taxonID	type	description	language	source
F061AC4CF9D058AFBE418F3B0F2E2961.taxon	description	Figs 1, 2	en	Triyutthachai, Natthawut, Fu, Long-Fei, Triboun, Pramote, Wei, Yi-Gang, Pornpongrungrueng, Pimwadee (2022): Three new species of Elatostema (Urticaceae) from Thailand. PhytoKeys 215: 37-50, DOI: http://dx.doi.org/10.3897/phytokeys.215.94591, URL: http://dx.doi.org/10.3897/phytokeys.215.94591
F061AC4CF9D058AFBE418F3B0F2E2961.taxon	diagnosis	Diagnosis. Elatostema kaweesakii is similar to E. atroviride W. T. Wang but differed by its large swollen rhizome, staminate inflorescences with 2 - 7 cm long peduncle, receptacle rectangular, glabrous, staminate flower with 5 ovate tepals, pistillate inflorescences with 5 - 10 mm long peduncle, receptacle rectangular, glabrous, pistillate flower without tepals, staminode present and achene smooth (Table 1).	en	Triyutthachai, Natthawut, Fu, Long-Fei, Triboun, Pramote, Wei, Yi-Gang, Pornpongrungrueng, Pimwadee (2022): Three new species of Elatostema (Urticaceae) from Thailand. PhytoKeys 215: 37-50, DOI: http://dx.doi.org/10.3897/phytokeys.215.94591, URL: http://dx.doi.org/10.3897/phytokeys.215.94591
F061AC4CF9D058AFBE418F3B0F2E2961.taxon	materials_examined	Type. Thailand. Loei: Nong Hin district, 17 ° 7.3 ' N, 101 ° 56.05 ' E, 360 m alt., 9 August 2022, Triyutth. 332 (holotype KKU!; isotypes AAU!, BKF!).	en	Triyutthachai, Natthawut, Fu, Long-Fei, Triboun, Pramote, Wei, Yi-Gang, Pornpongrungrueng, Pimwadee (2022): Three new species of Elatostema (Urticaceae) from Thailand. PhytoKeys 215: 37-50, DOI: http://dx.doi.org/10.3897/phytokeys.215.94591, URL: http://dx.doi.org/10.3897/phytokeys.215.94591
F061AC4CF9D058AFBE418F3B0F2E2961.taxon	description	Description. Perennial herbs, lithophytic, monoecious, rhizomatous. Rhizome rounded, 2 - 20 cm in diam., brownish. Stems 15 - 50 cm tall, simple or branched, succulent, greenish, glabrous. Stipules 2, persistent or sometimes caducous in reproductive stage, linear or lanceolate, 4 - 7 x 1 - 3 mm, membranous, glabrous. Nanophylls absent. Leaves distichous, alternate; petiole 0.5 - 1 cm long, greenish, glabrous; lamina asymmetrically ovate to elliptic, 5 - 20 x 3 - 10 cm, base oblique, margin serrate to dentate, apex acuminate, chartaceous; venation pinnate, major basal lateral veins present; upper and lower surfaces greenish, glabrous, cystoliths fusiform, 0.2 - 0.5 mm long. Staminate inflorescences axillary, solitary, capitate, 1.5 - 2.0 cm in diam.; peduncle 2 - 7 cm long, glabrous or pubescent; receptacle rectangular, 0.6 - 1.0 x 0.6 - 1.0 cm, chartaceous, glabrous; bracts ovate, connate at base, membranous, glabrous; bracteoles lanceolate, 0.6 - 0.8 x 0.2 - 0.3 mm, membranous, pubescent. Staminate flowers 80 - 100 per inflorescence, sessile to subsessile, glabrous; tepals 5, ovate, 1.0 - 1.5 x 1.0 - 1.5 mm, apex acuminate, membranous, pubescent; stamens 5, filaments 1.0 - 1.5 mm long, anthers 0.6 - 0.8 mm long. Pistillate inflorescences axillary, solitary, capitate 5 - 10 mm in diam., peduncle 0.5 - 1.0 cm long, glabrous; receptacle rectangular, 6 - 8 x 6 - 8 mm, chartaceous, glabrous; bracts ovate, 0.8 - 1.0 x 0.5 - 0.6 mm, membranous, pubescent; bracteoles lanceolate, 0.4 - 0.6 x 0.2 - 0.3 mm, membranous, pubescent. Pistillate flowers 40 - 100 per inflorescence, sessile to subsessile, glabrous; tepals absent; staminodes 3, oblong to linear, 0.2 - 0.3 mm long; ovary superior, ovoid, 0.3 - 0.5 mm long. Achenes ellipsoid, 0.8 - 1.2 mm long, brownish, smooth.	en	Triyutthachai, Natthawut, Fu, Long-Fei, Triboun, Pramote, Wei, Yi-Gang, Pornpongrungrueng, Pimwadee (2022): Three new species of Elatostema (Urticaceae) from Thailand. PhytoKeys 215: 37-50, DOI: http://dx.doi.org/10.3897/phytokeys.215.94591, URL: http://dx.doi.org/10.3897/phytokeys.215.94591
F061AC4CF9D058AFBE418F3B0F2E2961.taxon	distribution	Distribution. Currently known only from Loei province in northeastern Thailand.	en	Triyutthachai, Natthawut, Fu, Long-Fei, Triboun, Pramote, Wei, Yi-Gang, Pornpongrungrueng, Pimwadee (2022): Three new species of Elatostema (Urticaceae) from Thailand. PhytoKeys 215: 37-50, DOI: http://dx.doi.org/10.3897/phytokeys.215.94591, URL: http://dx.doi.org/10.3897/phytokeys.215.94591
F061AC4CF9D058AFBE418F3B0F2E2961.taxon	etymology	Etymology. This species is named in honor of Mr Kaweesak Keeratikiat, who first discovered the plants.	en	Triyutthachai, Natthawut, Fu, Long-Fei, Triboun, Pramote, Wei, Yi-Gang, Pornpongrungrueng, Pimwadee (2022): Three new species of Elatostema (Urticaceae) from Thailand. PhytoKeys 215: 37-50, DOI: http://dx.doi.org/10.3897/phytokeys.215.94591, URL: http://dx.doi.org/10.3897/phytokeys.215.94591
F061AC4CF9D058AFBE418F3B0F2E2961.taxon	materials_examined	Additional specimens examined. Thailand. Loei: Nong Hin district, 16 December 2013, S. Saengvirotjanapat 635 (QBG!), ibid., 20 April 2021, Triyutth. 329 (KKU!).	en	Triyutthachai, Natthawut, Fu, Long-Fei, Triboun, Pramote, Wei, Yi-Gang, Pornpongrungrueng, Pimwadee (2022): Three new species of Elatostema (Urticaceae) from Thailand. PhytoKeys 215: 37-50, DOI: http://dx.doi.org/10.3897/phytokeys.215.94591, URL: http://dx.doi.org/10.3897/phytokeys.215.94591
2CB20138A742526185B08DE6D21E5BCF.taxon	description	Figs 3, 4	en	Triyutthachai, Natthawut, Fu, Long-Fei, Triboun, Pramote, Wei, Yi-Gang, Pornpongrungrueng, Pimwadee (2022): Three new species of Elatostema (Urticaceae) from Thailand. PhytoKeys 215: 37-50, DOI: http://dx.doi.org/10.3897/phytokeys.215.94591, URL: http://dx.doi.org/10.3897/phytokeys.215.94591
2CB20138A742526185B08DE6D21E5BCF.taxon	diagnosis	Diagnosis. Elatostema rubricaule differs from other Thai Elatostema by its distinct sulcate and reddish stem with flattened and disk-like rhizome and chartaceous leaves with entire margin. It grows on seasonally moist sandstone rock (Table 1).	en	Triyutthachai, Natthawut, Fu, Long-Fei, Triboun, Pramote, Wei, Yi-Gang, Pornpongrungrueng, Pimwadee (2022): Three new species of Elatostema (Urticaceae) from Thailand. PhytoKeys 215: 37-50, DOI: http://dx.doi.org/10.3897/phytokeys.215.94591, URL: http://dx.doi.org/10.3897/phytokeys.215.94591
2CB20138A742526185B08DE6D21E5BCF.taxon	materials_examined	Type. Thailand. Buengkan: Wat A-Hong Silawas, 18 ° 25.47 ' N, 103 ° 28.2 ' E, 160 m alt., 13 September 2017, Triyutth. 201 (holotype KKU!; isotypes AAU!, BKF!, IBK!, K!).	en	Triyutthachai, Natthawut, Fu, Long-Fei, Triboun, Pramote, Wei, Yi-Gang, Pornpongrungrueng, Pimwadee (2022): Three new species of Elatostema (Urticaceae) from Thailand. PhytoKeys 215: 37-50, DOI: http://dx.doi.org/10.3897/phytokeys.215.94591, URL: http://dx.doi.org/10.3897/phytokeys.215.94591
2CB20138A742526185B08DE6D21E5BCF.taxon	description	Description. Perennial herbs, lithophytic, monoecious, rhizomatous. Rhizome flattened, disk-like, 1 - 3 cm in diam., brownish. Stems 5 - 25 cm tall, simple, sulcate, succulent, reddish, glabrous. Stipules 2, persistent or sometimes caducous in reproductive stage, linear or lanceolate, 2.0 - 2.5 x 0.5 - 1.5 mm, membranous, glabrous. Nanophylls absent. Leaves distichous, alternate; petiole 0.5 - 2.0 mm long, reddish, glabrous; lamina asymmetrically lanceolate to elliptic, 3 - 5 x 0.5 - 1.5 cm, base asymmetrical attenuate, margin entire, apex obtuse, chartaceous; venation pinnate, major basal lateral veins absent, lateral veins 5 - 7 pairs; upper surface greenish, glabrous, cystolith fusiform, 0.2 - 0.5 mm long; lower surface cinereous, glabrous, cystoliths fusiform, 0.2 - 0.5 mm long. Staminate inflorescences axillary, solitary, umbellate; peduncle 2 - 3 cm long, glabrous; receptacle absent; bracts lanceolate, 0.8 - 1.0 x 0.3 - 0.5 mm, membranous, pubescent; bracteoles lanceolate, 0.6 - 0.8 x 0.2 - 0.3 mm, membranous, pubescent. Staminate flowers 5 - 10 per inflorescence; pedicel 1 - 3 mm long, glabrous; tepals 5, ovate to oblong, 1.0 - 1.5 x 1.0 - 1.5 mm, apex obtuse, membranous, glabrous; stamens 5, filaments 1.5 - 2.0 mm long, anthers 0.6 - 1.0 mm long. Pistillate inflorescences axillary, solitary, capitate 5 - 8 mm in diam., sessile to subsessile, glabrous; receptacle elliptic, 2.0 - 2.5 x 1.0 - 1.5 mm, membranous, glabrous; bracts lanceolate, 0.4 - 0.6 x 0.2 - 0.3 mm, membranous, pubescent; bracteoles lanceolate, 0.3 - 0.4 x 0.2 - 0.3 mm, membranous, pubescent. Pistillate flowers 50 - 80 per inflorescence; pedicel 0.5 - 1.0 mm long, glabrous; tepals 5, lanceolate to oblong, 0.8 - 1.2 x 0.2 - 0.4 mm, membranous, pubescent; staminodes 5, linear, 0.2 - 0.4 x 0.1 - 0.2 mm; ovary superior, ovoid, 0.3 - 0.5 mm long. Achenes ellipsoid, 0.5 - 0.8 mm long, brownish, smooth.	en	Triyutthachai, Natthawut, Fu, Long-Fei, Triboun, Pramote, Wei, Yi-Gang, Pornpongrungrueng, Pimwadee (2022): Three new species of Elatostema (Urticaceae) from Thailand. PhytoKeys 215: 37-50, DOI: http://dx.doi.org/10.3897/phytokeys.215.94591, URL: http://dx.doi.org/10.3897/phytokeys.215.94591
2CB20138A742526185B08DE6D21E5BCF.taxon	distribution	Distribution. Currently known only from the type locality in northeastern Thailand.	en	Triyutthachai, Natthawut, Fu, Long-Fei, Triboun, Pramote, Wei, Yi-Gang, Pornpongrungrueng, Pimwadee (2022): Three new species of Elatostema (Urticaceae) from Thailand. PhytoKeys 215: 37-50, DOI: http://dx.doi.org/10.3897/phytokeys.215.94591, URL: http://dx.doi.org/10.3897/phytokeys.215.94591
2CB20138A742526185B08DE6D21E5BCF.taxon	etymology	Etymology. Latin ruber, red, and caulus, stem, alluding to stem color of the new species.	en	Triyutthachai, Natthawut, Fu, Long-Fei, Triboun, Pramote, Wei, Yi-Gang, Pornpongrungrueng, Pimwadee (2022): Three new species of Elatostema (Urticaceae) from Thailand. PhytoKeys 215: 37-50, DOI: http://dx.doi.org/10.3897/phytokeys.215.94591, URL: http://dx.doi.org/10.3897/phytokeys.215.94591
2CB20138A742526185B08DE6D21E5BCF.taxon	materials_examined	Additional specimens examined. Thailand. Buengkan: Wat A-Hong Silawas, 18 ° 25.47 ' N, 103 ° 28.2 ' E, 160 m alt., 28 April 2018, Triyutth. 269 (KKU!).	en	Triyutthachai, Natthawut, Fu, Long-Fei, Triboun, Pramote, Wei, Yi-Gang, Pornpongrungrueng, Pimwadee (2022): Three new species of Elatostema (Urticaceae) from Thailand. PhytoKeys 215: 37-50, DOI: http://dx.doi.org/10.3897/phytokeys.215.94591, URL: http://dx.doi.org/10.3897/phytokeys.215.94591
F1940737100D53FA90F59F9830F991BD.taxon	description	Figs 5, 6	en	Triyutthachai, Natthawut, Fu, Long-Fei, Triboun, Pramote, Wei, Yi-Gang, Pornpongrungrueng, Pimwadee (2022): Three new species of Elatostema (Urticaceae) from Thailand. PhytoKeys 215: 37-50, DOI: http://dx.doi.org/10.3897/phytokeys.215.94591, URL: http://dx.doi.org/10.3897/phytokeys.215.94591
F1940737100D53FA90F59F9830F991BD.taxon	diagnosis	Diagnosis. Elatostema saxatile is similar to E. bulbiferum Kurz, but differs by its flattened and disk-like rhizome, receptacle of pistillate inflorescences glabrous, staminode in pistillate flower 5, acute leaf apex and its sandstone habitat (Table 1).	en	Triyutthachai, Natthawut, Fu, Long-Fei, Triboun, Pramote, Wei, Yi-Gang, Pornpongrungrueng, Pimwadee (2022): Three new species of Elatostema (Urticaceae) from Thailand. PhytoKeys 215: 37-50, DOI: http://dx.doi.org/10.3897/phytokeys.215.94591, URL: http://dx.doi.org/10.3897/phytokeys.215.94591
F1940737100D53FA90F59F9830F991BD.taxon	materials_examined	Type. Thailand. Nakhon Phanom: Tad Kham Waterfall, 17 ° 57.228 ' N, 104 ° 9.6 ' E, 160 m alt., 13 September 2017, Triyutth. 200 (holotype KKU!; isotypes AAU!, BKF!, IBK!).	en	Triyutthachai, Natthawut, Fu, Long-Fei, Triboun, Pramote, Wei, Yi-Gang, Pornpongrungrueng, Pimwadee (2022): Three new species of Elatostema (Urticaceae) from Thailand. PhytoKeys 215: 37-50, DOI: http://dx.doi.org/10.3897/phytokeys.215.94591, URL: http://dx.doi.org/10.3897/phytokeys.215.94591
F1940737100D53FA90F59F9830F991BD.taxon	description	Description. Perennial herbs, lithophytic, monoecious, rhizomatous. Rhizome flattened, disk-like, 2 - 7 cm in diam., brownish. Stems 5 - 35 cm tall, simple or branched, succulent, greenish, glabrous. Stipules 2, persistent or sometimes caducous in reproductive stage, linear or lanceolate, membranous, glabrous. Nanophylls lanceolate to oblanceolate, 0.5 - 1.5 x 0.2 - 1.0 cm, chartaceous, glabrous, cystoliths fusiform. Leaves distichous, alternate; petiole 1 - 3 mm long, glabrous or puberulous; lamina asymmetrically elliptic to obovate, 1.5 - 7.0 x 0.5 - 3.0 cm, base oblique, margin serrate, apex acute, chartaceous; venation pinnate, major basal lateral veins absent, lateral veins 5 - 7 pairs; upper surface green, glabrous, cystolith fusiform, 0.2 - 0.5 mm long; lower surface greenish or cinereous, glabrous, cystolith fusiform, 0.2 - 0.5 mm long. Staminate inflorescences axillary, solitary, umbellate; peduncle 1 - 5 cm long, glabrous; receptacle absent; bracts lanceolate, 0.6 - 1.0 x 0.3 - 0.4 mm, membranous, pubescent; bracteoles lanceolate, 0.5 - 0.8 x 0.2 - 0.3 mm, membranous, pubescent. Staminate flowers 10 - 30 per inflorescence; pedicel 1.0 - 1.5 mm long, glabrous; tepals 5, ovate to oblong, 1.0 - 1.5 x 1.0 - 1.5 mm, apex obtuse, membranous, glabrous; stamens 5, filaments 1.0 - 1.5 mm long, anthers 0.6 - 1.0 mm long. Pistillate inflorescences axillary, solitary, capitate, 2 - 8 mm in diam., subsessile to pedunculate; peduncle 0.5 - 5.0 mm long, glabrous; receptacle elliptic, 2 - 4 mm in diam., membranous, glabrous; bracts lanceolate, 0.2 - 0.3 x 0.8 - 1.0 mm, membranous, pubescent; bracteoles lanceolate, 0.2 - 0.3 x 0.5 - 0.8 mm, membranous, pubescent. Pistillate flowers 20 - 50 per inflorescence; pedicel 0.5 - 1.5 mm long, glabrous; tepals 5, lanceolate, 0.8 - 1.2 x 0.2 - 0.4 mm, membranous, pubescent; staminodes 5, oblong, 0.1 - 0.2 x 0.1 - 0.2 mm; ovary superior, ovoid, 0.4 - 0.8 mm long. Achenes ellipsoid, 0.8 - 1.2 mm long, brownish, smooth.	en	Triyutthachai, Natthawut, Fu, Long-Fei, Triboun, Pramote, Wei, Yi-Gang, Pornpongrungrueng, Pimwadee (2022): Three new species of Elatostema (Urticaceae) from Thailand. PhytoKeys 215: 37-50, DOI: http://dx.doi.org/10.3897/phytokeys.215.94591, URL: http://dx.doi.org/10.3897/phytokeys.215.94591
F1940737100D53FA90F59F9830F991BD.taxon	distribution	Distribution. Northeastern Thailand.	en	Triyutthachai, Natthawut, Fu, Long-Fei, Triboun, Pramote, Wei, Yi-Gang, Pornpongrungrueng, Pimwadee (2022): Three new species of Elatostema (Urticaceae) from Thailand. PhytoKeys 215: 37-50, DOI: http://dx.doi.org/10.3897/phytokeys.215.94591, URL: http://dx.doi.org/10.3897/phytokeys.215.94591
F1940737100D53FA90F59F9830F991BD.taxon	etymology	Etymology. The specific epithet refers to the habitat of this plant that dwelling on the rock.	en	Triyutthachai, Natthawut, Fu, Long-Fei, Triboun, Pramote, Wei, Yi-Gang, Pornpongrungrueng, Pimwadee (2022): Three new species of Elatostema (Urticaceae) from Thailand. PhytoKeys 215: 37-50, DOI: http://dx.doi.org/10.3897/phytokeys.215.94591, URL: http://dx.doi.org/10.3897/phytokeys.215.94591
F1940737100D53FA90F59F9830F991BD.taxon	materials_examined	Additional specimens examined. Thailand. Buengkan: Chet Si Waterfall, 14 September 2017, Triyutth. 202 (KKU!), ibid., 27 April 2018, Triyutth. 266 (KKU!), 267 (KKU!), Chanean Waterfall, 14 September 2017, Triyutth. 203 (KKU!), Phu Wua, 21 May 2004, R. Pooma et al. 4201 (BKF!), 4202 (BKF!), 4191 (BKF!), ibid., 28 December 2011, M. Norsaengsri & N. Tathana 8707 (QBG!), ibid., 13 July 2016, Triyutth. 97 (KKU!), 98 (KKU!), Phu Sing, 26 August 2001, R. Pooma et al. 2694 (BKF!), Phu Tok, 27 April 2018, Triyutth. 268 (KKU!), Phu Tok Noi, 22 June 1995, C. Niyomdham 4448 (BKF!), ibid., 21 June 1997, C. Niyomdham 5052 (BKF!), Wat Tham Phra, Phu Wua, 20 May 2014, S. Sirimongkol et al. 593 (BKF!); Nakhon Phanom: Tad Kham Waterfall, 25 August 2001, R. Pooma et al. 2652 (BKF!), ibid., 13 September 2017, Triyutth. 199 (KKU!).	en	Triyutthachai, Natthawut, Fu, Long-Fei, Triboun, Pramote, Wei, Yi-Gang, Pornpongrungrueng, Pimwadee (2022): Three new species of Elatostema (Urticaceae) from Thailand. PhytoKeys 215: 37-50, DOI: http://dx.doi.org/10.3897/phytokeys.215.94591, URL: http://dx.doi.org/10.3897/phytokeys.215.94591
