal one-third. It is nearest in form to the aedeagus of E. arizonicus (Fig. 70), but that species has a much narrower and longer median lobe, which extends further beyond the slightly differently shaped parameres. Externally (Figs. 66, 67), the two species differ in color, E. costatus is reddish and E. arizonicus is brown, and E. costatus has the non-carinate elytral intervals smoother.</p> <p>Distribution. Currently known from Texas and Arkansas (Fig. 140).</p></div> 	https://treatment.plazi.org/id/3601F35E9E7C931FA3F8FE53FC05DBC1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7F931FA3F8FE1BFB2CDED0.text	3601F35E9E7F931FA3F8FE1BFB2CDED0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus burruyacu Oliva 1986	<div><p>Epimetopus burruyacu Oliva, 1986</p> <p>Figs. 68 (habitus), 72 (aedeagus), 126 (map)</p> <p>Epimetopus burruyacu Oliva, 1986: 7.</p> <p>Type material: Holotype (male): " ARGENTINA, Prov. Tucuman, Dpto. Burruyacú rio Salas 12.II.1982 Leg. R. Golbach; Epimetopus burruyacu A. Oliva 1984; HOLOTYPO; aedeagus drawn by P. D. Perkins; DIGITAL IMAGE captured 2012, P. D. Perkins " (MACN).</p> <p>Differential diagnosis. The habitus (Fig. 68) is similar to that of several species in the Costatus group; reliable determinations will be based on examination of the male genitalia. The aedeagus is somewhat similar in general to that of E. angustus (Figs. 28, 72); both have slender, slightly arching parameres. E. burruyacu differs from E. angustus in that the parameres are shorter compared with the length of the basal piece; also, the median lobe is wider and shorter. Externally, E. burruyacu is larger that E. angustus (ca. 1.55 vs. 1.31 mm), has the elytral carinae and pronotal sculpture slightly less developed, and has smaller elytral punctures (Figs. 24, 68).</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 1.55/0.78; head (width) 0.41; pronotum 0.55/0.56; elytra 1.00/0.78. Habitus and sculpture as illustrated (Fig. 68). Head piceous, dorsum red, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 3–4 facets between canthus and posterior margin. Protibiae slightly arcuate. Elytral granules linking punctures. Metaventral depression moderately deep, ca. six granules along base.</p> <p>Distribution. Currently known from two narrowly separated localities in Argentina (Fig. 126).</p></div> 	https://treatment.plazi.org/id/3601F35E9E7F931FA3F8FE1BFB2CDED0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E7F9310A3F8F978FB93DB8A.text	3601F35E9E7F9310A3F8F978FB93DB8A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus graniger (Mulsant 1851)	<div><p>Epimetopus graniger (Mulsant, 1851)</p> <p>Figs. 73–76 (habitus), 77 (aedeagus), 126 (map)</p> <p>Ceratoderus graniger Mulsant, 1851: 2.</p> <p>Epimetopus graniger (Mulsant): Lacordaire 1854: 465, 467.</p> <p>Epimetopus graniger (Mulsant): Balfour-Browne (1949: 18); catalog.</p> <p>Epimetopus graniger (Mulsant): Rocha (1969: 184).</p> <p>Type material: Holotype (male; examined): "Colombia" (MNHN). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.083336&amp;materialsCitation.latitude=4.6" title="Search Plazi for locations around (long -74.083336/lat 4.6)">New</a> collection record: Colombia: C. Amara, La Aguadita, 4° 36' N, 74° 5' W, 7 vii 1970, J. M. Campbell (1 CNC).</p> <p>Differential diagnosis. Distinguished from other members of the Mendeli group by the very ovate elytra (l/w ca. 1.71/1.55), the relatively small elytral punctures, and the multiple grooves of the pronotal hood carinae (Figs. 73–76). The aedeagus (Fig. 77) is not closely similar to other members of the Mendeli group. The distal part of the median lobe of the aedeagus differs from that of other members: the two apical processes are long, slender, and pointed; the hooks are large and claw-like.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 2.60/1.55; head (width) 0.73; pronotum 0.98/1.05; elytra 1.71/1.55. Habitus as illustrated (fig. 73). Dorsal surface piceous brown; elytra without hint of fascia; ventral surface piceous black, epipleura reddish brown; antennae reddish brown, maxillary palpi reddish with palpomere 4 brown; trochanters and bases of femora dark brown to brown, distal portions of femora and entire tibiae and tarsi reddish. Body widely elongate, widest at about midlength of elytra, body moderately convex in lateral view, elytra very transversely convex. Eye canthus incomplete, ca. 3 facets between canthus and posterior margin. Pronotum with hood carinae slightly arcuate, confluent anteriorly, grooved throughout, grooves multiple in posterior 1/2; primary lateral lobe with anterior angle roundly produced; secondary lateral lobe extremely small or absent. Elytron with four granulate carinae, third interrupted anteriorly for distance of ca. 3–4 punctures; first three carinae (from suture) very low and very weakly granulate; punctures round, small, each with moderately large granule at anterior margin, non-carinate intervals with irregular row of small granules. Protibiae straight or very slightly emarginate on inner margin, outer margin weakly arcuate. Metaventral depression with less dense granules than remainder of mesoventrite.</p> <p>Distribution. Currently known from one unambiguous site in Colombia (Fig. 126); the holotype locality data is only " Colombia."</p> <p>Notes. The holotype (Figs. 73–76), which had not previously been dissected, luckily proved to be a male. Two other specimens, one of each gender, were received with the holotype. Neither of these two specimens had any labels, but both were pinned with the same kind of very fine insect pin as the holotype. The second male, which I dissected, is conspecific with the holotype. The female probably is also conspecific. The female differs from the two males in the following: the body size is slightly smaller, the pronotal lateral lobes have the anterior part slightly more indented, the elytral granules are slightly less developed, and the circular transparent serial punctures of the elytra are smaller. The lateral margins of the elytra of the female, in the area slightly anterior to midlength, are straight or slightly emarginate, whereas in males the margins are clearly arcuate; this is probably a sexual dimorphism.</p></div> 	https://treatment.plazi.org/id/3601F35E9E7F9310A3F8F978FB93DB8A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E739315A3F8FF73FB94DBB2.text	3601F35E9E739315A3F8FF73FB94DBB2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus angulatus Balfour-Browne 1949	<div><p>Epimetopus angulatus Balfour-Browne, 1949</p> <p>Figs. 78 (habitus), 79, 80 (aedeagus), 126 (map)</p> <p>Epimetopus angulatus Balfour-Browne 1949: 16.</p> <p>Epimetopus angulatus: Oliva (1986: 8); misidentification.</p> <p>Epimetopus angulatus: Fikáček, et al. (2011: 489).</p> <p>Type material: Holotype (male): " Type [red disc]; Yungas Valley. Bolivia. 12–13.vi.1937; H. E. Hinton Collector; Epimetopus angulatus Type! J. Balfour-Browne det.; Hinton Coll. B.M. 1939-583" (BMNH).</p> <p>Differential Diagnosis. The dorsal habitus of E. angulatus is very similar to that of E. peruvianus; E. angulatus is slightly larger (ca. 2.73 vs. 2.62 mm) but otherwise very similar (Figs. 78, 82). Reliable determinations will be based on examination of the aedeagi of the two species, which are markedly dissimilar (Figs. 78, 80, 82). The aedeagus could be compared with that of E. coleuncus (Fig. 85), but differs from that in several respects, including the following: 1) the aedeagus is wider, the lateral processes of the median lobe (the part contacting the parameres) being notably larger; 2) the median lobe apical part is narrower and is emarginate laterally, whereas in E. coleuncus the apical part is wider and arcuate laterally, 3) the hooks rest in a slot on each side of the median lobe, whereas in E. coleuncus the hooks each rest in a groove on the dorsal surface of the median lobe; 4) the parameres are shaped differently; and 5) the basal piece is longer in E. coleuncus. The aedeagus of E. angulatus differs markedly from E. mendeli in overall shape, and many details.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 2.73/1.43; head (width) 0.68; pronotum 0.95/1.00; elytra 1.82/1.43. Habitus as illustrated (Fig. 78). Head and pronotum dark brown to piceous; elytra brown to dark brown, without fascia; maxillary palpi light brown; venter and coxae piceous, remainder of legs reddish brown. Eye canthus incomplete, ca. 3 facets between canthus and posterior margin. Pronotum with hood carinae slightly arcuate, confluent at anterior tip, grooved throughout; primary lateral lobe with anterior angle produced and acute; secondary lateral lobe extremely small or absent. Elytron with four granulate carinae, third interrupted anteriorly for distance of ca. 3–4 punctures; punctures round, small, each with moderately large granule at anterior margin, non-carinate intervals with irregular row of granules. Protibiae straight or very slightly emarginate on inner margin, outer margin weakly arcuate. Metaventral depression small, ca. 8 granules along base.</p> <p>Distribution. Currently known only from the type locality in Bolivia (Fig. 126).</p> <p>Notes. Oliva (1986; fig. 4), illustrated an aedeagus supposedly of E. angulatus. However, it is not, and appears instead to perhaps be the aedeagus of E. coleuncus. The aedeagus of the holotype of E. angulatus was illustrated by Fikáček et al. (2011), and a detail of the apex of the same aedeagus is illustrated herein (Fig. 79). Because of the error, and the fact that other species have a habitus similar to that E. angulatus, the distribution records given by Oliva (1986) must be considered invalid until being confirmed by the dissection of males.</p> <p>Epimetopus mendeli Fiká č ek, Barclay &amp; Perkins, 2011</p> <p>Fig. 126 (map)</p> <p>Epimetopus mendeli Fikáček, Barclay &amp; Perkins, 2011: 480.</p> <p>New collection records: Peru: Cusco, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.166664&amp;materialsCitation.latitude=-13.066667" title="Search Plazi for locations around (long -71.166664/lat -13.066667)">Pil</a> [l]copata, premontane moist forest, elev. 600 m, 13° 4' S, 71° 10' W, 8–10 xii 1979, J. B. Heppner (1 USNM); same locality, elev. 600 m, 13° 4' S, 71° 10' W, 11–14 xii 1979, J. B. Heppner (2 USNM); Prov. Quispicanchis, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-70.76667&amp;materialsCitation.latitude=-13.233334" title="Search Plazi for locations around (long -70.76667/lat -13.233334)">Quincemil</a> [Quince Mil], 13° 14' S, 70° 46' W, 6–11 x 1976, Robert Gordon (1 USNM); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.333336&amp;materialsCitation.latitude=-14.183333" title="Search Plazi for locations around (long -71.333336/lat -14.183333)">San Pedro</a> (at km 152), MV light, elev. 1430 m, 14° 11' S, 71° 20' W, 30 viii 1989, R. A. Faitoute et al. (8 USNM); Santa Isabel, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.3&amp;materialsCitation.latitude=-13.0" title="Search Plazi for locations around (long -71.3/lat -13.0)">Cosnipata Valley</a>, 13° 0' S, 71° 18' W, 9 xii 1951, Felix Woytkowski (1 FMNH); Madre de Dios, Hostel Erica (near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.23333&amp;materialsCitation.latitude=-12.883333" title="Search Plazi for locations around (long -71.23333/lat -12.883333)">Salvacion</a>), elev. 550 m, 12° 53' S, 71° 14' W, 3–5 ix 1989, R. A. Faitoute et al. (1 USNM).</p> <p>Differential diagnosis. The dorsal habitus is similar to that of E. angulatus, but E. mendeli is considerably larger (ca. 3.10 vs. 2.73 mm), and has more elongate elytra (l/w ratio ca. 1.40 vs. 1.26), and a distinctive elytral fascia (see Figs. in Fikáček et al., 2011). The aedeagus of E. mendeli differs markedly from E. angulatus in overall shape, and many details (Figs. 79, 80). The aedeagus of E. mendeli is somewhat similar to that of E. coleuncus. It differs from E. coleuncus in the following: 1) the shape and length of the parameres, which are shorter and lack the subapical lateral sinuations; 2) the shape of the apical part of the median lobe (more sharply rounded at the tips); and 3) in having narrower lateral processes of the median lobe.</p> <p>Distribution. Currently known only from Peru (Fig. 126).</p> <p>Notes. For a detailed description see Fikáček et al. 2011.</p> <p>Epimetopus flavicaptus Fiká č ek, Barclay &amp; Perkins, 2011</p> <p>Figs. 81 (habitus), 126 (map)</p> <p>Epimetopus flavicaptus Fikáček, Barclay &amp; Perkins, 2011: 488.</p> <p>Type material: Holotype (male): Ecuador, Napo Province [see data in Fikáček, et al. 2011].</p> <p>Differential diagnosis. Distinguished from other members of the Mendeli group by the transverse or slightly emarginate anterior margin of the pronotal hood (Fig. 81). The aedeagus is illustrated by Fikáček et al. (2011). The distal part of the median lobe is clearly diagnostically different than that of the other members of the Mendeli group.</p> <p>Distribution. Currently known only from the type locality in Ecuador (Fig. 126).</p></div> 	https://treatment.plazi.org/id/3601F35E9E739315A3F8FF73FB94DBB2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E749309A3F8F8C6FC5CDAB9.text	3601F35E9E749309A3F8F8C6FC5CDAB9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus peruvianus Perkins 2012	<div><p>Epimetopus peruvianus, new species</p> <p>Figs. 82 (habitus), 83 (aedeagus), 128 (map)</p> <p>Type Material. Holotype (male): Peru: Ayacucho, La Mar, Santa Rosa, elev. 640 m, 13° 0' S, 73° 41' W, 19–25 ix 1976, Robert Gordon (USNM). Paratypes: Same data as holotype (2 USNM); same data, except 8–15 ix 1976 (10 USNM).</p> <p>Differential Diagnosis. The dorsal habitus of E. peruvianus is very similar to that of E. angulatus (Figs. 78, 82); E. peruvianus is slightly smaller (ca. 2.62 vs. 2.73 mm). Reliable determinations will be based on examination of the aedeagi of the two species (Figs. 79, 80, 82). The aedeagus clearly differs from that of other members of the Mendeli group: the distal part of the median lobe is deeply divided into two processes that diverge from one another; the membranous part of the median lobe is very wide; and the parameres are wide at the base, but abruptly narrowed at about midlength, and from there to the apices are parallel.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 2.62/1.34; head (width) 0.64; pronotum 0.90/0.94; elytra 1.76/1.34. Habitus as illustrated (Fig. 82). Head and pronotum piceous, maxillary palpi light brown; elytra brown to dark brown except yellowish fascia slightly past midlength (very faint in some specimens); venter and coxae piceous, remainder of legs dark brown. Eye canthus incomplete, ca. 3 facets between canthus and posterior margin. Pronotum with hood carinae arcuate, confluent at anterior tip, grooved throughout; primary lateral lobe with anterior angle produced; secondary lateral lobe very small, almost absent. Elytron with four granulate carinae, third interrupted anteriorly for distance of ca. 2–3 punctures; 4th carina slightly higher than others; punctures round, moderately large, each with small granule at anterior margin, non-carinate intervals with irregular row of small granules. Protibiae rather long and slender, emarginate on inner margin in distal ½. Metaventral depression moderately deep and wide, granules sparse, rather widely separated, ca. seven granules along base.</p> <p>Distribution. Currently known only from the type locality in Peru (Fig. 128).</p> <p>Etymology. Named in reference to the geographical distribution.</p></div> 	https://treatment.plazi.org/id/3601F35E9E749309A3F8F8C6FC5CDAB9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E69930BA3F8F8C9FC22DBC1.text	3601F35E9E69930BA3F8F8C9FC22DBC1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus coleuncus Perkins 2012	<div><p>Epimetopus coleuncus, new species</p> <p>Figs. 84 (habitus), 85 (aedeagus), 127 (map)</p> <p>Type Material. Holotype (male): Argentina: Tucuman, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-64.75&amp;materialsCitation.latitude=-26.5" title="Search Plazi for locations around (long -64.75/lat -26.5)">Rio Salas</a>, departamento de Burruyacu, 26° 30' S, 64° 45' W, 12 ii 1982, Golbach (MACN). Paratypes (7): Same data as holotype (6 MACN); Bolivia: Tarija, Ing. Bermejo, 21° 32' S, 64° 44' W, 14–28 ii 1969, R. Golbach (1 MACN).</p> <p>Differential Diagnosis. Differing externally from other members of the Mendeli group by the pit-like depression at the base of the fused pronotal carinae (Fig. 84). The aedeagus differs from that of E. angulatus (Figs. 79, 80, 85) in several respects, including the following: 1) it is not as wide, the lateral processes of the median lobe being notably smaller; 2) the median lobe apical part is wider, arcuate laterally, whereas in E. angulatus the apical part is emarginate laterally; 3) the hooks each rest in a groove on the dorsal surface of the median lobe, whereas in E. angulatus the hooks rest in a slot on each side; 4) the parameres are shaped differently; and 5) the basal piece is longer in E. coleuncus. It differs from E. mendeli in 1) the shape and length of the parameres, 2) the shape of the apical part of the median lobe (more rounded at the tips), and 3) in having wider lateral processes of the median lobe.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 2.74/1.45; head (width) 0.69; pronotum 0.95/1.05; elytra 1.90/1.45. Habitus as illustrated (Fig. 84). Head and pronotum piceous, maxillary palpi brown; elytra dark brown without hint of fascia; venter black, coxae and basal ½ of femora piceus, remainder of legs dark brown. Eye canthus incomplete, ca. 3 facets between canthus and posterior margin. Pronotum with hood carinae arcuate, completely confluent in anterior ½, grooved throughout; primary lateral lobe with anterior angle produced and acute; secondary lateral lobe very small. Elytron with four carinae, third interrupted anteriorly for distance of ca. 3–4 punctures; first three carinae (from suture) very low and very weakly granulate; 4th carina higher and more strongly granulate; punctures round, large, each with minute granule at anterior margin, non-carinate intervals with unilinear row of minute, well separated granules. Protibiae straight or very slightly emarginate on inner margin, outer margin weakly arcuate. Metaventral depression shallow, ca. six granules along base.</p> <p>Etymology. Named in reference to the sheathed hooks of the median lobe of the aedeagus.</p> <p>Distribution. Currently known from Argentina and Bolivia (Fig. 127).</p></div> 	https://treatment.plazi.org/id/3601F35E9E69930BA3F8F8C9FC22DBC1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E6D930FA3F8FF73FE4AD979.text	3601F35E9E6D930FA3F8FF73FE4AD979.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus thermarum Schwarz & Barber 1917	<div><p>Epimetopus thermarum Schwarz &amp; Barber, 1917</p> <p>Figs. 86 (habitus), 87, 91 (aedeagus), 133 (map)</p> <p>Epimetopus thermarum Schwarz &amp; Barber, 1917: 130.</p> <p>Epimetopus thermarum: Balfour-Browne (1949: 13); key.</p> <p>Epimetopus thermarum: Rocha (1969: 178); Figs.</p> <p>Type material: Holotype (female; examined): “Hot Sprgs Ar 26.6.01; Schwarz &amp; Barber Coll; Epimetopus thermarum S &amp; B 1917; Type No. 21052 U.S. N.M” (USNM). Many paratypes with same locality data (USNM).</p> <p>New collection records (219 specimens): Belize: Cayo District, nr Teakettle Bank, Pook’s Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-88.85&amp;materialsCitation.latitude=17.216667" title="Search Plazi for locations around (long -88.85/lat 17.216667)">Sweep</a>, 17° 13' N, 88° 51' W, 6 i 2003, C. R. Bartlett (1 SEMC); Stann Creek District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-88.48333&amp;materialsCitation.latitude=16.8" title="Search Plazi for locations around (long -88.48333/lat 16.8)">Nr Armenia</a> at river, 16° 48' N, 88° 29' W, 7 i 2003, C. R. Bartlett (2 SEMC); Costa Rica: Guanacaste, 18.7 km NW Bagaces, Hwy 1, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.36667&amp;materialsCitation.latitude=10.65" title="Search Plazi for locations around (long -85.36667/lat 10.65)">Rio Carana</a>, elev. 91 m, 10° 39' N, 85° 22' W, 4 vi 1973, Erwin &amp; Hevel (28 USNM); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.2&amp;materialsCitation.latitude=10.033334" title="Search Plazi for locations around (long -85.2/lat 10.033334)">Est. Manglar Ju Villa Nandayure</a>, at light, 10° 2' N, 85° 12' W, 11 xii 2002, W. Porras (1 SEMC); Puntarenas, Corcovado National Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.583336&amp;materialsCitation.latitude=8.4" title="Search Plazi for locations around (long -83.583336/lat 8.4)">Sirena Stn.</a>, ex UV light, elev. 5 m, 8° 24' N, 83° 35' W, 28 vi 2000, Z. H. Falin (CR1ABF00 038) (3 SEMC); Guatemala: Baja Verapaz, 4 mi. S. Rabinal, stream, transition xeric-tropical, 15° 3' N, 90° 27' W, 10 vi 1974, M. E. &amp; P. D. Perkins (1 MCZ); El Progresso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-90.066666&amp;materialsCitation.latitude=14.85" title="Search Plazi for locations around (long -90.066666/lat 14.85)">El Progresso</a>, interm. desert stream, 14° 51' N, 90° 4' W, 2 vi 1974, M.E. &amp; P.D. Perkins (1 MCZ); El Progresso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-90.066666&amp;materialsCitation.latitude=14.85" title="Search Plazi for locations around (long -90.066666/lat 14.85)">Km.</a> 69 on C. A. 9, 14° 51' N, 90° 4' W, 2 vii 1974, W. E. Steiner (2 MCZ); Mexico: Baja California Sur, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-109.76667&amp;materialsCitation.latitude=23.366667" title="Search Plazi for locations around (long -109.76667/lat 23.366667)">1.5 mi. NW Miraflores</a>, collected at blacklight, elev. 213 m, 23° 22' N, 109° 46' W, 28–29 x 1968, E. L. Sleeper &amp; F. J. Moore (2 CAS); 13 mi. up Canyon San Bernardo fr. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-110.01667&amp;materialsCitation.latitude=23.383333" title="Search Plazi for locations around (long -110.01667/lat 23.383333)">Boca de la Sierra—San</a> Bernardo, elev. 609 m, 23° 23' N, 110° 1' W, 6 viii 1971, H. G. Real &amp; R. E. Main (1 CAS); 3 mi N Miraflores, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-113.23333&amp;materialsCitation.latitude=28.083334" title="Search Plazi for locations around (long -113.23333/lat 28.083334)">Boca de la Sierra</a>, 28° 5' N, 113° 14' W, 22 vi 1997, W. D. Shepard (1 NMW); 5 mi. S. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-113.23333&amp;materialsCitation.latitude=28.016666" title="Search Plazi for locations around (long -113.23333/lat 28.016666)">Miraflores</a>, 28° 1' N, 113° 14' W, 10 vii 1938, Michelbacher &amp; Ross (1 FMNH); 5 mi W <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-109.95&amp;materialsCitation.latitude=23.733334" title="Search Plazi for locations around (long -109.95/lat 23.733334)">San Bartolo</a>, 23° 44' N, 109° 57' W, 13 vii 1938, Michelbacher &amp; Ross (1 FMNH, 9 CAS); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-110.21667&amp;materialsCitation.latitude=24.083334" title="Search Plazi for locations around (long -110.21667/lat 24.083334)">8 mi. SE La Paz</a>, collected at blacklight, elev. 304 m, 24° 5' N, 110° 13' W, 13 x 1968, E. L. Sleeper &amp; F. J. Moore (1 CAS); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-112.28333&amp;materialsCitation.latitude=27.283333" title="Search Plazi for locations around (long -112.28333/lat 27.283333)">9 mi. N San Lucas</a>, collected at blacklight, elev. 304 m, 27° 17' N, 112° 17' W, 24–25 x 1968, E. L. Sleeper &amp; F. J. Moore (1 CAS); 9 mi. up Canyon San Bernardo fr. Boca de la Sierra (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-109.95&amp;materialsCitation.latitude=23.383333" title="Search Plazi for locations around (long -109.95/lat 23.383333)">Potrero</a>), elev. 457 m, 23° 23' N, 109° 57' W, 2 viii 1971, H. G. Real &amp; R. E. Main (2 FMNH, 98 CAS); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-109.833336&amp;materialsCitation.latitude=23.466667" title="Search Plazi for locations around (long -109.833336/lat 23.466667)">Sa. Victoria</a>, 7 mi. W Santiago, collected at blacklight, 23° 28' N, 109° 50' W, 30–31 x 1968, E. L. Sleeper &amp; F. J. Moore (10 CAS); Sa. Victoria, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-109.833336&amp;materialsCitation.latitude=23.466667" title="Search Plazi for locations around (long -109.833336/lat 23.466667)">El Chorro</a>, collected at blacklight, elev. 304 m, 23° 28' N, 109° 50' W, 29–30 x 1968, E. L. Sleeper &amp; F. J. Moore (4 CAS); Chiapas, Chintul, Rio Chintul (trib. R. Grijalva), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-93.58517&amp;materialsCitation.latitude=17.2425" title="Search Plazi for locations around (long -93.58517/lat 17.2425)">Rt.</a> 187, 17° 14.55' N, 93° 35.11' W, 20 ix 1966, Bueno, Barba, Contreras (1 USNM); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-93.73333&amp;materialsCitation.latitude=16.65" title="Search Plazi for locations around (long -93.73333/lat 16.65)">Municipio de Cintalapa La Mina</a>, elev. 914 m, 16° 39' N, 93° 44' W, 14 ix 1981, D. E. &amp; P. M. Breedlove and C. G. Whitefield (3 CAS); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-91.98333&amp;materialsCitation.latitude=17.516666" title="Search Plazi for locations around (long -91.98333/lat 17.516666)">Palenque</a>, 17° 31' N, 91° 59' W, 22–23 vi 1969, D. Bright &amp; J. M. Campbell (6 CNC); Jalisco, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-104.933334&amp;materialsCitation.latitude=19.466667" title="Search Plazi for locations around (long -104.933334/lat 19.466667)">25 mi NW of Barra de Navidad</a>, intermittent stream, scrub tropics, 19° 28' N, 104° 56' W, 22 vii 1974, M. E. &amp; P. D. Perkins (1 MCZ); SE of Chamela, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.066666&amp;materialsCitation.latitude=19.533333" title="Search Plazi for locations around (long -105.066666/lat 19.533333)">Rio Cuitzmala</a>, elev. 55 m, 19° 32' N, 105° 4' W, 6 i 2005, W. D. Shepard (1 SEMC); Nayarit, 32 miles S. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.3&amp;materialsCitation.latitude=22.016666" title="Search Plazi for locations around (long -105.3/lat 22.016666)">Acaponeta</a>, 22° 1' N, 105° 18' W, 24 xi 1948, H. B. Leech (1 CAS); Sinaloa, 21 mi. E. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.916664&amp;materialsCitation.latitude=23.2" title="Search Plazi for locations around (long -105.916664/lat 23.2)">Villa Union</a>, 23° 12' N, 105° 55' W, 25 vii 1964, H. F. Howden (2 CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-106.066666&amp;materialsCitation.latitude=23.283333" title="Search Plazi for locations around (long -106.066666/lat 23.283333)">Concordia</a>, 23° 17' N, 106° 4' W, 1–30 vii 1974, D. Chandler (1 USNM); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-107.38333&amp;materialsCitation.latitude=24.8" title="Search Plazi for locations around (long -107.38333/lat 24.8)">Culiacan</a>, 24° 48' N, 107° 23' W, 16 vii 1963, P. J. Spangler (1 USNM); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-106.4&amp;materialsCitation.latitude=23.233334" title="Search Plazi for locations around (long -106.4/lat 23.233334)">Mazatlan</a>, blk. lite trap, 23° 14' N, 106° 24' W, 17–23 vii 1963, P. J. Spangler (2 USNM); Sonora, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-109.03333&amp;materialsCitation.latitude=27.016666" title="Search Plazi for locations around (long -109.03333/lat 27.016666)">6 mi. W Alamos</a>, 27° 1' N, 109° 2' W, 14 viii 1963, Werner &amp; Nutting (1 USNM); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-108.933334&amp;materialsCitation.latitude=27.016666" title="Search Plazi for locations around (long -108.933334/lat 27.016666)">Alamos</a>, 27° 1' N, 108° 56' W, 12 viii 1960, P. H. Arnaud, Jr., E. S. Ross, D. C. Rentz (1 CAS); same locality, 27° 1' N, 108° 56' W, 17 ix 1980, Robert Gordon (1 USNM); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-108.85&amp;materialsCitation.latitude=26.95" title="Search Plazi for locations around (long -108.85/lat 26.95)">Rio Cuchujaqui</a>, 7 mi. SE Alamos, black lite, 26° 57' N, 108° 51' W, 25 x 1972, E. M. Fisher (2 MCZ); Panama: Panama Province, Parque Nac. Soberania, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.666664&amp;materialsCitation.latitude=9.066667" title="Search Plazi for locations around (long -79.666664/lat 9.066667)">Pipeline Rd.</a>, ex. light, 9° 4' N, 79° 40' W, 9 vi 1993, M. L. Jameson (2 SEMC); U.S.A.: Texas, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.666664&amp;materialsCitation.latitude=29.983334" title="Search Plazi for locations around (long -95.666664/lat 29.983334)">Cypress Mills</a> [no date or collector], 29° 59' N, 95° 40' W, (1 MCZ); Venezuela: Aragua, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-66.76667&amp;materialsCitation.latitude=10.116667" title="Search Plazi for locations around (long -66.76667/lat 10.116667)">Ocumare</a>, 10° 7' N, 66° 46' W, 19–20 ii 1969, P. &amp; P. Spangler (5 USNM); Barinas, Pte. Parangula, 8 km S. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-70.416664&amp;materialsCitation.latitude=8.683333" title="Search Plazi for locations around (long -70.416664/lat 8.683333)">Barinitas</a>, 8° 41' N, 70° 25' W, 18 ii 1976, C. M. &amp; O. S. Flint, Jr. (1 USNM); Guarico, ~ 15 km S. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.35335&amp;materialsCitation.latitude=9.772017" title="Search Plazi for locations around (long -67.35335/lat 9.772017)">San</a> Juan, stream margin &amp; rock pools, elev. 255 m, 9° 46.321' N, 67° 21.201' W, 3 i 2006, A.E.Z. Short (AS- 06-005) (3 AEZS); Trujillo, Mcp. Rafael Rangel, La Gira, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-70.73333&amp;materialsCitation.latitude=9.333333" title="Search Plazi for locations around (long -70.73333/lat 9.333333)">Qda. La Amarilla</a>, elev. 520 m, 9° 20' N, 70° 44' W, 10–12 vi 1994, J. Camacho, M. Garcia (1 MALUZ); Mcp. Rafael Rangel, La Gira, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-70.73333&amp;materialsCitation.latitude=9.333333" title="Search Plazi for locations around (long -70.73333/lat 9.333333)">Qda. La Amarilla</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-70.73333&amp;materialsCitation.latitude=9.333333" title="Search Plazi for locations around (long -70.73333/lat 9.333333)">Trampa</a> interceptacion, elev. 520 m, 9° 20' N, 70° 44' W, 20–22 v 1995, J. Camacho, M. Garcia (3 MALUZ); Mcpio. Rafael Rangel, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-70.73333&amp;materialsCitation.latitude=9.333333" title="Search Plazi for locations around (long -70.73333/lat 9.333333)">La</a> Gira, Trampa Interceptacion, elev. 520 m, 9° 20' N, 70° 44' W, 16–17 ix 1995, M. Garcia, D. Contreras (2 MALUZ); Same locality, 23 x 1997, J. Camacho, M. Garcia, E. Gomez (3 MALUZ); Zulia, Mcpio. Lagunillas, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.23333&amp;materialsCitation.latitude=10.116667" title="Search Plazi for locations around (long -71.23333/lat 10.116667)">Zona de Reserva de Burro Negro</a>, elev. 250 m, 10° 7' N, 71° 14' W, 29–31 vii 1995, J. Comacho, M. Garcia (3 MALUZ).</p> <p>Differential diagnosis. The dorsal habitus of E. thermarum is quite similar to that of E. balfourbrownei (Figs. 86, 92); differing therefrom in the larger size (ca. 2.73 vs. 2.16 mm) and the slightly more elongate elytra which are slightly more convex and have a more abrupt posterior declivity. However, reliable determinations will require examination of the aedeagus. The aedeagus of E. thermarum is recognized by the following combination of characters: 1) the very long and very slender dorsal processes; 2) the slender median lobe, which apically is comparatively long and with narrow, pointed tips; and 3) the parameres bend outward slightly and have truncate tips. There is some variation in aedeagal form (Figs. 87, 91) across the rather large geographical distribution of E. thermarum, which ranges from Arizona to Venezuela (Fig. 133). This variation mostly involves the degree to which the median lobe distal part is narrowed, elongated, and produced in sharp tips. The most extreme variant in this respect is from Venezuela, but other specimens from Venezuela, which have a shape more like those from Panama (Fig. 91), have been seen.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 2.73/1.45; head (width) 0.68; pronotum 0.91/1.05; elytra 1.86/1.45. Habitus as illustrated (Fig. 86). Head piceous, maxillary palpi light brown; pronotum dark brown to piceous; elytra dark brown except yellowish large Vshaped fascia near midlength, and at apical 1/3 of elytra, which joins fascia laterally; venter piceous; legs rufous. Pronotum with hood carinae very slightly arcuate, almost straight, converging and confluent anteriorly, grooved throughout; secondary lateral lobe very small. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 3–4 punctures; punctures round or oval, large, each with small granule at anterior margin, non-carinate intervals usually with irregular row of granules. Protibiae slightly emarginate on inner margin, almost straight. Metaventral depression moderately deep and wide, ca. seven granules along base.</p> <p>Distribution. The most widely distributed species of Epimetopus, currently known from Venezuela north to Texas and southern Arizona (Fig. 133).</p> <p>Notes. Specimens generally have an irregular row of granules on the non-carinate elytral intervals. These granules are usually somewhat larger in specimens from Arizona than in specimens from areas south. A small percentage of specimens from Costa Rica and Venezuela lack these granules. There is little variation in size throughout the range.</p></div> 	https://treatment.plazi.org/id/3601F35E9E6D930FA3F8FF73FE4AD979	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E6F930FA3F8FC83FAC9DD83.text	3601F35E9E6F930FA3F8FC83FAC9DD83.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus balfourbrownei Rocha 1969	<div><p>Epimetopus balfourbrownei Rocha, 1969</p> <p>Figs. 92, 93 (habitus), 89, 90 (aedeagus), 127 (map)</p> <p>Epimetopus balfourbrownei Rocha 1969: 180.</p> <p>Epimetopus alvarengai Rocha 1969: 183; Oliva 1986.</p> <p>Epimetopus leechi Rocha 1969: 182; new synonymy.</p> <p>Type material: Holotype (male): Brazil, Mato Grosso, " Jacaré P.N. Xingu M. T. XI-1965, M. Alvarenga; [male symbol]; 3; [label with measurements]; Epimetopus balfourbrownei A. A. Rocha det.; HOLOTYPO [red label]" (MSP).</p> <p>Epimetopus leechi (new synonymy): Holotype (male): " BRAZIL: Matto Grosso side of Rio Araguaia, Santa Isabel; VIII-10 to 20–57 Borys Malkin; 2; [male symbol]; [label with measurements]; HOLOTIPO [red]; Epimetopus leechi A. A. Rocha det.; California Academy of Sciences Type No. 10192/" (CAS).</p> <p>New collection records: Four specimens with same locality data as holotype of E. balfourbrownei (3 FMNH, 1 USNM).</p> <p>Differential diagnosis. The dorsal habitus of E. balfourbrownei is quite similar to that of E. thermarum (Figs. 86, 92); differing therefrom in the smaller size (ca. 2.16 vs. 2.73 mm) and the slightly less elongate elytra which are slightly less convex and have a less abrupt posterior declivity. However, reliable determinations will require examination of the aedeagus. The aedeagus is similar to that of E. thermarum, but notably differs in the following: 1) the aedeagus is considerably shorter (ca. 0.65 vs. 0.80 mm); 2) the dorsal processes are wider, stouter; 3) the median lobe distal part that extends beyond the tips of the dorsal processes is shorter, wider, and less acutely pointed apically; and 4) the basal piece is shorter. The aedeagus of the holotype of E. leechi (Fig. 89) is virtually identical to the aedeagus of the holotype of E. balfourbrownei (Fig. 90); E. leechi is therefore herein synonomized with E. balfourbrownei.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 2.16/1.17; head (width) 0.62; pronotum 0.78/0.90; elytra 1.38/1.17. Habitus as illustrated (Figs. 92, 93). Head piceous, maxillary palpi light brown; pronotum dark brown to piceous; elytra dark brown except yellowish large V-shaped fascia near midlength, and very faintly yellowish apically; venter piceous; legs rufous. Pronotum with hood carinae straight, converging and confluent anteriorly, grooved throughout; secondary lateral lobes very small, almost absent. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 3–4 punctures; punctures round or oval, large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae slightly emarginate on inner margin, almost straight. Metaventral depression moderately deep and wide, ca. nine granules along base.</p> <p>Distribution. Currently known from two rather narrowly separated localities in Brazil (Fig. 127).</p></div> 	https://treatment.plazi.org/id/3601F35E9E6F930FA3F8FC83FAC9DD83	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E6E9300A3F8F911FBA3DBC1.text	3601F35E9E6E9300A3F8F911FBA3DBC1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus arcuatus Perkins 2012	<div><p>Epimetopus arcuatus, new species</p> <p>Figs. 94 (habitus), 88 (aedeagus), 128 (map)</p> <p>Type Material. Holotype (male): Paraguay: Dep. Concepcion, Arroyo Toro Paso, Unterlauf (contrib. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-56.766666&amp;materialsCitation.latitude=-23.85" title="Search Plazi for locations around (long -56.766666/lat -23.85)">Rio Apa</a>), seepage, 23° 51' S, 56° 46' W, 24 x 2002, U. Drechsel (NMW). Paratype: Same data as holotype (1 NMW).</p> <p>Differential Diagnosis. The dorsal habitus (Fig. 94) differs from that of other members of the Thermarum group by the elytral carinae which, in dorsal view, appear arcuate and, in lateral view, appear sinuate posteriorly.</p> <p>The elytral punctures are quite large and round. The pronotal lateral lobes are slightly more angulate that those of the other members of the group. The aedeagus is similar to that of E. thermarum (Figs. 87, 88), but notably differs in the following: 1) the median lobe is wider, and less acutely pointed apically; 2) the lateral margins of the parameres are virtually parallel, not bending outward; and 3) the median lobe distal part that extends beyond the tips of the dorsal processes is shorter (best seen in lateral view).</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 2.11/1.11; head (width) 0.55; pronotum 0.75/0.86; elytra 1.34/1.11. Habitus as illustrated (Fig. 94). Head piceous, maxillary palpi light brown; pronotum brown; elytra brown to dark brown except very faint lighter fascia in front of middle, and one behind middle (specimen may be slightly teneral); venter and coxae piceous, remainder of legs brown to reddish brown. Head (male) with sharply upturned clypeus. Pronotum with hood carinae slightly arcuate, converging and confluent anteriorly, grooved throughout; secondary lateral lobe small. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 3 punctures; punctures round or oval, large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae with inner margin straight, outer margin arcuate. Metaventral depression shallow, moderately wide, ca. seven granules along base.</p> <p>Etymology. Named in reference to the arcuate elytral carinae.</p> <p>Distribution. Currently known only from the type locality in Paraguay (Fig. 128).</p></div> 	https://treatment.plazi.org/id/3601F35E9E6E9300A3F8F911FBA3DBC1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E639305A3F8FF73FB92DBC1.text	3601F35E9E639305A3F8FF73FB92DBC1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus clypeatus Perkins 2012	<div><p>Epimetopus clypeatus, new species</p> <p>Figs. 95 (habitus), 97 (aedeagus), 126 (map)</p> <p>Type Material. Holotype (male): Guyana: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-58.683334&amp;materialsCitation.latitude=4.6666665" title="Search Plazi for locations around (long -58.683334/lat 4.6666665)">Region</a> 8, Iwokrama Forest, 1 km W Kurupukan, Iwokrama Field Stn., ex. mercury vapor/UV light, elev. 60 m, 4° 40' N, 58° 41' W, 20 v 2001, R. Brooks, Z. Falin (GUY1BF01 001) (SEMC).</p> <p>Differential Diagnosis. The dorsal habitus (Fig. 95) is very similar to that of other members of the Thermarum group; the pronotal lateral lobes are slightly smaller than is usual in the group. Males are distinguished from other members of the group by the distinctively upturned and medially pointed clypeus. The aedeagus (Fig. 97) differs from that of other members of the group in several respects: 1) the dorsal processes converge slightly, and at the tips are produced in a sharp point directed medially; 2) the median lobe distal part is widened and is deeply cleft in the midline; and 3) each paramere is produced in a point apically, and bear two microsetae on the point.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 1.92/1.05; head (width) 0.53; pronotum 0.68/0.73; elytra 1.30/1.05. Head piceous, maxillary palpi light brown; pronotum brown; elytra brown except very faint lighter fascia near middle, (specimen may be slightly teneral); venter and coxae piceous, remainder of legs brown to reddish brown. Head (male) with sharply upturned clypeus. Pronotum with hood carinae very slightly arcuate, converging and confluent anteriorly, grooved throughout; secondary lateral lobe small. Elytron with four strong carinae, weakly granulate, third interrupted anteriorly for distance of ca. 3 punctures; punctures round or oval, small, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae with inner margin straight, outer margin arcuate. Metaventral depression moderately wide, ca. seven granules along base.</p> <p>Etymology. Named in reference to the modified male clypeus.</p> <p>Distribution. Currently known only from the type locality in Guyana (Fig. 126).</p></div> 	https://treatment.plazi.org/id/3601F35E9E639305A3F8FF73FB92DBC1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E649304A3F8FF73FBA6D9AB.text	3601F35E9E649304A3F8FF73FBA6D9AB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus surinamensis Perkins 2012	<div><p>Epimetopus surinamensis, new species</p> <p>Figs. 96 (habitus), 98 (aedeagus), 126 (map)</p> <p>Type Material. Holotype (male): Suriname: Sipaliwini District, Camp 3, Wehepai; 2010 CI-RAP <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-56.697685&amp;materialsCitation.latitude=2.3629334" title="Search Plazi for locations around (long -56.697685/lat 2.3629334)">Survey</a>, sandy forest creek, elev. 237 m, 2° 21.776' N, 56° 41.861' W, 4–6 ix 2010, Short &amp; Kadosoe (SR10-0904-01A) (SEMC). Paratypes: Same data as holotype (2 SEMC). 77</p> <p>Differential Diagnosis. The dorsal habitus of E. surinamensis is very similar to that of E. clypeatus (Figs. 95, 96); differing therefrom in the larger size (ca. 2.44 vs. 1.92 mm), and the unmodified male clypeus. The aedeagi of the two species differ markedly (Figs. 97, 98). The aedeagus differs from that of other members of the Thermarum group in several respects: 1) the dorsal processes are relatively large, arch away from one another, and do not have ridges on the tips; 2) the median lobe distal part is notably widened, and strongly sclerotized on each side of the median slit; and 3) the parameres gradually decrease in width, to a narrow tip.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 2.44/1.32; head (width) 0.64; pronotum 0.88/0.96; elytra 1.60/1.32. Habitus as illustrated (Fig. 96). Head piceous, maxillary palpi light brown; pronotum dark brown to piceous; elytra dark brown except yellowish at Vshaped fascia near midlength, and second fascia at posterior declivity; venter and coxae black, remainder of legs brown to reddish brown. Pronotum with hood carinae slightly arcuate, converging and confluent anteriorly, grooved throughout; secondary lateral lobe small. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 3–4 punctures; punctures round or oval, large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae slightly emarginate on inner margin, outer margin arcuate. Metaventral depression moderately wide, ca. seven granules along base, shallow.</p> <p>Etymology. Named in reference to the geographical distribution.</p> <p>Distribution. Currently known only from the type locality in Suriname (Fig. 126).</p></div> 	https://treatment.plazi.org/id/3601F35E9E649304A3F8FF73FBA6D9AB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E649306A3F8FA09FBFFDBEA.text	3601F35E9E649306A3F8FA09FBFFDBEA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus plaumanni : Oliva 1986	<div><p>Epimetopus plaumanni (Costa Lima)</p> <p>Figs. 99 (habitus), 103 (aedeagus), 126 (map)</p> <p>Georyssus plaumanni Costa Lima, 1954: 177.</p> <p>Epimetopus plaumanni: Rocha, 1969: 179.</p> <p>Epimetopus plaumanni: Oliva, 1986: 11.</p> <p>Type material: Holotype (male): “ Brasilien, Nova Teutonia, 27° 11’B–52° 23’ l, 300–500 m, IX 1952, Fritz Plaumann; Georyssus plaumanni n. sp. Costa Lima det.; TIPO [red]; Epimetopus plaumanni n. comb. Costa Lima 1954, A. A. Rocha det.;10.989; HOLOTIPO [red]” (MSP).</p> <p>New collection record: Brazil: Santa Catarina, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-52.4&amp;materialsCitation.latitude=-27.05" title="Search Plazi for locations around (long -52.4/lat -27.05)">Nova Teutonia</a>, 27° 3' S, 52° 24' W, 1–30 i 1971, F. Plaumann (1 MCZ).</p> <p>Differential diagnosis. The dorsal habitus of E. plaumanni (Fig. 99) is very similar to that of other members of the Plaumanni group. It differs from the others in the larger body size (ca. 2.50 mm) and the fact that males do not have the clypeus upturned. Dissection of males will be necessary for reliable identifications. The aedeagus is similar in size and general proportions to that of E. vianai (Figs. 103, 104); it differs therefrom in the following: 1) the dorsal processes are shaped differently, being smaller and not having the apices turning outward; 2) the length of the median lobe above the dorsal processes is greater; 3) the apex of the median lobe is shaped differently; and 4) the apices of the parameres are shaped differently, in E. plaumanni being produced in a point on the medial side, a character not seen in the other species. There are also differences in the internal structure of the median lobe in the two species, E. plaumanni having the internal sac narrower and much more densely spiculate.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 2.50/1.25; head (width) 0.65; pronotum 0.88/0.96; elytra 1.69/1.25. Habitus as illustrated (Fig. 99). Head piceous, maxillary palpi light brown; pronotum dark brown to piceous; elytra dark brown except yellowish fascia near midlength; venter piceous; legs rufous. Pronotum with hood carinae very slightly arcuate, converging and confluent anteriorly, grooved throughout; secondary lateral very small, almost absent. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 3 punctures; punctures round or oval, large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae slightly emarginate on inner margin, almost straight. Metaventral depression moderately deep and wide, ca. nine granules along base.</p> <p>Distribution. Currently known only from the type locality in Brazil (Fig. 126).</p></div> 	https://treatment.plazi.org/id/3601F35E9E649306A3F8FA09FBFFDBEA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E199379A3F8FF73FEE3DE92.text	3601F35E9E199379A3F8FF73FEE3DE92.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus vianai Balfour-Browne 1949	<div><p>Epimetopus vianai Balfour-Browne, 1949</p> <p>Figs. 100 (habitus), 104 (aedeagus), 126 (map)</p> <p>Epimetopus vianai Balfour-Browne, 1949: 14.</p> <p>Epimetopus vianai: Oliva, 1986: 12.</p> <p>Type material: Holotype (male; examined): "CORDOBA— ARGENTINA Dep. de Calamuchita " El Sauce " XII - 1938, Manuel J. Viana; 17- [green label]; Type [red disc]; Brit. Mus. 1966-473; Epimetopus vianai Type J. Balfour- Browne det." (BMNH).</p> <p>Differential diagnosis. The dorsal habitus of E. vianai (Fig. 100) is very similar to that of other members of the Plaumanni group. It differs from the others in having a slightly more elongate pronotum. Males have the clypeus upturned, as do E. vulpinus and E. multiportus. Examination of the male genitalia will be essential for reliable determinations. The aedeagus is similar in size and general proportions to that of E. plaumanni (Figs. 103, 104); it differs therefrom in the following: 1) the dorsal processes are shaped differently, being larger and having the apices turning outward and sharply pointed at the tip, a character not seen in the other species; 2) the length of the median lobe above the dorsal processes is less; 3) the apex of the median lobe is shaped differently, being more deeply bifurcate in E. vianai; and 4) the apices of the parameres are shaped differently, in E. vianai being more truncate, and in E. plaumanni being produced in a point on the medial side. There are also differences in the internal structure of the median lobe in the two species, E. vianai having the internal sac wider and much less densely spiculate.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 2.44/1.20; head (width) 0.62; pronotum 0.88/0.90; elytra 1.60/1.20. Habitus as illustrated (Fig. 100). Head piceous, maxillary palpi light brown; pronotum dark brown to piceous; elytra dark brown except yellowish, large V-shaped fascia near midlength, and faintly yellowish at apical 1/3 of elytra; venter and coxae piceous, remainder of legs reddish brown; legs rufous. Pronotum with hood carinae very slightly arcuate, almost straight, converging and confluent anteriorly, grooved throughout; secondary lateral very small, almost absent. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 3 punctures; punctures round or oval, large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae straight on inner margin, outer margin weakly arcuate. Metaventral depression moderately large, ca. 9 granules along base.</p> <p>Distribution. Currently known from Argentina (Fig. 126). Oliva (1986) reported specimens from several Provinces in Argentina; these specimens were not seen during this revision, and therefore could not be confirmed as E. vianai.</p> </div>	https://treatment.plazi.org/id/3601F35E9E199379A3F8FF73FEE3DE92	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E199378A3F8FAB0FB9ADB09.text	3601F35E9E199378A3F8FAB0FB9ADB09.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus vulpinus Perkins 2012	<div><p>Epimetopus vulpinus, new species</p> <p>Figs. 101 (habitus), 105 (aedeagus), 126 (map)</p> <p>Type Material. Holotype (male): Brazil: Rio Grande do Sul, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-52.333332&amp;materialsCitation.latitude=-31.766666" title="Search Plazi for locations around (long -52.333332/lat -31.766666)">Pelotas</a>, 31° 46' S, 52° 20' W, 28 xi 1952, C. Biezanko (USNM). Paratype: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-54.383335&amp;materialsCitation.latitude=-33.533333" title="Search Plazi for locations around (long -54.383335/lat -33.533333)">Uruguay</a>: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-54.383335&amp;materialsCitation.latitude=-33.533333" title="Search Plazi for locations around (long -54.383335/lat -33.533333)">Lavalleja</a>, Rio Cebollati, Picada de Rodriguez, 33° 32' S, 54° 23' W, 5 i 1957, C. S. Carbonell (1 USNM).</p> <p>Differential Diagnosis. The dorsal habitus of E. vulpinus (Fig. 101) is very similar to that of other members of the Plaumanni group. It differs from the others in having slightly more ovate elytra, and usually a larger V-shaped fascia on the elytra. It is the smallest member of the group (ca. 2.17 mm). Males have the clypeus upturned, as do E. vianai and E. multiportus. Examination of the male genitalia will be essential for reliable determinations. The aedeagus (Fig. 105) is much broader (in both views) than others in the species group; the dorsal processes are shaped differently and are broader basally than those of other species; and the median lobe above the bases of the dorsal processes is not parallel-sided.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 2.17/1.19; head (width) 0.59; pronotum 0.81/0.87; elytra 1.44/1.19. Habitus as illustrated (Fig. 101). Head piceous, maxillary palpi light brown; pronotum dark brown to piceous; elytra dark brown except yellowish on carinae in basal ¼, large V-shaped fascia near midlength, and at apical ¼ of elytra; venter piceous; legs rufous. Pronotum with hood carinae very slightly arcuate, almost straight, converging and confluent anteriorly, grooved throughout; secondary lateral very small, almost absent. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 3 punctures; punctures round or oval, large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae slightly emarginate on inner margin, almost straight. Metaventral depression moderately deep and wide, ca. nine granules along base.</p> <p>Etymology. Named in reference to the dorsal processes of the median lobe of the aedeagus, which are shaped like fox ears.</p> <p>Distribution. Currently known from southeastern Brazil and Uruguay (Fig. 126).</p></div> 	https://treatment.plazi.org/id/3601F35E9E199378A3F8FAB0FB9ADB09	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E1B937DA3F8F8ECFBA1D856.text	3601F35E9E1B937DA3F8F8ECFBA1D856.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus multiportus Perkins 2012	<div><p>Epimetopus multiportus, new species</p> <p>Figs. 102 (habitus), 106 (aedeagus), 127 (map)</p> <p>Type Material. Holotype (male): Uruguay: 90 km SW Artigas, 31° 1' S, 57° 52' W, 27–30 ix 2001, R. Linek (NMW). Paratypes (32): Paraguay: Dep. Concepcion, Arroyo Toro Paso, Unterlauf (contrib. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-56.766666&amp;materialsCitation.latitude=-23.85" title="Search Plazi for locations around (long -56.766666/lat -23.85)">Rio Apa</a>), seepage, 23° 51' S, 56° 46' W, 24 x 2002, U. Drechsel (19 NMW); Uruguay: 90 km SW Artigas, 31° 1' S, 57° 52' W, 27–30 ix 2001, R. Linek (13 NMW).</p> <p>Differential Diagnosis. The dorsal habitus of E. multiportus (Fig. 102) is very similar to that of other members of the Plaumanni group. Males have the clypeus upturned, as do E. vianai and E. vulpinus. Examination of the male genitalia will be essential for reliable determinations. The aedeagus (Fig. 106) is smaller than that of other members of the species group (ca. 0.66 vs. 0.76 mm), and the apices of the dorsal processes fit into indentations in the median lobe, a character not seen in the other species.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 2.30/1.13; head (width) 0.59; pronotum 0.80/0.85; elytra 1.48/1.13. Habitus as illustrated (Fig. 102). Head piceous, maxillary palpi light brown; pronotum dark brown to piceous; elytra dark brown except yellowish, large V-shaped fascia near midlength, and at apical ¼ of elytra; venter and coxae piceous, remainder of legs dark brown. Pronotum with hood carinae very slightly arcuate, converging and confluent anteriorly, grooved throughout; secondary lateral very small, almost absent. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 3 punctures; punctures round or oval, large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae with inner margin straight, outer arcuate. Metaventral depression moderately deep, ca. seven granules along base.</p> <p>Distribution. Currently known from Paraguay and Uruguay (Fig. 127).</p> <p>Etymology. Named in reference to the transparent, porthole-like elytral punctures.</p></div> 	https://treatment.plazi.org/id/3601F35E9E1B937DA3F8F8ECFBA1D856	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E1D937DA3F8FBDAFC17DDB4.text	3601F35E9E1D937DA3F8FBDAFC17DDB4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus lanceolatus Perkins 2012	<div><p>Epimetopus lanceolatus, new species</p> <p>Figs. Figs. 107 (habitus), 109 (aedeagus), 128 (map)</p> <p>Type Material. Holotype (male): Brazil: Mato Grosso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-57.683334&amp;materialsCitation.latitude=-16.066668" title="Search Plazi for locations around (long -57.683334/lat -16.066668)">Caceres</a>, MT., 16° 4' S, 57° 41' W, 1–30 xii 1955, M. Alvarenga (MSP). Paratype: Brazil: Mato Grosso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-57.683334&amp;materialsCitation.latitude=-16.066668" title="Search Plazi for locations around (long -57.683334/lat -16.066668)">Caceres</a>, MT., 16° 4' S, 57° 41' W, 2–4 xii 1955, M. Alvarenga (1 CAS).</p> <p>Differential Diagnosis. Differentiated from E. lanceolatulus (Figs. 107, 108) by the much larger size (ca. 3.63 vs. 2.61 mm), the longer and more sharply arcuate pronotal hood, the different spearhead-like shape formed by the pronotal carinae, the presence of a pronotal ridge leading posteriorly from the posterior margin of the pronotal hood, and more robust elytra. In addition, the pronotal hood carinae are anteriorly confluent in E. lanceolatus, and separated in E. lanceolatulus. The aedeagus is notably broader than that of E. lanceolatulus (Figs. 109, 110), but despite the great difference in size of adults of the two species the aedeagi are of approximately the same length. Also, the parameres are shaped differently in the two species.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 3.63/2.03; head (width) 0.94; pronotum 1.34/1.44; elytra 2.44/2.03. Habitus as illustrated (Fig. 107). Head dark brown to piceous, maxillary palpi light brown, dorsum dark brown except light brown narrow transverse fascia on elytra, venter and coxae dark brown to piceous, remainder of legs brown, abdominal ventrites brown, margin slightly lighter. Pronotum with primary lateral lobes large, angulate, secondary lobes small; hood anterior margin rather sharply arcuate; hood carinae anteriorly converging, confluent at anterior tip, together forming very sharp angle; pronotal ridges strong. Elytra with carinae strong, except third carina absent in short area subbasally; non-carinate intervals each with an irregular row of small granules. Metaventral depression moderately deep and wide, ca. eight large granules along base.</p> <p>Etymology. Named in reference to the shape of the aedeagus.</p> <p>Distribution. Currently known only from the type locality (Fig. 128).</p></div> 	https://treatment.plazi.org/id/3601F35E9E1D937DA3F8FBDAFC17DDB4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E1C937EA3F8F8C3FC09DBC1.text	3601F35E9E1C937EA3F8F8C3FC09DBC1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus lanceolatulus Perkins 2012	<div><p>Epimetopus lanceolatulus, new species</p> <p>Figs. 108 (habitus), 110 (aedeagus), 128 (map)</p> <p>Type Material. Holotype (male): Brazil: Mato Grosso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-57.683334&amp;materialsCitation.latitude=-16.066668" title="Search Plazi for locations around (long -57.683334/lat -16.066668)">Caceres</a>, MT., 16° 4' S, 57° 41' W, 1–30 xii 1955, M. Alvarenga (MSP). Paratypes (11): Brazil: Mato Grosso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-56.45&amp;materialsCitation.latitude=-17.533333" title="Search Plazi for locations around (long -56.45/lat -17.533333)">Corumba</a>, 17° 32' S, 56° 27' W, 14–19 xii 1919 (3 MCZ); Paraguay: Ita, 25° 29' S, 57° 21' W, 12 iv 1980, Spangler et al. (6 USNM); Chaco, P. N. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-57.35&amp;materialsCitation.latitude=-25.483334" title="Search Plazi for locations around (long -57.35/lat -25.483334)">Defensores del Chaco</a>,</p> <p>Madrejon, at black light, 20° 38' S, 59° 52' W, 10 xii 1981, J. A. Kochalka (1 MCZ); Dep. San Pedro, [Estancia] Carumbé, 25° 33' S, 56° 40' W, 1–30 i 1971, R. Goldbach (1 MACN).</p> <p>Differential Diagnosis. Differentiated from E. lanceolatus (which see) by the much smaller body size (ca. 2.61 vs. 3.63 mm), several characters of the pronotum and elytra, and form of the aedeagus.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 2.61/1.33; head (width) 0.72; pronotum 0.85/1.04; elytra 1.74/1.33. Habitus as illustrated (Fig. 108). Head dark brown to piceous, maxillary palpi light brown, dorsum dark brown except light brown narrow transverse fascia on elytra, venter dark brown to piceous, legs dark brown. Pronotum with primary lateral lobes large, more angulate anteriorly than laterally, secondary lobes small; hood anterior margin not sharply arcuate; hood carinae very straight, not confluent at anterior tip; pronotal ridges only moderately strong. Elytra with carinae strong, except third carina absent in short area subbasally; non-carinate intervals without granules. Metaventral depression moderately deep, wide, ca. 11 granules along base; granules lateral to depression very coarse.</p> <p>Etymology. Named in reference to the shape of the aedeagus, and relatively small size, compared to E. lanceolatus.</p> <p>Distribution. Currently known from Brazil and Paraguay (Fig. 128).</p></div> 	https://treatment.plazi.org/id/3601F35E9E1C937EA3F8F8C3FC09DBC1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E119370A3F8FD56FCC3DB09.text	3601F35E9E119370A3F8FD56FCC3DB09.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus trogoides (Sharp 1875)	<div><p>Epimetopus trogoides (Sharp, 1875)</p> <p>Figs. 111 (habitus), 112 (aedeagus), 126 (map)</p> <p>Sepidulum trogoides Sharp 1874: 249.</p> <p>Epimetopus trogoides: Bedel 1880: 73; catalog.</p> <p>Epimetopus trogoides: Balfour-Browne (1949: 13, fig.1); misidentification.</p> <p>Epimetopus trogoides: Rocha (1969: 176, fig.13); misidentification.</p> <p>Epimetopus trogoides: Oliva (1986: 9); misidentification.</p> <p>Type material: Holotype (male): "Type H. T. [red disc]; Amer. mer.; Sharp Coll. 1905-313; Aedeagus drawn by P. D. Perkins 1993" (BMNH).</p> <p>New collection records: Brazil: Mato Grosso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-57.683334&amp;materialsCitation.latitude=-16.066668" title="Search Plazi for locations around (long -57.683334/lat -16.066668)">Caceres</a>, MT., 16° 4' S, 57° 41' W, 2–4 xii 1955, M. Alvarenga (1 CAS); Jacaré, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-57.7&amp;materialsCitation.latitude=-15.266666" title="Search Plazi for locations around (long -57.7/lat -15.266666)">Parque Nacional do Xingu</a>, 15° 16' S, 57° 42' W, 1–30 xi 1965, M. Alvarenga &amp; Werner (1 FMNH). Sao Paulo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.65&amp;materialsCitation.latitude=-22.733334" title="Search Plazi for locations around (long -47.65/lat -22.733334)">Piracicaba</a>, collected at black light, 22° 44' S, 47° 39' W, 7 x 1965, C. A. Triplehorn (2 USNM).</p> <p>Differential diagnosis. This species is very similar externally to E. clandestinus (Figs. 111, 115); it is larger (ca. 2.88 vs. 2.50 mm) and the pronotal lateral lobes are shaped slightly differently. Reliable determinations will require dissection of males. The aedeagus is superficially similar to that of E. deceptus (Figs. 112, 114). It differs in several respects, including the following: 1) the parameres apically are narrowed and pointed, have the lateral margin curved subapically, and have very thin walls; 2) the dorsal processes are longer; 3) the distance from the apex of the dorsal processes to the tip of the median lobe is shorter; 4) the tip of the median lobe is more acute; 5) the distance that the apex of the paramere extends beyond the tip of the dorsal process is less; 6) the basal piece is less arcuate; and 7) the oriface of the basal piece is larger.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 2.88/1.53; head (width) 0.80; pronotum 0.95/1.20; elytra 2.53/1.53. Habitus as illustrated (Fig. 111). Head dark brown to piceous, maxillary palpi light brown; dorsum dark brown with lighter fascia at about midlength of elytra; venter and coxae piceous, remainder of legs reddish brown. Pronotum with hood carinae straight, converging and very narrowly separated anteriorly; secondary lateral lobe moderately large. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 4–5 punctures; punctures round or oval, large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae slightly emarginate on inner margin, outer margin arcuate. Metaventral depression small, ca. 7 granules along base.</p> <p>Distribution. Currently known (reliable determinations) from three localities in Brazil (Fig. 126). According to Sharp (1875), in the original description, the holotype is from “South America, or possibly Mexico."</p> <p>Notes. This species has a long history of misidentifications. These errors are the result of the external similarity of several species related to E. trogoides, combined with the failure to examine the holotype aedeagus, or uncritical examination of the aedeagus of the holotype. Balfour-Browne (1949; fig. 1) illustrated an aedeagus supposedly of E. trogoides. However, it is not, and may represent the aedeagus of E. lanceolatulus n. sp., but the detail is inadequate to be sure. Rocha (1969; fig. 13) illustrated an aedeagus supposedly of E. trogoides. However, it is not, nor is it the same species as illustrated by Balfour-Browne (1949); the detail of the illustration is inadequate to positively determine what other species it represents. Likewise, Oliva (1986; fig. 5), illustrated an aedeagus supposedly of E. trogoides. However, it also is not, and appears instead to be the aedeagus of E. lanceolatulus. The distribution records given by Rocha (1969) and Oliva (1986) for E. trogoides are therefore invalid until proven, by careful examination of the aedeagi, to actually be E. trogoides. The aedeagus of the holotype of E. trogoides is illustrated herein (Fig. 112).</p> </div>	https://treatment.plazi.org/id/3601F35E9E119370A3F8FD56FCC3DB09	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E139374A3F8F917FA9CDBEA.text	3601F35E9E139374A3F8F917FA9CDBEA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus deceptus Perkins 2012	<div><p>Epimetopus deceptus, new species</p> <p>Figs. 113 (habitus), 114 (aedeagus), 129 (map)</p> <p>Type Material. Holotype (male): Brazil: Mato Grosso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-57.683334&amp;materialsCitation.latitude=-16.066668" title="Search Plazi for locations around (long -57.683334/lat -16.066668)">Caceres</a>, MT., 16° 4' S, 57° 41' W, d 16° 4' S, 57° 41' W, 2–4 xii 1955, M. Alvarenga (MSP). Paratypes (24): Brazil: Same data as holotype (1 MSP, 6 CAS); Mato Grosso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-57.683334&amp;materialsCitation.latitude=-16.066668" title="Search Plazi for locations around (long -57.683334/lat -16.066668)">Caceres</a>, MT., 16° 4' S, 57° 41' W, 1–30 xii 1955, M. Alvarenga (12 MSP); Pantanal, 17 km POCONE <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-56.633335&amp;materialsCitation.latitude=-16.266666" title="Search Plazi for locations around (long -56.633335/lat -16.266666)">Pousada</a> (farm), PORTAL PARADISO (B10), 16° 16' S, 56° 38' W, 1 viii 2000, Jan Ruzicka (5 NMPC).</p> <p>Differential Diagnosis. In dorsal habitus (Fig. 113), this species is very similar to E. trogoides and E. clandestinus, including having the carinae of the pronotal hood not confluent with one another anteriorly. It differs from E. fimbriatus in having fewer and larger tubercles on the pronotum, smaller body size (ca. 2.73 vs. 3.59 mm), and a very different aedeagus (Figs. 114, 122). The aedeagus is superficially similar to that of E. trogoides (Fig. 112). It differs in several respects, including the following: 1) the parameres apically are wider and rounded, having the lateral margin slightly emarginate subapically, and having the walls thicker, especially the medial walls; 2) the dorsal processes are shorter, more distinctly knobbed and arcuate apically, and in repose fitting into grooves in the median lobe; 3) the distance from the apex of the dorsal processes to the tip of the median lobe is greater; 4) the tip of the median lobe is more rounded; 5) the distance that the apex of the paramere extends beyond the tip of the dorsal process is greater; 6) the basal piece is more arcuate; and 7) the oriface of the basal piece is smaller.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 2.73/1.39; head (width) 0.75; pronotum 0.89/1.14; elytra 1.91/1.39. Habitus as illustrated (Fig. 113). Head dark brown to piceous, maxillary palpi light brown; dorsum dark brown with lighter fascia at about midlength of elytra; venter and coxae piceous, remainder of legs brown to reddish brown. Pronotum with hood carinae straight, converging and narrowly separated anteriorly; secondary lateral lobe moderately large; sometimes with very small third lateral lobe near base. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 3–4 punctures; punctures round or oval, large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae slightly emarginate on inner margin, outer margin arcuate. Metaventral depression moderately wide, shallow, ca. nine granules along base.</p> <p>Etymology. Named in reference to the similarity to E. trogoides and E. clandestinus.</p> <p>Distribution. Currently known from two narrowly separated localities in southwestern Brazil (Fig. 129).</p></div> 	https://treatment.plazi.org/id/3601F35E9E139374A3F8F917FA9CDBEA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E179376A3F8FF73FF24D9AA.text	3601F35E9E179376A3F8FF73FF24D9AA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus clandestinus Perkins 2012	<div><p>Epimetopus clandestinus, new species</p> <p>Figs. 115 (habitus), 116 (aedeagus), 128 (map)</p> <p>Type Material. Holotype (male): Brazil: Mato Grosso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-57.683334&amp;materialsCitation.latitude=-16.066668" title="Search Plazi for locations around (long -57.683334/lat -16.066668)">Caceres</a>, MT., 16° 4' S, 57° 41' W, 16° 4' S, 57° 41' W, 2–4 xii 1955, M. Alvarenga (MSP). Paratypes (6): Same data as holotype (1 CAS, 1 MSP); Brazil: Mato Grosso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-57.683334&amp;materialsCitation.latitude=-16.066668" title="Search Plazi for locations around (long -57.683334/lat -16.066668)">Caceres</a>, MT., 16° 4' S, 57° 41' W, 1–30 xii 1955, M. Alvarenga (3 MSP); Venezuela: Barinas, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-70.98333&amp;materialsCitation.latitude=7.5333333" title="Search Plazi for locations around (long -70.98333/lat 7.5333333)">Rio Caparo Res. Station</a>, 32 km E El Canton, seasonal forest, black light, 7° 32' N, 70° 59' W, 3–5 ii 1978, J. B. Heppner (1 USNM).</p> <p>Differential Diagnosis. This species is very similar externally to E. trogoides (Figs. 111, 115); it is smaller (ca. 2.50 vs. 2.88 mm) and the pronotal lateral lobes are shaped slightly differently. Reliable determinations will require dissection of males. The aedeagus is superficially similar to that of E. trogoides (Figs. 112, 116). It differs in several respects, including the following: 1) the parameres apically are less pointed, subapically are narrower, and the medial margin is distinctly bisinuate; 2) the dorsal processes are shorter and more arcuate; 3) the distance from the apex of the dorsal processes to the tip of the median lobe is much shorter; 4) the apex of the paramere does not attain the level of the tip of the dorsal process; 6) the basal piece is more arcuate; and 7) the oriface of the basal piece is smaller.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 2.50/1.26; head (width) 0.72; pronotum 0.80/1.06; elytra 1.70/1.26. Habitus as illustrated (Fig. 115). Head dark brown to piceous, maxillary palpi light brown; dorsum dark brown with lighter fascia at about midlength of elytra; venter and legs dark brown. Pronotum with hood carinae straight, converging and very narrowly separated anteriorly; secondary lateral lobe moderately large. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 3–4 punctures; punctures round or oval, large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae slightly emarginate on inner margin, outer margin arcuate. Metaventral depression moderately wide, ca. 10 granules along base, shallow.</p> <p>Distribution. Currently known from the type locality in Brazil, and one questionable locality in Venezuela (Fig. 128).</p> <p>Notes. The one record from Venezuela is based on a male. The aedeagus of this specimen does not significantly differ from that of the specimens from Brazil. However, the distance separating these two populations is so great that one must question whether the specimen was mislabeled. More material is needed to verify this record.</p></div> 	https://treatment.plazi.org/id/3601F35E9E179376A3F8FF73FF24D9AA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E169376A3F8FBF8FC5CDDDB.text	3601F35E9E169376A3F8FBF8FC5CDDDB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus venezuelensis Perkins 2012	<div><p>Epimetopus venezuelensis, new species</p> <p>Figs. 117 (habitus), 121 (aedeagus), 126 (map)</p> <p>Type Material. Holotype (male): Venezuela: Apure State, ca. 1 km N. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.64484&amp;materialsCitation.latitude=7.169367" title="Search Plazi for locations around (long -67.64484/lat 7.169367)">Rio Claro</a>, roadside swale, elev. 52 m, 7° 10.162' N, 67° 38.69' W, 4 i 2006, A.E.Z.Short (AS-06-009) (MIZA). Paratypes (33): Venezuela: Same data as holotype (29 SEMC); Apure, Hato El Frio, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-68.9&amp;materialsCitation.latitude=7.8166666" title="Search Plazi for locations around (long -68.9/lat 7.8166666)">Fundo Ceibote</a>, elev. 100 m, 7° 49' N, 68° 54' W, 20 v 1975, C. J. Rosales (1 MIZA); Cojedes, Hato Mata Clara, cr. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-68.35&amp;materialsCitation.latitude=9.283334" title="Search Plazi for locations around (long -68.35/lat 9.283334)">El Baul</a>, 9° 17' N, 68° 21' W, 1–3 iii 1981, F. Fernandes (2 MIZA); Guarico, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.433334&amp;materialsCitation.latitude=8.933333" title="Search Plazi for locations around (long -67.433334/lat 8.933333)">Calabozo</a>, 8° 56' N, 67° 26' W, 7 ii 1969, P. &amp; P. Spangler (1 USNM).</p> <p>Differential Diagnosis. This species differs from all others in the Trogoides group by the entirely dull, shagreened dorsal surface. It is otherwise similar in dorsal habitus to E. fimbriatus (Figs. 117, 119). It differs therefrom in some subtle characters, such as the less denticulate pronotal lateral lobes, and a slightly narrower body form. The aedeagi of the two species differ markedly, especially the shape of the apex of the median lobe, and the fact that the dorsal processes of E. venezuelensis fit into grooves in the median lobe (Figs. 121, 122). The aedeagus shows some very weak superficial similarity to that of E. deceptus (Fig. 114); however, the parameres and dorsal processes are shaped very differently in the two species.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 3.50/1.79; head (width) 0.94; pronotum 1.18/1.44; elytra 2.41/1.79. Habitus as illustrated (Fig. 117). Head dark brown to piceous, maxillary palpi light brown; dorsum dark brown with lighter fascia at about midlength of elytra and another, less distinct, at posterior declivity; venter and coxae piceous, remainder of legs reddish brown. Entire dorsal surface dull, matt. Pronotum with hood carinae straight, converging and narrowly separated anteriorly; secondary lateral lobe moderately large; smaller, third lateral lobe near base. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 3 punctures; punctures round or oval, large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae slightly emarginate on inner margin, outer margin arcuate. Metaventral depression small, ca. 7 granules along base.</p> <p>Distribution. Currently known only from Venezuela (Fig. 126).</p> <p>Etymology. Named in reference to the geographical distribution.</p></div> 	https://treatment.plazi.org/id/3601F35E9E169376A3F8FBF8FC5CDDDB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E09936BA3F8F8E4FBFFDBA2.text	3601F35E9E09936BA3F8F8E4FBFFDBA2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus tridens Perkins 2012	<div><p>Epimetopus tridens, new species</p> <p>Figs. 118 (habitus), 123 (aedeagus), 126 (map)</p> <p>Type Material. Holotype (male): Brazil: Sao Paulo, Campos do Jordão, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.6&amp;materialsCitation.latitude=-22.733334" title="Search Plazi for locations around (long -45.6/lat -22.733334)">Parque do Estado Rio Galharada</a>, 22° 44' S, 45° 36' W, 17 x 1992, Exp. MZSP (MSP). Paratypes: Same data as holotype (3 MSP).</p> <p>Differential Diagnosis. This distinctive species (Fig. 118) is differentiated from other members of the Trogoides group by several characters: 1) the longer and more sharply rounded pronotal hood; 2) the anteriorly confluent pronotal hood carinae; 3) the very small, almost absent, secondary pronotal lateral lobes, 4) the relatively deep pronotal depressions; and 5) the comparatively small elytral punctures. All carinae are well developed, and the carinae of the pronotal hood are confluent and raised anteriorly, thereby differing from all other members of the Trogoides group. Also, males have an upturned clypeus, a character not present in the others species in the group. The aedeagus (Fig. 123), which has a very long and slender median lobe, and slender parameres, cannot be confused with any other known Epimetopus. It is similar in general plan to that of E. fimbriatus (Fig. 122), a species with well developed secondary lateral lobes of the pronotum.</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 3.16/1.63; head (width) 0.82; pronotum 1.26/1.21; elytra 2.08/1.63. Habitus as illustrated (Fig. 118). Head dark brown to piceous, maxillary palpi light brown; dorsum dark brown to reddish brown with lighter V-shaped fascia at about midlength of elytra; venter and coxae reddish brown, remainder of legs ochraceous. Head (male) with clypeus upturned. Pronotum with hood carinae straight, confluent and raised anteriorly; secondary lateral lobe very small, almost absent; all ridges well raised, depressions deep. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 4–5 punctures; punctures round or oval, not large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae slightly emarginate on inner margin in distal 1/2, outer margin arcuate. Metaventral depression moderately wide, ca. 14 granules along base, shallow.</p> <p>Etymology. Named in reference to the trident-like median lobe of the aedeagus.</p> <p>Distribution. Currently known only from the type locality in Brazil (Fig. 126).</p></div> 	https://treatment.plazi.org/id/3601F35E9E09936BA3F8F8E4FBFFDBA2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E0A936FA3F8F8C1FBC7DDA0.text	3601F35E9E0A936FA3F8F8C1FBC7DDA0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus fimbriatus Perkins 2012	<div><p>Epimetopus fimbriatus, new species</p> <p>Figs. 119 (habitus), 122 (aedeagus), 126 (map)</p> <p>Type Material. Holotype (male): Brazil: Mato Grosso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.666668&amp;materialsCitation.latitude=-11.616667" title="Search Plazi for locations around (long -50.666668/lat -11.616667)">São Felix</a>, 11° 37' S, 50° 40' W, 1–30 vi 1961, M. Alvarenga (MSP). Paratype: Brazil: Mato Grosso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.666668&amp;materialsCitation.latitude=-10.833333" title="Search Plazi for locations around (long -50.666668/lat -10.833333)">Barra do Tapirapé</a>, 10° 50' S, 50° 40' W, 1–30 xi 1964, B. Malkin (1 MSP).</p> <p>Differential Diagnosis. Habitus similar to that of E. deceptus (Figs. 113, 119); differing therefrom in the smaller and denser pronotal tubercles, the larger denticles on the anterior angles of the elytra, and the larger size (ca. 3.59 v. 2.73 mm). The aedeagi of the two species are marked dissimilar (Figs. 114, 122). The aedeagus can perhaps be compared with that of E. tridens (Fig. 123); they have a similar general plan, but differ markedly in proportions. In E. fimbriatus the median lobe is much wider, as are the parameres and the basal piece; also, the dorsal processes of the median lobe are located more proximally, and the basal piece is markedly more arcuate. The habitus of E. fimbriatus and E. tridens are markedly dissimilar (Figs. 118, 119).</p> <p>Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 3.59/1.79; head (width) 0.94; pronotum 1.18/1.50; elytra 2.50/1.79. Habitus as illustrated (Fig. 119). Head dark brown to piceous, maxillary palpi light brown; dorsum dark brown with lighter fascia at about midlength of elytra; venter and coxae dark reddish brown, remainder of legs ochraceous. Pronotum with hood carinae straight, converging and well separated anteriorly; secondary lateral lobe moderately large; sometimes with smaller, third lobe near base. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 3–4 punctures; punctures round or oval, large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae slightly emarginate on inner margin, outer margin arcuate. Metaventral depression moderately deep and wide, ca. eight large granules along base.</p> <p>Etymology. Named in reference to the fimbriate lateral margins of the pronotum and anterior angles of the elytra. Distribution. Currently known only from Mato Grosso, Brazil (Fig. 126).</p></div> 	https://treatment.plazi.org/id/3601F35E9E0A936FA3F8F8C1FBC7DDA0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
3601F35E9E019360A3F8FF73FBFFDB52.text	3601F35E9E019360A3F8FF73FBFFDB52.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epimetopus tuberculatus Rocha 1969	<div><p>Epimetopus tuberculatus Rocha, 1969</p> <p>Figs. 124 (habitus), 120 (aedeagus), 126 (map)</p> <p>Epimetopus tuberculatus Rocha, 1969: 185.</p> <p>Epimetopus tuberculatus: Oliva, 1986: 10.</p> <p>Type material: Holotype (male): “ Arinos M. Gerais — Brasil, 6–8.XI.1964, Exp. Dep. Zool.; 1; [label with measurements]; Epimetopus tuberculatus A. A. Rocha det.; HOLOTIPO [red]” (MSP).</p> <p>Differential diagnosis. Immediately differentiated from all other members of the genus by the markedly tuberculate dorsum and pronotal sculpture (Fig. 124). Although the outline of the pronotum somewhat resembles that of members of the Trogoides group, the aedeagus is markedly different, much less complex and lacking dorsal processes.</p> <p>Distribution. Currently known only from the type locality in Brazil (Fig. 126).</p></div> 	https://treatment.plazi.org/id/3601F35E9E019360A3F8FF73FBFFDB52	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Perkins, Philip D.	Perkins, Philip D. (2012): 3531. Zootaxa 3531: 1-95
