taxonID	type	description	language	source
3364878DFFBCFFCEFF06FDB2FEEBF915.taxon	materials_examined	Type species: Chelonoderus basilewskyi Hoberlandt, 1956	en	Heiss, Ernst, Grebennikov, Vasily (2015): Review and three new species of the flat bug genus Neochelonoderus Hoberlandt, 1967 from East Africa (Hemiptera: Heteroptera: Aradidae). Zootaxa 3947 (4): 581-589, DOI: 10.11646/zootaxa.3947.4.9
3364878DFFBCFFCEFF06FDB2FEEBF915.taxon	description	Redescription. Apterous, medium sized species 5.2 – 8.1 in body length. Body elongate oval (Figs. 1 – 8), surface dull with tubercles, ridges and pattern of apodemal markings on abdomen. Head. Longer than, or as long as, width of head between lateral margins of eyes; genae projecting anteriorly over clypeus, reaching apex of antennomere I, apices of variable shape; antennae slender, antennomere I clubshaped and thickest, II shortest, III longest, IV pyriform with pilose apex; antenniferous lobes prominent, projecting anterolaterally; eyes small, substylate; postocular lobes rounded or angulate adherent to eyes or exserted, posteriorly constricted to narrow neck with usually 2 pairs of larger round tubercles, posterior ones at lower level; vertex with 2 rows of round tubercles and smooth ovate callosities laterad; labium arising from slit-like atrium, as long as head, labial groove closed posteriorly. Pronotum. Subrectangular, wider than long; collar ring-like; disk with 2 (1 + 1) median large round vertical tubercles separated by longitudinal groove and 2 (1 + 1) sublateral round tubercles of similar size (Figs. 7, 8), or elongate ridges of same height (Figs. 1 – 6); posterolateral angles with round tubercle. Mesonotum. Strongly transverse, triangularly produced posteriorly, separated from metanotum by distinct suture; disk with median elevation which continues as ridge along thorax and abdomen, flanked but separated by deep groove and by 2 (1 + 1) vertical tubercles. Metanotum. Consisting of 2 (1 + 1) ovate sclerites with median roundish elevations, triangularly extending toward median elevation where they meet; posterior margin ill-defined and fused to mtg I. Mtg I + II. Mtg I medially raised and fused to mtg II, fusion marked by ill-defined transverse suture. Mtg II fully separated from tergal plate (mtg III) (Figs. 2, 4, 7, 8) or medially fused (Figs. 1, 3, 5, 6), then with rhomboidal median depression extending from mtg II – III. Abdomen. Tergal plate with longitudinal ridge carinate on mtg IV, depressed on mtg III, V – VI; lateral parts with deep apodemal impressions, deltg II and III not fused, separated by suture, surface of deltg III – VII with 2 apodemal impressions and sublateral carinae or tubercles, tergite VII of female deeply depressed at middle, raised posteriorly (Figs. 2, 3, 6, 8); tergite VII of males medially raised for reception of globular posteriorly conical pygophore (Figs. 1, 4, 5, 7). Venter. Metathoracic scent gland canal openings oval with carinate margins, visible from above; spiracles II – VII placed far from lateral margins of vltg II – VII, VIII terminal on ptg VIII visible from above. Legs. Femora moderately incrassate at middle; tibiae cylindrical, tarsi 2 - segmented, claws with setiform curved pulvilli.	en	Heiss, Ernst, Grebennikov, Vasily (2015): Review and three new species of the flat bug genus Neochelonoderus Hoberlandt, 1967 from East Africa (Hemiptera: Heteroptera: Aradidae). Zootaxa 3947 (4): 581-589, DOI: 10.11646/zootaxa.3947.4.9
3364878DFFBFFFCBFF06F96EFCE6FE47.taxon	materials_examined	Material examined. HOLOTYPE, male (CNC): “ Tanzania, Kimboza / forest / 7 ° 01 ’ 18 ’’ S, 37 ° 46 ’ 15 ’’ E / 8. i. 2012, 288 m / sift 24, V. Grebennikov // CNCCOLVG / 0 0 0 0 3935 // CNCCOLVG / 00004757 ”. PARATYPES, 3 females: “ Tanzania, Kimboza / forest / 7 ° 01 ’ 18 ’’ S, 38 ° 48 ’ 28 ’’ E / 8. i. 2012, 263 m / sift 25, V. Grebennikov // CNCCOLVG / 0 0 0 0 3936 // CNCCOLVG / 00004758 ” (CNC); “ Tanzania, Kimboza / forest / 7 ° 01 ’ 21 ’’ S, 37 ° 48 ’ 29 ’’ E / 9. i. 2012, 217 m / sift 24, V. Grebennikov // CNCCOLVG / 0 0 0 0 3942 // CNCCOLVG / 00004759 ” (CEHI); “ Tanzania, Kimboza / forest / 7 ° 01 ’ 21 ’’ S, 37 ° 48 ’ 29 ’’ E / 9. i. 2012, 217 m / sift 24, V. Grebennikov // CNCCOLVG / 00003943 ” (CEHI).	en	Heiss, Ernst, Grebennikov, Vasily (2015): Review and three new species of the flat bug genus Neochelonoderus Hoberlandt, 1967 from East Africa (Hemiptera: Heteroptera: Aradidae). Zootaxa 3947 (4): 581-589, DOI: 10.11646/zootaxa.3947.4.9
3364878DFFBFFFCBFF06F96EFCE6FE47.taxon	diagnosis	Diagnosis. Easily recognized by the wide body, the anteriorly diverging genae and the characters mentioned in the key.	en	Heiss, Ernst, Grebennikov, Vasily (2015): Review and three new species of the flat bug genus Neochelonoderus Hoberlandt, 1967 from East Africa (Hemiptera: Heteroptera: Aradidae). Zootaxa 3947 (4): 581-589, DOI: 10.11646/zootaxa.3947.4.9
3364878DFFBFFFCBFF06F96EFCE6FE47.taxon	description	Description. DNA barcode GenBank accessions: KJ 427723 (holotype), KJ 427726, KJ 427728, KJ 427729 (paratypes). Body surface subglabrous, colouration ferrugineous; left and right middle leg missing. Head. Wider than long (1.7 / 1.55); produced genae diverging anteriorly, apices rounded, with distinct basal tubercle; antenniferous lobes large, posteriorly excised, inner face angulate; antennae relatively short, 1.07 x as long as width of head, antennomeres I / II / III / IV = 0.55 / 0.275 / 0.6 / 0.4; eyes placed on anterolateral angle of wide postocular lobes which is subangularly rounded and converging posteriorly, with 2 – 3 pairs of round tubercles visible from above. Pronotum. About 2.5 x as wide as long (2.1 / 0.85); disk with prominent median tubercles and sublateral ridges; lateral margins angularly expanded their structure as of N. talaus n. sp. Mesonotum. Strongly transverse (2.55 / 0.85); median elevation triangular without longitudinal sulcus; sublateral conical elevations lower than those of pronotum; lateral margins raised and bisinuate apically. Metanotum. Posteriorly fused to mtg I, fusion line indistinct; round sublateral elevations lower than those of mesonotum. Mtg I + II. Mtg I fused to mtg II but fusion line distinct; median raised ridge with longitudinal furrow; mtg II only laterally separated from mtg III, fused at middle, rhomboidal depression of median ridge continues uninterrupted on mtg III. Abdomen. Median elevation longitudinally depressed; triangular deltg II posterolaterally expanded, posterolateral angles of deltg III – VII roundly produced, decreasing in size from deltg II – VII. Female. Similar to male (Fig. 6); tergite VII trapezoidal, depressed at middle, carinate posteriorly; ptg VIII visible as small round projections. Measurements. HOLOTYPE male: body length 6.8; length of antennae 1.83; width of abdomen across tergite V 3.25. Paratypes female: body length 6.8, 7.0, 7.3; length of antennae 1.9; ratio length of antennae / width of head 1.06; width of abdomen across tergite III 3.65, V 3.75.	en	Heiss, Ernst, Grebennikov, Vasily (2015): Review and three new species of the flat bug genus Neochelonoderus Hoberlandt, 1967 from East Africa (Hemiptera: Heteroptera: Aradidae). Zootaxa 3947 (4): 581-589, DOI: 10.11646/zootaxa.3947.4.9
3364878DFFBFFFCBFF06F96EFCE6FE47.taxon	etymology	Etymology. This species epithet is a Latinized Greek mythical name of Areius, an Argonaut, brother of Argonaut Talaus and first cousin of Jason; noun in apposition.	en	Heiss, Ernst, Grebennikov, Vasily (2015): Review and three new species of the flat bug genus Neochelonoderus Hoberlandt, 1967 from East Africa (Hemiptera: Heteroptera: Aradidae). Zootaxa 3947 (4): 581-589, DOI: 10.11646/zootaxa.3947.4.9
3364878DFFB9FFCAFF06F989FE0CFEAA.taxon	discussion	Discussion. A female labelled: “ Zaire / Prov. Kivu, Irangi / II 1986 E. Heiss (CEHI) shares all essential characters of basilewskyi (structure of head with spatuliform genae, ratio length of antennae / width of head, and elevations of pro- and mesonotum); we therefore assign it to this species. The distance between the localities of our specimen and the type of N. basilewskyi (“ Usumbura at 800 m in Burundi and “ Irangi at 820 m in North Kivu Province, DRC) is about 240 km (Fig. 9). The localities are separated by the Rift Valley depression along Tanganyika and Kivu Lakes, which are connected by the Rusizi River. Although it is questionable whether an apterous species can be found on both sides of the Rift Valley depression, we accept this for now as a temporary hypothesis pending further investigation.	en	Heiss, Ernst, Grebennikov, Vasily (2015): Review and three new species of the flat bug genus Neochelonoderus Hoberlandt, 1967 from East Africa (Hemiptera: Heteroptera: Aradidae). Zootaxa 3947 (4): 581-589, DOI: 10.11646/zootaxa.3947.4.9
3364878DFFB9FFCAFF06F989FE0CFEAA.taxon	description	Measurements. HOLOTYPE, male (MRAC), not examined, description from Hoberlandt (1956): Body length 7.6; head length / width 1.71 / 1.79; length of antennae 2.10, length of antennomeres I / II / III / IV = 0.61 / 0.38 / 0.72 / 0.49; ratio length of antennae / width of head 1.23; pronotum l / w 0.84 / 2.36; width of abdomen 3.15. Female from Irangi: Length 8.0; head l / w 1.8 / 1.75; length of antennae 2.12, antennomeres I / II / III / IV = 0.65 / 0.3 / 0.72 / 0.45; ratio length of antennae / width of head 1.21; pronotum l / w 0.9 / 2.25; width of abdomen across tergite III 3.85, IV 3.95.	en	Heiss, Ernst, Grebennikov, Vasily (2015): Review and three new species of the flat bug genus Neochelonoderus Hoberlandt, 1967 from East Africa (Hemiptera: Heteroptera: Aradidae). Zootaxa 3947 (4): 581-589, DOI: 10.11646/zootaxa.3947.4.9
3364878DFFB8FFC9FF06FE48FBCDFB6F.taxon	materials_examined	Material examined. Holotype, male (CEHI): “ Zambia 4.11.0 2 / Solwezi / 50 – 100 km West / lg. Wachtel .. Paratype, female (CEHI): “ Zambia NW Prov. / Mufumbwe to Kasempa / 22 XI 2003 / K. Werner & Smrz.	en	Heiss, Ernst, Grebennikov, Vasily (2015): Review and three new species of the flat bug genus Neochelonoderus Hoberlandt, 1967 from East Africa (Hemiptera: Heteroptera: Aradidae). Zootaxa 3947 (4): 581-589, DOI: 10.11646/zootaxa.3947.4.9
3364878DFFB8FFC9FF06FE48FBCDFB6F.taxon	diagnosis	Diagnosis. Larger species (body length 7.1 – 8.1), recognized and distinguished from other species by the more slender subparallel habitus, the shape of genae and the structure of median ridge extending from mesonotum to mtg. VI.	en	Heiss, Ernst, Grebennikov, Vasily (2015): Review and three new species of the flat bug genus Neochelonoderus Hoberlandt, 1967 from East Africa (Hemiptera: Heteroptera: Aradidae). Zootaxa 3947 (4): 581-589, DOI: 10.11646/zootaxa.3947.4.9
3364878DFFB8FFC9FF06FE48FBCDFB6F.taxon	description	Description. Body surface subglabrous and colouration ferrugineous (Fig. 7) or matte and dark brown (Fig. 8). Head. Distinctly longer than wide (1.8 / 1.47); genae finger-like and converging anteriorly leaving wide cleft between them; antenniferous lobes diverging anteriorly, apex narrowly rounded, with small tubercle at base of inner face; antennae long and slender, 1.4 x as long as width of head, antennomeres I / II / III / IV = 0.575 / 0.35 / 0.7 / 0.45; eyes oval, exserted and stylate; rectangular carinate expansion of postocular lobes separated anteriorly from eye and posteriorly from round basal tubercle by deep incisions. Pronotum. Strongly transverse (2.05 / 0.77); disk with 4 prominent conical tubercles; lateral margins angularly expanded and carinate; transverse carina of posterior margin interrupted at middle by deep median groove, lateral apices formed by large round tubercles. Mesonotum. Disk with narrow subparallel ridge with longitudinal sulcus, laterally depressed and 2 small conical tubercles; lateral margins raised and granulate. Metanotum. Posteriorly fused to mtg I; median ridge narrow extending to posterior margin of mtg II split by longitudinal furrow; disk deeply depressed laterally of median ridge, oval rugosities moderately elevated. Mtg I + II. Fused but fusion line discernible; disks depressed laterally; mtg II separated from tergal plate and mtg III by distinct transverse suture. Abdomen. The split median ridge continues on mtg III, followed by narrower keel on mtg IV, surface of median elevation widened on mtg IV and depressed on mtg V – VI; lateral margins of deltg carinate and granulate with row of larger tubercles along inner side of deltg II and III, posteroexterior angles of deltg II laterally protruding, those of deltg V – VII with round tubercles. Female. Similar to male but of larger size (Fig. 7); tergite VII depressed at middle, carinate and elevated posteriorly; tergite VIII bilobate, narrow tergite IX protruding posteriorly. Measurements. Holotype male: body length 7.1 length of antennae 2.07; width of abdomen across tergite III 2.7, V 2.8. Paratype female: body length 8.1; head l / w 1.95 / 1.65; pronotum l / w 0.9 / 2.15; length of antennae 2.22, antennomere I / II / III / IV = 0.6 / 0.35 / 0.8 / 0.47; ratio length of antennae / width of head 1.35; width of abdomen across tergite III 3.4, V 3.45.	en	Heiss, Ernst, Grebennikov, Vasily (2015): Review and three new species of the flat bug genus Neochelonoderus Hoberlandt, 1967 from East Africa (Hemiptera: Heteroptera: Aradidae). Zootaxa 3947 (4): 581-589, DOI: 10.11646/zootaxa.3947.4.9
3364878DFFB8FFC9FF06FE48FBCDFB6F.taxon	etymology	Etymology. This interesting species is dedicated to the late Ludvik Hoberlandt, who recognized this genus and contributed significantly to the knowledge of African Aradidae.	en	Heiss, Ernst, Grebennikov, Vasily (2015): Review and three new species of the flat bug genus Neochelonoderus Hoberlandt, 1967 from East Africa (Hemiptera: Heteroptera: Aradidae). Zootaxa 3947 (4): 581-589, DOI: 10.11646/zootaxa.3947.4.9
3364878DFFB8FFC9FF06FE48FBCDFB6F.taxon	discussion	Discussion. Although the geographical distance between the localities of the two type specimens is about 180 km, the similar peculiar body structures argue for their assignment to the same species.	en	Heiss, Ernst, Grebennikov, Vasily (2015): Review and three new species of the flat bug genus Neochelonoderus Hoberlandt, 1967 from East Africa (Hemiptera: Heteroptera: Aradidae). Zootaxa 3947 (4): 581-589, DOI: 10.11646/zootaxa.3947.4.9
3364878DFFBBFFC9FF06FB1CFB76F945.taxon	discussion	Discussion. The holotype female labelled: “ Congo, Parc National de l’Upemba: Kasivabilenga 700 m, 29 – 30 X 1947 (MRAC). The size of this specimen is smaller and can be distinguished at once from other congeners by the acute anteriorly diverging antenniferous lobes and the narrow contiguous genae, however all share the generic thoracic and abdominal features. The type locality, a lowland forest reserve in south-eastern part of DRC distant from the type localities of basilewskyi and the new Tanzanian species, challenges the generic assignment of this species.	en	Heiss, Ernst, Grebennikov, Vasily (2015): Review and three new species of the flat bug genus Neochelonoderus Hoberlandt, 1967 from East Africa (Hemiptera: Heteroptera: Aradidae). Zootaxa 3947 (4): 581-589, DOI: 10.11646/zootaxa.3947.4.9
3364878DFFBBFFC9FF06FB1CFB76F945.taxon	description	Measurements. HOLOTYPE, female, not seen, description from Hoberlandt (1967): Body length 6.8; head l / w 1.52 / 1.53; length of antennae 1.82, antennomeres I / II / III / IV = 0.5 / 0.34 / 0.64 / 0.34; ratio length of antennae / width of head 1.19 (incorrectly given as 1.40 in the original description); pronotum l / w 0.72 / 1.78.	en	Heiss, Ernst, Grebennikov, Vasily (2015): Review and three new species of the flat bug genus Neochelonoderus Hoberlandt, 1967 from East Africa (Hemiptera: Heteroptera: Aradidae). Zootaxa 3947 (4): 581-589, DOI: 10.11646/zootaxa.3947.4.9
3364878DFFBBFFC8FF06F927FE19FC6A.taxon	materials_examined	Material examined. HOLOTYPE, male (CNC): “ Tanzania, Pugu Hills / at reservoir S 6.91012 ° / E 39.09678 °, 166 m / 19. i. 2013, sift 50 / V. Grebennikov leg. // CNCCOLVG / 0 0 0 0 5711 // CNCCOLVG / 0 0 0 0 5712 // ”.	en	Heiss, Ernst, Grebennikov, Vasily (2015): Review and three new species of the flat bug genus Neochelonoderus Hoberlandt, 1967 from East Africa (Hemiptera: Heteroptera: Aradidae). Zootaxa 3947 (4): 581-589, DOI: 10.11646/zootaxa.3947.4.9
3364878DFFBBFFC8FF06F927FE19FC6A.taxon	diagnosis	Diagnosis. The smallest species of the genus (body length 5.2), recognized and distinguished from other species by the unique combination of characters: transverse suture separating mtg II and III uninterrupted at middle, protruding genae spatuliform, head as long as wide, median ridge of mesonotum triangular.	en	Heiss, Ernst, Grebennikov, Vasily (2015): Review and three new species of the flat bug genus Neochelonoderus Hoberlandt, 1967 from East Africa (Hemiptera: Heteroptera: Aradidae). Zootaxa 3947 (4): 581-589, DOI: 10.11646/zootaxa.3947.4.9
3364878DFFBBFFC8FF06F927FE19FC6A.taxon	description	Description. DNA barcode GenBank accession: KJ 427725 (holotype). Body surface subglabrous, colouration ferrugineous; left foreleg, right middle leg and distal part of left antenna missing. Head. As long as wide (1.2 / 1.2); spatuliform genae longer than antennomere I, slightly cleft at middle; antenniferous lobes with subparallel lateral margins, apex widely rounded, round tubercle at its base opposite to basolateral tubercle of clypeus; antennae 1.19 x as long as width of head, length of antennomeres I / II / III / IV = 0.4 / 0.275 / 0.45 / 0.35; eyes exserted and substylate; postocular lobes with angulate lateral extension and 3 pairs of round tubercles visible from above. Pronotum. Much wider than long (1.5 / 0.62); disk with 2 prominent round tubercles and 2 (1 + 1) elevated anteriorly converging sublateral ridges; lateral margins with round tubercle then carinate and converging anteriorly; carinate posterior margin terminating laterally in larger round tubercle, separated from disk by deep groove. Mesonotum. About 3 x as wide as long (1.85 / 0.6); lateral margins delimited by 2 round tubercles; median elevation triangular with longitudinal sulcus, sublateral conical tubercles prominent. Metanotum. Posteriorly fused to mtg I, fusion line ill-defined; sublateral conical elevations prominent. Mtg I + II. Fused but fusion line distinct; mtg I and mtg II medially elevated as part of continuous ridge reaching from mesonotum to mtg VI, with longitudinal impression at middle, laterally with deep apodemal markings; mtg II separated from mtg III by uninterrupted transverse suture. Abdomen. Longitudinally elevated at middle with elongate depression, short median carina developed on mtg IV; lateral margins of deltg I + II carinate, posterolateral angles of deltg II – VII with round lateral tubercle. Measurements. Body length 5.2; length of antennae 1.42; width of abdomen across tergite II 2.15, IV 2.2.	en	Heiss, Ernst, Grebennikov, Vasily (2015): Review and three new species of the flat bug genus Neochelonoderus Hoberlandt, 1967 from East Africa (Hemiptera: Heteroptera: Aradidae). Zootaxa 3947 (4): 581-589, DOI: 10.11646/zootaxa.3947.4.9
3364878DFFBBFFC8FF06F927FE19FC6A.taxon	etymology	Etymology. This species epithet is a Latinized Greek mythical name of Talaus, an Argonaut, son of Bias and Pero; noun in apposition.	en	Heiss, Ernst, Grebennikov, Vasily (2015): Review and three new species of the flat bug genus Neochelonoderus Hoberlandt, 1967 from East Africa (Hemiptera: Heteroptera: Aradidae). Zootaxa 3947 (4): 581-589, DOI: 10.11646/zootaxa.3947.4.9
