taxonID	type	description	language	source
336E87BE2905FFA2E3E9B8E4FD20F8BD.taxon	description	(Figs 1 – 5)	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
336E87BE2905FFA2E3E9B8E4FD20F8BD.taxon	etymology	Etymology. Humeralis, from Latin humero (shoulder), refers to the prominent humeri of the elytra with dense yellow scales.	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
336E87BE2905FFA2E3E9B8E4FD20F8BD.taxon	diagnosis	Diagnosis. Frons (Fig. 3) with narrow light yellow spots on each side of inner margin of eye and middle of vertex. Pronotum (Figs 1 and 4) with posteriorly oriented median tubercle and with longitudinal pale yellow stripe divided obliquely into two beyond anterobasal carinae, forming an inverted V-shaped pattern reaching posterior margin of pronotum; each side of yellow stripe with irregular black spot formed by black scales. Elytral humeri (Figs 1 – 2) with tufted pale yellow scales.	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
336E87BE2905FFA2E3E9B8E4FD20F8BD.taxon	description	Description, male. Measurements (n = 2): TL = 2.9 – 3.4 mm (average = 3.1 mm); RL = 0.6 – 0.7 mm (average = 0.65 mm); RAW = 0.5 mm; RBW = 0.4 mm; PL = 0.9 – 1.0 mm (average = 0.95 mm); PW = 1.1 – 1.2 mm (average = 1.15 mm); EL = 1.8 – 2.0 mm (average = 1.9 mm); EW = 0.7 mm. Vestiture. Integument black (Figs 1 – 5). Dorsal vestiture (Figs 1 – 5) with predominant semi-decumbent blackbrownish scales intermingled with decumbent yellow and whitish scales, moderately denser on pronotum and elytra. Pronotum and elytra with semi-decumbent black scales on tufted tubercles, lateral base of pronotum and middle of elytra. Frons (Fig. 3) with narrow yellow stripe on each side of inner margin of eye and middle of vertex, with some sparse yellow scales along rostrum. Antennae with some sparse elongate and white setae, shorter and more concentrated on club. Pronotum with three tubercles tufted with elongate and black scales; median tubercle (Figs 4 – 5) with longitudinal pale yellow stripe posteriorly extended, divided obliquely in two beyond anterobasal carinae, forming an inverted V-shaped pattern reaching posterior margin of pronotum; each side of yellow stripe with irregular black spot formed by black scales. Scutellar shield (Fig. 1) with anteriorly oriented, elongate, suberect and sparse brownish-black scales. Elytra (Figs 1 – 2) with sparse decumbent white to pale yellow scales; humeri with tufted pale yellow scales, on interstria 1 with alternate dark and pale yellow vestiture, darker areas longer than yellow gap; basal tubercles with sparse pale yellow scales on anterior surfaces, with pair of irregular black stripes, oblique and posteriorly extended from the basal tubercles, diverging and reaching outer margin in middle of elytra; with a second pair of stripes having same shape, on interstria 1 in middle of elytra (divided or not) diverging to outer margin but only reaching half of elytra in four-fifths of its length. Ante-apical tubercle tufted with black scales, irregularly reaching sutural margin; without any black transverse stripe. Ventral side and legs (Figs 1 – 2) with decumbent and white scales, moderately dense; femora and tibiae with two white ring-shaped transverse stripes. Morphology. Head 1.5 times as wide as width of rostrum at base. Head and rostrum strongly punctate-corrugate. Rostrum (Figs 1 – 3) moderately long, 1.2 to 1.4 times longer than wide, 1.5 times longer than head length, slightly depressed near frons, with prominent longitudinal carinae. Antennae (Fig. 1) with segments I and II moderately thick; III 1.5 times longer than II; IV – VIII progressively smaller, club robust. Pronotum (Figs 1 and 4) slightly wider than long, subplane and punctate-corrugate, with three tubercles at transverse midline, the central tubercle evidently more developed. Ante-basal carinae almost straight, 3 - fragmented. Elytra (Figs 1 – 2) almost three times longer than wide; with prominent humeral callus, epipleura vertical and flattened; basal gibbosities formed by elongate and narrow interstriae 1 – 3, slightly more developed and subrounded at top; posteriorly to the middle with one tubercle moderately developed on interstria 3; with two ante-apical tubercles less developed, moderately acute at apex of interstriae 3 and 5. Macropterous. Ventrites I – III subequal in length at middle, each one slightly longer than IV. Ventrite V 1 / 3 times longer than IV, amply impressed at disc, subtruncate at apex. Female. Unknown.	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
336E87BE2905FFA2E3E9B8E4FD20F8BD.taxon	biology_ecology	Biology and distribution. Hylotribus humeralis sp. nov. occurs during the Southern Hemisphere summer, in December. It inhabits high altitude areas of PARNASO, in Rio de Janeiro State, around 2,120 m. It occurs in the “ Campos de Altitude ”.	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
336E87BE2905FFA2E3E9B8E4FD20F8BD.taxon	discussion	Remarks. Hylotribus humeralis sp. nov. (Figs 1 – 5) is easily distinguished from the rest of this group by the vestiture of the pronotum (Fig. 4). The median tubercle is clothed by a longitudinal pale yellow stripe that is posteriorly oriented and obliquely divided into two, and which passes the anterobasal carinae and forms an inverted V-shaped pattern reaching the posterior margin of the pronotum (present otherwise only in Hylotribus queirozi sp. nov., which lacks the elytral humeri with lateral tufted pale yellow scales). Each side of the yellow stripe has an irregular black spot formed by black scales. Additionally, on the elytra, the humeri have tufted pale yellow scales, the basal tubercles are slightly more developed than the two ante-apical tubercles along interstria 3 (developed and robust basal and ante-apical tubercles occur in H. fluminensis, and ante-apical tubercles are more developed than the basal tubercles in H. sublimis and H. gauchus), and there are no secondary tubercles or white tuft of pilosity on interstria 3 or 5 (H. fluminensis, H. sublimis and H. gauchus have white tufts of pilosity on interstria 3 and 5). Furthermore, a black transverse stripe between the ante-apical tubercles of the elytra is present in H. fluminensis, H. sublimis and H. gauchus, but lacking in H. humeralis.	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
336E87BE2905FFA2E3E9B8E4FD20F8BD.taxon	materials_examined	Type material. Holotype: BRAZIL. Rio de Janeiro. Teresópolis. Parque Nacional da Serra dos Órgãos. PVE Pt. 14 A, 22 ° 27 ’ 37.3 ” S, 43 ° 01 ’ 39.3 ” W, 2127 m, 1 male, XII. 2014, R. Monteiro col. (DZRJ). Paratype: BRAZIL. Rio de Janeiro. Teresópolis. Parque Nacional da Serra dos Órgãos. PVE Pt. 14 B, 22 ° 27 ’ 37 ” S, 43 ° 01 ’ 37.4 ” W, 2125 m, 1 male, XII. 2014, R. Monteiro col. (DZRJ).	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
336E87BE2903FFA1E3E9BA17FCEAFEA5.taxon	description	(Figs 6 – 10)	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
336E87BE2903FFA1E3E9BA17FCEAFEA5.taxon	etymology	Etymology. Nanico is a popular Brazilian word. From greek nános means ‘ dwarf’, or Latin nanus + - ico, which refers to the one that has the smaller appearance or the low body stature, like that of a dwarf. The name is a noun in apposition.	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
336E87BE2903FFA1E3E9BA17FCEAFEA5.taxon	diagnosis	Diagnosis. Dorsum of rostrum (Fig. 8), frons and anterior middle of vertex with dense yellow and decumbent scales. Pronotum (Figs 6 and 9) with moderately denser yellow scales on median third, forming longitudinal medial stripe reaching lateral tubercles of pronotum and which is larger anteriorly and with a subtriangular black spot. Femora (Fig. 7) with dark integumental area at proximal two thirds, and on tibia at middle; femora and tibia with dense yellow scales, tibiae with brownish scales at middle third. Ventral vestiture (Fig. 7) with each side of ventrites I – IV with subrounded dark spot with dense brownish scales. Elytra (Figs 6 – 7) with humeral callus on interstria 8 and slight swelling on slightly costate interstria 7; basal gibbosities moderately wide and acute at top, subequal in size to ante-apical tubercles of interstria 3; interstria 5 slightly swollen in middle; apical declivity with a moderately developed tubercle on each of interstriae 4 and 6, these moderately acute at apex, and linked by an intumescence that reaches interstria 9 at apex.	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
336E87BE2903FFA1E3E9BA17FCEAFEA5.taxon	description	Description, female. Measurements (n = 2): TL = 2.9 – 3.3 mm (average = 3.1 mm); RL = 0.5 – 0.6 mm (average = 0.55 mm); RAW = 0.6 – 0.7 mm (average = 0.65 mm); RBW = 0.5 – 0.6 mm (average = 0.55 mm); PL = 1.1 – 1.3 mm (average = 1.2 mm); PW = 1.2 – 1.5 mm (average = 1.35 mm); EL = 1.8 – 1.9 mm (average = 1.85 mm); EW = 0.8 mm. Vestiture. Integument black (Figs 6 – 10), antennae, tibiae and tarsi brown; antennal club yellowish-brown. Dorsum of rostrum (Fig. 8), frons and anterior middle of vertex with dense yellow and decumbent scales. Posterior edge of head subglabrous. Antennae with some elongate and sparse light yellow setae, shorter and more concentrated on club. Pronotum (Figs 6 and 9) with moderately denser yellow scales on median third, forming a longitudinal middle stripe reaching lateral tubercles of pronotum and anterobasal carina, and which is larger anteriorly and with a subtriangular black spot; lateral tubercles tufted with suberect black-brownish scales, median tubercle with yellow and brownish suberect scales; posterior declivity of pronotum with large and well defined stripe, almost quadrangular, clothed by yellowish scales; each side with irregular black spots. Scutellar shield (Fig. 6) with elongate, suberect sparse brownish-black scales, anteriorly oriented. Elytra (Figs 6 – 7) with yellow to white vestiture denser on anterior two thirds; humeri whiter, with tufted and denser pale yellow scales; interstria 1 with alternate dark and pale yellow vestiture, darker areas longer than yellow gaps; basal tubercles tufted with pale yellow and black-brownish scales; ante-apical tubercles with a distinct transverse dark stripe between them; apical third of elytra and pygidium with dense black-brownish scales. Femora (Fig. 7) with dark integumental area at proximal two thirds, and on tibia at middle; femora and tibia with dense yellow scales, tibiae with brownish scales at median third. Ventral vestiture (Fig. 7) with moderately dense yellow scales, each side of ventrites I – IV with subrounded dark spot with dense brownish scales. Ventrite V with suberect black-brownish scales. Morphology. Head (Fig. 8) 1.3 times as wide as width of rostrum at base. Head and rostrum microsculpture obliterated by vestiture. Rostrum short, as wide as long, as long as head length, slightly depressed near frons, with prominent longitudinal carinae. Antennae (Fig. 6) with segments I and II moderately thick; III as long as II; IV – VIII progressively shorter, VIII subequal to half of IX, club robust. Pronotum (Figs 6 and 9) slightly wider than long, coarsely punctate, the punctures partially obliterated by vestiture; with three tubercles at transverse midline, the central tubercle slightly more developed, others at sides almost indistinct. Ante-basal carinae 2 - fragmented, clearly sinuous and elongate at middle, with lateral segment short and curved. Prosternum subplane and punctate-corrugate. Scutellar shield (Fig. 6) subtriangular. Elytra (Figs 6 – 7) with humeral callus on interstria 8, and slight swelling on slightly costate interstria 7; basal gibbosities on interstria 3 moderately wide and acute at top, subequal in size with ante-apical tubercles; interstria 5 slightly swollen in middle; apical declivity with two moderately developed tubercles on interstriae 4 and 6, these moderately acute at apex and linked by an intumescence that reaches interstria 9 at apex. Apterous (checked in one specimen with raised elytra). Ventrites I – III at middle subequal in length; IV very short. Ventrite V two times longer than IV, slightly truncate at apex. Male. Unknown.	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
336E87BE2903FFA1E3E9BA17FCEAFEA5.taxon	biology_ecology	Biology and distribution. Hylotribus nanico sp. nov. occurs during the Southern Hemisphere spring (November) and autumn (April), possibly occurring also in summer. It inhabits high altitude areas of PARNASO, in Rio de Janeiro State, between 1,812 m and 1,935 m. It occurs in the high-montane forest.	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
336E87BE2903FFA1E3E9BA17FCEAFEA5.taxon	discussion	Remarks. Hylotribus nanico sp. nov. (Figs 6 – 10) differs from other species in the genus in the diagnostic characters cited above, but mainly by the vestiture of rostrum and frons, the apical declivity with two moderately developed tubercles on interstriae 4 and 6 that are moderately acute at the apex and linked by an intumescence that reaches interstria 9 at apex, and the subrounded dark spot clothed with dense brownish scales on each side of ventrites I – IV.	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
336E87BE2903FFA1E3E9BA17FCEAFEA5.taxon	materials_examined	Type material. Holotype: BRAZIL. Rio de Janeiro. Teresópolis. Parque Nacional da Serra dos Órgãos. PVE Pt. 13 B, 22 ° 27 ’ 14.3 ” S, 43 ° 01 ’ 13.2 ” W, 1935 m, 1 female, IV. 2015, R. Monteiro col. (DZRJ). Paratype: BRA- ZIL. Rio de Janeiro. Teresópolis. Parque Nacional da Serra dos Órgãos. PVE Pt. 12 A, 22 ° 27 ’ 18.2 ” S, 43 ° 00 ’ 58.9 ” W, 1812 m, 1 female, XI. 2015, R. Monteiro col. (DZRJ).	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
336E87BE2900FFAFE3E9BC3FFC03FD11.taxon	description	(Figs 11 – 15)	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
336E87BE2900FFAFE3E9BC3FFC03FD11.taxon	etymology	Etymology. The name is in honor to Fernando Ávila Queiroz, a fellow friend and coleopterist. Fernando develops research with Anthribidae, having described six species of Hylotribus with the last author, besides helping in the identification of the material used in this study. The name is a singular genitive and masculine.	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
336E87BE2900FFAFE3E9BC3FFC03FD11.taxon	diagnosis	Diagnosis. Dorsum of rostrum (Fig. 13), frons and vertex with moderately dense and decumbent yellow scales, denser on base of rostrum, near inner ocular margin and middle of vertex; each side of vertex with dark area with sparse decumbent brown scales, reaching dorso-internal margin of eyes. Pronotum (Fig. 14) with sparse yellow scales, more sparse on lateral areas; with longitudinal narrow stripe formed by pale yellow scales extending from median tubercle, becoming divided past anterobasal carinae and forming an inverted V-shaped pattern reaching posterior margin of pronotum; with three tubercles tufted with suberect black-brownish and yellow scales, the median tubercle slightly more developed; anterior third of pronotum, each side of median tubercle and posterior third of pronotum with dark irregular spots, the last past the anterobasal carina. Elytra (Figs 11 – 12) with basal gibbosities slightly more developed than ante-apical tubercles; without transverse stripe on apical third, but with two small dark spots between suture and post-median tubercles. Pygidium with narrow longitudinal stripe, clothed with dense whitish yellow scales, each side with sparse and slightly longer brown scales.	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
336E87BE2900FFAFE3E9BC3FFC03FD11.taxon	description	Description, male. Measurements (n = 1): TL = 3 mm; RL = 0.57 mm; RAW = 0.57 mm; RBW = 0.5 mm; PL = 0.96 mm; PW = 1.2 mm; EL = 2 mm; EW = 0.8 mm. Vestiture. Integument black (Figs 11 – 15), antennae, tibia and tarsus brown. Dorsum of rostrum (Fig. 13), frons and vertex with moderately dense yellow and decumbent scales, denser on base of rostrum, near inner ocular margin and at middle of vertex; each side of vertex with dark area with sparse brown and decumbent scales, reaching dorsointernal margin of eyes. Antennae with some elongate and sparse light yellow setae, shorter and more concentrated on club. Pronotum (Figs 14 – 15) with sparse yellow scales, more sparse on lateral areas; with longitudinal narrow stripe formed by pale yellow scales extending from median tubercle, divided past anterobasal carinae and forming an inverted V-shaped pattern reaching posterior margin of pronotum; with three tubercles tufted with suberect black-brownish and yellow scales, the median tubercle slightly more developed; anterior third and posterior third of pronotum, and on each side of median tubercle, with dark irregular spots, the last extending past the anterobasal carina. Scutellar shield (Fig. 11) with elongate anteriorly oriented suberect sparse whitish yellow scales. Elytra (Figs 11 – 12) with yellow to whitish yellow scales, denser on lateral areas and on posterior third; humeri with whitish yellow scales; interstria 1 with few alternate dark and whitish yellow spots, with darker spots longer than yellow gaps; interstria 5 with small white spot at midline; all tubercles tufted with pale yellow and black-brownish scales; apical third without transverse stripe, with two small dark spots between suture and post-median tubercles. Pygidium with narrow longitudinal stripe, clothed with dense whitish yellow scales, each side with sparse, slightly longer brown scales. Legs with dense whitish yellow scales; tibiae with two rings of whitish yellow scales at middle. Ventral vestiture (Fig. 12) with moderate dense yellow scales. Morphology. Head (Fig. 13) 1.3 times as wide as width of rostrum at base. Head and rostrum microsculptured, sometimes obliterated by vestiture. Rostrum short, as long as wide, slightly longer than head length, slightly depressed near frons, with prominent median longitudinal carina and pair of vestigial lateral carinae. Antennae (Fig. 12) with segments I and II slightly thick; III as long as II; IV – VIII progressively smaller, VIII subequal to half of IX; club robust. Pronotum (Figs 14 – 15) slightly wider than long, coarsely punctate and partially obliterated by vestiture; with three tubercles at transverse midline, the central tubercle slightly more developed, others at sides almost indistinct. Ante-basal carinae 3 - fragmented, clearly sinuous and elongate at middle, lateral segment short and curved. Prosternum subplane and punctate-corrugate. Scutellar shield (Fig. 11) subtriangular. Elytra (Figs 11 – 12) with humeral callus on interstria 8 and slight swelling on interstria 7; with basal gibbosities on interstria 3, these slightly more de- veloped than ante-apical tubercles; interstria 5 slightly swollen; ante-apical tubercles on interstriae 5 and 7 smaller and moderately acute at apex. Macropterous. Ventrite I, at middle, two times longer than each of II – IV, which are progressively shorter. Ventrite V one third longer than IV, impressed on disc, slightly truncate at apex. Female. Unknown.	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
336E87BE2900FFAFE3E9BC3FFC03FD11.taxon	biology_ecology	Biology and distribution. Hylotribus queirozi sp. nov. occurs during the Southern Hemisphere summer (December). It inhabits high elevation areas of PARNASO, in Rio de Janeiro State, at 1,941 m. It occurs in the highmontane forest.	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
336E87BE2900FFAFE3E9BC3FFC03FD11.taxon	discussion	Remarks. Hylotribus queirozi sp. nov. (Figs 11 – 15) differs from the other species by the diagnostic characters cited above, but mainly by having the pronotum with a longitudinal narrow anteriorly oriented pale yellow stripe extending from the median tubercle and divided past the anterobasal carinae forming an inverted V-shaped pattern reaching the posterior margin of the pronotum. This character is otherwise present only in H. humeralis sp. nov. However, the elytral humeri of H. humeralis sp. nov. have lateral tufted pale yellow scales, these lacking in H. queirozi sp. nov. It is also similar to H. plaumanni Queiroz & Mermudes, 2014, but has the basal gibbosity slightly larger than the ante-apical tubercles (H. plaumanni has the ante-apical tubercles larger than the basal gibbosity).	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
336E87BE2900FFAFE3E9BC3FFC03FD11.taxon	materials_examined	Type material. Holotype: BRAZIL. Rio de Janeiro. Teresópolis. Parque Nacional da Serra dos Órgãos. PVE Pt. 13 A, 22 ° 27 ’ 17.8 ” S, 43 ° 01 ’ 12.2 ” W, 1941 m, 1 male, XII. 2014 (DZRJ).	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
336E87BE290EFFABE3E9BFB3FD58FD85.taxon	description	(Figs 16 – 29)	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
336E87BE290EFFABE3E9BFB3FD58FD85.taxon	description	Male. Pygidium (Fig. 22) slightly wider than long, and weakly sinuous at apical margin. Ventrite V (Fig. 21) at least 1.5 times longer than IV, impressed on disc; apical margin slightly rounded. Terminalia (Figs 23 – 27): Tergite VIII (Fig. 23) largely membranous at middle; sides subparallel and slightly constricted at middle, apical margin slightly sinuous; sternite VIII (Fig. 23) strongly transverse and membranous, with apical lobes strongly pigmented and pronounced. Sternite IX (Fig. 23) with apodeme almost 2.5 x as long as arms. Tegmen (Figs 24 – 25) with apodeme as long as sclerotized ring, this with angles of pre-apical flange individualized (not bilobate); apex of parameres rounded. Penis (Figs 26 – 27) elongate, strongly curved in apical fifth; body almost 2.5 x as long as apodemes; with sclerotized bridge between apodemes; tectum two times longer than wide, strongly acuminate in apical half; pedon strongly curved, with apodemes sinuous. Internal sac (Fig. 27) slightly longer than apodemes, distinctly spiculate in proximal half, lacking sclerotized pieces. Previously known distribution. Type locality: Seara (300 – 600 m), in Santa Catarina State (Queiroz & Mermudes 2014). Updated distribution. Minas Gerais. Itamonte, in the Parque Nacional de Itatiaia, at around 2,100 m elevation. Rio de Janeiro. Teresópolis, on the Parque Nacional de Serra dos Órgãos, from 1,799 to 2,140 m. Santa Catarina. Seara. Seasonality. This species seems to occur between September and May, during the Southern Hemisphere spring, summer and autumn. Variability. H. plaumanni was described as having a V-shaped yellowish stripe with obtuse angle on the frons, and a prominent longitudinal carina on the rostrum. These characters were also used in the key to species of Hylotribus, in order to separate H. plaumanni from H. frontispeltastes and H. involucer (Queiroz & Mermudes 2014). In the specimens of H. plaumanni we observed that the V-shaped stripe was highly variable (Figs 18 – 20). It could either form an acute angle with the longitudinal carina (Figs 18 – 19), forming a diamond with another inverted Vshaped stripe on the vertex (Fig. 18), or instead be smudged (Fig. 19) and not forming a V-shaped pattern (Fig. 20). The longitudinal carina on the rostrum is always distinct, but a pair of lateral carinae can be slightly elevated or vestigial. Wing variation within the species ranges from macropterous to apterous (Figs 28 – 29), and was verified in at least three specimens with the elytra partially elevated (allowing examination of wings): 1. The paratype of H. plaumanni (apterous) from Seara, locality with altitude variation of 300 – 600 m in Santa Catarina, South of Brazil; 2. A specimen compared with the type-series, from Teresópolis, Parque Nacional da Serra dos Órgãos, Trilha da Pedra do Sino, PRE-PVE Pt. 5, 22 ° 27 ’ 16.7 ” S, 43 ° 01 ’ 13.7 ” W, 1,961 m (macropterous, Fig 28), Southeastern Brazil (Rio de Janeiro). 3. A third specimen, also compared with the type-series, from Teresópolis, Parque Nacional da Serra dos Órgãos, PVE Pt. 14 B, 22 ° 27 ’ 37 ” S, 43 ° 01 ’ 37.4 ” W, 2125 m (micropterous, Fig. 29), Southeastern Brazil (Rio de Janeiro).	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
336E87BE290EFFABE3E9BFB3FD58FD85.taxon	materials_examined	Material examined. BRAZIL. Minas Gerais. Itamonte. Parque Nacional de Itatiaia. Setor Brejo da Lapa, Brejo da Lapa, PNI-M 1 A, 22 ° 21 ’ 32.4 ” S, 44 ° 44 ’ 14.0 ” W, 2142 m, 1 male, 3. XII. 2015 - 15. I. 2016; PNI-M 1 B, 22 ° 21 ’ 34.1 ” S, 44 ° 44 ’ 9.4 ” W, 2162 m, 1 female, 3. XII. 2015 - 15. I. 2016, Monné col. (MNRJ). Rio de Janeiro. Teresópolis. Parque Nacional da Serra dos Órgãos. Trilha da Pedra do Sino. PRE-PVE Pt. 5, 22 ° 27 ’ 16.7 ” S, 43 ° 01 ’ 13.7 ” W, 1961 m, 1 female, 04. I. 2014, R. Monteiro col. (DZRJ); Teresópolis. Parque Nacional da Serra dos Órgãos. PVE Pt. 12 B, 22 ° 27 ’ 17,4 ” S, 43 ° 00 ’ 57,3 ” W, 1799 m, 1 male, IX. 2015; PVE Pt. 13 B, 22 ° 27 ’ 14.3 ” S, 43 ° 01 ’ 13.2 ” W, 1935 m, 1 female, V. 2015; PVE Pt. 14 B, 22 ° 27 ’ 37 ” S, 43 ° 01 ’ 37.4 ” W, 2125 m, 1 female, III. 2015; PVE Pt. 14 A, 22 ° 27 ’ 37.3 ” S, 43 ° 01 ’ 39.3 ” W, 2127 m, 1 female, X. 2015; PVE Pt. 15 B, 22 ° 27 ’ 37,4 ” S, 43 ° 01 ’ 42,9 ” W, 2140 m, 1 male, XII. 2014, R. Monteiro col. (DZRJ).	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
336E87BE290AFFABE3E9BFDFFF01FBEC.taxon	description	(Figs 30 – 31)	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
336E87BE290AFFABE3E9BFDFFF01FBEC.taxon	description	Updated distribution. Minas Gerais. Itamonte, on the Parque Nacional de Itatiaia, around 2,150 m. São Paulo. Campos do Jordão. Seasonality. This species seems to occur during the Southern Hemisphere spring and summer, between October and January.	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
336E87BE290AFFABE3E9BFDFFF01FBEC.taxon	materials_examined	Material examined. BRAZIL. Minas Gerais. Itamonte. Parque Nacional de Itatiaia. Setor Brejo da Lapa, Brejo da Lapa, PNI-M 1 A, 22 ° 21 ’ 32.4 ” S, 44 ° 44 ’ 14.0 ” W, 2142 m, 1 female, 3. XII. 2015 - 15. I. 2016, Monné col. (MNRJ).	en	Roza, André Silva, Mermudes, José Ricardo Miras (2019): Three new species of Hylotribus Jekel, 1860 from Brazil, with new records and discussion of species relationships and wing variation within the genus (Coleoptera Anthribidae). Zootaxa 4711 (1): 140-156, DOI: 10.11646/zootaxa.4711.1.5
