taxonID	type	description	language	source
2F3187B8FFF2FF84889C65EEFDF4F8CC.taxon	diagnosis	Revised diagnosis Chanbria are medium-sized eremobatids, with mature individuals ranging from 14 to 27 mm in length when measured from the distal tip of the abdomen to the distal-most portion of the chelicerae. They are generally a uniform light tan to yellowish in colour, with some individuals taking on a darker, sandy reddish brown that might be dependent on substrate association. Coloration is interspecifically variable among some species, notably among the pedipalps and legs of C. serpentinus, C. plicatus, and C. mapemes, hence these taxa are noticeably darker in overall coloration than C. regalis. Papillae are present on the palpal tarsi in all species; however, this character is absent in specimens of C. plicatus and C. regalis, possibly owing to habitat association. Leg I possess a single terminal claw (Supporting Information, Fig. S 1 A, B). The female genital opercula vary both inter- and intraspecifically (Fig. 7 A – D), with simple lobate, thin, angular, triangular, or broad anterior pillars, with laterally curving lobes. Dentition on the FF of the male chelicerae is heavily reduced or absent distally, but present proximally in C. regalis and C. mapemes (Fig. 7). The MF of the male chelicerae displays a prominent, recurved proximal tooth, with MSM, an MM, and MST, except in C. plicatus. Female chelicerae are noticeably anteroposteriorly elongate and dorsoventrally slim in comparison to other eremobatids. Females also display between seven and nine teeth on the FF (Supporting Information, Fig. S 2). Except for C. mapemes, Chanbria lack ctenidia. Synapomorphies for Chanbria include a distinctively shaped MF of the male chelicerae that forms the shape of an exponential curve (Supporting Information, Fig. S 1 C, D) and extreme morphologies of the FF not observed in any other eremobatid genus, such as sigmoidal shapes, modified folded tips, or FF in a positive slope position with respect to the manus. Pedipalpal tarsi possess a high density of setae. Legs are also slim and long (e. g. Fig. 3 B).	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFF2FF84889C65EEFDF4F8CC.taxon	discussion	Note on spine-like setae Chanbria brookharti (Fig. 8) and C. mapemes (Fig 9) are the first observed Chanbria species known to have enlarged, moveable palpal spines. Similar palpal spines have been found in other eremobatids and is a current synapomorphy for the genus Horribates Muma, 1962 (Muma, 1989). In eremobatids, such spines are motile, having a ~ 45 ° range of movement between their extended state perpendicular to the pedipalp and their relaxed state. Histological cross-sections of such spines indicate that haemolymph pressure might be responsible for extending and retracting these spines in a hydraulic fashion. Likewise, the presence of pores at the tip of these specialized spines that open into the subcutaneous tissue suggests a sensory function (Garcia EL, Laudier D, Cushing PE, unpublished data). In some eremobatid specimens that possess these spines, a notched base is present which allows the spines to be ‘ locked’ in an extended position. Similar moveable spines are also present on the appendages of other solifuge families (WGarcia EL, pers. obs.) and arachnid groups, such as in the lycosid spider Zoropsis spinimana (Dufour, 1820), which uses these spines to aid in subduing prey during capture (Eggs et al. 2015). Eremobatid solifuges have been observed capturing and manipulating prey with their pedipalps (Willemart et al. 2011), hence the palpal spines of C. brookharti, C. mapemes, and other eremobatids that possess these structures might play a similar role in subduing prey.	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFF5FF998A636430FA79FE08.taxon	description	(Figs 6 A – F, 7 A – D)	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFF5FF998A636430FA79FE08.taxon	materials_examined	Type material: Male holotype and one male paratype from Twentynine Palms, CA, USA. Female allotype from Palm Springs, CA, USA (AMNH). Holotype examined. Other material examined: México: Baja California: Ensenada: South of Punta Final, 29.743587 ° N, 114.292885 ° W, 25 April 2019, one ♂, H. Munillo; K. Munguia; E. Lopez, DMNS (DMNS ZA. 44237); USA: Arizona: Pima Co.: South of Punta Final, 29.743587 ° N, 114.292885 ° W, 25 April 2019, one ♀, H. Munillo; K. Munguia; E. Lopez, DMNS (DMNS ZA. 44238); Organ Pipe National Monument, 32.035706 ° N, 112.857283 ° W, 472 m, 23 May 1959, one ♂, K. E. Lucas, CAS (CASENT 9033503); California: Imperial Co.: near Ogilby, 32.814682 ° N, 114.841597 ° W, 545 m, 23 May 1959, three ♂, two ♀, V. Roth, CAS (CASENT 9033512); Salton Sea, 33.1872 ° N, 115.8376 ° W, July 2001, one ♀, USGS San Diego, Pitfall Trap Array, DMNS (DMNS ZA. 16444); Salton Sea, 33.1802 ° N, 115.8379 ° W, August 2001, one ♂, one ♀, USGS San Diego, Pitfall Trap Array, DMNS (DMNS ZA. 16445); Salton Sea, 33.1867 ° N, 115.8457 ° W, August 2001, one ♀, USGS San Diego, Pitfall Trap Array, DMNS (DMNS ZA. 16447); Salton Sea, 33.1974 ° N, 115.8374 ° W, 761 m, August 2001, one ♂, USGS San Diego, Pitfall Trap Array, DMNS (DMNS ZA. 16456); Salton Sea, 33.1872 ° N, 115.8376 ° W, 763 m, April 2001, one ♂, USGS San Diego, Pitfall Trap Array, DMNS (DMNS ZA. 16457); Salton Sea, 33.1868 ° N, 115.8431 ° W, 780 m, July 2001, one ♂, USGS San Diego, Pitfall Trap Array, DMNS (DMNS ZA. 16458); Salton Sea, 33.1734 ° N, 115.84 ° W, 80 7 m, July 2001, one ♂, USGS San Diego, Pitfall Trap Array, DMNS (DMNS ZA. 16459); Salton Sea, 33.1734 ° N, 115.84 ° W, July 2001, one ♂, USGS San Diego, Pitfall Trap Array, DMNS (DMNS ZA. 16460); Salton Sea, 33.1872 ° N, 115.8376 ° W, June 2001, one ♂, USGS San Diego, Pitfall Trap Array, DMNS (DMNS ZA. 16461); Salton Sea, 33.1802 ° N, 115.8379 ° W, April 2002, one ♀, USGS San Diego, Pitfall Trap Array, DMNS (DMNS ZA. 16462); Salton Sea, 33.197 ° N, 115.8452 ° W, August 2001, one ♂, USGS San Diego, Pitfall Trap Array, DMNS (DMNS ZA. 16463); Salton Sea, 33.1872 ° N, 115.8376 ° W, August 2001, one ♀, USGS San Diego, Pitfall Trap Array, DMNS (DMNS ZA. 16464); Salton Sea, 33.173 ° N, 115.8518 ° W, July 2001, one ♂, USGS San Diego, Pitfall Trap Array, DMNS (DMNS ZA. 16465); Salton Sea, 33.2039 ° N, 115.8407 ° W, August 2001, one ♀, USGS San Diego, Pitfall Trap Array, DMNS (DMNS ZA. 16466); 3 miles down dirt road off Olgilby Road headed N road on left, 32.7965 ° N, 114.871 ° W, 9 May 2001, one ♀, M. Hedin; D. Wood, DMNS (DMNS ZA. 16467); Algodones Sand Dunes Cahuilla Ranger Station Gecho Road just S of Highway 78, 32.94 ° N, 115.13 ° W, 8 October 2004, one ♀, Wendell R. Icenogle, DMNS (DMNS ZA. 17219); Algodones Sand Dunes Ted Kipf Road 11 / 2 & S of Highway 78 (GIAMIS), 32.76 ° N, 114.84 ° W, 61 m, 8 September 2005, one ♀, Wendell R. Icenogle, sand dunes below low Palo Verde tree limb, DMNS (DMNS ZA. 17248); Algodones sand dunes Ted Kipf Road 1.5 miles S Highway 78 Chamis, 33.02 ° N, 115.1 ° W, 207 m, 9 September 2005, one ♂, Wendell R. Icenogle, DMNS (DMNS ZA. 17279); Salton Sea, 33.20293 ° N, 115.84905 ° W, 98 m, August 2001, one ♂, USGS San Diego, Pitfall Trap Array, DMNS (DMNS ZA. 19092); Highway 78 at junction San Felipe Creek 10.5 miles W. junction Highway 86, 33.12563 ° N, 116.0443 ° W, 195 m, 26 September 2009, one ♂, Wendell R. Icenogle, Alcohol Sprayed, DMNS (DMNS ZA. 25437); Highway 78 at junction San Felipe Creek, 10.5 miles W. junction Highway 86, 33.12563 ° N, 116.0443 ° W, 195 m, 17 June 2010, one ♂, Wendell R. Icenogle, Alcohol Sprayed, At Lights, DMNS (DMNS ZA. 25439); Highway 78 at junction, San Felipe Creek, 10.5 miles W. junction Highway 86, 33.1256 ° N, 116.0443 ° W, 109 m, 7 June 2008, one ♂, Wendell R. Icenogle, Alcohol Sprayed, At Lights, DMNS (DMNS ZA. 25442); Highway 78 at junction, San Felipe Creek, 10.5 miles W. junction Highway 86, 33.1256 ° N, 116.0443 ° W, 7 m, 1 May 2008, one ♂, Wendell R. Icenogle, Alcohol Sprayed, At Lights, DMNS (DMNS ZA. 25443); Imperial Sand Dunes, 32.80883 ° N, 114.89218 ° W, 545 m, 15 May 2021, one ♂, one ♀, Paula E. Cushing; E. L. Garcia, Pitfall trap, DMNS (DMNS ZA. 42066); Imperial Sand Dunes, 32.8105 ° N, 114.89265 ° W, 545 m, 15 May 2021, one ♂, one ♀, Paula E. Cushing; E. L. Garcia, DMNS (DMNS ZA. 42071); San Bernardino Co.: Pisgah Crater, 34.756044 ° N, 116.38388 ° W, 392 m, 22 August 1968, one ♂, W. Savary; R. M. Haradon, CAS (CASENT _ NoNumber); Pisgah Lava Flow, 34.756044 ° N, 116.38388 ° W, 472 m, 1 August 1974, one ♂, B. Banta, CAS (CASENT 9033500); Pisgah Lava Flow, 34.756044 ° N, 116.38388 ° W, 472 m, 28 July 1956, one juvenile, B. Banta, CAS (CASENT 9033508); Pisgah Lava Flow, 34.756044 ° N, 116.38388 ° W, 488 m, 24 May 1960, one ♂, B. Banta, CAS (CASENT 9033511); Marine Corps Air Ground Combat Center, 34.2994 ° N, 116.32067 ° W, August 1999, one ♀, USGS San Diego, Pitfall Trap Array, DMNS (DMNS ZA. 16442); Marine Corps Air Ground Combat Center, 34.2998 ° N, 116.3003 ° W, July 1999, one ♀, USGS San Diego, Pitfall Trap Array, DMNS (DMNS ZA. 16443); Marine Corps Air Ground Combat Center, 34.3177 ° N, 116.26689 ° W, 816 m, October 1999, one ♀, USGS San Diego, Pitfall Trap Array, DMNS (DMNS ZA. 16448); Marine Corps Air Ground Combat Center, 34.2834 ° N, 116.2998 ° W, 780 m, July 1999, one ♂, USGS San Diego, Pitfall Trap Array, DMNS (DMNS ZA. 16449); Joshua Tree National Park, 33.90925 ° N, 115.85617 ° W, May 2000, one ♀, Jack Brookhart, DMNS (DMNS ZA. 16450); Marine Corps Air Ground Combat Center, 34.2826 ° N, 116.2414 ° W, August 1999, one ♂, USGS San Diego, Pitfall Trap Array, DMNS (DMNS ZA. 16451); Maine Corps Air Ground Combat Center, 34.3177 ° N, 116.26689 ° W, August 2000, one ♂, USGS San Diego, Pitfall Trap Array, DMNS (DMNS ZA. 16452); Marine Corps Air Ground Combat Center, 34.2998 ° N, 116.3003 ° W, 766 m, June 2001, one juvenile, USGS San Diego, Pitfall Trap Array, DMNS (DMNS ZA. 16453); Marine Corps Air Ground Combat Center, 34.2994 ° N, 116.32067 ° W, 804 m, July 1999, one ♂, USGS San Diego, Pitfall Trap Array, DMNS (DMNS ZA. 16454); Joshua Tree National Park, 33.90925 ° N, 115.85617 ° W, 539 m, May 2000, one juvenile, USGS San Diego, Pitfall Trap Array, DMNS (DMNS ZA. 16455); Twentynine Palms junction Utah Trail & Amboy Road, 34.1652 ° N, 116.0368 ° W, 207 m, 2 August 2006, one ♂, Wendell R. Icenogle, Sand dunes nearby, ground below yellowish street light, DMNS (DMNS ZA. 17246); 29 Palms West of Surprise Springs, 34.18 ° N, 116.167 ° W, 1 – 30 June 2007, three ♀, three juveniles, G. Pratt, DMNS (DMNS ZA. 22756); Coachella Valley at Train Station 1 / 2 mile S Junction Inoian Cyn Drive and Highway 10, 33.897367 ° N, 116.54835 ° W, 24 May 2007, one ♂, Wendell R. Icenogle, DMNS (DMNS ZA. 24696); Wonder Valley, Amboy Road, 8.5 miles E. Twentynine Palms (at Adobe Road junction), 34.1657 ° N, 115.9037 ° W, 7 m, 2 May 2007, one ♂, Wendell R. Icenogle, Alcohol Sprayed, DMNS (DMNS ZA. 25444); Coachella Valley, Amtrak Station, 5 miles S junction Indian Canyon Drive and Highway 10, 33.89736 ° N, 116.54835 ° W, 487 m, 7 September 2009, one ♂, Wendell R. Icenogle, DMNS (DMNS ZA. 25456); Twentynine Palms, Amboy Road 1 mile E junction Adobe Road, 34.164917 ° N, 116.038167 ° W, 550 m, 19 September 2011, one ♀, one juvenile, Wendell R. Icenogle, Alcohol Sprayed, DMNS (DMNS ZA. 28407); Twentynine Palms, AmboyRoad 1 mileEjunctionAdobeRoad, 34.164917 ° N, 116.038167 ° W, 550 m, 25 July 2011, one ♂, Wendell R. Icenogle, Alcohol Sprayed, DMNS (DMNS ZA. 28408); Twentynine Palms, Amboy Road 1 mile E junction Adobe Road, 34.164917 ° N, 116.038167 ° W, 550 m, 29 August 2011, one ♀, Wendell R. Icenogle, Alcohol Sprayed, DMNS (DMNS ZA. 28409); Twentynine Palms, Amboy Road 1 mile E junction Adobe Road, 34.164917 ° N, 116.038167 ° W, 550 m, 17 October 2011, one ♀, Wendell R. Icenogle, Alcohol Sprayed, DMNS (DMNS ZA. 28410); Twentynine Palms, Amboy Road 1 mile E junction Adobe Road, 34.164917 ° N, 116.038167 ° W, 550 m, 1 August 2009, one ♂, Wendell R. Icenogle, Alcohol Sprayed, DMNS (DMNS ZA. 28411); Amboy Road, 8.5 miles E. of Twentynine Palms (at Adobe Road Junction), 34.1657 ° N, 115.903383 ° W, 81 m, 30 July 2009, one ♀, Wendell R. Icenogle, On pavement below lights on building, DMNS (DMNS ZA. 29497); Junction Amboy Road and Utah Trail, 34.09915 ° N, 116.02208 ° W, 82 m, 21 July 2009, one ♂, Wendell R. Icenogle, DMNS (DMNS ZA. 33730); Amboy Road just W junction Utah Trail, 34.16458 ° N, 116.0382 ° W, 472 m, 26 August 2017, one ♂, Wendell R. Icenogle, At lights, On sand drifts and pavement, DMNS (DMNS ZA. 42121); Amboy Road just W junction Utah Trail, 34.16458 ° N, 116.0382 ° W, 472 m, 26 April 2017, one ♂, Wendell R. Icenogle, At lights, On sand drifts and pavement, DMNS (DMNS ZA. 42127); Amboy Road just W junction Utah Trail, 34.16458 ° N, 116.0382 ° W, 472 m, 3 September 2018, one ♂, Wendell R. Icenogle, At lights, On sand drifts and pavement, DMNS (DMNS ZA. 42128); Amboy Road, 8.5 miles E of Twentynine Palms, Wonder Valley Community Center (Blower Road), 34.16567 ° N, 115.9038 ° W, 472 m, 9 May 2007, one ♂, Wendell R. Icenogle, At lights, DMNS (DMNS ZA. 42176); Amboy Road, 8.5 miles E of Twentynine Palms, Wonder Valley Community Center (Blower Road), 34.16567 ° N, 115.9038 ° W, 17 May 2007, one ♂, Wendell R. Icenogle, At lights, DMNS (DMNS ZA. 42177); San Diego County: Amboy Road, 8.5 miles E of Twentynine Palms, Wonder Valley Community Center (Blower Road), 34.16567 ° N, 115.9038 ° W, 3 June 2007, one ♂, Wendell R. Icenogle, At lights, DMNS (DMNS ZA. 42178); Amboy Road, 8.5 miles E of Twentynine Palms, Wonder Valley Community Center, Blower Road, 34.16567 ° N, 115.9038 ° W, 2 May 2010, one ♂, Wendell R. Icenogle, At lights, DMNS (DMNS ZA. 42179); Junction of Palm Canyon Road and Borrego Valley Road, Borrego Springs, 33.255782 ° N, 116.374497 ° W, 208 m, no date, three ♂, one ♀, W. Savary; R. M. Haradon, CAS (CASENT 9033507); Ocotillo Circle, just N of junction Palm Canyon Drive, 33.25758 ° N, 116.385083 ° W, 279 m, 27 August 2008, one ♂, Wendell R. Icenogle, Alcohol Sprayed, DMNS (DMNS ZA. 25435); Ocotillo Circle, just N of junction Palm Canyon Drive, 33.257583 ° N, 116.385083 ° W, 195 m, 16 September 2009, one ♀, Wendell R. Icenogle, Alcohol Sprayed, DMNS (DMNS ZA. 25436); Ocotillo Circle, just N junction Palm Canyon Drive, 33.257583 ° N, 116.385083 ° W, 7 m, 19 August 2010, one ♂, Wendell R. Icenogle, Alcohol Sprayed, DMNS (DMNS ZA. 25440); Highway 78, Ocotillo Wells Ranger Station (State Vehicular Recreation Area), 33.155 ° N, 116.168 ° W, 195 m, 12 September 2008, one ♂, Wendell R. Icenogle, Alcohol Sprayed, DMNS (DMNS ZA. 25441); Ocotillo Circle, just N junction Palm Canyon Drive, 33.25758 ° N, 116.385083 ° W, 7 m, 12 June 2008, one ♂, Wendell R. Icenogle, Alcohol Sprayed, DMNS (DMNS ZA. 25438); Riverside County: 8 miles NW Palm Springs, 33.912332 ° N, 116.644096 ° W, 472 m, 1 May 1959, one ♂, M. Wasbauer, CAS (CASENT 9033506); Dos Palmas Reserve (San Andres Palm Oasis) 5 miles E Salton Sea State Rec Area headquarters Highway 111, 33.51 ° N, 115.89 ° W, 3 April 2004, one ♂, Wendell R. Icenogle, path to Palm Oasis, DMNS (DMNS ZA. 17210); Coachella Valley, Palm Springs Amtrak Station 1 / 2 miles S junction Indian Canyon Drive Highway I- 10, 33.8974 ° N, 116.5483 ° W, 21 May 2006, one ♀, Wendell R. Icenogle, Sand dunes, DMNS (DMNS ZA. 17213); Coachella Valley Palm Springs Amtrak Station 1 / 2 mile S junction Indian Canyon Drive and Highway I- 10, 33.8974 ° N, 116.5484 ° W, 549 m, 21 September 2006, one ♀, Wendell R. Icenogle, Netted, Sand dunes nearby, on concrete walkway below yellowish lights, DMNS (DMNS ZA. 17284); Coachella Valley Palm Springs Amtrak Station 1 / 2 mile S junction Indian Canyon Drive and Highway I- 10, 33.8974 ° N, 116.5484 ° W, 107 m, 24 May 2006, one ♂, Wendell R. Icenogle, Sand dunes nearby, pavement below yellowish lights, DMNS (DMNS ZA. 17285); Cottonwood Road, 1 / 10 mile S Junction Snow Creek Road, 33.8897725 ° N, 116.6841503 ° W, 545 m, 1 May 2011, one ♂, Wendell R. Icenogle, DMNS (DMNS ZA. 42018); Coachella Valley Palm Springs, Amtrak Station 1 / 2 mile S junction Indian Canyon Drive and Highway I- 10, 33.8974 ° N, 116.54584 ° W, 71 m, 24 May 2006, one ♀, Wendell R. Icenogle, DMNS (DMNS ZA. 17235); Coachella Valley Palm Springs Nature Station 1 / 2 mile S junction Indian Canyon Drive and Highway I- 10, 33.8974 ° N, 116.5484 ° W, 3 m, 4 September 2006, one ♀, Wendell R. Icenogle, Sand dunes nearby, on ground beside concrete walkway under yellowish lights, DMNS (DMNS ZA. 17244); Utah: Washington Co.: West of Gunlock Road, ~ 160 feet, ~ 1.8 miles south of Gunlock State Park, 37.2297 ° N, 113.780503 ° W, 1056 m, 19 July 2020, one ♂, Zach J. Valois, At lights, DMNS (DMNS ZA. 42397).	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFF5FF998A636430FA79FE08.taxon	diagnosis	Diagnosis Chanbria regalis can be distinguished from C. serpentinus by the presence of fixed finger dentition on the male chelicerae and by a more robust base of the fixed finger. This species can be distinguished from all other Chanbria by a tapering sigmoidal male FF. Individuals are generally absent of dark pigmentation.	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFF5FF998A636430FA79FE08.taxon	description	Measurements All measurements are reported as averages, and ranges are in parentheses. Males (n = 17): CL: 6.096 (3.154 – 7.829); CH: 2.060 (1.189 – 2.589); FFH: 0.587 (0.356 – 0.709); tip of MF to MM: 1.726 (1.078 – 2.407); tip of MF to MP: 2.041 (1.309 – 2.994); PL: 24.629 (20.880 – 29.289); LI: 12.506 (10.063 – 15.665); LIV: 21.618 (16.587 – 27.282); PPW: 4.079 (3.126 – 5.319); PPL: 3.014 (2.261 – 3.793); TL: 22.289 (17.467 – 27.841). Females (n = 9): CL: 6.06 (3.367 – 7.354); CH: 2.154 (1.1555 – 2.612); FFH: 0.682 (0.333 – 0.939); tip of MF to MD: 1.644 (0.733 – 2.124); tip of MF to MP: 1.830 (1.384 – 2.596); tip of FF to FD: 0.905 (0.759 – 1.020); PL: 20.403 (17.190 – 23.738); LI: 11.633 (10.335 – 13.182); LIV: 21.268 (18.046 – 23.454); PPW: 4.385 (3.832 – 5.200); PPL: 2.977 (2.219 – 4); TL: 23.467 (19.818 – 26.549); GOL: 1.401 (1.244 – 1.557); GOW: 0.619 (0.492 – 0.746); GOD: 1.930 (1.7425 – 2.117). Redescription: males Coloration: Overall coloration light tan. Legs infuscate distally near ends of tibiae, femurs, and tarsi. Pedipalps darken near end of the tibiae. Palpal tibia, metatarsi, and tarsi darker reddish orange. Propeltidium darkens slightly along margins. FF and MF of the chelicerae darken distally. Chelicera: FF sigmoidal, with dentition present ventrally. In some populations, dentition is reduced. Between three and four teeth present on MF. MP tooth largest, prominent, and recurved. When present, MSD tooth is smallest. Fondal notch absent. Fondal teeth vary in length. Redescription: females Coloration: Coloration as in males. Chelicera: FF with seven to nine teeth. The following FF states were recorded among C. regalis females: seven teeth with FF formula FD- 2 FSD-FM- 2 FSM-FD, eight teeth with FD- 2 FSD-FM- 3 FSM-FP, or nine teeth with FD- 3 FSD-FM- 3 FSM-FP. The last two states are the most common. Moveable finger with four teeth. MP tooth largest, followed by MM tooth, with two smaller MSM teeth between them. Many of the chelicerae examined have FST and MST. Operculum: Opercular morphology is highly variable between individual females. AMNH allotype with rounded opercular plates resembling isosceles triangles. Other morphologies present include elongate tapered lobes in specimen DMNS ZA. 16442, broad anterior pillars with laterally curving lobes resembling an L-shape in specimen DMNS ZA. 17213, or elongate, ovoid lobes as in specimen DMNS ZA. 44238 (Fig. 7).	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFF5FF998A636430FA79FE08.taxon	distribution	Distribution Sonoran Desert. This species is wide ranging, occurring from the southwestern corner of Utah to the Sonoran Desert / Peninsular Desert in Baja California. It is restricted primarily to the eastern side of the Colorado River. One record is from the western side on the southern end of the distribution in Pima County, Arizona, suggesting that they might occur on both sides of the southern end of the river.	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFF5FF998A636430FA79FE08.taxon	discussion	Discussion Individuals of this widespread species are notable for a high level of intraspecific morphological variation in both the male chelicerae and the female genital operculum (Fig. 7 A – D), and these structures have been used historically for species delimitation in eremobatids (Muma 1951, 1962). Such variability, especially in the relative lengths of the fondal teeth and general shape of the FF of the male chelicerae, challenges previous taxonomic assumptions about the usefulness of these ‘ sexually selected’ structures for delimiting species within the genus. Chanbria regalis and C. rectus were differentiated by Muma (1962) based on the lack of curvature of the dorsal profile of the male FF (Supporting Information, Fig. S 5) and the presence or absence of palpal scopula (papillae). However, the results of the PCAs from the SNP and mitochondrial dataset, the small K 80 genetic distance, the relatively recent divergence time, and the lack of a distinct geographical barrier support the synonymization of these two species. More sampling from populations across the C. regalis range is necessary to determine whether the variable morphological traits we observed are stable across the geographical range or are aberrant traits. Given the little divergence in genetic data and inconsistent morphological traits, we are therefore in favour of a single-species hypothesis that maintains subtle variation of traits across populations. Owing to this, our genetic results, and the level of intraspecific variation that we observe within this clade, in addition to our observation of the type material, we suggest the synonymization of Chanbria tehachapianus with C. regalis. This decision is motivated by the nearly identical morphology that C. tehachapianus shares with individuals of C. regalis, and the diagnostics used to erect the species [i. e. constricted base at the FF and lack of scopula (also known as papillae; Muma 1962)] are insufficient to maintain the species status confidently. Additionally, a lack of known specimens other than the holotype identified as C. tehachapianus and the lack of a distinct geographical barrier between its collection locality and the known range of C. regalis are evidence in favour of synonymizing these two species. The high degree of variable morphology in the opercula of female C. regalis suggests that mating complementarity in C. regalis is not dependent on the shape of the genital opercular plate, and instead internal structures might be more important for facilitating mating complementarity than external structures, as noted in other studies (Garcia et al. 2024). Alternatively, this variation might simply be explained by the recent divergence and is an example of characters that have yet to reach fixation.	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFE8FF9A88F262F5FE63F9FE.taxon	description	(Figs 6 I, J, 7 E, F)	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFE8FF9A88F262F5FE63F9FE.taxon	materials_examined	Type material: Male holotype from Tucson, AZ, USA (AMNH). Examined. Other material examined: USA: Arizona: Pima Co.: Posta Quemado Canyon 8 miles N Vail, came to UV light, 32.0543 ° N, 110.634 ° W, 1045 m, 30 May 1968, one ♀, F. Werner, DMNS (DMNS ZA. 17735); Catalina State Park, 32.42445 ° N, 110.92271 ° W, 822 m, 27 April 2011, one ♂, P. E. Cushing; G. Selby, At Lights, DMNS (DMNS ZA. 26433); Catalina State Park; Campsite A and B, 32.42445 ° N, 110.92271 ° W, 822 m, 4 June 2011, one ♀, P. E. Cushing; P. Casto; B. E. Trierweiler; C. G. Olds, Lantern Light, DMNS (DMNS ZA. 26852); 4020 S Via del Pica Maderos, 31.8542 ° N, 110.9937 ° W, 886 m, 13 July 2011, one ♀, W. R. Savary, Casual, DMNS (DMNS ZA. 28202); Catalina State Park, campsite A and B, 32.42445 ° N, 110.92271 ° W, 822 m, 6 Jun 2011, one ♂, C. Ho, Casual, DMNS (DMNS ZA. 17648); Catalina State Park Campgrounds A and B; 32.42630 − 110.92372, 32.4263 ° N, 110.92372 ° W, 825 m, 13 June 2018, five ♂, two ♀, P. E. Cushing; F. Channiago; L. Herrera; R. R. Jones, At lights, DMNS (DMNS ZA. 38728); Catalina State Park Campgrounds A and B, 32.4263 ° N, 110.92372 ° W, 825 m, 13 June 2018, one ♂, P. E. Cushing; F. Channiago; L. Herrera; R. R. Jones, At lights, DMNS (DMNS ZA. 38729); Catalina State Park Campgrounds A and B, 32.4263 ° N, 110.92372 ° W, 825 m, 12 July 2018, one ♂, P. E. Cushing; F. Channiago; L. Herrera; R. R. Jones, At lights, DMNS (DMNS ZA. 38833); La Paz County: Ehenberg, Comber Boulevard at Colorado River, 33.61639 ° N, 114.52361 ° W, August 1974, one ♀, W. B. Warner, ASU (ASU _ NoNumber).	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFE8FF9A88F262F5FE63F9FE.taxon	diagnosis	Diagnosis Chanbria serpentinus can be distinguished from other Chanbria species by the male chelicerae with an elongate sigmoidally shaped FF lacking dentition. It is also noticeably darker and smaller in overall body size compared with the geographically close species, C. regalis.	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFE8FF9A88F262F5FE63F9FE.taxon	description	Measurements All measurements are reported as averages, and ranges are in parentheses. Males (n = 9): CL: 6.153 (5.164 – 7.829); CH: 2.03761 (1.641 – 2.589); FFH: 0.555 (0.443 – 0.694); tip of MF to MM: 1.658 (1.351 – 2.05606); tip of MF to MP: 2.262 (1.544 – 2.994); PL: 23.914 (20.800 – 29.217); LI: 12.980 (10.667 – 15.665); LIV: 21.755 (18.133 – 27.282); PPW: 3.957 (3.2 – 5.319); PPL: 2.797 (2.135 – 3.689); TL: 22.275 (18.4 – 27.840). Females (n = 4): CL: 5.710 (5.171 – 6.498); CH: 1.957 (1.761 – 2.205); FFH: 0.636 (0.543 – 0.685); tip of MF to MD: 1.197 (0.925 – 1.386); tip of MF to MP: 1.969 (1.758 – 2.143); PL: 18.067 (15.916 – 19.733); LI: 10.584 (8.236 – 12.110); LIV: 19.360 (17.404 – 21.312); PPW: 2.577 (2.397 – 2.839); PPL: 3.701 (3.2 – 4.271); TL: 21.094 (19.385 – 24.933); GOL: 1.275 (1.122 – 1.499); GOW: 1.0547 (0.856 – 1.358); GOD: 1.681 (1.415 – 1.985). Redescription: males Coloration: Overall coloration sandy brown. Appendages darken distally near ends of tibiae and tarsi. Propeltidium with darker lateral margins. Cheliceral manus darkens proximally. Chelicera: FF is sigmoidal, lacks dentition. Four teeth are present on MF. MP tooth largest, prominent, and recurved, followed by MM tooth, then posterior MSM tooth. Fondal notch absent. Fondal teeth reduced. Redescription: females Coloration: Coloration as in males. Chelicera: FF with eight or nine teeth, formula FD- 2 FSD-FM- 3 FSM-FP or FD- 3 FSD-FM- 3 FSM-FP. Moveable finger with four teeth. MP tooth largest, followed by MD tooth, with two smaller MSD teeth between them. Operculum: Two morphologies were observed in the opercular plates of C. serpentinus females. One is like those observed in some C. regalis females, with rounded plates that resemble scalene triangles, and the other consists of more broadly lobate plates with tapered anterior edges.	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFE8FF9A88F262F5FE63F9FE.taxon	distribution	Distribution Sonoran Desert. This species is apparently restricted to the eastern side of the Colorado River.	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFE8FF9A88F262F5FE63F9FE.taxon	discussion	Discussion Chanbria serpentinus females are nearly identical to C. regalis females, with C. serpentinus being distinguishable from C. regalis by the smaller size, darker coloration, and geographical location. Like C. regalis, the genital opercula of C. serpentinus females are intraspecifically variable and therefore not wholly informative for species identification.	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFEBFF9B8B8B6547FF51FE68.taxon	description	(Figs 6 M, N, 8) urn: lsid: zoobank. org: act: 5671 F 240 - 4 BF 6 - 4 AA 0 - A 56 E-B 05581 A 32440	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFEBFF9B8B8B6547FF51FE68.taxon	materials_examined	Type material: Male holotype (DMNS ZA. 36968) from Dalquest DesertResearchStation, BrewsterCounty, TX, USA. Examined. Type material at the DMNS with accession number DMNS 2009 - 110.	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFEBFF9B8B8B6547FF51FE68.taxon	etymology	Etymology The specific epithet is a patronym in honour of John (Jack) Odell Brookhart, who has dedicated> 50 years of research to this arachnid order, profoundly contributing vast amounts of knowledge to the group. This dedication is also an expression of deep gratitude and appreciation of Jack’s mentorship, training, advice, and unconditional friendship that he has given selflessly to any aspiring solifugologist who crossed his path. Jack is as rare a person as is this solifuge species is to science.	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFEBFF9B8B8B6547FF51FE68.taxon	diagnosis	Diagnosis Chanbria brookharti can be distinguished from other Chanbria species by the large protuberant structure on the FF tips of the male chelicerae. This structure is folded dorsodistally, laterally round, and FF is on a diagonal slope with respect to the manus, unlike the folded FF tip of C. plicatus male chelicerae.	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFEBFF9B8B8B6547FF51FE68.taxon	description	Measurements Male holotype: CL: 5.135; CH: 1.722; FFH: 0.426; tip of MF to MD: 1.561; tip of MF to MP: 1.986; PL: 17.291; LI: 13.245; LIV: 18.371; PPW: 3.602; PPL: 2.261; TL: 18.259. Description: male Coloration: Overall coloration very light tan, nearly translucent in some places, possibly owing to degradation in ethanol of the single known specimen. Propeltidium uniform in colour. Tibiae darken distally, femurs darken proximally, and palpal metatarsi darken distally. Palpal tarsi dark. Cheliceral manus uniform in colour. FF and MF darken distally into a red-orange. Protuberant structure on dorsal FF red-orange. Chelicera: FF morphology unique among eremobatids. The most prominent feature is the modified tip, displaying a proportionally large, dorsodistally oriented, mesally folded protuberant structure. When viewed retrolaterally, this protuberance is anteriorly rounded and posteriorly tapered, with a pinhole near the distal margin. When viewed dorsally, the dorsal margin of FF extends straight near the manus before undulating in a sigmoidal ectally then mesally. Modified tip of FF folds to form distal parabolic cup-like structure. Ventral dentition on FF highly reduced. Dentition on MF is highly reduced and triangular, whereas in the other Chanbria they are strongly or subtly recurved. MP tooth is largest, recurved, and similar in stature to other Chanbria species. The proximal MM tooth is the second largest. There are MST present before the MM. The cheliceral manus is more ovular than in other Chanbria, lacking the dorsal hump observed in other species. Pedipalp: Male. Approximately 10 pairs of enlarged moveable spines present on ventral surface of the palpal fibulae. Females unknown.	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFEBFF9B8B8B6547FF51FE68.taxon	distribution	Distribution Distribution: Chihuahuan Desert. This species is known from only a single locality at Dalquest Research Station in Brewster County, TX, USA.	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFEBFF9B8B8B6547FF51FE68.taxon	discussion	Discussion Based on the male cheliceral morphology, this specimen is best placed in Chanbria. The specimen was collected serendipitously via a long-term pitfall trap from 2 August to 27 September 2007. We were unable to extract viable DNA for UCEs from this specimen, probably owing to its long-term exposure to solar radiation and changing environmental conditions. Several attempts, during different seasons, have been made to capture more representatives of this species, without success. This species is elusive and was found in a canyon near Alamo Springs and might be endemic to this canyon. Owing to the remote location where this specimen was found with respect to the other previously described Chanbria species, we believe the unique features and geographical location of this specimen merits new species status.	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFEAFF9B8BF262D6FAD6F866.taxon	description	(Figs 6 O, P, 7 H, 9) urn: lsid: zoobank. org: act: 203 EC 990 - AF 19 - 4255 - BA 04 - 3 C 8 B 09 DBE 62 D	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFEAFF9B8BF262D6FAD6F866.taxon	materials_examined	Type material: Male holotype (IBUNAM CNANSO 106) from Sierra Mojada, Coahuila, México. Paratypes and allotypes (one ♂, two ♀) from the same locality, collected on the same date by D. Sissom, E. González-S., B. Hendrixon, S. Grant. Type and paratype material deposited at the Instituto de Biología, Universidad Nacional Autónoma de México (UNAM) arachnology collection with the accession numbers CNAN-T 01867 (holotype) and CNAN-T 01868 (paratypes). Specimens were examined by the third author and photographs by the other two.	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFEAFF9B8BF262D6FAD6F866.taxon	etymology	Etymology This species epithet is inspired both by the indigenous Cocoyome word ‘ mapeme’, meaning high rock or elevated hill, and by the Bolsón de Mapimí, a large internal basin surrounded by lower-elevation mountain ranges (Facio García, 2020). This basin is in the northwest of the Mexican Plateau, from which the type and paratype species were collected. The Cocoyome tribe was once a distinct indigenous group that inhabited this desert basin in present-day México.	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFEAFF9B8BF262D6FAD6F866.taxon	diagnosis	Diagnosis Chanbria mapemes is morphologically distinguished from the other Chanbria by not having the characteristic sigmoidshaped FF as in C. regalis and C. serpentinus males. Instead, this species possesses an FF that is elongate, thin, and is virtually uniform in height along the extent of the length of the finger, except where it tapers towards the tip. Tip of FF is constricted. There are six teeth present on FF. Moveable finger, medial tooth (MM) is large and a similar size to the moveable finger, proximal tooth (MP), whereas in the other species it is reduced or absent. The geographical location of this species is remote from the other type localities and is the southernmost record for the genus.	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFEAFF9B8BF262D6FAD6F866.taxon	description	Measurements Male holotype: CL: 5; CH: 1.64; FFH: 0.3; tip of MF to MM: 1.42; tip of MF to MP: 1.84; PL: 15.13; LI: 12.3; LIV: 18.75; PPW: 3.2; PPL: 2.05; TL: 17.75. Male paratype: CL: 4.45; CH: 1.4; FFH: 0.3; tip of MF to MM: 1.04; tip of MF to MP: 1.72; PL: 15.13; LI: 12.3; LIV: 18.75; PPW: 3.2; PPL: 2.05; TL: 14.88. Female allotypes (2): CL: 4.15, 4.05; CH: 1.58, 1.55; FFH: 0.38, 0.2; tip of MF to MM: 1.02, 0.76; tip of MF to MP: 1.46, 1.20; PL: 11.65, 11.1; LI: 10.25, 9.4; LIV: 14.14, 13.2; PPW: 2.8, 2.75; PPL: 1.65, 1.25; TL: 13.875, 18.875. Description: males Coloration: Overall coloration is light beige with dark pigmentation. Propeltidium is predominantly dark, with anterior margins with dark pigmentation, light coloration in the centre that darkens distally towards the edges. Pedipalps are almost entirely dark, except for a light-coloured patch at the proximal end. Cheliceral manus uniform in coloration. FF and MF darken distally into a red-orange towards the tip. Legs I and II have no pigmentation. Legs III and IV have dark pigmentation on the distal ends of femora and proximal ends of metatarsi. Chelicera: FF is long, slim, and nearly uniform in height. Tip of FF terminates in a sharp curved tip and is constricted, with a prominent indentation on the ectal side of the structure. Narrow line of stout setae on the mesal side of the FF along most of the length. Dentition present on FF, with six distinct teeth starting from the FP to FD. MF dentition is like that of C. brookharti, such that the MSD are triangular and there are MST present; however, the difference is that the MM is larger than any of the Chanbria species, and it is also slight recurved. Manus is ovoid like C. brookharti, with more of a dorsal hump, but less than that observed in the other species. Pedipalp: Pedipalps are dark, with light coloration at the proximal end. Enlarged, moveable palpal spines are also present on the ventral side, as in C. brookharti. Papillae present in the males examined. Ctenidia: Four ‘ stiletto-like’ abdominal ctenidia. Description: females Coloration: Same as males. Chelicera: Entire chelicerae are elongate and dorsoventrally slender. FF tooth formula is as follows: FST-FD- 2 FSD-FM- 2 FSM-FP, with seven principal teeth, not counting the teeth before the FD. Moveable finger dentition has four principal teeth, with the tooth formula MST-MM- 2 MSM-MP. Tips of chelicerae and tips of teeth darken at tips. Operculum: Genital plates are scalene triangle shaped, with nearly horizontal sides at the posterior. The lateral side of opercula is the longest of the three sides.	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFEAFF9B8BF262D6FAD6F866.taxon	distribution	Distribution Chihuahuan Desert, Mexican plateau.	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFEAFF9B8BF262D6FAD6F866.taxon	discussion	Discussion Thisspeciesisanother Chanbria thatisendemictotheChihuahuan Desert; however, this species is the southernmost record for the genus thus far. Owing to the remote location, which is in a basin between two mountain ranges, coupled with the distinct morphology in comparison to the other species, we believe this evidence favours elevating this specimen to new species status.	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFEDFF9C8BAF639BFBF1F863.taxon	description	(Figs 6 K, L, 7 G)	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFEDFF9C8BAF639BFBF1F863.taxon	materials_examined	Type material: Holotype male from Mercury, NV, USA, 15 July 1960, coll. Elden Beck (AMNH). Allotype female from Mercury, NNV, USA, on 6 July 1960, coll. Elden Beck (AMNH). Examined male holotype from images, but unable to examine female allotype. Non-type material examined: USA: Nevada: Churchill Co.: Sand Mountain, 22 miles SE of Fallon, 39.291389, − 118.415556, 1 – 8 July 2011, one ♂, W. B. Warner, Pitfall Trap Array, ASU (ASU _ NoNumber); Soda Lake ~ 1 mile NW Fallon, 39.518885, − 118.868331, 23 July 1965, one ♂, T. Briggs; K. Hom; V. Lee, CAS (CASENT 9033966); Clark Co.: trail off Calico Basin Road, 36.144061, − 115.4088627, 1060 m, 1 June 2021, one ♂, one ♀, E. L. Garcia, DMNS (DMNS ZA. 41887); Lincoln County: Corrola Dunes, ~ 23 miles South US- 93 on old Alamo Road, 37.02029, − 115.19195, 1006 m, 24 May 2014, one ♂, Zach J. Valois, Wind-blown dune field / high dunes, DMNS (DMNS ZA. 33144); Corrola Dunes, ~ 23 miles South US- 93 on old Alamo Road, 37.02029, − 115.19195, 1006 m, 24 May 2014, one ♂, Zach J. Valois, Wind-blown dune field / high dunes, DMNS (DMNS ZA. 33145); Mineral Count: junction of Highways 95 and 360, 38.27833, − 118.1025, 1340 m, 26 June – 20 July 2018, one ♂, one juvenile, M. E. Irwin; G. R. Ballmer, Malaise Trap, CAS (CASENT 9094952); Nye County: Mercury in can trap in Rock Valley, 36.633, − 116.313, 870 m, 10 July 1965, one ♂, M. H. Muma, DMNS (DMNS ZA. 16071); Nevada Test Site, Mercury, Jackass Flats, off Jackass Flats Road, 36.63619, − 116.0756, 878 m, 30 June 2011, one ♂, P. E. Cushing, DMNS (DMNS ZA. 27726); 12 miles NW Tonopah Crescent Dune, 38.229722, − 117.335, 30 April – 7 July 2011, one ♂, W. B. Warner, Pitfall Trap Array, ASU (ASU _ NoNumber).	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFEDFF9C8BAF639BFBF1F863.taxon	diagnosis	Diagnosis Chanbria plicatus can be distinguished from other Chanbria by the distinct mesally folded tip of the male cheliceral FF that forms a concavity on the mesal side and by the presence of a deep, thin, proximally upturned fondal notch. Chanbria plicatus is the only Chanbria species in which the male chelicerae have a fondal notch and flagellar groove. It is also noticeably darker in coloration than C. regalis and C. brookharti.	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFEDFF9C8BAF639BFBF1F863.taxon	description	Measurements All measurements are reported as averages, and ranges are in parentheses. Males (n = 5): CL: 4.057 (3.701 – 4.739); CH: 1.854 (1.619 – 2.285); FFH: 0.413 (0.378 – 0.457); tip of MF to MM: 1.412 (1.294 – 1.614); tip of MF to MP: 1.487 (1.440 – 1.534); PL: 18.114 (16.116 – 21.553); LI: 10.671 (9.753 – 11.411); LIV: 18.070 (16.159 – 20.240); PPW: 3.115 (2.856 – 3.744); PPL: 2.240 (2.030 – 2.789); TL: 16.713 (14.423 – 18.735). Females (n = 2): CL: 4.969 (4.716 – 5.222); CH: 1.702 (1.599 – 1.80531); FFH: 0.555 (0.504 – 0.608); tip of MF to MP: 1.7485 (1.695 – 1.802); tip of MF to MM: 1.859 (1.695 – 2.079); PL: 15.096 (14.481 – 15.711); LI: 9.239 (8.024 – 10.455); LIV: 17.220 (15.442 – 18.997); PPW: 3.372 (3.165 – 3.578); PPL: 2.175 (2.060 – 2.290); TL: 18.382 (17.729 – 19.037); GOL: 1.362 (1.359 – 1.367); GOW: 0.700 (0.643 – 0.756); GOD: 1.518 (1.515 – 1.523). Redescription: males Coloration: Dorsally sandy brown, ventrally light tan. Femurs and tarsi uniform sandy brown, tibiae lighten proximally. Propeltidium with darker anterior margin. Cheliceral manus sandy brown with lighter margins. FF and MF darken distally. Chelicera: FF with distinct tip that folds over to the flagellar groove (FG) on mesal side. Height of FF greater than in other Chanbria. FF lacks dentition. Dorsal margin of the FF is shaped like an elongate, sigmoidal wave. The proximal tooth on MF is large, prominent, slightly recurved. distal teeth on MF present in each specimen but highly reduced, with fondal notch deep, thin, upturned dorsally. Fondal teeth prominent, with tooth I in both ectal and mesal row longest. Shape of cheliceral manus is unique among Chanbria species, being less dorsoventrally compressed and with a more prominent dorsal hump from base of FF and subsequently tapering near posterior portion of manus. Redescription: females Coloration: Same as in males. Chelicera: FF with seven teeth, with formula FD- 2 FSD-FM- 2 FSM-FP. Moveable finger with four teeth. MP tooth is largest, followed by MM tooth, with two smaller MSM teeth between them. A diastema, a wide gap between two teeth, exists between first and second fondal tooth in each row. Operculum: Opercular plates in the shape of elongate tapered lobes. Plates are tapered anteriorly and lobed posteriorly.	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFEDFF9C8BAF639BFBF1F863.taxon	distribution	Distribution Nevada, USA. Mojave Basin and Range (Great Basin) Province, occurring on the east side of the Colorado River.	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
2F3187B8FFEDFF9C8BAF639BFBF1F863.taxon	discussion	Discussion This species is similar in size and has dark pigmentation on the propeltidium, pedipalps, and legs, as in C. serpentinus. Collection records indicate that this species has affinities to dune-like or sandy habitats in the Mojave Basin and Range. Owing to the similarities in habitat preference, dense setation on pedipalps, strongly procurved MF in males, and elongated chelicerae in females, we suggest that Eremochelis plicatus Muma, 1962 should be moved into Chanbria as Chanbria plicatus. This new combination is strongly supported by phylogenomic and morphological evidence, and this species is likely to be the earliest diverging member of Chanbria.	en	Garcia, Erika L., Hansen, Quincy G., Castillo, Jaír R. (2024): A phylogenomic approach to a taxonomic revision: a combination, new synonymies, and a description of two new species within the camel spider genus Chanbria Muma, 1951 (Solifugae: Eremobatidae). Zoological Journal of the Linnean Society 202 (4): 1-25, DOI: 10.1093/zoolinnean/zlae037, URL: https://doi.org/10.1093/zoolinnean/zlae037
