taxonID	type	description	language	source
2B7E87C3FFEFE61A4F9CFA3C75B4BFE5.taxon	type_taxon	Type species. Galathea pusilla Henderson, 1885, by original designation.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFEFE61A4F9CFA3C75B4BFE5.taxon	diagnosis	Diagnosis. (Modified after Macpherson & Baba 2011; Schnabel & Ahyong 2019) Carapace dorsally unarmed or with few epigastric spines, rarely with parahepatic spines, with distinct transverse ridges; lateral margin spinose. Rostrum triangular, dagger-shaped, leaf-like or bottle-shaped, usually with well-developed supraocular basal and subapical spine on each side, margin usually minutely serrated. Lateral orbital spine always present. Thoracic sternite 3 with posterior margin widely or narrowly contiguous to sternite 4. Telson subdivision incomplete, composed of ten plates. Eyes movable; cornea usually as wide as peduncle. Antennular article 1 with 3 – 5 distal spines (nearly always including double distolateral spine). Antennal article 1 with strong ventral distomesial process usually overreaching antennal article 4. Mxp 3 ischium with distal spines, crista dentata with row of spines along entire margin; merus much shorter than ischium, with strong spines on flexor and extensor distal margins. G 1 absent; G 2 present. P 1 spinose, fingers distally with row of spines along curved margin, tip excavated, spooned.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFE2E6194F9CF8FC7164BC2F.taxon	description	(Figs. 2, 11 A, 54 A)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFE2E6194F9CF8FC7164BC2F.taxon	materials_examined	Type material. Holotype. Chesterfield Islands, EBISCO Stn CP 2507, 24.7167 ° S, 159.7167 ° EE, 286 m, 7 October 2005: ov. F 2.4 mm (MNHN-IU- 2014 - 13796). Paratypes. Chesterfield Islands. EBISCO Stn CP 2493, 24.7333 ° S, 159.71667 ° E, 285 – 545 m, 6 October 2005: 1 ov. F 3.1 mm (MNHN-IU- 2019 - 2692). — KANADEEP Stn CP 4930, 25.1333 ° S, 159.9167 ° E, 300 m, 3 September 2017: 1 M 2.1 mm (MNHN-IU- 2017 - 2521). — Stn CP 4934, 25.0667 ° S, 159.91667 ° E, 290 – 300 m, 3 – 4 September 2017: 2 M 2.4 – 3.0 mm, 2 ov. F 2.2 – 2.9 mm (MNHN-IU- 2017 - 2529). — Stn DW 4940, 25.4833 ° S, 159.8167 ° E, 310 – 320 m, 4 September 2017: 1 M 2.6 mm, 1 F 2.1 mm (MNHN-IU- 2017 - 2634). Chesterfield Islands. EBISCO Stn DW 4943, 25.4167 ° S, 159.8500 ° E, 300 – 310 m, 4 September 2017: 3 M 2.4 – 2.9 mm (MNHN-IU- 2017 - 2646). Other material. Philippines. MUSORSTOM 1 Stn 18, 13.9500 ° N, 120.2833 ° E, 150 – 159 m, 21 March 1976: 1 ov. F 2.0 mm (MNHN-IU- 2014 - 13867). Papua New Guinea. KAVIENG Stn DW 4412, 02.5500 ° S, 150.6667 ° E, 500 – 600 m, 27 August 2014: 1 M 2.5 mm (MNHN-IU- 2014 - 10017). Vanuatu. MUSORSTOM 8 Stn CP 1132, 15.6333 ° S, 167.0500 ° E, 161 – 182 m, 11 October 1994: 1 M 2.0 mm, 2 ov. F 2.0 – 2.1 mm (MNHN-IU- 2016 - 424). — MUSORSTOM 8 Stn CP 1133, 15.6500 ° S, 167.050 ° E, 174 – 210 m, 11 October 1994: 1 M 2.9 mm, 1 ov. F 2.4 mm (MNHN-IU- 2019 - 2700). Chesterfield Islands. EBISCO Stn CP 4953, 24.1667 ° S, 159.6833 ° E, 270 – 320 m, 5 September 2017: 1 M 1.9 mm, 1 broken (MNHN-IU- 2019 - 2688). New Caledonia. LIFOU Stn 1648, 20.9017 ° S, 167.0550 ° E, 150 – 200 m, 7 November 2000: 1 M 1.7 mm (MNHN- IU- 2019 - 2676).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFE2E6194F9CF8FC7164BC2F.taxon	etymology	Etymology. From the name Acastus, an Argonaut, son of Pelias and Anaxibia (Phylomache). The name is considered a substantive in apposition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFE2E6194F9CF8FC7164BC2F.taxon	description	Description. Carapace: as long as or slightly longer than broad; transverse ridges with dense short setae, and few scattered iridescent thick long setae. Gastric region flattened, with 4 transverse ridges: epigastric ridge indistinct, with 5 spines (1 median and 2 pairs of spines laterally); anterior protogastric ridge not medially interrupted (interrupted only in the paratype MNHN-IU- 2014 - 10017), nearly extending laterally to carapace margin; anterior mesogastric ridge not medially interrupted, laterally interrupted by anterior branch of cervical groove, and continuing uninterrupted to first branchial spine; anterior metagastric ridge scale-like, sometimes followed by short scattered scales on posterior metagastric region. Mid-transverse ridge not interrupted, slightly medially depressed, preceded by shallow or indistinct cervical groove, followed by 2 not interrupted, or minutely interrupted ridges, interspersed with 1 short lateral ridge and sometimes few, short scattered scales. Lateral margins slightly convex, with 7 spines: first anterolateral spine well-developed, reaching anteriorly to level of lateral orbital spine, second spine (hepatic) small, slightly dorsomesially from lateral margin, and followed by 5 branchial spines (3 anterior and 2 posterior). Rostrum triangular, horizontal, dorsally flattish or slightly concave, 1.8 – [2.0] × as long as broad, length 0.4 – [0.5] and breadth 0.2 – [0.3] that of carapace; lateral margins smooth and straight, with well-developed supraocular basal spines, subapical spines absent. Pterygostomian flap ending in acute tooth, upper margin smooth. Thoracic sternum: 1.1 as wide as long. Sternite 3 moderately broad, [2.1] 2.0 – 3.0 × as wide as long, anterior margin convex, with or without median feeble excavation. Sternite 4 widely contiguous to sternite 3; surface not depressed in midline, smooth; greatest width [2.7] 2.0 – 3.0 × that of sternite 3, 2.0 – [3.0] × as wide as long. Pleon: Elevated ridges with short setae and a few scattered long setae. Tergites 2 and 3 with anterior and posterior transverse elevated ridges; tergite 4 with anterior transverse ridge; tergites 5 – 6 smooth. Eye: Eyestalk length about 1.0 – [1.3] × broader than long, peduncle distally setose, not distinctly expanded proximally; maximum corneal diameter [1.0] 1.0 × rostrum width, as wide as eyestalk. Antennule: Article 1 1.3 × longer than wide, with 5 spines: distomesial spine well-developed; proximal lateral spine small, always present. Antenna: Article 1 with prominent mesial process, distally falling well short of lateralmost antennular spine. Article 2 with distinct distal spines laterally and mesially. Article 3 often with small distomesial spine. Article 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus 0.4 – [0.6] × length of ischium, with well-developed distal spine on extensor margin and 2 strong spines on flexor margin. P 1: 3.5 – 3.6 (males), [2.7] – 2.8 (females) × carapace length; subcylindrical, scales with dense short setae and long stiff setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus [1.2] 1.1 – 1.3 length of carapace, 1.7 × as long as carpus. Carpus 2.2 – [2.4] × as long as wide. Palm 1.0 – [1.1] × carpus length, [1.8] – 2.4 × as long as broad. Fingers subequal in length to palm; fixed finger unarmed; movable finger with well-developed basal spine. P 2 – 4: Slender, subcylindrical, moderately setose and spinose. Meri successively shorter posteriorly: P 3 merus 0.7 × length of P 2 merus, P 4 merus 0.9 × length of P 3 merus. P 2 merus, 0.8 – 0.9 × carapace length, 5.5 – 6.0 × as long as broad, 1.1 – 1.4 × as long as P 2 propodus; P 3 merus 5.4 – 5.5 × as long as broad, 1.4 × as long as P 3 propodus; P 4 merus [4.9] – 5.4 × as long as broad, 1.1 – 1.4 × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, with distal spine; flexor margins of P 2 – 4 irregular, each with distal spine; P 4 lateral surface with median row of 4 small spines, absent in P 2 – 3. Carpi unarmed on extensor margin on P 3 – 4, distal spine prominent on P 2 – 3, minute on P 4; row of small spines below extensor margin on lateral surface of P 2 – 3, unarmed on P 4; flexor margin unarmed. Propodi moderately slender, [4.8 – 5.4] 4.8 – 6.0 × as long as broad; extensor margin irregular, unarmed or with 2 – 4 proximal spines; flexor margin with 2 – 4 slender movable spines in addition to distal pair. Dactyli [0.6] – 0.7 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 6 – 7 movable spines. Eggs: Ov. F carried approximately 5 – 12 eggs of 0.3 – 0.4 mm diameter. Live colour. Base colour of carapace and pleon orange. Rostrum light orange. P 1 light orange, fingers with reddish strip, distal tips whitish. P 2 – 4 light orange, propodi and dactyli with orange bands. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFE2E6194F9CF8FC7164BC2F.taxon	distribution	Distribution. Philippines, Papua New Guinea, Vanuatu, Chesterfield Islands and New Caledonia, from 150 to 600 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFE2E6194F9CF8FC7164BC2F.taxon	discussion	Remarks. Phylladiorhynchus acastus belongs to the group of species that has 5 epigastric spines, the rostrum margins are straight and without subapical spines, 3 spines on the anterior branchial margin, and the Mxp 3 merus with two prominent spines along the flexor margin. Phylladiorhynchus acastus closely resembles to P. argus, from French Polynesia, New Caledonia and Chesterfield Islands, and P. paula, from the southwestern Indian Ocean (see the differences under the Remarks of P. paula). The specimen from Papua New Guinea has the anterior protogastric ridge of the carapace interrupted and a pattern of setae slightly different than specimens from the Phillipines, Vanuatu and New Caledonia. Since there is a single specimen from this population and we were not able to obtain molecular data, we have identified this specimen as P. acastus until additional material can be collected from Papua New Guinea. Furthermore, the specimens from the Chesterfield Islands (KANADEEP, Stn CP 4934, Stn DW 4943) have the extensor margin of the P 2 – 4 propodi armed with 2 – 4 proximal spines, whereas this margin is unarmed in the other species, suggesting that this character could be variable. The genetic divergences between P. acastus and other species were always higher than 12 % (COI) and 3 % (16 S), P. paula was the closest clade. The two sequences of P. acastus from New Caledonia and Chesterfield Islands diverged 0.15 % for COI and 0 % for 16 S. The specimens of P. acastus range in size from 1.7 to 2.9 mm postorbital carapace length.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFE1E61E4F9CFAC9711ABEC2.taxon	description	(Figs. 3, 11 B – C)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFE1E61E4F9CFAC9711ABEC2.taxon	materials_examined	Type material. Holotype. Western Australia, S end Ashmore Reef, Stn 134 / K 13, 12.2960 ° S, 123.0272 ° E, 12 m, 30 September 2013: M 2.4 mm (WAM C 55691). Paratypes. Western Australia, Vulcan Shoal, Stn 146 / K 13, 12.7993 ° S, 124.2667 ° E, 19 m, 6 October 2013: 1 ov. F 2.2 mm (WAM C 55694). — Eugene McDermont Shoal, Stn 147 / K 13, 13.0769 ° S, 124.5835 ° E, 22 m, 6 October 2013: 1 M 2.8 mm (WAM C 55695). — Imperieuse Reef. Stn 157 / K 14 - T 2, 17.5070 ° S, 118.9658 ° E, 12 – 15 m, 5 October 2014: 1 M 1.8 mm, 2 ov. F 2.1 – 2.2 mm (WAM C 53891). — Rowley Shoals, Clerke Reef, Stn 154 / K 14, 17.2799 ° S, 119.3766 ° E, no depth, 4 October 2014: 1 M 2.4 mm, 1 F 2.6 mm (WAM C 53887). Other material. Western Australia, W Point Ashmore, Stn 135 / K 13, 12.2437 ° S, 122.2437 ° E, no depth, 1 October 2013: 1 M 2.2 mm, 1 ov. F 2.4 mm (WAM C 53888). — Rowley Shoals, Clerke Reef, Stn 151 / K 14, 17.2519 ° S, 119.3590 ° E, 12 – 20 m, 2 October 2014: 1 M 2.4 mm, 1 F 2.1 mm (WAM C 56280).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFE1E61E4F9CFAC9711ABEC2.taxon	etymology	Etymology. From the name Amphion, an Argonaut, son of Hyperasius and Hypso. The name is considered a substantive in apposition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFE1E61E4F9CFAC9711ABEC2.taxon	description	Description. Carapace: as long as or slightly broader than long, sexually dimorphic (wider on females) 0.9 – 1.0 (males), [0.8] – 0.9 (females) × as long as broad; transverse ridges with dense short setae and few scattered thick setae. Gastric region slightly convex with 4 transverse ridges: epigastric ridge distinct with 2 median spines and some lateral short scales, often followed by small scattered scales; anterior protogastric ridge not medially interrupted, nearly extending laterally to carapace margin, often followed by small scattered scales; anterior mesogastric ridge not medially interrupted, continuing to first branchial spine; anterior metagastric often medially interrupted, sometimes followed by short scattered scales on posterior metagastric region. Mid-transverse ridge not interrupted, slightly medially depressed, cervical groove indistinct, followed by 2 not interrupted or minutely interrupted ridges, interspersed with 2 short lateral ridges and sometimes few, short scattered scales. Lateral margins convex, with 6 – 7 distinct spines: first anterolateral spine well-developed, reaching anteriorly to level of lateral orbital spine, second spine (hepatic) well-developed, slightly dorsomesially from lateral margin, and followed by 4 – 5 branchial spines (3 anterior and 1 – 2 posterior). Rostrum leaf-like, horizontal, dorsally flattish or slightly concave, sexually dimorphic (shorter and wider in females) [1.6] – 1.8 (males), 1.2 – 1.4 (females) × as long as broad, length [0.4] 0.3 – 0.5 and breadth 0.2 – [0.3] that of carapace; lateral margins smooth and convex, with well-developed supraocular basal spines and small subapical spines. Pterygostomian flap ending in blunt tooth, upper margin smooth. Thoracic sternum: Slightly wider than long. Sternite 3 moderately broad, 2.3 – [2.5] × as wide as long, anterior margin convex, anterolaterally rounded. Sternite 4 widely contiguous to sternite 3; anterolaterally smooth, surface depressed in midline, smooth; greatest width 2.4 – [3.0] × that of sternite 3, 2.5 – [3.0] × as wide as long. Pleon: Elevated ridges with short setae and a few scattered long setae. Tergites 2 – 3 with anterior and posterior transverse elevated ridges; tergite 4 with anterior transverse ridge, posterior transverse ridge absent; tergites 5 – 6 smooth. Eye: Eyestalk length about 0.9 – [1.1] × broader than long, peduncle distally setose, not distinctly expanded proximally; cornea expanded distally, maximum corneal diameter 0.8 – [0.9] × rostrum width, as wide as eyestalk. Antennule: Article 1 slightly longer than wide, with 5 distal spines: distomesial spine well-developed; proximal lateral spine small, always present. Antenna: Article 1 with prominent mesial process, distally falling well short of lateralmost antennular spine. Article 2 with well-developed distomesial and distolateral spines. Article 3 with distomesial spine. Article 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus 0.8 × length of ischium, with well-developed distal spine on extensor and flexor margins. P 1: 3.0 – [3.2] (males), 2.0 – 2.2 (females) × carapace length; subcylindrical, spiny and with dense short setae and long stiff setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus 0.8 – [1.2] length of carapace, [1.6] 1.5 – 1.8 × as long as carpus. Carpus 1.7 – [2.2] × as long as wide. Palm [1.1] – 1.2 × carpus length, 1.4 – [2.0] × as long as broad. Fingers 0.8 × palm length; fixed finger with 1 – 2 basal spines; movable finger with 3 spines. P 2 – 4: stout, setose and spinose. Meri successively shorter posteriorly: P 3 merus 0.8 – [0.9] × length of P 2 merus, P 4 merus [0.9] 0.9 × length of P 3 merus. P 2 merus, 0.6 × carapace length, [3.0] – 3.8 × as long as broad, 1.1 – [1.2] × as long as P 2 propodus; P 3 merus 2.8 – [3.2] × as long as broad, 1.0 – [1.2] × as long as P 3 propodus; P 4 merus 3.5 × as long as broad, 1.1 × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, unarmed; flexor margin irregular, with distal spine on P 2 – 3, absent or small on P 4. Carpi with 2 – 4 spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, absent on P 4; row of granules below extensor margin on lateral surface of P 2 – 4; flexor margin unarmed. Propodi stout, [4.0 – 4.3] 3.9 – 4.6 × as long as broad; extensor margin irregular; flexor margin with 3 – 4 slender movable spines in addition to distal pair. Dactyli [0.5 – 0.7] 0.5 – 0.6 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with dactylar spines at basis of 5 – 6 movable spines. Eggs: Ov. F carried approximately 10 – 25 eggs of 0.3 – 0.5 mm diameter. Live colour. Unknown. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFE1E61E4F9CFAC9711ABEC2.taxon	distribution	Distribution. Western Australia, from 12 to 22 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFE1E61E4F9CFAC9711ABEC2.taxon	discussion	Remarks. Phylladiorhynchus amphion belongs to the group of species having 2 epigastric spines, 1 hepatic spine, 3 spines on anterior branchial margin of the carapace and dactylar spines along the flexor margin of P 2 – 4 dactyli. The specimens range from 1.8 to 2.8 mm postorbital carapace length. The closest relative is P. joannotae from French Polynesia, Guam Island, Papua New Guinea, Vanuatu and New Caledonia (see the differences under the Remarks of this species). The two sequences of P. amphion from Western Australia diverged 1.8 % for COI and 1.2 % for 16 S.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFE5E6024F9CFF2971D5BA9E.taxon	description	(Fig. 4, 11 D) Phylladiorhynchus ikedai. — Baba, 1991: 485 fig. 4 a, b (in part, only material from MUSORSTORM 4, Stn 238 [MNHN Ga- 2043], MUSORSTORM 6, Stn DW 485 [MNHN Ga- 2045] and CALSUB, Pl 16 [MNHN Ga- 2046]).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFE5E6024F9CFF2971D5BA9E.taxon	materials_examined	Type material. Holotype. New Caledonia. EXBODI Stn DW 3785, 22.2500 ° S, 167.1667 ° E, 386 – 387 m, 02 September 2011: M 2.5 mm (MNHN-IU- 2011 - 7659). Paratypes. New Caledonia. MUSORSTOM 4 Stn CP 238, 22.2167 ° S, 167.2333 ° E, 500 – 510 m, 2 October 1985: 1 ov. F 3.5 mm (MNHN-IU- 2014 - 23836 (Ga- 2043 )). — MUSORSTOM 6 Stn DW 485, 21.3913 ° S, 167.9883 ° E, 380 m, 23 February 1989: 1 F 3.2 mm (MNHN-IU- 2014 - 23835 (Ga- 2045 )). — CALSUB Pl 16, 20.6300 ° S, 167.0450 ° E, 500 m, 7 March 1989: 1 M 3.2 mm (MNHN-IU- 2013 - 19941 (Ga- 2046 )). Other material. New Caledonia. KANACONO Stn DW 4778, 23.0500 ° S, 168.3000 ° E, 170 – 248 m, 28 August 2016: 1 M 2.9 mm (MNHN-IU- 2016 - 488). Chesterfield Islands. KANADEEP Stn CP 4985, 20.8167 ° S, 160.9500 ° E, 480 – 540 m, 10 September 2017: 1 M 3.0 mm (MNHN-IU- 2017 - 3138). French Polynesia. BENTHAUS Stn DW 2006, 22.4500 ° S, 151.3167 ° W, 350 – 450 m, 24 November 2002: 1 M 3.4 mm (MNHN-IU- 2014 - 13883).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFE5E6024F9CFF2971D5BA9E.taxon	etymology	Etymology. From the name Argus, an Argonaut, son of Phrixus; builder of the boat Argo. The name is considered a substantive in apposition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFE5E6024F9CFF2971D5BA9E.taxon	description	Description. Carapace: As long as or slightly longer than broad; transverse ridges with dense short setae, and few scattered thick setae. Gastric region flattened with some transverse ridges: epigastric ridge indistinct, with 5 spines (1 median and 2 pairs of spines laterally); anterior protogastric ridge not medially interrupted, nearly extending laterally to carapace margin; anterior mesogastric ridge widely or minutely interrupted, laterally interrupted by anterior branch of cervical groove, continuing to first branchial spine; anterior metagastric ridge scale-like. Mid-transverse ridge not interrupted, medially depressed, preceded by distinct cervical groove, followed by 2 not interrupted or minutely interrupted ridges, interspersed with 1 short lateral ridge and few, short scattered scales. Lateral margins slightly convex, with 7 spines: first anterolateral spine well-developed, reaching end of lateral orbital spine, second spine (hepatic) small to well-developed, slightly dorsomesially from lateral margin, and followed by 5 branchial spines (3 anterior well-developed subequal branchial spines and 2 well developed posterior branchial spines decreasing in size posteriorly). Rostrum triangular, horizontal, dorsally flattish or slightly concave, [1.5] 1.2 – 1.6 × as long as broad, length [0.35] 0.3 – 0.4 and breadth [0.25] 0.2 – 0.3 that of carapace; lateral margins smooth and straight, with well-developed supraocular basal spines, subapical spines absent. Pterygostomian flap ending in blunt tooth, upper margin smooth. Thoracic sternum: About as wide as long. Sternite 3 sharply broad, [4.0] – 4.2 × as wide as long, anterior margin convex with a median deep notch flanked by 2 lobes. Sternite 4 widely contiguous to sternite 3; surface depressed in midline, smooth; greatest width 2.3 – [2.6] × that of sternite 3, [2.6] – 2.8 × as wide as long. Pleon: Transverse ridges with short setae and scattered few setae. Tergite 2 with anterior and posterior transverse elevated ridges; tergites 3 and 4 with anterior transverse ridge; tergites 5 – 6 smooth. Eye: Eyestalk length about [1.2] × broader than long, peduncle distally setose, not distinctly expanded proximally; maximum corneal diameter [1.2] × rostrum width, as wide as eyestalk. Antennule: Article 1 1.5 × longer than wide, with 5 spines: distomesial spine small; proximal lateral spine small, always present. Antenna: Article 1 with prominent mesial process, distally falling well short of lateralmost antennular spine. Article 2 with distinct distal spines laterally and mesially. Articles 3 and 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus 0.6 × length of ischium, with well-developed distal spine on extensor margin and 2 strong spines on flexor margin. P 1: [3.1] 3.5 (males), 2.8 (females) × carapace length; subcylindrical, spiny and with scattered long stiff setae and few plumose setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus [1.2] 1.1 – 1.3 length of carapace, 1.7 – [2.5] × as long as carpus. Carpus [2.0] – 3.0 × as long as wide. Palm [1.4] 1.1 – 1.5 × carpus length, [1.8] 1.6 – 2.0 × as long as broad. Fingers 0.7 – [1.0] × palm length; fixed finger unarmed; movable finger with well-developed basal spine. P 2 – 4: Slender, subcylindrical, moderately setose and spinose. Meri successively shorter posteriorly: P 3 merus 0.7 – [0.8] times length of P 2 merus, P 4 merus [0.7] – 0.8 × length of P 3 merus. P 2 merus, [0.8] or as long as carapace length, [5.5] – 7.4 × as long as broad, [1.3] – 1.6 × as long as P 2 propodus; P 3 merus [6.0] – 6.2 × as long as broad, [1.3] – 1.4 × as long as P 3 propodus; P 4 merus [4.5] – 5.9 × as long as broad, [1.0] – 1.3 × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular with small distal spine; flexor margins of P 2 – 4 irregular, each with distal spine; P 4 lateral surface with median row of 3 small spines, absent in P 2 – 3. Carpi with 3 or 4 spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, smaller on P 4; row of small spines below extensor margin on lateral surface of P 2 – 3, unarmed on P 4; flexor margin unarmed. Propodi slender, [6.1 – 7.0] 6.1 – 7.8 × as long as broad; extensor margin irregular, usually unarmed or armed with proximal spine on P 2 – 3; flexor margin with 2 – 4 slender movable spines in addition to distal pair. Dactyli [0.6 – 0.7] 0.6 – 0.7 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 7 – 8 movable spines. Eggs: Ov. F (MNHN-IU- 2014 - 23836) 12 eggs of 0.3 – 0.4 mm diameter. Live colour. Unknown. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFE5E6024F9CFF2971D5BA9E.taxon	distribution	Distribution. French Polynesia, New Caledonia and Chesterfield Islands from 170 to 510 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFE5E6024F9CFF2971D5BA9E.taxon	discussion	Remarks. Phylladiorhynchus argus belongs to the group of species that has 5 epigastric spines, the rostrum margin straight, the subapical spines of the rostrum absent, 3 spines on the anterior branchial margin, and the Mxp 3 merus with two prominent spines along the flexor margin. The specimens range from 2.9 to 3.5 mm postorbital carapace length. Phylladiorhynchus argus closely resembles P. acastus from Philippines, Papua New Guinea, Vanuatu, Chesterfield Islands and New Caledonia, and P. paula, from southwestern Indian Ocean (see the differences under the Remarks of P. paula).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFFAE6034F9CFC6C7502BE6E.taxon	description	(Figs. 5, 11 E, 30 G – I, Q, R)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFFAE6034F9CFC6C7502BE6E.taxon	materials_examined	Type material. Holotype. Western Australia. Ningaloo Reef. 22.6083 ° S, 113.6249 ºE, 10 m, 01 May 2009: M 2.1 mm (UF 22296). Paratypes. Western Australia. Ningaloo Reef, 22.6083 ° S, 113.6249 ° E, 10 m, 1 May 2009: 1 ov. F 2.1 mm (UF 22401). — 22.681 ° S, 113.6304 ° E, 15 m, rubble, 20 May 2006: 1 M 2.4 mm (UF 27886). — Norwegian Channel, 22.6006 ° S, 113.6274 ° E, 23 m (in rubble), 28 May 2006: 1 ov. F 2.1 mm (UF 27601). Other material. Western Australia. Dampier. Stn DA 3 / 99 / 50, 20.5473 ° S, 116.4455 ° E, 10 – 20 m, 31 August 1999: 1 M 2.4 mm, 2 ov. F 2.2 – 2.4 mm (WAM C 27667). — Vulcan Shoal, Stn 146 / K 13, 12.7993 ° S, 124.2667 ° E, 19 m, 6 October 2013: 1 M 2.4 mm (WAM C 55694 - 2).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFFAE6034F9CFC6C7502BE6E.taxon	etymology	Etymology. From the name Asclepius, an Argonaut, son of Apollo and Coronis or Arsinoe. The name is considered a substantive in apposition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFFAE6034F9CFC6C7502BE6E.taxon	description	Description. Carapace: Usually broader than long (0.8 – [0.9] × as long as wide); transverse ridges with dense short setae and thick iridescent setae. Gastric region slightly convex with some transverse ridges: epigastric ridge distinct with 2 median spines and some lateral short scales, followed by small short scales on posterior epigastric region; anterior protogastric ridge not medially interrupted, nearly extending laterally to carapace margin, often followed by uninterrupted posterior protogastric ridge or short scales; anterior mesogastric ridge not medially interrupted, laterally interrupted by anterior branch of cervical groove, and continuing uninterrupted to first branchial spine; anterior metagastric ridge not medially interrupted, followed by posterior scale-like metagastric ridge. Midtransverse ridge not interrupted, medially depressed, followed by shallow or indistinct cervical groove, followed by 2 – 3 not interrupted or minutely interrupted ridges, interspersed with 2 interrupted riges and few short scales. Lateral margins convex, with 6 distinct spines: first anterolateral spine well-developed, reaching anteriorly to level of lateral orbital spine, second spine (hepatic) well-developed, slightly dorsomesially from lateral margin, and followed by 4 – 5 branchial spines (3 anterior and 1 – 2 posterior). Rostrum dagger-like, horizontal dorsally flattish or slighly concave [1.8] – 2.0 × as long as broad, length 0.5 and breadth 0.2 – [0.3] that of carapace; lateral margins serrated and straight or nearly straight, with well-developed supraocular basal spines and subapical spines. Pterygostomian flap with anterior spine, upper margin smooth. Thoracic sternum: As wide as long. Sternite 3 quadrangular, [1.4] – 2.1 × as wide as long, anterior margin convex, with a blunted median projection, anterolaterally rounded. Sternite 4 widely contiguous to sternite 3; anterolaterally smooth, surface depressed in midline, smooth; greatest width 2.6 – [3.1] × that of sternite 3, [2.6] – 3.3 × as wide as long. Pleon: Elevated ridges with short setae and a few scattered long setae. Tergite 2 – 4 with anterior and posterior transverse elevated ridges; tergites 5 – 6 smooth. Eye: Eyestalk length about 0.9 – [1.1] × broader than long, peduncle distally setose, not distinctly expanded proximally, with few short transverse striae on lateral surfaces; cornea expanded distally, maximum corneal diameter [0.8] – 1.0 × rostrum width, [0.8] as long as eyestalk. Antennule: Article 1 longer than wide, with 5 distal spines: distomesial spine well-developed; proximal lateral spine small, always present. Antenna: Article 1 with prominent mesial process, distally falling well short of lateralmost antennular spine. Article 2 with well-developed distomesial and distolateral spines. Article 3 with small to distinct mesial spine. Article 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus 0.8 × length of ischium, with well-developed distal spine on extensor and flexor margins. P 1: [2.3] – 2.7 (males), 2.0 – 2.1 (females) × carapace length; subcylindrical, spiny and with long stiff setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus [0.8] – 1.0 length of carapace, [2.0] – 2.3 × as long as carpus. Carpus 1.2 – [1.3] × as long as wide. Palm [1.3] 1.3 × carpus length, [1.4] – 1.5 × as long as broad. Fingers [0.9] – 1.0 × palm length; fixed finger with 0 – 1 basal spines; movable finger often with 1 basal spine. P 2 – 4 (P 3 lost in holotype): Setose and spinose. Meri successively shorter posteriorly: P 3 merus 0.9 × length of P 2 merus, P 4 merus 0.8 – 0.9 × length of P 3 merus. P 2 merus, 0.6 – [0.7] × carapace length, 3.3 – 3.8 × as long as broad, 1.1 – 1.2 × as long as P 2 propodus; P 3 merus 3.6 – 3.7 × as long as broad, 1.1 – 1.2 × as long as P 3 propodus; P 4 merus 3.7 – 4.0 × as long as broad, 1.0 – 1.1 × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, unarmed; flexor margin irregular, with distal spine on P 2 – 3, absent or small on P 4. Carpi with 2 – 4 small spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, absent on P 4; row of granules below extensor margin on lateral surface of P 2 – 4; flexor margin unarmed. Propodi stout, [4.0 – 4.5] 3.8 – 4.5 × as long as broad; extensor margin irregular; flexor margin with 3 – 6 slender movable spines in addition to distal pair. Dactyli [0.6] – 0.7 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with dactylar spines at basis of 5 – 6 movable spines. Eggs: Ov. F carried approximately 8 – 15 eggs of 0.4 – 0.5 mm diameter. Live colour. Unknown. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFFAE6034F9CFC6C7502BE6E.taxon	distribution	Distribution. Western Australia, from 8 to 45 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFFAE6034F9CFC6C7502BE6E.taxon	discussion	Remarks. Phylladiorhynchus asclepius belongs to the group of species having 2 epigastric spines, 1 hepatic spine, 3 spines on anterior branchial margin, rostrum dagger-like (margin straights or nearly straight) and dactylar spines along the flexor margin of the P 2 – 4 dactyli. This species complex includes the following species: P. asclepius, P. euryalus, from Australia (Queensland), New Caledonia and the Chesterfield Islands, P. lini, and P. spinosus Schnabel & Ahyong, 2019 from Australia, New Caledonia and New Zealand. The Western Australian material tentatively referred to P. spinosus by Schnabel & Ahyong (2019) is referable to the new species (Ahyong, pers. com.). These species are morphologically very similar but genetically very distinct (see the differences under the Remarks of P. spinosus). The examined specimens of P. asclepius range from 2.1 to 2.4 mm postorbital carapace length. The two sequences of P. asclepius from Western Australia diverged 0.6 % for COI and 0.2 % for 16 S.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFFCE6044F9CFF297114BD42.taxon	description	(Fig. 11 F)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFFCE6044F9CFF297114BD42.taxon	materials_examined	Material examined. New Zealand. NIWA 76630, Stn. TAN 1108 / 166, North Canterbury, 43.099 – 43.095 ° S, 173.446 – 173.444 ° E, 79 m, 26 May 2011: 4 M 4.4 – 5.2 mm, 1 ov. F 4.1 mm, 7 F 4.0 – 4.8 mm. Donated to MNHN (MNHN-IU- 2019 - 2596).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFFCE6044F9CFF297114BD42.taxon	diagnosis	Diagnosis (modified from Schnabel & Ahyong 2019). Rostrum lateral margins convex; subapical spines present. Epigastric region with 4 spines; parahepatic spines on protogastric region absent. Anterior metagastric ridge continuous, not medially interrupted; hepatic spine present. Thoracic sternite 3 anterior margin biconcave, with obtuse median projection. Pleonal tergite 3 with anterior and posterior transverse ridge. Antennular article 1 with 5 distal spines, lateralmost spine always distinct. Antennal article 1 mesial process distally overreaching peduncle and usually reaching second lateral antennular spine; article 2 with lateral spine distinctly larger than mesial spine; article 3 usually unarmed or with minute mesial spine only. Flexor margin of Mxp 3 merus with one prominent spine only. P 2 – 4 dactylus extensor margin without upright spines at bases of movable spines. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFFCE6044F9CFF297114BD42.taxon	distribution	Distribution. New Zealand continental shelf, from the Snares to the Three King Islands and northwards to Norfolk Island, and westward to southern Australia (New South Wales to South Australia), between 15 and 366 m (from Schnabel & Ahyong 2019).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFFCE6044F9CFF297114BD42.taxon	discussion	Remarks. The species closely resembles P. nui Schnabel & Ahyong, 2019, from New Zealand and Eastern Australia and P. integrus (Benedict, 1902) from Japan to Chesterfield Islands (see the differences under the Remarks of P. integrus).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFFCE6054F9CFB117198BC65.taxon	description	(Figs. 6, 11 G)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFFCE6054F9CFB117198BC65.taxon	materials_examined	Type material. Holotype. Saudi Arabia, Yanbu, Ras Majiz, highly dissected reef, 38.2761 ° N, 23.7725 ° E, 4 – 8 m, 3 March 2014: M 2.5 mm (UF 40205).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFFCE6054F9CFB117198BC65.taxon	etymology	Etymology. From Bahamut, a Persian sea monster supporting the structure that holds up the earth. The name is a substantive in apposition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFFCE6054F9CFB117198BC65.taxon	description	Description. Carapace: As long as or slightly longer than broad; transverse ridges with dense short setae and some scattered iridescent thick long setae. Gastric region slightly convex with 4 transverse ridges: epigastric ridge distinct with 2 median spines, short scales laterally; anterior protogastric ridge not medially interrupted, nearly extending laterally to carapace margin; anterior mesogastric ridge not medially interrupted, laterally continuing to first branchial spine; anterior metagastric ridge not medially interrupted, followed by median scale. Mid-transverse ridge not interrupted, medially depressed, cervical groove indistinct, followed by 2 not interrupted or minutely interrupted ridges, interspersed with 1 short lateral ridge and few short scales. Lateral margins convex, with 5 distinct spines: first anterolateral spine well-developed, reaching anteriorly to level of lateral orbital spine, second spine (hepatic) small, slightly dorsomesially from lateral margin, and followed by 3 branchial spines (2 anterior and 1 posterior). Rostrum leaf-like, horizontal, dorsally slightly concave, [1.4] × as long as broad, length [0.4] and breadth [0.2] that of carapace; lateral margins minutely serrated and convex, with well-developed supraocular basal spines, subapical spines absent. Pterygostomian flap ending in sharp tooth, upper margin smooth. Thoracic sternum: As wide as long. Sternite 3 moderately broad, [1.7] × as wide as long, anterior margin serrated and convex, with a median projection. Sternite 4 widely contiguous to sternite 3; anterolaterally serrated, surface flattish, smooth; greatest width [3.2] × that of sternite 3, [2.9] × as wide as long. Pleon: Elevated ridges with short setae and a few scattered long setae. Tergite 2 with anterior and posterior transverse elevated ridges; tergites 3 – 4 with anterior transverse ridge only; tergites 5 – 6 smooth. Eye: Eyestalk length about [1.1] × broader than long, peduncle distally setose, not distinctly expanded proximally; cornea expanded distally, posterior margin irregular, maximum corneal diameter [1.1] × rostrum width, as wide as or wider than eyestalk. Antennule: Article 1 slightly longer than wide, with 5 distal spines: distomesial spine well-developed; proximal lateral spine small, obsolescent. Antenna: Article 1 with prominent mesial process, nearly reaching end of lateralmost antennular spine. Article 2 with well-developed distomesial and distolateral spines. Articles 3 and 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus [0.8] × length of ischium, with well-developed distal spine on extensor and flexor margins. P 1: [3.3] (male) × carapace length; subcylindrical, spiny and with long stiff setae, without plumose setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus [1.1] length of carapace, [1.6] × as long as carpus. Carpus [1.8] × as long as wide. Palm [1.3] × carpus length, [2.1] × as long as broad. Fingers [0.7] × palm length; fixed finger unarmed; movable finger with small basal spine. P 2 – 4: Slender, densely setose and spinose. Meri successively shorter posteriorly: P 3 merus [0.9] × length of P 2 merus; P 4 merus [0.9] × length of P 3 merus. P 2 merus [4.7] × as long as broad, [1.3] × as long as P 3 propodus; P 3 merus [3.4] × as long as broad, [1.2] × as long as P 3 propodus; P 4 merus [3.6] × as long as broad, [1.1] × as long as P 4 propodus; extensor margin of P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, unarmed; flexor margin irregular, with distal spine on P 3, distal spine absent in P 4. Carpi with 1 – 3 spines on extensor margin on P 3, unarmed on P 4; distal spine prominent on P 3, smaller on P 4; granules below extensor margin on lateral surface of P 3 – 4; flexor margin unarmed. Propodi stout, [4.5 – 4.6] × as long as broad; extensor margin irregular, armed with 2 – 4 spines; flexor margin with 3 – 4 slender movable spines in addition to distal pair. Dactyli [0.6] × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 4 movable spines. Eggs: No data. Live colour. Unknown. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFFCE6054F9CFB117198BC65.taxon	distribution	Distribution. Red Sea, from 4 to 8 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFFCE6054F9CFB117198BC65.taxon	discussion	Remarks. Phylladiorhynchus bahamut agrees with the species recorded and illustrated by Lewinsohn (1982) from the Gulf of Aqaba, who also pointed out that it might be a different species than P. integrirostris. Phylladiorhynchus bahamut belongs to the group of species having 2 median epigastric spines, the hepatic margin with a small spine and 2 spines on the anterior branchial margin. The closest species is P. peneleos (see the differences under the Remarks of this species).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFFDE6084F9CFA7471C4B897.taxon	description	(Figs. 7, 11 H, 54 B – C)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFFDE6084F9CFA7471C4B897.taxon	materials_examined	Type material. Holotype. Papua New Guinea, PAPUA NIUGINI Stn PR 76, 05.0267 ° S, 145.7983 ° E, 2 – 15 m, 21 November 2012: M 2.5 mm (MNHN- IU- 2014 - 13813) Paratypes. Papua New Guinea. PAPUA NIUGINI Stn PS 07, 05.1800 ° S, 145.8300 ° E, 13 m, 8 November 2012: 1 ov. F 1.8 mm (MNHN-IU- 2019 - 2670). — Stn PB 13, 05.2967 ° S, 145.7817 ° E, 13 m, 12 November 2012: 1 M 2.3 mm, 1 F 2.4 mm (MNHN-IU- 2019 - 2667). — Stn PB 29, 05.3000 ° S, 145.7683 ° E, 17 m, 27 November 2012: 1 ov. F 1.6 mm (MNHN-IU- 2019 - 2641). — Stn PB 33, 05.1517 ° S, 145.8283 ° E, 30 m, 3 December 2012: 1 F 2.4 mm (MNHN-IU- 2019 - 2668). — Stn PS 08, 05.18395 ° S, 145.80718 ° E, 8 m, 30 December 2012: 1 M 1.4 mm (MNHN- IU- 2014 - 13865). — Stn PB 39, 05.2650 ° S, 145.7850 ° E, 18 – 27 m, 6 December 2012: 2 F 2.1 – 2.3 mm, 1 parasitized (MNHN-IU- 2019 - 2638). New Caledonia, Touho. 20.6101 ° S, 165.2242 ° E, 5 – 7 m, September 1993: 1 M 3.1 mm, 1 ov. F 1.7 mm (MNHN- IU- 2014 - 13861). Vanuatu. SANTO Stn ZR 12, 15.6117 ° S, 167.0333 ° E, 2 – 30 m, 5 October 2006: 1 M 2.0 mm (MNHN-IU- 2014 - 13808), 1 M 2.2 mm (MNHN-IU- 2019 - 2643). Other material. Papua New Guinea. KAVIENG Stn KB 16, 02.5767 ° S, 150.7717 ° E, 13 – 14 m, 7 June 2014: 1 M parasitized 1.8 mm (MNHN-IU 2014 - 13619). — Stn KB 20, 02.7533 ° S, 150.6950 ° E, 8 m, 9 June 2014: 1 ov. F 2.3 mm (MNHN-IU- 2014 - 2311). Vanuatu. SANTO Stn FB 43, 15.4733 ° S, 167.2483 ° E, 19 m, 30 September 2006: 1 M 2.1 mm (MNHN-IU- 2019 - 2642). — Stn FB 68, 15.5900 ° S, 166.9950 ° E, 11 m, 11 October 2006: 1 M 2.5 mm (MNHN-IU- 2014 - 13875). — Stn FB 72, 15.6017 ° S, 166.9750 ° E, 16 m, 12 October 2006, 1 M 1.6 mm (MNHN-IU 2014 - 13810). — Stn DB 20, 15.5083 ° S, 167.0233 ° E, 20 – 25 m, 15 September 2006: 2 M 1.3 – 2.0 mm (MNHN- IU- 2019 - 2630). New Caledonia. Lagon Est. Stn 0607, 22.2017 ° S, 167.0417 ° E, 48 – 54 m, 5 August 1986: 1 ov. F 2.8 mm (MNHN-IU- 2014 - 13740).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFFDE6084F9CFA7471C4B897.taxon	etymology	Etymology. Named after Any Barbé, mother of one of the authors, for her unremitting support.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFFDE6084F9CFA7471C4B897.taxon	description	Description. Carapace: as long as or slightly longer than broad; transverse ridges with dense short setae. Gastric region slightly convex, with 4 transverse ridges: epigastric ridge distinct with 2 median spines; anterior protogastric ridge not medially interrupted, nearly extending laterally to carapace margin, sometimes followed by a few short scales; anterior mesogastric ridge not medially interrupted, laterally continuing to first branchial spine, followed by some short lateral scales; anterior metagastric ridge not medially interrupted, followed by a short median scale. Mid-transverse ridge not interrupted, medially depressed, cervical groove indistinct, followed by 2 – 3 not interrupted or minutely interrupted ridges, interspersed with 2 short lateral ridge and sometimes few, short scattered scales. Lateral margins nearly straight or slightly convex, with 4 distinct spines: first anterolateral spine well-developed, reaching anteriorly to level of lateral orbital spine, hepatic margin unarmed; anterolateral spine followed by 3 branchial spines (2 anterior and 1 posterior). Rostrum leaf-like, horizontal, dorsally flattened, [1.2] 1.1 – 1.5 × as long as broad, length [0.3] – 0.4 and breadth [0.2] – 0.3 that of carapace; lateral margins serrated and convex, with well-developed supraocular basal spines, subapical spines absent. Pterygostomian flap ending in blunt tooth, upper margin smooth. Thoracic sternum: As wide as long. Sternite 3 moderately broad, [1.8] – 2.0 × as wide as long, anterior margin serrated and slightly convex or straight. Sternite 4 widely contiguous to sternite 3; anterolaterally serrated, surface depressed in midline, smooth; greatest width 3.3 – [3.4] × that of sternite 3, 2.5 – [2.6] × as wide as long. Pleon: Elevated ridges with short setae and a few scattered long setae. Tergite 2 with anterior and posterior transverse elevated ridges; tergites 3 – 4 with anterior transverse ridge; tergites 5 – 6 smooth. Eye: Eyestalk length about 0.9 – [1.0] × broader than long, peduncle distally setose, not distinctly expanded proximally; maximum corneal diameter 0.7 × rostrum width, as wide as eyestalk. Antennule: Article 1 slightly longer than wide, with 5 distal spines: distomesial spine well-developed; proximal lateral spine small, always present. Antenna: Article 1 with prominent mesial process, distally falling well short of lateralmost antennular spine. Article 2 with well-developed distomesial and distolateral spines. Article 3 often with distomesial spine. Article 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus 0.6 – [0.7] × length of ischium, with well-developed distal spine on extensor and flexor margins. P 1 (lost in holotype): Slender, 2.8 – 3.9 (males), 2.6 – 2.7 (females) × carapace length; subcylindrical, spiny and with long stiff setae and dense thick and plumose setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus 0.9 – 1.4 length of carapace, 1.5 – 1.7 × as long as carpus. Carpus 3.0 – 3.4 × as long as wide. Palm 1.1 × carpus length, 2.9 – 3.0 × as long as broad. Fingers 0.6 – 0.7 × palm length; fixed finger unarmed; movable finger with a basal spine. P 2 – 4 (P 3 lost in holotype): Slender, densely setose and spinose. Meri successively shorter posteriorly: P 3 merus 0.6 × length of P 2 merus, P 4 merus 0.9 × length of P 3 merus. P 2 merus, 0.7 – [0.8] × carapace length, [4.0] 3.9 – 4.2 × as long as broad, [1.0] 1.1 × as long as P 2 propodus; P 3 merus 4.2 × as long as broad, 1 – 1.1 × as long as P 3 propodus; P 4 merus 4.2 – 4.7 [4.5] × as long as broad, [1.0] – 1.1 × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, unarmed; flexor margin irregular, with distal spine on P 2 – 4, P 4 lateral surface with median row of 2 spines. Carpi with 1 – 3 minute spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, smaller on P 4; row of small granules below extensor margin on lateral surface of P 2 – 4; flexor margin unarmed. Propodi slender, [5.5 – 6.5] 5.2 – 7.2 × as long as broad; extensor margin irregular, armed with 1 – 4 spines on proximal half; flexor margin with 3 – 4 slender movable spines in addition to distal pair. Dactyli 0.6 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 5 – 6 movable spines. Eggs: Ov. F carried approximately 10 – 20 eggs of 0.3 – 0.5 mm diameter. Live colour. Carapace and pleon with reddish-brownish or orange patches. P 1 whitish, covered by some reddish-brownish spots basally. P 2 – 4 light whitish with reddish-brownish stripes. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFFDE6084F9CFA7471C4B897.taxon	distribution	Distribution. Papua New Guinea, Vanuatu, New Caledonia from 2 to 54 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFFDE6084F9CFA7471C4B897.taxon	discussion	Remarks. Phylladiorhynchus barbeae belongs to the group of species having 2 median epigastric spines, the hepatic margin unarmed and 2 spines on the anterior branchial margin. The specimens range from 1.4 to 3.1 mm postorbital carapace length. The closest species is P. pepei from Madagascar (see the differences under the Remarks of this species). The four sequences of P. barbeae from New Caledonia, Papua New Guinea and Vanuatu diverged 0.2 % for COI and 0 % for 16 S.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFF0E6084F9CFE71744DBD7D.taxon	diagnosis	Diagnosis. Rostrum triangular, subapical spines absent or, if present, obsolescent, lateral margins straight. Carapace with epigastric ridge armed with 5 spines; anterior branchial margin with 3 spines; anterior protogastric ridge not interrupted medially, without scale behind median epigastric spine; anterior mesogastric and metagastric ridges not medially interrupted. Anterior margin of thoracic sternite 3 nearly transverse with median and lateral projections. Antennule article 1 with 5 distal spines; distomesial spine minute. Antennal article 2 with distolateral spine longer than distomesial; article 3 with minute distomesial spine. Flexor margin of Mxp 3 merus with 2 prominent spines. P 2 – 4 dactylus extensor margin without upright spines at bases of movable spines.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFF0E6084F9CFE71744DBD7D.taxon	distribution	Distribution. Only known from the type locality, Andaman Sea, at 77 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFF0E6084F9CFE71744DBD7D.taxon	discussion	Remarks. The species is only known by the holotype male and it is characterized by the presence of 5 epigastric spines, 1 small hepatic spine, and 3 spines on the anterior branchial margin. The closest relative is P. ikedai (Miyake & Baba, 1965) from Bonin Islands, Japan. However, they can be easily differentiated by the shape of thoracic sternite 3, with the anterior margin nearly transverse, and median and lateral projections, in P. bengalensis, whereas the sternite has the anterior margin convex, with feeble median excavation in P. ikedai.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFF0E60E4F9CFB167518BC81.taxon	description	(Figs. 8, 11 I)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFF0E60E4F9CFB167518BC81.taxon	materials_examined	Type material. Holotype. Chesterfield Islands. KANADEEP Stn DW 4969, 21.0000 ° S, 158.5000 ° E, 500 – 750 m, 8 September 2017: M 2.6 mm (MNHN-IU- 2017 - 3816) Paratypes. Chesterfield Islands. KANADEEP Stn DW 4969, 21.0000 ° S, 158.5000 ° E, 500 – 750 m, 8 September 2017: 1 M 3.0 mm (MNHN-IU- 2019 - 2690). — Stn DW 4993, 4.2500 ° S, 166.9667 ° E, 560 – 610 m, 13 September 2017: 1 ov. F 2.5 mm (MNHN-IU- 2017 - 3830)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFF0E60E4F9CFB167518BC81.taxon	etymology	Etymology. Named after Philippe Bouchet, head of conservation of the general mollusk collection of the Muséum national d’Histoire naturelle, Paris.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFF0E60E4F9CFB167518BC81.taxon	description	Description. Carapace: as long as or longer than broad ([1.0] – 1.2 × as long as broad); transverse ridges with dense short setae, without long setae. Gastric region flattened, with 3 transverse ridges: epigastric ridge distinct with 4 spines; anterior protogastric ridge not medially interrupted, nearly extending laterally to carapace margin; anterior mesogastric ridge medially interrupted, laterally interrupted by cervical groove; anterior metagastric area with a few scales, otherwise absent. Mid-transverse ridge not interrupted, medially depressed, preceded by distinct cervical groove, followed by 2 not interrupted or minutely interrupted ridges, interspersed with 2 short lateral ridges and few, short scattered scales. Lateral margins straight or slightly convex, with 6 – 7 spines: first anterolateral spine welldeveloped, overreaching anteriorly level of lateral orbital spine, second spine (hepatic) well-developed, slightly dorsomesially from lateral margin, and followed by 4 – 5 branchial spines (3 anterior and 1 – 2 posterior). Rostrum subtriangular, horizontal, dorsally flattish or slightly concave, 0.8 – [1.3] × as long as broad, length 0.2 – [0.3] and breadth [0.3] 0.3 that of carapace; lateral margins smooth and slightly concave, with well-developed supraocular basal spines, and minute subapical spines. Orbit sharply excavated. Pterygostomian flap ending in blunt tooth; upper margin smooth. Thoracic sternum: As wide as long. Sternite 3 sharply broad, [3.5] – 4.5 × as wide as long, anterior margin slightly concave, or straight, with median feeble excavation. Sternite 4 widely contiguous to sternite 3; surface depressed in midline, smooth; greatest width 3.0 × that of sternite 3, 2.7 – [2.9] × as wide as long. Pleon: Elevated ridges with a few scattered short setae. Tergite 2 with anterior and posterior transverse elevated ridges; tergites 3 – 4 with anterior transverse ridge; tergites 5 – 6 smooth. Eye: Eyestalk length as broad as long, peduncle distally setose, not distinctly expanded proximally; maximum corneal diameter 1.0 – [1.1] × rostrum width, as wide as eyestalk. Antennule: Article 1 1.5 × longer than wide, with 5 distal spines: distomesial spine well-developed; proximal lateral spine small. Antenna: Article 1 with prominent mesial process, distally clearly not reaching lateralmost antennular spine. Article 2 with minute distal spines laterally and mesially. Articles 3 and 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus [0.6] – 0.7 × length of ischium, with well-developed distal spine on extensor margin and 2 strong spines on flexor margin. P 1: [3.9] × carapace length (male), 2.9 (female); subcylindrical, spiny and with scattered long stiff setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, mesial spines usually stronger than others. Merus 1.2 – [1.6] length of carapace, [2.1] – 2.3 × as long as carpus. Carpus [2.7] – 3.5 × as long as wide. Palm 1.1 – [1.3] × carpus length, [2.6] – 2.7 × as long as broad. Fingers [0.6] – 0.8 × palm length; fixed finger unarmed; movable finger with basal spine. P 2 – 4: Slender, subcylindrical, moderately setose and spinose. Meri successively shorter posteriorly: P 3 merus 0.7 – [0.8] × length of P 2 merus, P 4 merus 0.8 – [0.9] × length of P 3 merus. P 2 merus, 0.9 × carapace length, [7.2] 7.0 – 8.5 × as long as broad, [1.3] – 1.4 × as long as P 2 propodus; P 3 merus [6.5] – 9.0 × as long as broad, [1.1] – 1.3 × as long as P 3 propodus; P 4 merus [5.4] 5.0 – 7.1 × as long as broad, as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, distal spine absent; flexor margins of P 2 – 3 irregular, with distal spine, absent in P 4. Carpi with 1 – 2 spines on extensor margin on P 2 – 3, distal spine prominent, unarmed on P 4; row of small spines below extensor margin on lateral surface of P 2 – 3. Propodi slender, [5.5 – 7.0] 7.1 – 9.0 × as long as broad; extensor margin irregular, usually armed with distinct spine on proximal half of P 2; flexor margin with 4 slender movable spines in addition to distal pair. Dactyli 0.5 – [0.6] × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 7 – 10 movable spines. Eggs: Ov. F (MNHN-IU- 2017 - 3830) carried 25 eggs of 0.3 – 0.5 mm diameter. Live colour. Unknown. Genetic data. COI, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFF0E60E4F9CFB167518BC81.taxon	distribution	Distribution. Chesterfield Islands, from 500 to 750 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFF0E60E4F9CFB167518BC81.taxon	discussion	Remarks. Phylladiorhynchus boucheti is characterized by the presence of a triangular rostrum, 4 epigastric spines, 3 spines on the anterior branchial margin of the carapace and Mxp 3 merus with 2 spines on the flexor margin. The specimens range from 2.5 to 3.0 mm postorbital carapace length. Phylladiorhynchus boucheti closely resembles to P. iphiclus from French Polynesia and Vanuatu (see the differences under the Remarks of this species).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFF6E60D4F9CFA5D7504BB7E.taxon	description	(Figs. 9, 11 J) Phylladiorhynchus ikedai. — Baba, 1991, 485, fig. 4 a (in part, only specimens from MUSORSTOM 6, Stn CP 401 [MNHN Ga- 2044]). Records requiring verification:	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFF6E60D4F9CFA5D7504BB7E.taxon	materials_examined	Type material. Holotype. Indonesia, KARUBAR Stn DW 18, 05.3000 ° S, 133.0167 ° E, 205 – 212 m, 24 October 1991: M 3.0 mm (MNHN-IU- 2014 - 13803). Paratypes. Indonesia. KARUBAR Stn DW 18, 05.3000 ° S, 133.0167 ° E, 205 – 212 m, 25 October 1991: 1 F 2.7 mm (MNHN-IU- 2019 - 2697). New Caledonia. MUSORSTOM 6 Stn CP 401, 20.7025 ° S, 167.0058 ° E, 270 m, 14 February 1989: 1 M 2.3 mm (MNHN-IU- 2014 - 23834 (Ga- 2044 )). Chesterfield Islands. EBISCO Stn DW 2547, 21.1000 ° S, 158.6000 ° E, 356 – 438 m, 11 October 2005: 1 F 2.1 mm (MNHN-IU- 2016 - 449). Other material. New Caledonia. KANADEEP Stn DW 5025, 20.3833 ° S, 158.6667 ° E, 350 – 420 m, 21 September 2017: 1 M 2.6 mm (MNHN-IU- 2017 - 2754). EXBODI Stn CP 3829, 22.0333 ° S, 167.0833 ° E, 350 – 360 m, 8 September 2011: 1 ov. F 2.4 mm (MNHN-IU- 2019 - 2694). French Polynesia. TARASOC Stn DW 3503, 17.5667 ° S, 149.3000 ° W, 350 m, 25 October 2009: 2 ov. F 2.1 – 2.2 mm (MNHN-IU- 2014 - 13735).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFF6E60D4F9CFA5D7504BB7E.taxon	etymology	Etymology. From the name Butes, an Argonaut, son of Coronus. The name is considered a substantive in apposition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFF6E60D4F9CFA5D7504BB7E.taxon	description	Description. Carapace: as long as or slightly longer than broad; transverse ridges with dense short setae. Gastric region flattened with 4 transverse ridges: epigastric ridge indistinct, with few plumose setae, with 4 – 5 spines (rarely with 1 median spine, usually with one median produced scale furnished with thick plumose setae, and 2 pairs of spines laterally); anterior protogastric ridge not medially interrupted, nearly extending laterally to carapace margin; anterior mesogastric ridge not medially interrupted, or minutely interrupted, laterally interrupted by anterior branch of cervical groove, continuing uninterrupted to first branchial spine; anterior metagastric ridge scale-like. Mid-transverse ridge not interrupted, preceded by shallow or undistinct cervical groove, followed by 2 not interrupted or minutely interrupted ridges, interspersed with 1 – 2 short lateral ridges and sometimes few, short scattered scales. Lateral margins slightly convex, with 7 spines: first anterolateral spine well-developed, reaching or exceeding level of lateral orbital spine, second spine (hepatic) small, slightly dorsomesially from lateral margin, and followed by 5 branchial spines (3 anterior and 2 posterior). Rostrum leaf-like, horizontal, dorsally flattish or slightly concave, [1.6] 1.5 – 1.8 × as long as broad, length [0.4] – 0.5 and breadth 0.3 that of carapace; lateral margins smooth and slightly convex, with well-developed supraocular basal spines, subapical spines minute or absent. Pterygostomian flap with anterior small spine; upper margin smooth. Thoracic sternum: As wide as or slightly longer than wide. Sternite 3 quadrangular, 1.2 – [1.7] × as wide as long, anterior margin straight or slightly convex, moderately produced anterolaterally. Sternite 4 widely contiguous to sternite 3; surface not depressed in midline, smooth; greatest width 2.4 – [3.0] × that of sternite 3, 1.7 – [2.2] × as wide as long. Pleon: Elevated ridges with short setae and few scattered long setae. Tergite 2 – 3 with anterior and posterior transverse elevated ridges; tergite 4 with anterior transverse ridge; tergites 5 – 6 smooth. Eye: Eyestalk length about 1.1 × broader than long, peduncle distally setose, not distinctly expanded proximally; maximum corneal diameter [1.1] × rostrum width, as wide as eyestalk. Antennule: Article 1 1.3 × longer than wide, with 5 spines: distomesial spine well-developed; proximal lateral spine small, always present. Antenna: Article 1 with prominent mesial process, nearly reaching end of lateralmost antennular spine. Article 2 with distinct distal spines laterally and mesially. Article 3 with small distomesial spine. Article 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus 0.5 × length of ischium with well-developed distal spine on extensor margin and 2 strong spines on flexor margin. P 1 (lost in holotype): 3.4 – 3.6 (males and females) × carapace length; slender, subcylindrical, spiny and with scattered long stiff setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus 1.3 – 1.4 length of carapace, 1.9 – 2 × as long as carpus. Carpus 3.1 – 3.3 × as long as wide. Palm 1.3 – 1.4 × carpus length, 3.0 – 3.6 × as long as broad. Fingers unarmed, 0.7 – 0.8 × palm length. P 2 – 4: Slender, subcylindrical, moderately setose and spinose. Meri successively shorter posteriorly: P 3 merus 0.8 – [0.9] × length of P 2 merus, P 4 merus [0.8] – 0.9 × length of P 3 merus. P 2 merus, [0.8] – 0.9 × carapace length, [4.5] – 5.5 × as long as broad, 1.2 – [1.3] × as long as P 2 propodus; P 3 merus [4.5] – 5.0 × as long as broad, 1.1 – [1.3] × as long as P 3 propodus; P 4 merus [4.0] – 5.1 × as long as broad, [1.0] – 1.2 × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, with small distal spine; flexor margins of P 2 – 4 irregular, each with distal spine; P 4 lateral surface with 1 – 2 small spines, absent in P 2 – 3. Carpi with 3 or 4 spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, smaller on P 4; row of small spines below extensor margin on lateral surface of P 2 – 3, unarmed on P 4; flexor margin unarmed. Propodi moderately slender, [5.5 – 6.0] 5.5 – 7.5 × as long as broad; extensor margin irregular, usually armed with 2 – 4 small proximal spines on P 2 – 3; flexor margin with 2 – 4 slender movable spines in addition to distal pair. Dactyli 0.6 – 0.7 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 5 – 8 movable spines. Eggs: Ov. F carried approximately 5 – 15 eggs of 0.4 – 0.6 mm diameter. Live colour. Unknown. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFF6E60D4F9CFA5D7504BB7E.taxon	distribution	Distribution. New Caledonia, Chesterfield Islands, Indonesia (Kei Islands), French Polynesia, from 205 to 438 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFF6E60D4F9CFA5D7504BB7E.taxon	discussion	Remarks. Phylladiorhynchus butes belongs to the group of species having the epigastric ridge with 4 spines and often one median process or scale with thick plumose setae (rarely with 5 spines). The specimens range from 2.1 to 3.0 mm postorbital carapace length. Phylladiorhynchus butes resembles P. cepheus, from French Polynesia, and P. maestratii, from New Caledonia (see the differences under the Remarks of P. maestratii). The three sequences of P. butes from French Polynesia, Indonesia and New Caledonia diverged 0.3 % for COI and 0.3 % for 16 S.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFF5E6334F9CFD197327BAD8.taxon	description	(Figs. 10, 11 K)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFF5E6334F9CFD197327BAD8.taxon	materials_examined	Type material. Holotype. French Polynesia. BENTHAUS Stn DW 2009, 22.5333 ° S, 151.3333 ° W, 320 – 450 m, 24 November 2002: ov. F 3.4 mm (MNHN-IU- 2014 - 13860). Paratypes. French Polynesia. BENTHAUS Stn DW 1945, 23.8167 ° S, 147.7000 ° W, 120 – 500 m, 17 November 2002: 1 M 2.4 mm (MNHN-IU- 2014 - 13858). Other material. French Polynesia. BENTHAUS Stn DW 2006, 22.4500 ° S, 151.3167 ° W, 350 – 450 m, 24 November 2002: 1 M 2.7 mm, 1 ov. F 2.2 mm (MNHN-IU- 2014 - 13882). — Stn DW 2009, 22.5333 ° S, 151.3333 ° W, 320 – 450 m, 24 November 2002: 1 ov. F 2.5 mm (MNHN-IU- 2019 - 2686), 1 ov. F 3.1 (MNHN-IU- 2014 - 13732).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFF5E6334F9CFD197327BAD8.taxon	etymology	Etymology. From the name Cepheus, an Argonaut, son of Coronus. The name is considered a substantive in apposition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFF5E6334F9CFD197327BAD8.taxon	description	Description. Carapace: [1.1] – 1.2 × as long as broad, transverse ridges with dense short setae. Gastric region flattened, with 3 – 4 transverse ridges: epigastric ridge indistinct, with a few plumose setae, and 4 – 5 spines (rarely 1 median spine, usually 1 median produced process or scale with thick setae, and 2 pairs of spines laterally); anterior protogastric ridge not medially interrupted; anterior mesogastric ridge not medially interrupted, laterally interrupted by cervical groove; anterior metagastric ridge scale-like, otherwise absent, with short scattered scales in anterior metagastric area. Mid-transverse ridge not interrupted, preceded by shallow or undistinct cervical groove, followed by 2 uninterrupted or minutely interrupted ridges, interspersed with 1 – 2 short lateral ridges and few, short scattered scales. Lateral margins straight, with 7 spines: first anterolateral spine well-developed, reaching or exceeding level of lateral orbital spine, second spine (hepatic) small, slightly dorsomesially from lateral margin, and followed by 5 branchial spines (3 anterior and 2 posterior). Rostrum leaf-like to sub-triangular, horizontal, dorsally flattish or slightly concave [1.5] – 1.9 × as long as broad, length [0.4] – 0.5 and breadth 0.2 – [0.3] that of carapace; lateral margins smooth and slightly convex, with well-developed supraocular basal spines and minute subapical spines. Pterygostomian flap ending in sharp tooth; upper margin smooth. Thoracic sternum: As wide as long. Sternite 3 moderately broad [2.3] 2.0 – 2.5 × as wide as long, anterior margin convex. Sternite 4 widely contiguous to sternite 3; surface depressed in midline, smooth; greatest width 2.4 – [2.8] × that of sternite 3, 3.0 × as wide as long. Pleon: Elevated ridges with short setae. Tergite 2 with anterior and posterior transverse elevated ridges; tergites 3 – 4 with anterior transverse ridge; tergites 5 – 6 smooth. Eye: Eyestalk length about 1.0 – [1.1] × broader than long, peduncle distally setose, not distinctly expanded proximally; maximum corneal diameter [0.8] – 0.9 × rostrum width, as wide as eyestalk. Antennule: Article 1.4 × longer than wide, with 5 distal spines: distomesial spine well-developed; proximal lateral spine small, always present. Antenna: Article 1 with prominent mesial process, nearly reaching end of lateralmost antennular spine. Article 2 unarmed, sometimes with minute distolateral spine or granule. Articles 3 and 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus 0.5 – [0.6] × length of ischium at midlength, extensor margin with well-developed distal spine, flexor margin with 2 strong spines. P 1: slender 4.5 – 5.0 × carapace length (males), [3.7] 3.5 – 3.9 (females); subcylindrical, spiny and with scattered long stiff setae and dense plumose thick setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, mesial spines usually stronger than others. Merus [1.4] 1.3 – 1.9 length of carapace, [2.1] – 2.5 × as long as carpus. Carpus [3.3] 2.4 – 3.3 × as long as wide. Palm [1.3] – 1.4 × carpus length, [3.9] 3.3 – 4.6 × as long as broad. Fingers [0.7] 0.6 – 0.8 × palm length; fixed and movable fingers each with basal spine. P 2 – 4: Slender, subcylindrical, moderately setose and spinose. Meri successively shorter posteriorly: P 3 merus [0.5] – 0.6 × length of P 2 merus, P 4 merus 0.9 – [1.0] × length of P 3 merus. P 2 merus, [0.8] – 1.0 × carapace length, [5.4] – 6.3 × as long as broad, [1.1] – 1.3 × as long as P 2 propodus; P 3 merus [5.1] – 5.3 × as long as broad, 1.2 × as long as P 3 propodus; P 4 merus 5.5 – [5.9] × as long as broad, as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, distal spine absent; flexor margins with distal spine in P 2 – 3, absent in P 4; P 4 lateral surface with 3 small spines, absent in P 2 – 3. Carpi with 8 – 9 small spines on extensor margin on P 2 – 3 (serrated), with 2 small spines on P 4; distal spine prominent on P 2, smaller or absent in P 3 – 4; flexor margin unarmed. Propodi slender, [7.5 – 8.0] 7.8 – 8.0 × as long as broad; extensor margin irregular, usually unarmed; flexor margin with 2 – 3 slender movable spines in addition to distal pair. Dactyli 0.4 – 0.5 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 6 – 7 movable spines. Eggs: Ov. F carried approximately 30 – 40 eggs of 0.3 – 0.4 mm diameter. Live colour. Unknown. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFF5E6334F9CFD197327BAD8.taxon	distribution	Distribution. French Polynesia, from 230 to 500 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFF5E6334F9CFD197327BAD8.taxon	discussion	Remarks. Phylladiorhynchus cepheus belongs to the group that present the epigastric ridge with 4 spines and often one median process or scale with thick plumose setae (rarely with 5 spines), 3 spines on the anterior branchial margin of the carapace and the Mxp 3 merus with two prominent spines along the flexor margin. The specimens range from 2.4 to 3.4 mm postorbital carapace length. Phylladiorhynchus cepheus closely resembles to P. butes, from New Caledonia, Chesterfield Islands, Indonesia and French Polynesia, and P. maestratii, from New Caledonia (see the differences under the Remarks of P. maestratii).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFCBE6374F9CFBB3740BBBB6.taxon	description	(Figs. 11 L, 12) Phylladiorhynchus ikedai. — Baba, 1991, 485 (in part only specimens from MUSORSTOM 4, Stn 151 [MNHN Ga- 2042] and CALSUB, Pl 18 [MNHN Ga- 2047]). Records requiring verification:	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFCBE6374F9CFBB3740BBBB6.taxon	materials_examined	Type material. Holotype. Indonesia. KARUBAR Stn DW 18, 05.3000 ° S, 133.0167 ° E, 205 – 212 m, 24 October 1991: M 3.0 mm (MNHN-IU- 2014 - 13846). Paratypes. Philippines. MUSORSTOM 2 Stn CP 01, 14.0000 ° N, 120.3000 ° E, 188 – 198 m, 20 November 1980: 1 M 2.1 mm (MNHN-IU- 2014 - 13801). Indonesia. KARUBAR Stn DW 18, 05.3000 ° S, 133.0167 ° E, 205 – 212 m, 24 October 1991: 1 F 2.8 mm (MNHN- IU- 2016 - 497). — Stn DW 50, 07.9833 ° S, 133.0333 ° E, 184 – 186 m, 30 October 1991: 1 F 1.7 mm (MNHN-IU- 2014 - 13850). Papua New Guinea. BIOPAPUA Stn DW 3770, 05.5667 ° S, 51.5333 ° E, 220 – 294 m, 16 October 2010: 1 M 2.5 mm (MNHN-IU- 2011 - 2157). Other material. New Caledonia. MUSORSTOM 4 Stn DW 151, 19.1167 ° S, 163.3667 ° E, 200 m, 14 September 1985: 1 M 2.2 mm (MNHN-IU- 2014 - 23833 (Ga- 2042 )). — CALSUB Pl 18, 22.7667 ° S, 167.3333 ° E, 200 – 300 m, 29 March 1989: 1 M 2.3 mm (MNHN-IU- 2014 - 23837 (Ga- 2047 )). — Lagon Nord. Stn DW 1151, 19.0167 ° S, 163.4500 ° E, 270 – 280 m, 28 October 1989: 1 M 3.2 mm (MNHN-IU- 2014 - 13798). — EXBODI Stn CP 3833, 22.0333 ° S, 167.0667 ° E, 325 – 332 m, 8 September 2011: 1 M 2.4 mm (MNHN-IU- 2019 - 2695). — KANACONO Stn CP 4656, 22.6667 ° S, 167.2000 ° E, 219 – 244 m, 10 August 2016: 1 F 2.4 mm (MNHN-IU- 2017 - 11637). — Stn CP 4665, 22.8667 ° S, 167.3333 ° E, 535 – 563 m, 11 August 2016: 1 M 1.9 mm, 2 F 2.4 – 2.5 mm (MNHN- IU- 2017 - 3954).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFCBE6374F9CFBB3740BBBB6.taxon	etymology	Etymology. From the name Eneus, an Argonaut, son of Caeneus. The name is considered a substantive in apposition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFCBE6374F9CFBB3740BBBB6.taxon	description	Description. Carapace: Moderately robust, 0.9 – [1.0] × as long as broad; transverse ridges with dense short setae, and few scattered thick iridescent setae. Gastric region convex (upraised dorsally) with some transverse ridges: epigastric ridge indistinct, with 5 spines (1 median and 2 pairs of spines laterally) followed by median scale behind epigastric ridge, often armed with very long thick plumose setae; anterior protogastric ridge medially interrupted, followed by 1 median scale in posterior protogastric area; anterior mesogastric ridge not medially interrupted, laterally interrupted by anterior branch of cervical groove; anterior metagastric ridge not medially interrupted, and followed by a few scales on posterior metagastric region. Mid-transverse ridge medially depressed, not interrupted, preceded by distinct cervical groove, followed by 2 not interrupted or minutely interrupted ridges, interspersed with 1 short lateral ridge. Lateral margins clearly convex, with 7 spines: first anterolateral spine well-developed, reaching end of lateral orbital spine, second spine (hepatic) small, slightly dorsomesially from lateral margin, and followed by 5 branchial spines (3 anterior and 2 posterior). Rostrum leaf-like, horizontal, dorsally flattish or slightly concave, [1.3] – 1.5 × as long as broad, length 0.3 – [0.4] and breadth 0.2 – [0.3] that of carapace; lateral margins serrated and convex, with well-developed supraocular basal spines, subapical spines minute or obsolescent. Pterygostomian flap ending in anterior spine, upper margin serrated, with series of uprised striae. Thoracic sternum: As wide as long or slightly wider than long. Sternite 3 moderately broad, [2.5] 2.0 – 3.0 × as wide as long, anterior margin convex with a median deep notch. Sternite 4 widely contiguous to sternite 3; surface depressed in midline, smooth; greatest width 2.6 – [2.7] × that of sternite 3, 2.3 – [2.6] × as wide as long. Pleon: Elevated ridges with short setae and a few scattered long setae. Tergites 2 – 3 with anterior and posterior transverse elevated ridges; tergite 4 with anterior transverse ridge; tergites 5 – 6 smooth. Eye: Eyestalk length about 1.2 – 1.3 × broader than long, peduncle distally setose, not distinctly expanded proximally; maximum corneal diameter 0.75 × rostrum width, as wide as eyestalk. Antennule: Article 1 slightly longer than wide, with 5 spines: distomesial spine small; proximal lateral spine small, always present. Antenna: Article 1 with prominent mesial process, distally clearly not reaching lateralmost antennular spine. Article 2 with distinct distomesial and distolateral spines. Articles 3 and 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus 0.5 – [0.6] × length of ischium, with well-developed distal spine on extensor margin and 2 strong spines on flexor margin. P 1: [3.1] 3.0 – 3.3 (males), 2.8 – 2.9 (females) × carapace length; subcylindrical, spiny and with dense short setae and scattered long stiff setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus [1.2] 1.1 – 1.3 length of carapace, [2.0] – 2.7 × as long as carpus. Carpus 1.8 – [2.3] × as long as wide. Palm 1.2 – [1.3] × carpus length, [2.2] – 2.7 × as long as broad with parallel rows of spines running along lateral and mesial margin and scattered spines on dorsal and ventral surfaces. Fingers [0.7] 0.7 – 0.9 × palm length fixed finger unarmed; movable finger with well-developed basal spine. P 2 – 4: Slender, subcylindrical, moderately setose and spinose. Meri successively shorter posteriorly: P 3 merus 0.8 – [1.0] × length of P 2 merus, P 4 merus [lost in holotype] 0.9 × length of P 3 merus. P 2 merus, [0.7] – 0.8 × carapace length, 4.9 – [5.0] × as long as broad, 1.1 – [1.2] × as long as P 2 propodus; P 3 merus [6.0] – 7.0 × as long as broad, [1.1] – 1.3 × as long as P 3 propodus; P 4 merus 7.0 × as long as broad, as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, with small distal spine; flexor margins of P 2 – 4 irregular, each with distal spine; P 4 lateral surface with median row of 4 small spines, absent in P 2 – 3. Carpi with 2 or 3 spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, smaller on P 4; row of small spines below extensor margin on lateral surface of P 2 – 3, unarmed on P 4; flexor margins of all P 2 – 4 irregular, each with distal spine. Propodi slender, [7.5 – 8.5] 7.0 – 10.0 × as long as broad; extensor margin irregular, usually unarmed; flexor margin with 3 – 4 slender movable spines in addition to distal pair. Dactyli [0.7] – 0.8 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 6 – 7 movable spines. Eggs: No data. Live colour. Unknown. Genetic data. COI, mini-barcode fragment (158 bp).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFCBE6374F9CFBB3740BBBB6.taxon	distribution	Distribution. Philippines, Indonesia, Papua New Guinea and New Caledonia, from 184 to 563 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFCBE6374F9CFBB3740BBBB6.taxon	discussion	Remarks. Phylladiorhynchus eneus belongs to the group of species having the epigastric ridge with 5 spines, 3 spines on the anterior branchial margin of the carapace and the Mxp 3 merus with 2 prominent spines along the flexor margin. This group contains the following species: P. acastus from the Philippines, Papua New Guinea, Vanuatu, Chesterfield Islands and New Caledonia, P. argus, from French Polynesia, New Caledonia and Chesterfield Islands, P. paula, from the southwestern Indian Ocean, and P. erebus Schnabel & Ahyong, 2019 from Norfolk and Kermadec Ridges. However, the new species can be distinguished from the other species of the group by the following characters: - One median scale behind the median epigastric spine in P. eneus, whereas this scale is absent in the other species. - The anterior protogastric ridge is medially interrupted or scale-like in P. eneus, whereas this ridge is not interrupted in the other species. - The P 2 – 4 propodi are more slender (7 – 10 × longer than wide) in P. eneus, whereas these propodi are 4 – 7 × longer than wide in the other species. - The anterior margin of the pterygostomian flap is serrated margin in P. eneus whereas smooth in the other species. The specimens of P. eneus range in size range from 1.9 to 3.2 mm postorbital carapace length. The genetic divergences between P. eneus and other species were always higher than 18 % (COI, based on the 158 bp minibarcode). The two sequences of P. eneus from New Caledonia diverged by 0.6 %.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFCFE6374F9CFD447384BCA3.taxon	diagnosis	Diagnosis (modified from Schnabel & Ahyong 2019). Rostrum triangular, subapical spines obsolescent, lateral margins straight. Carapace with transverse row of 5 epigastric spines; parahepatic spines on protogastric region absent; anterior protogastric ridge usually medially interrupted; anterior metagastric ridge scale-like; anterior branchial margin with 3 spines. Pleonal tergite 3 without posterior transverse ridge. Thoracic sternite 3 anterior margin sinuous, with shallow median concavity. Antennular article 1 with 5 or 6 distal spines: distomesial spine distinctly shorter than upper distolateral spine. Antennal article 1 mesial process distally falling well short of second lateral antennular spine; article 2 distolateral and distomesial spines subequal; article 3 with small to distinct mesial spine, laterally unarmed. Mxp 3 merus with 2 prominent spines on flexor margin. P 2 – 4 dactylus extensor margin without upright spines at bases of movable spines. Genetic data. COI, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFCFE6374F9CFD447384BCA3.taxon	distribution	Distribution. New Zealand, Norfolk and Kermadec ridges, at 290 – 506 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFCFE6374F9CFD447384BCA3.taxon	discussion	Remarks. This species is characterized by the presence of 5 epigastric spines, 3 spines on the anterior branchial margin and the anterior protogastric ridge usually medially interrupted. The species is closely related to P. acastus from the Philippines, Papua New Guinea, Vanuatu, Chesterfield Islands and New Caledonia, P. argus, from French Polynesia, New Caledonia and Chesterfield Islands, P. paula, from SW Indian Ocean, and P. eneus from Indonesia to New Caledonia (see the differences under the Remarks of P. eneus).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFCFE63A4F9CF9B370D8BAD6.taxon	description	(Figs. 13, 21 A, 30 A – C, M – N)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFCFE63A4F9CF9B370D8BAD6.taxon	materials_examined	Type material. Holotype. Australia. Queensland, Heron Is. 23.4733 ° S, 151.9505 ° E, no depth, 17 November 2006: ov. F 2.8 mm (UF 25255). Paratypes. Australia. Queensland, Heron Island. 23.4418 ° S, 151.9004 ° E, no depth, 16 November 2006: 1 M 1.8 mm (UF 34732). — 23.5545 ° S, 152.0339 ° E, no depth, 16 November 2006: 1 F 1.1 mm (UF 26300). — 23.4733 ° S, 151.9505 ° E, no depth, 17 November 2006: 1 F 1.8 mm (UF 25213), 2 M 1.6 – 1.8 mm (UF 25216), 1 ov. F 2.8 mm UF 25255), 2 ov. F 2.1 – 2.3 mm (UF 25215), 1 ov. F 2.7 mm (UF 25222). New Caledonia. Grand Recif Sud. Stn 296, 22.6833 ° S, 166.722 ° E, 26 m, November 1984: 2 F 2.3 – 3.5 mm (MNHN-IU- 2016 - 489). Other material. New Caledonia. Ouen Is. Prony Bay. Stn 101, 22.5167 ° S, 166.5983 ° E, 18 m, August 1984: 1 ov. F 2.8 mm (MNHN-IU- 2019 - 2618). — Grand Recif Sud. Stn 297, 22.6483 ° S, 116.7600 ° E, 30 m, November 1984: 2 M 3.2 – 3.3 mm (MNHN-IU- 2019 - 2612). — Stn 304, 22.6633 ° S, 166.7983 ° E, 27 m, November 1984: 1 M 3.2 mm, 2 ov. F 3.5 – 3.6 mm (MNHN-IU- 2019 - 2617). — Stn 338, 22.7450 ° S, 166.8183 ° E, 32 m, November 1984: 1 M 3.2 mm (MNHN-IU- 2019 - 2611). — Stn 291, 22.6400 ° S, 166.7283 ° E, 31 m, November 1984: 1 M 2.7 mm (MNHN- IU- 2019 - 2609). — Passe Sarcelle. 35 – 40 m, 1 ov. F 2.7 mm (MNHN-IU- 2019 - 2610). — Lagon. Stn 336, 22.6917 ° S, 166.8567 ° E, 26 m, November 1984: 2 M 2.3 – 3.4 mm (MNHN-IU- 2014 - 13911). — Recif Serrez. 7 m, 7 September 1992: 1 F 3.4 mm (MNHN-IU- 2019 - 2615). — LIFOU Stn 1410, 20.9450 ° S, 167.0516 ° E, 2 – 4 m, 25 November 2000: 1 M 2.1 mm (MNHN-IU- 2019 - 2604). Chesterfield Islands. CORAIL 2 Stn DW 101, 19.1500 ° S, 158.4366 ° E, 37 m, 27 July 1988: 1 M 2.3 mm (MNHN-IU- 2016 - 480 (Ga- 2053 )). — Stn CP 23, 20.5100 ° S, 161.0600 ° E, 80 – 83 m, 22 July 1988: 2 ov. F 2.0 – 2.1 mm (MNHN-IU- 2016 - 479 (Ga- 2048 )). — Stn CP 24, 20.4566 ° S, 161.0783 ° E, 74 – 75 m, 22 July 1988: 1 ov. F 1.9 mm (MNHN-IU- 2016 - 471 (Ga- 2049 )). — Stn DW 33,19.4166 ° S, 158.8683 ° E, 52 m, 23 July 1988: 1 ov. F 2.1 mm (MNHN-IU- 2016 - 478).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFCFE63A4F9CF9B370D8BAD6.taxon	etymology	Etymology. From the name Euryalus, an Argonaut, son of Mecisteus. The name is considered a substantive in apposition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFCFE63A4F9CF9B370D8BAD6.taxon	description	Description. Carapace: As long as or slightly longer than broad; transverse ridges with dense short setae and thick iridescent setae. Gastric region slightly convex with some transverse ridges: epigastric ridge distinct with 2 median spines and some lateral short scales, followed by small short scales on posterior epigastric region; anterior protogastric ridge not medially interrupted, nearly extending laterally to carapace margin, often followed by uninterrupted posterior protogastric ridge or short scales; anterior mesogastric ridge not medially interrupted, laterally interrupted by anterior branch of cervical groove, and continuing uninterrupted to first branchial spine, often followed by small scales; anterior metagastric ridge not medially interrupted, followed by posterior scalelike metagastric ridge. Mid-transverse ridge not interrupted, medially depressed, followed by shallow or indistinct cervical groove, followed by 2 – 3 not interrupted or minutely interrupted ridges, interspersed with 2 interrupted ridges and few short scales. Lateral margins convex, with 6 distinct spines: first anterolateral spine well-developed, reaching anteriorly to level of lateral orbital spine, second spine (hepatic) well-developed, slightly dorsomesially from lateral margin, and followed by 4 branchial spines (3 anterior and 1 posterior). Rostrum dagger-like, horizontal dorsally flattish or slighly concave [1.5] – 1.9 × as long as broad, length [0.4] – 0.5 and breadth [0.3] 0.3 that of carapace; lateral margins serrated and straight or nearly straight, with well-developed supraocular basal spines and subapical spines. Pterygostomian flap with anterior spine, upper margin smooth. Thoracic sternum: As wide as long. Sternite 3 moderately broad, 2.2 – [2.3] × as wide as long, anterior margin convex, with blunted median projection, anterolaterally rounded to produced. Sternite 4 widely contiguous to sternite 3; anterolaterally smooth, surface depressed in midline, smooth; greatest width [2.7] – 3.2 × that of sternite 3, [2.5] – 4.0 × as wide as long. Pleon: Elevated ridges with short setae and a few scattered long setae. Tergite 2 – 4 with anterior and posterior transverse elevated ridges; tergites 5 – 6 smooth. Eye: Eyestalk length about 0.9 – [1.1] × broader than long, peduncle distally setose, not distinctly expanded proximally, with few short transverse striae on lateral surfaces; cornea expanded distally, maximum corneal diameter 0.8 – [0.9] × rostrum width, [0.8] as long as eyestalk. Antennule: Article 1 longer than wide, with 5 distal spines: distomesial spine well-developed; proximal lateral spine prominent, reaching half-length of first lateral spine and overreaching distomesial spine, always present. Antenna: Article 1 with prominent mesial process, distally falling well short of lateralmost antennular spine. Article 2 with well-developed distomesial and distolateral spines. Article 3 and 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus [1.1] × length of ischium, with well-developed distal spine on extensor and flexor margins. P 1: Moderately slender, 2.6 – 3.1 (males), 2.0 – [2.2] (females) × carapace length; subcylindrical, spiny and with long stiff setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus [0.7] – 1.3 length of carapace, [1.4] – 1.6 × as long as carpus. Carpus 1.5 – [2.2] × as long as wide. Palm [1.0] – 1.1 × carpus length, [2.0] – 2.1 × as long as broad. Fingers 0.9 – [1.1.] × palm length; fixed finger with 0 – 1 basal spines; movable finger often with basal spines or granules P 2 – 4: Setose and spinose. Meri successively shorter posteriorly: P 3 merus [0.8] – 0.9 × length of P 2 merus, P 4 merus [0.8] – 0.9 × length of P 3 merus. P 2 merus, 0.6 – [0.7] × carapace length, 3.6 – [4.3] × as long as broad, 1.1 – [1.4] × as long as P 2 propodus; P 3 merus [3.6] – 3.7 × as long as broad, 1.1 – [1.4] × as long as P 3 propodus; P 4 merus 3.2 – [3.6] × as long as broad, 1.1 × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, unarmed; flexor margin irregular, with distal spine on P 2 – 3, absent or small on P 4. Carpi with 2 – 4 small spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, absent on P 4; row of granules below extensor margin on lateral surface of P 2 – 4; flexor margin unarmed. Propodi stout, 3.0 – 4.0 × as long as broad; extensor margin irregular; flexor margin with 3 – 6 slender movable spines in addition to distal pair. Dactyli 0.6 – [0.7] × length of propodi, ending in incurved, strong, sharp spine; flexor margin with dactylar spines at basis of 4 – 6 movable spines. Eggs: Ov. F carried approximately 9 – 20 eggs of 0.5 – 0.6 mm diameter. Live colour. Unknown. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFCFE63A4F9CF9B370D8BAD6.taxon	distribution	Distribution. Australia, Queensland (Heron Islands), New Caledonia and Chesterfield Islands, from 2 to 83 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFCFE63A4F9CF9B370D8BAD6.taxon	discussion	Remarks. Phylladiorhynchus euryalus belongs to the species-group having 2 epigastric spines, 1 hepatic spine, 3 spines on anterior branchial margin, rostrum dagger-like (margin straight or nearly straight) and dactylar spines along the flexor margin of the P 2 – 4 dactyli. This species complex includes the following species: P. asclepius, from Western Australia, P. euryalus, from Queensland, New Caledonia and Chesterfield Islands, P. lini, from Taiwan, and P. spinosus Schnabel & Ahyong, 2019, from New Zealand and New Caledonia. These species are morphologically very similar although genetically very distinct (see the differences under the Remarks of P. spinosus). The specimens of P. euryalus range from 1.6 to 3.6 mm postorbital carapace length. The six sequences of P. euryalus from Australia (Queensland) and New Caledonia diverged 1.3 % for COI, four sequences diverged 0.7 % for 16 S.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFC2E63B4F9CFBA47240BE6E.taxon	description	(Figs. 14, 21 B, 27 A – B, 54 D)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFC2E63B4F9CFBA47240BE6E.taxon	materials_examined	Type material. Holotype. French Polynesia. Society Islands, Moorea Island, NE of Tareu Pass, 17.4836 ° S, 149.8581 ° W, 22 m, 24 July 2006: ov. F 2.2 (UF 9766). Paratypes. Cook Islands. Rarotonga Island, Matavera, 21.226 ° S, 159.7293 ° W, 9 – 24 m, 26 June 2006: 1 ov. F 2.0 mm (UF 9232). French Polynesia. Society Islands. Moorea Is. 17.506 ° S, 149.759 ° W, 15 – 22 m, 27 May 2006: 1 ov. F 1.8 mm (UF 9471), 1 ov. F 2.0 (UF 9603). — 17.4836 ° S, 149.8581 ° W, 22 m, 24 July 2006: 1 M 1.7 (UF 9756). — 17.4768 ° S, 149.8327 ° W, 5 August 2006: 2 M 1.0 – 2.2 mm, 1 ov. F 2.1 mm (UF 14832). — 17.4764 ° S, 149.8327 ° W, 4 – 7 m, 10 November 2008: 1 M 2.3 mm (UF 16338), 1 ov. F 2.3 mm (UF 16370), 1 M 2.4 mm, 1 ov. F 2.1 mm, 1 F 1.7 mm (UF 18384), 1 M 2.4 mm (UF 16399). — French Polynesia. BENTHAUS Stn DW 1926, 24.6360 ° S, 146.0136 ° W, 50 – 90 m, 13 November 2002: 1 M 2.2 mm, 1 ov. F 2.6 mm (MNHN- IU- 2019 - 2592). Other material. French Polynesia. Society Islands. Moorea Is. Maharepa, 2 m, 11 October 2008: 1 ov. F 2.3 mm (UF 15431). — on Pocillopora, June 2009: 1 M 2.0 mm (UF 37850). — 17.484 ° S, 149.9264 ° W, 19 – 20 m, 3 November 2009: 1 M 1.8 mm (UF 23879). — 17.4756 ° S, 149.8425 ° E, 13 – 17 m, 5 December 2009: 1 ov. F 2.2 mm (UF 24183). — 17.4764 ° S, 149.8316 ° W, 13 m, 31 January 2012: 4 M 1.4 – 2.1 mm, 9 ov. F 1.7 – 2.1 mm, 2 F 1.2 – 1.9 mm (UF 33774). — 17.4759 ° S, 149.8419 ° W, 13 m, 4 February 2012: 10 M 1.4 – 2.3 mm, 9 ov. F 1.6 – 2.3 mm, 1 F 1.9 mm (UF 33968), 1 F 2.2 mm (UF 33966), 1 M 2.0 mm (UF 33967). Gambier Islands. Totegegie Airport, 23.0776 ° S, 134.8884 ° W, 22.5 m, 6 February 2013: 1 ov. F 2.3 mm (UF 35486), 1 M 2.2 mm (UF 35473), 1 F 1.8 mm (UF 35479), 1 ov. F 2.5 mm (UF 35481), 1 M 2.3 mm (UF 35485).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFC2E63B4F9CFBA47240BE6E.taxon	etymology	Etymology. The new species is named after our colleague Gustav Paulay from the Florida Museum of Natural History for his valuable contribution to marine research.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFC2E63B4F9CFBA47240BE6E.taxon	description	Description. Carapace: Slightly broader than long; transverse ridges with dense short setae and long and thick iridescent setae. Gastric region slightly convex with some transverse ridges: epigastric ridge distinct with 2 spines, short scales laterally; anterior protogastric ridge not medially interrupted, nearly extending laterally to carapace margin, sometimes followed by small scales; anterior mesogastric ridge not medially interrupted, laterally continuing to first branchial spine, sometimes followed by some short lateral scales; anterior metagastric ridge scalelike, often followed by some short small median scale posteriorly. Mid-transverse ridge not interrupted, medially depressed, cervical groove indistinct, followed by 2 not interrupted or minutely interrupted ridges, interspersed with 1 short lateral ridge and few short scales. Lateral margins slightly convex, with 6 – 7 distinct spines: first anterolateral spine well-developed, reaching anteriorly to level of lateral orbital spine, second spine (hepatic) small, slightly dorsomesially from lateral margin, and followed by 5 branchial spines (3 anterior and 1 – 2 posterior). Rostrum leaf-like, horizontal, dorsally concave, [1.1] – 1.4 × as long as broad, length [0.3] – 0.4 and breadth 0.2 – [0.3] that of carapace; lateral margins serrated and strongly convex, with well-developed supraocular basal spines and small subapical spines (tridentiform). Pterygostomian flap ending in blunt tooth, upper margin smooth. Thoracic sternum: As wide as long. Sternite 3 moderately broad, 1.8 – [1.9] × as wide as long, anterior margin serrated and convex, anterolaterally rounded. Sternite 4 widely contiguous to sternite 3; anterolaterally serrated, surface depressed in midline, smooth; greatest width [3.3] × that of sternite 3, [2.5] × as wide as long. Pleon: Elevated ridges with short setae and a few scattered long setae. Tergites 2 – 3 with anterior and posterior transverse elevated ridges; tergite 4 with anterior transverse ridge only; tergites 5 – 6 smooth. Eye: Eyestalk length about 1.0 – [1.1] × broader than long, peduncle distally setose, not distinctly expanded proximally; cornea expanded distally, maximum corneal diameter [0.6] – 0.7 × rostrum width, narrower than eyestalk (0.8 – 0.9 maximum peduncle width). Antennule: Article 1 slightly longer than wide, with 4 distal spines: distomesial spine well-developed; proximal lateral spine always absent. Antenna: Article 1 with prominent mesial process distally falling well short or overreaching lateralmost antennular spine. Article 2 with well-developed distomesial and distolateral spines. Article 3 with distomesial spine. Article 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus 0.7 – [0.8] × length of ischium, with well-developed distal spine on extensor and flexor margins. P 1: [2.8] – 3.0 (males), 1.9 – 2.0 (females) × carapace length; subcylindrical, spiny and with long stiff setae and iridescent setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus 0.7 – [1.0] length of carapace, [1.6] – 1.9 × as long as carpus. Carpus [1.5] – 1.7 × as long as wide. Palm [1.1] – 1.2 × carpus length, [1.5] – 1.6 × as long as broad. Fingers 0.8 × palm length fixed finger with 2 small basal spines, movable finger with basal spine or unarmed. P 2 – 4: Stout, setose, with iridiscent setae, and spinose. Meri successively shorter posteriorly: P 3 merus 0.9 × length of P 2 merus, P 4 merus [0.8] – 0.9 × length of P 3 merus. P 2 merus, [0.7] – 0.9 × carapace length, 3.1 – [3.4] × as long as broad, 1.1 – [1.2] × as long as P 2 propodus; P 3 merus 3.4 – [3.8] × as long as broad, 1.1 – [1.2] × as long as P 3 propodus; P 4 merus 3.2 × as long as broad, 1.0 – 1.1 × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, unarmed; flexor margin irregular, with distal spine on P 2 – 3, distal spine absent in P 4. Carpi with 1 – 3 spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, smaller on P 4; granules below extensor margin on lateral surface of P 2 – 4; flexor margin unarmed. Propodi stout, [4.0] 3.6 – 4.3 × as long as broad; extensor margin irregular, armed with 2 – 4 spines; flexor margin with 3 – 6 slender movable spines in addition to distal pair. Dactyli [0.6] – 0.7 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 5 – 6 movable spines. Eggs: Ov. F carried approximately 8 – 20 eggs of 0.4 – 0.6 mm diameter. Live colour. Base colour of carapace and pleon whitish or beige, with small brownish patches. P 1 whitish or beige with scattered brownish spots. P 2 – 4 whitish, with brown bandings. Genetic data. COI, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFC2E63B4F9CFBA47240BE6E.taxon	distribution	Distribution. French Polynesia, Society Islands and Cook Islands, between 2 and 90 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFC2E63B4F9CFBA47240BE6E.taxon	discussion	Remarks. Phylladiorhynchus gustavi belongs to the group of species having 2 epigastric spines, 1 hepatic spine and 3 spines on anterior branchial margin. The specimens range from 1.2 to 2.6 mm postorbital carapace length. The closest species is P. laureae, from from Japan, Mariana Islands, American Samoa, Papua New Guinea, Vanuatu and New Caledonia (see the differences under the Remarks of that species). The two sequences of P. gustavi from French Polynesia diverged 0.4 % for COI.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFC1E63E4F9CFF2973E4BBEE.taxon	description	(Figs. 15, 21 C)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFC1E63E4F9CFF2973E4BBEE.taxon	materials_examined	Type material. Holotype. Chesterfield Islands. KANADEEP Stn DW 4960, 23.0667 ° S, 159.4667 ° E, 310 m, 6 September 2017: F 2.1 mm (MNHN-IU- 2017 - 2534). Paratypes. Chesterfield Islands. EBISCO no Stn details: 1 ov. F 1.8 mm (MNHN-IU- 2014 - 13851). KANADEEP Stn DW 4950, 24.0833 ° S, 159.7000 ° E, 330 – 500 m, 5 September 2017: 1 M 2.2 mm (MNHN-IU- 2017 - 2515). — Stn CP 4953, 24.1667 ° S, 159.6833 ° E, 270 – 320 m, 5 September 2017: 4 M 1.0 – 1.4 mm, 2 ov. F 1.6 – 1.8 mm, 6 F 1.4 – 1.8 mm (MNHN-IU- 2017 - 3326). Other material. Chesterfield Islands. KANADEEP Stn DW 4961, 23.0333 ° S, 159.4666 ° E, 300 – 430 m, 6 September 2017: 1 F 2.2 mm (MNHN-IU- 2017 - 2729). New Caledonia. KANACONO Stn CP 4673, 22.7833 ° S, 167.4500 ° E, 244 – 285 m, 13 August 2016: 1 ov. F 1.9 mm (MNHN-IU- 2017 - 11724).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFC1E63E4F9CFF2973E4BBEE.taxon	etymology	Etymology. From the Greek hepta, seven, and akantha, spine, in reference to the seven spines on the dorsal surface of the carapace.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFC1E63E4F9CFF2973E4BBEE.taxon	description	Description. Carapace: Robust or massive, [0.8] – 0.9 × as long as broad; transverse ridges elevated, serrated, with few short setae, and few scattered long thick iridescent setae. Gastric region convex (upraised dorsally) with transverse scale-like ridges: epigastric ridge indistinct, with 5 spines (1 median and and 2 pairs of spines laterally), followed by 1 scale behind median epigastric spine; anterior protogastric ridge undistinct or scale like, not continuing laterally to carapace margin, armed with 1 parahepatic spine on each side, behind lateralmost epigastric spines, posterior protogastric area scale-like; anterior mesogastric ridge indistinct with some scales or scale-like, laterally continuing with few scales, posterior mesogastric ridge scale-like; anterior metagastric ridge scale-like, followed by small scale (s) on posterior metagastric region. Mid-transverse ridge not interrupted, medially depressed, serrated laterally, preceded by distinct cervical groove, followed by 2 uninterrupted or minutely interrupted ridges, interspersed with 1 short lateral ridge and some scales. Lateral margins clearly convex, with 7 spines: first anterolateral spine well-developed, not reaching anteriorly end of strong lateral orbital spine, second spine (hepatic) well-developed, and followed by 5 branchial spines (3 anterior and 2 posterior). Rostrum leaf-like to subtriangular, dorsally sharply concave in anteroposterior midline, [1.1] 1.0 – 3.0 × as long as broad, length [0.4] – 0.4 and breadth [0.3] 0.3 that of carapace; lateral margins serrated and convex, with small supraocular basal spines, subapical spines absent. Pterygostomian flap ending in anterior spine, upper margin smooth, with series of uprised ridges. Thoracic sternum: 0.9 × as long as wide. Sternite 3 moderately broad, [2.4] – 2.6 × as wide as long, anterior margin straight, or slightly convex. Sternite 4 widely contiguous to sternite 3; surface depressed in midline, smooth; greatest width 2.6 – [2.7] × that of sternite 3, 2.5 × as wide as long. Pleon: Elevated ridges upraised dorsally, with a few scattered short setae. Tergites 2 – 3 with anterior and posterior transverse elevated ridges; tergite 4 with anterior transverse ridge; tergites 5 – 6 smooth. Eye: Eyestalk length about [1.5] 1.3 – 1.6 × broader than long, peduncle distally setose, not distinctly expanded proximally; maximum corneal diameter [0.5] – 0.6 × rostrum width, narrower than eyestalk. Antennule: Article 1 1.4 longer than wide, with 5 spines, distomesial spine well-developed, proximal lateral spine small. Antenna: Article 1 with prominent mesial process, distally falling well short of lateralmost antennular spine. Article 2 with small distal spines laterally and mesially. Article 3 with minute or distinct distomesial and distolateral spines. Article 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus [0.5] × length of ischium at midlength, with 0 – [1] distal spine on extensor margin and 2 spines on flexor margin. P 1: [2.2] (female) (lost in most specimens) × carapace length; subcylindrical, spiny and with scattered long stiff setae and scattered plumose setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus as long as carapace. Carpus [1.9] – 2.0 × as long as wide. Palm 1.2 – [1.3] × carpus length, [2.3] × as long as broad. Fingers unarmed, [0.7] – 0.8 × palm length. P 2 – 4: (lost in most specimens) Stout, subcylindrical, moderately setose and spinose. Meri successively shorter posteriorly: P 3 merus [0.8] × length of P 2 merus, P 4 merus [0.8] × length of P 3 merus. P 2 merus, 0.6 × carapace length, 4.4 × as long as broad, [0.9] × as long as P 2 propodus; P 3 merus [5] × as long as broad, [1.2] × as long as P 3 propodus; P 4 merus [4.6] × as long as broad, [0.9] × as long as P 4 propodus; extensor margin of P 2 – P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin with 2 spines, distal spine absent; flexor margins of P 2 – 4 irregular, with distal spine; P 2 – 4 lateral surface with short striae. Carpi with 1 – 2 spines on extensor margin on P 2 – 3, gap between the marginal and the distal spine, serrated on P 4; distal spine prominent on P 2 – 4; P 2 – 4 flexor margins irregular, with distal spine. Propodi moderately slender, [6.8] – 7.0 × as long as broad, extensor margin irregular, flexor margin with 2 – 3 slender movable spines in addition to distal pair. Dactyli 0.6 – 0.7 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 5 – 6 movable spines. Eggs. Ov. F carried approximately 5 – 15 eggs of 0.5 – 0.6 mm diameter. Colour. After some months in ethanol: light orange with reddish spots and patches remaining in carapace, rostrum and pleon. Genetic data. COI, mini-barcode fragment (158 bp).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFC1E63E4F9CFF2973E4BBEE.taxon	distribution	Distribution. New Caledonia and Chesterfield Islands, depth 244 – 500 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFC1E63E4F9CFF2973E4BBEE.taxon	discussion	Remarks. Phylladiorhynchus heptacanthus belongs to the group of species having 5 epigastric spines, 3 spines on the anterior branchial margin and a leaf-like rostrum. The closest species is P. eneus, from Indonesia, Philippines, Papua New Guinea and New Caledonia. However, P. heptacanthus is easily distinguished from this species by the presence of parahepatic spines, being absent in P. eneus. Furthermore, the rostrum supraocular basal spines are small in P. heptacanthu s, whereas they are well-developed in P. eneus. The specimens of P. heptacanthus range from 1.0 to 2.2 mm postorbital carapace length. The COI divergence (mini-barcode fragment) between P. eneus and P. heptacanthus was 22 %.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFC6E63D4F9CFC897284BC12.taxon	description	(Figs. 16, 21 D) Records requiring verification:	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFC6E63D4F9CFC897284BC12.taxon	materials_examined	Type material. Holotype. Reunion Island, MD 32 Stn CP 43, 21.3450 ° S, 55.4483 ° E, 73 – 77 m, 18 August 1982: M 1.9 mm (MNHN-IU- 2016 - 496). Paratypes. Reunion Island. MD 32 Stn CP 43, 21.3450 ° S, 55.4483 ° E, 73 – 77 m, 18 August 1982: 1 M 1.7 mm, 1 ov. F 1.6 mm (MNHN-IU- 2016 - 495).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFC6E63D4F9CFC897284BC12.taxon	etymology	Etymology. From the name Hylas, an Argonaut, son of Theiodamas and Menodice. The name is considered a substantive in apposition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFC6E63D4F9CFC897284BC12.taxon	description	Description. Carapace: 0.9 × as long as broad; transverse ridges with some short setae, and few scattered iridescent thick long setae. Gastric region slightly convex with some transverse ridges: epigastric ridge indistinct, with 3 spines (1 median and 2 spines laterally) and some outer granules, continuing with few scales; anterior protogastric ridge not medially interrupted, laterally interrupted with few scales to carapace margin, posterior protogastric ridge scale-like; anterior mesogastric ridge scale-like, laterally interrupted by anterior branch of cervical groove, laterally continuing with few scales, posterior mesogastric area with few scales; anterior metagastric not medially interrupted, followed by small scales on posterior metagastric region. Mid-transverse ridge not interrupted, medially depressed, preceded by a shallow or indistinct cervical groove, followed by 2 not interrupted or minutely interrupted ridges, interspersed with 3 short lateral ridges. Lateral margins slighly convex, with 7 spines: first anterolateral spine welldeveloped, reaching end of lateral orbital spine, second spine (hepatic) minute, and followed by 5 branchial spines (3 anterior and 2 posterior). Rostrum leaf-like, horizontal, dorsally flattish, 1.1 – [1.2] × as long as broad, length 0.4 and breadth 0.3 that of carapace; lateral margins serrated and convex, with well-developed supraocular spines, lateral margins of supraocular spines serrated, subapical spines distinct. Orbit sharply excavated. Pterygostomian flap ending in anterior spine, upper margin serrated. Thoracic sternum: As wide as long. Sternite 3 moderately broad, 2.0 – [2.6] × as wide as long, anterior margin convex, and with median feeble excavation. Sternite 4 widely contiguous to sternite 3; surface depressed in midline, smooth; greatest width 2.7 – [2.9] × that of sternite 3, 2.3 – [2.8] × as wide as long. Pleon: Elevated ridges with a few scattered short setae. Tergites 2 – 3 with anterior and posterior transverse elevated ridges; tergite 4 with anterior transverse ridge only; tergites 5 – 6 smooth. Eye: Eyestalk length about 1.1 × broader than long, peduncle distally setose, not distinctly expanded proximally; maximum corneal diameter 0.8 × rostrum width, as wide as eyestalk. Antennule: Article 1 1.3 × longer than wide, with 5 spines: distomesial spine well-developed; proximal lateral spine minute. Antenna: Article 1 with prominent mesial process, distally clearly not reaching lateralmost antennular spine. Article 2 with small distomesial and distolateral spines. Article 3 sometimes with small distomesial spine. Article 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus [0.7] – 0.8 × length of ischium, with 1 median and 1 distal spine on extensor margin, 1 – 3 much larger spines on flexor margin. P 1: [2.3] (male) × carapace length; subcylindrical, spiny, with scattered long stiff setae and few thick setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus [0.9] length of carapace, [2.0] × as long as carpus. Carpus [2.1] × as long as wide. Palm [1.1] × carpus length, [1.9] × as long as broad. Fingers [0.9] × palm length; unarmed. P 2 – 4: Subcylindrical, moderately setose and spinose. Meri successively shorter posteriorly: P 3 merus [0.8] × length of P 2 merus, P 4 merus [0.7] × length of P 3 merus. P 2 merus, [0.7] – 0.8 × carapace length, [4.9] – 5.7 × as long as broad, [1.2] – 1.4 × as long as P 2 propodus; P 3 merus [4.4] × as long as broad, [1.3] × as long as P 3 propodus; P 4 merus [3.3] × as long as broad, [1.0] × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, with small distal spine; flexor margins of P 2 – 4 serrated, each with distal spine; P 4 lateral surface with median row of 3 small spines. Carpi with 2 or 3 spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, smaller on P 4; row of small spines below extensor margin on lateral surface of P 2 – 3, unarmed on P 4; flexor margins P 2 – 4 irregular, each with distal spine. Propodi stout, [4.4 – 4.8] 4.0 – 5.0 × as long as broad; extensor margin serrated, unarmed; flexor margin with 2 – 4 slender movable spines in addition to distal pair. Dactyli 0.7 – 0.8 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 5 movable spines. Eggs. Ov. F (MNHN-IU- 2016 - 495) carried 5 eggs of 0.3 mm diameter. Live colour. Unknown. Genetic data. No data.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFC6E63D4F9CFC897284BC12.taxon	distribution	Distribution. Indian Ocean, Reunion Island, probably Red Sea, at 73 – 77 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFC6E63D4F9CFC897284BC12.taxon	discussion	Remarks. The specimens identified and illustrated by Tirmizi (1966) as P. pusillus are very similar to the new species, P. hylas, although they are different from P. pusillus (see also Lewinshon 1969; Schnabel & Ahyong 2019). Unfortunately, we have not examined this material and the status of these specimens should be resolved in future studies. Phylladiorhynchus hylas has 3 spines on the epigastric ridge and closely resembles P. koumac, from New Caledonia, and P. pulchrus, from French Polynesia, Phillipines, and Vanuatu based on the 3 spines on the anterior branchial margin, gastric region strongly convex, the leaf-like rostrum (margins clearly convex and subapical spines present), and elevated ridges upraised dorsally. However, they can be easily differentiated by the number of epigastric spines, 3 spines in P. hylas and 5 spines in the other species. The specimens of P. hylas range from 1.6 to 1.9 mm postorbital carapace length.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFC5E6224F9CFAE772DEBE6E.taxon	description	(Fig. 17)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFC5E6224F9CFAE772DEBE6E.taxon	materials_examined	Type material. Holotype. French Polynesia. TARASOC Stn DW 3393, 15.8200 ° S, 148.2800 ° W, 800 m, 7 October 2009: F 2.5 mm (MNHN-IU- 2014 - 13842).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFC5E6224F9CFAE772DEBE6E.taxon	etymology	Etymology. From the name Idas, an Argonaut, son of Aphareus and Arene. The name is considered a substantive in apposition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFC5E6224F9CFAE772DEBE6E.taxon	description	Description. Carapace: Slightly longer than broad; transverse ridges with short setae. Gastric region flattened, with 4 transverse ridges: epigastric ridge indistinct, with 3 spines (1 median and 2 lateral spines) and some outer granules; anterior protogastric ridge medially interrupted, nearly extending laterally to carapace margin; anterior mesogastric ridge medially interrupted, laterally interrupted by anterior branch of cervical groove; anterior metagastric ridge indistinct, with few small scales in the anterior metagastric area. Mid-transverse ridge not interrupted, slightly medially depressed, preceded by shallow or indistinct cervical groove, followed by 2 uninterrupted or minutely interrupted ridges, interspersed with 1 short lateral ridge. Lateral margins slightly convex, with 6 spines: first anterolateral spine well-developed, reaching end of lateral orbital spine, second spine (hepatic) minute but distinct, slightly dorsomesially from lateral margin, and followed by 4 branchial spines (2 anterior spines, first well-developed, followed by a second obsolescent to undistinct spine, and 2 posterior spines). Rostrum triangular, horizontal, dorsally flattish or slightly concave, [1.1] × as long as broad, length [0.3] and breadth [0.3] that of carapace; lateral margins smooth and slightly concave proximally, with well-developed supraocular spines, subapical spines absent. Pterygostomian flap ending in round, upper margin smooth. Thoracic sternum: As wide as long. Sternite 3 broad, [3.0] × as wide as long, anterior margin convex, with a median feeble excavation, moderately produced anterolaterally. Sternite 4 widely contiguous to sternite 3; surface not depressed in midline, smooth; greatest width [2.5] × that of sternite 3, [2.8] × as wide as long. Pleon: Elevated ridges with short setae and with few scattered long setae. Tergite 2 with anterior and posterior transverse elevated ridges; tergites 3 and 4 with anterior transverse ridge; tergites 5 – 6 smooth. Eye: Eyestalk length about as broad as long, peduncle distally setose, not distinctly expanded proximally; maximum corneal diameter [1.4] × rostrum width, as wide as eyestalk. Antennule: Article 1 more than 1.5 × longer than wide, with 5 spines: distomesial spine well-developed; proximal lateral spine small. Antenna: Article 1 with prominent mesial process, nearly reaching lateralmost antennular spine end. Article 2 with distinct distomesial and distolateral spines. Articles 3 and 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus [0.7] × length of ischium at middle length, with well-developed distal spine on extensor margin and 2 strong spines on flexor margin (median spine distinctly curved). P 1: [3.8] × carapace length; subcylindrical, spiny and with several plumose setae on merus, and carpus and long stiff setae on palm and fingers; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus [1.5] length of carapace, [2.4] × as long as carpus. Carpus [3.2] × as long as wide. Palm [1.2] × carpus length, [3.2] × as long as broad. Fingers subequal in length to palm; fixed finger unarmed; movable finger with well-developed basal spine. P 2 – 4: Slender, subcylindrical, moderately setose and spinose. Meri successively shorter posteriorly: P 3 merus [0.6] × length of P 2 merus, P 4 merus [0.9] × length of P 3 merus. P 2 merus, as long as carapace length, [9.2] × as long as broad, [1.4] × as long as P 2 propodus; P 3 merus [8.3] × as long as broad, [1.2] × as long as P 3 propodus; P 4 merus [7] × as long as broad, [1.1] × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, with distal spine; flexor margins of P 2 – 4 irregular, each with distal spine; P 4 lateral surface with median row of 4 small spines, absent in others. Carpi armed on extensor margin with 1 spine on P 2, unarmed on P 3 – 4, distal spine prominent on P 2 – 4. Propodi sharply slender, [10.0 – 10.1] × as long as broad; extensor margin irregular, usually unarmed; flexor margin with 2 – 4 slender movable spines in addition to distal pair. Dactyli [0.5] × length of propodus, ending in incurved, strong, sharp spine; flexor margin with 5 – 6 movable spines. Eggs. No data. Live colour. Unknown. Genetic data. No data.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFC5E6224F9CFAE772DEBE6E.taxon	distribution	Distribution. French Polynesia, 800 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFC5E6224F9CFAE772DEBE6E.taxon	discussion	Remarks. Phylladiorhynchus idas resembles P. kermadecensis Schnabel & Ahyong, 2019, from New Zealand and P. punctatus from New Caledonia. However, the species can be distinguished by the following characters: - The carapace has 5 epigastric spines in P. kermadecensis and P. punctatus, whereas there are 3 spines and a few granules in P. idas. - The anterior protogastric ridge is not medially interrupted or with a small median gap in P. kermadecensis and P. punctatus, whereas the anterior protogastric ridge is medially interrupted in P. idas. - The anterior metagastric ridge is medially interrupted in P. kermadecensis whereas this ridge is scale-like in P. idas. - Posterior branchial margin with one spine in P. kermadecensis, whereas there are 2 spines in P. idas and P. punctatus. - The extensor margin of the P 2 – 3 propodus is usually armed with 2 proximal spines in P. kermadecensis and P. punctatus, whereas these margins are usually unarmed in P. idas. - The flexor margin of the P 2 – 4 dactyli have 10 – 11 movable spines in P. kermadecensis, whereas there are 5 – 6 movable spines in P. idas and 6 – 7 in P. punctatus.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD8E6204F9CFF297542BD36.taxon	description	Records requiring verification:	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD8E6204F9CFF297542BD36.taxon	diagnosis	Diagnosis. Rostrum triangular, subapical spines absent or, if present, obsolescent, lateral margins straight. Carapace with epigastric ridge armed with 5 spines; anterior branchial margin with 3 spines; anterior protogastric ridge not interrupted medially, without scale behind median epigastric spine; anterior mesogastric and metagastric ridges not medially interrupted. Pleonal tergite 3 without posterior transverse ridge. Anterior margin of thoracic sternite 3 moderately convex sometimes with feeble median excavation Antennule article 1 with 4 distal spines (double distolateral spine absent). Antennal article 2 with distolateral spine longer than distomesial; article 3 with distinct distomesial spine. Flexor margin of Mxp 3 merus with 2 prominent spines. P 2 – 4 dactylus extensor margin without upright spines at bases of movable spines.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD8E6204F9CFF297542BD36.taxon	distribution	Distribution. Only known from the type locality, Bonin Islands, Japan.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD8E6204F9CFF297542BD36.taxon	discussion	Remarks. Pending the verification of some previous records, P. ikedai is only known from the male holotype. The species is characterized by the presence of 5 epigastric spines, 1 hepatic spine, 3 anterior branchial spines, the anterior metagastric ridge not medially interrupted and the antennular article 1 with 4 spines only. The existence of some closely related species, e. g. P. argus, P. butes, P. eneus, etc., suggests that some occurrences of P. ikedai remain to be confirmed. Unfortunately, we have not studied the specimens from Maldives (Baba 1977), Japan (Fukuoka) (Baba 2005) and South China Sea (Dong & Li 2013), recommending further revision to confirm their identity.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD8E6274F9CFB217445B85E.taxon	description	(Figs. 21 E, 37 A – B)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD8E6274F9CFB217445B85E.taxon	materials_examined	Material examined. Hawaiian Islands, French Frigate Shoals. 27.765 ° N, 166.169 ° W, 32 m, 17 October 2006: 1 M 2.1 mm, 2 F 1.7 – 2.0 mm (UF 13155). — 23.7695 ° N, 166.2608 ° W, 10 m, 21 October 2006: 1 ov. F 2.4 mm (UF 12407). — Cory Pittman, October 2006: 1 ov. F 1.7 mm (UF 13148). — Stn FFS- 1121, 23.63835 ° N, 166.18005 ° W, 12 m, 5 September 2013: 1 M 1.7 mm (UF 54390). — Stn FFS- 1123, 23.63835 ° N, 166.18005 ° W, 12 m, 5 September 2013: 1 F 1.6 mm (UF 54392). — Stn FFS- 288, 23.63835 ° N, 166.18005 ° W, 12 m, 5 September 2013: 1 M 1.9 mm, 1 F 1.5 mm (UF 54396). — Stn FFS- 353, 23.63835 ° N, 166.18005 ° W, 12 m, 5 September 2013: 1 ov. F 1.9 mm (UF 54393). — Stn FFS- 354, 23.63835 ° N, 166.18005 ° W, 12 m, 5 September 2013: 1 M 2.0 mm (UF 54391). — Stn FFS- 438, 23.63835 ° N, 166.18005 ° W, 12 m, 5 September 2013: 2 M 1.2 – 1.6 mm, 1 ov. F 1.8 mm (UF 54389). — Stn FFS- 440, 23.63835 ° N, 166.18005 ° W, 12 m, 05 September 2013: 1 M 1.4 mm (UF 54395). — Stn FFS- 479, 23.62792 ° N, 166.13538 ° W, 10 m, 5 September 2013: 1 F 2.0 mm (UF 54397). — Stn FFS- 559, 23.62792 ° N, 166.13538 ° W, 10 m, 5 September 2013: 2 F 1.5 – 1.6 mm (UF 54394). — Stn FFS- 692, 23.7906 ° N, 166.2320 ° W, 9 m, 6 September 2013: 2 M 2.2 – 2.3 mm (UF 54339). — Stn FFS- 693 23.7906 ° N, 166.232 ° W, 9 m, 6 September 2013: 1 M 2.0 mm (UF 54337). — Stn FFS- 735, 23.8061 ° N, 166.2306 ° W, 8 m, 6 September 2013: 1 ov. F 2.3 mm (UF 54340). — Stn FFS- 736: 23.8061 ° N, 166.2306 ° W, 8 m, 6 September 2013: 2 F 1.4 – 1.9 mm (UF 54348). — Stn FFS- 737, 23.8061 ° N, 166.2306 ° W, 8 m, 6 September 2013: 1 ov. F 2.2 mm (UF 54341). — Stn FFS- 788, 23.8061 ° N, 166.2306 ° W, 8 m, 6 September 2013: 1 M 1.7 mm, 2 F 1.6 – 2.1 mm (UF 54344). — Stn FFS- 789, 23.8061 ° N, 166.2306 ° W, 8 m, 6 September 2013: 1 specimen broken (UF 54338). — Stn FFS- 821, 23.8061 ° N, 166.2306 ° W, 8 m, 6 September 2013: 1 M 1.9 mm, 1 ov. F 2.1 mm, 2 F 1.7 – 2.0 mm (UF 54345). — Stn FFS- 822, 23.8061 ° N, 166,2306 ° W, 8 m, 6 September 2013: 1 M broken (UF 54347). — Stn FFS- 851, 23.8365 ° N, 166.2668 ° W, 9 m, 7 September 2013: 1 ov. F 2.2 mm (UF 54336). — Stn FFS- 882, 23.8365 ° N, 166.2668 ° W, 9 m, 7 September 2013: 1 ov. F 2.3 mm (UF 54342). — Stn FFS- 91923.8365 ° N, 166.2668 ° W, 9 m, 7 September 2013: 2 M 1.4 – 2.2 mm, 2 ov. F 1.9 – 2.3 mm (UF 54343). — Hawaii Island. Stn HAW- 1453, 20.26824 ° N, 155.86038 ° W, 13 m, 15 August 2016: 1 M 1.7 mm (UF 54335). — Stn HAW- 443, 19.03821667 ° N, 155.88255 ° W, 15 m, 4 August 2013: 2 M 1.8 – 2.0 mm, 3 ov. F 1.7 – 2.0 mm, 3 F 1.6 – 1.7 mm (UF 54360). — Stn HAW- 507, 19.03821667 ° N, 155.88255 ° W, 15 m, 4 August 2013: 7 M 1.6 – 2.3 mm, 1 ov. F 1.8 mm, 2 F 1.7 – 1.9 mm (UF 54357). — Stn HAW- 509, 19.03821667 ° N, 155.88255 ° W, 15 m, 4 August 2013: 2 ov. F 2.2 – 2.4 mm, 2 F 1.4 – 1.5 mm (UF 54358). — Stn HAW- 510, 19.03821667 ° N, 155.88255 ° W, 15 m, 4 August 2013: 1 ov. F 2.0 mm, 2 F 1.8 – 2.1 mm (UF 54359). — Stn HAW- 593, 19.03821667 ° N, 155.88255 ° W, 15 m, 4 August 2013: 3 M 1.6 – 2.2 mm, 3 F 1.5 – 1.8 mm (UF 54363). — Stn HAW- 595, 19.03821667 ° N, 155.88255 ° W, 15 m, 4 August 2013: 1 M 1.6 mm, 1 F 1.8 mm (UF 54362). — Stn HAW- 596, 19.03821667 ° N, 155.88255 ° W, 15 m, 4 August 2013: 1 M 1.1 mm, 1 F 1.7 mm (UF 54356). — Stn HAW- 692, 18.96859682 ° N, 155.730732 ° W, 15 m, 7 August 2013: 6 M 1.4 – 2.0 mm, 4 ov. F 1.2 – 1.8 mm, 5 F 1.0 – 1.9 mm (UF 54364). — Stn HAW- 740, 18.96859682 ° N, 155.730732 ° W, 15 m, 7 August 2013: 1 M 2.0 mm, 1 F 1.4 mm (UF 54367). — Stn HAW- 743, 18.96859682 ° N, 155.730732 ° W, 15 m, 7 August 2013: 2 M 1.2 – 2.1 mm, 2 ov. F 2.0 – 2.2 mm, 2 F 1.6 – 1.8 mm (UF 54366). — Stn HAW- 803, 18.96859682 ° N, 155.730732 ° W, 15 m, 7 August 2013: 3 M 1.8 – 2.2 mm, 3 ov. F 1.8 – 2.1 mm, 2 F 1.7 – 1.8 mm (UF 54368). — Stn HAW- 805, 18.96859682 ° N, 155.730732 ° W, 15 m, 7 August 2013: 1 F 1.4 mm (UF 54365). — Stn HAW- 978, 18.96859682 ° N, 155.730732 ° W, 15 m, 7 August 2013: 1 M 1.9 mm (UF 54369). — Stn HAW- 980, 19.03821667 ° N, 155.88255 ° W, 15 m, 4 August 2013: 3 M 1.8 – 2.1 mm, 2 ov. F 1.7 – 2.0 mm, 2 F 1.4 – 1.5 mm, 1 larvae (UF 54361). — Stn HAW- 981, 19.03821667 ° N, 155.88255 ° W, 15 m, 4 August 2013: 1 F 1.2 mm (UF 54355). — Kauai Island. Stn KAU- 526, 22.16405324 ° N, 159.298801 ° W, 14 m, 13 August 2013: 2 M 1.6 – 2.1 mm, 1 F 2.3 mm (UF 54372). — Stn KAU- 622, 22.16405324 ° N, 159.298801 ° W, 14 m, 13 August 2013: 1 M 1.8 mm (UF 54373). — Stn KAU- 623, 22.16405324 ° N, 159.298801 ° W, 14 m, 13 August 2013: 2 M 1.2 – 2.0 mm, 2 ov. F 1.6 – 2.0 mm, 1 F 1.9 mm (UF 54375). — Stn KAU- 625, 22.16405324 ° N, 159.298801 ° W, 14 m, 13 August 2013: 2 M 1.6 – 2.1 mm, 2 ov. F 1.9 – 2.0 mm (UF 54374). — Stn KAU- 652, 22.16684307 ° N, 159.6800101 ° W, 13 m, 15 August 2013: 2 M 1.5 – 1.8 mm, 1 ov. F 1.9 mm (UF 54377). — Stn KAU- 772, 22.16684307 ° N, 159.6800101 ° W, 13 m, 15 August 2013: 1 F 1.1 mm (UF 54378). — Stn KAU- 774, 22.16684307 ° N, 159.6800101 ° W, 13 m, 15 August 2013: 1 M 1.9 mm, 3 F 1.1 – 1.9 mm (UF 54376). — Stn KAU- 831, 21.8897599 ° N, 159.6088704 ° W, 12 m, 17 August 2013: 1 M 1.5 mm, 1 ov. F 1.2 mm, 2 F 1.6 – 1.8 mm, 1 postlarvae 1.0 mm (UF 54385). — Stn KAU- 832, 21.8897599 ° N, 159.6088704 ° W, 12 m, 17 August 2013: 6 M 1.2 – 1.9 mm, 1 ov. F 1.9 mm (UF 54381). — Stn KAU- 835, 21.8897599 ° N, 159,6088704 ° W, 12 m, 17 August 2013: 1 M 1.8 mm (UF 54384). — Stn KAU- 922, 21.8897599 ° N, 159.6088704 ° W, 12 m, 17 August 2013: 5 M 1.6 – 1.9 mm, 1 ov. F 1.9 mm, 2 F 1.3 – 2.0 mm, 2 postlarvae 1.0 mm (UF 54382). — Stn KAU- 923, 21.8897599 ° N, 159.6088704 ° W, 12 m, 17 August 2013: 1 M 1.8 mm, 1 ov. F 1.9 mm, 1 F 1.6 mm (UF 54383). — Stn KAU- 924, 21.8897599 ° N, 159.6088704 ° W, 12 m, 17 August 2013: 1 M 1.4 mm (UF 54379). — Stn KAU- 944, 21.8897599 ° N, 159.6088704 ° W, 12 m, 17 August 2013: 1 F 1.6 mm (UF 54380). — Kure Atoll. Stn KUR- 301, 28.416767 ° N, 178.378433 ° W, 14 m, 14 July 2013: 4 M 1.7 – 2.0 mm (UF 54544). — Stn KUR- 301, 28.416767 ° N, 178.378433 ° W, 14 m, 14 July 2013: 1 ov. F 2.1 mm (UF 54352). — Stn KUR- 348, 28.38230758 ° N, 178.3244794 ° W, 10 m, 14 July 2013: 12 M 1.2 – 2.4 mm, 3 ov. F 1.9 – 2.2 mm, 9 F 1.1 – 2.2 mm (UF 54353). — Stn KUR- 476, 28.38230758 ° N, 178.3244794 ° W, 10 m, 14 July 2013: 6 M 1.5 – 2.3 mm, 3 ov. F 1.5 – 2.3 mm, 2 F 1.8 – 2.2 mm (UF 54354). — Lisianski Island. Stn LIS- 212, 26.07841509 ° N, 173.9970114 ° W, 15 m, 12 September 2013: 1 F 1.8 mm (UF 54413). — Stn LIS- 213, 26.07841509 ° N, 173.9970114 ° W, 15 m, 12 September 2013: 1 M 1.7 mm (UF 54415). — Stn LIS- 239, 25.94462 ° N, 173.95361 ° W, 15 m, 13 September 2013: 2 M 1.7 – 2.0 mm, 1 F 1.8 mm (UF 54421). — Stn LIS- 260, 26.07841509 ° N, 173.9970114 ° W, 15 m, 12 September 2013: 1 M 1.8 mm, 1 ov. F 1.9 mm (UF 54417). — Stn LIS- 301, 26.07841509 ° N, 173.9970114 ° W, 15 m, 12 September 2013: 1 ov. F 2.3 mm, 1 F 2.2 mm (UF 54416). — Stn LIS- 302, 26.07841509 ° N, 173.9970114 ° W, 15 m, 12 September 2013: 1 M 1.7 mm, 2 ov. F 1.6 – 2.0 mm, 1 F 2.0 mm (UF 54414). — Stn LIS- 326, 25.94462 ° N, 173.95361 ° W, 15 m, 13 September 2013: 3 M 1.8 – 2.3 mm, 1 ov. F 1.4 mm, 1 F 1.4 mm (UF 54418). — Stn LIS- 336, 25.94462 ° N, 173.95361 ° W, 15 m, 13 September 2013: 2 M 1.3 – 1.8 mm, 1 F 1.4 mm (UF 54419). — Stn LIS- 340, 25.94462 ° N, 173.95361 ° W, 15 m, 13 September 2013: 1 ov. F 2.1 mm, 1 F 2.2 mm (UF 54420). — Maui Island. Airport Beach, Halimeda bed, 6 – 17 m, 15 March 2009: 1 M 1.9 mm (UF 20156). — Stn MAI- 196, 20.76226207 ° N, 155.9797715 ° W, 12 m, 9 August 2013: 1 M 2.2 mm (UF 54371). — Stn MAI- 236, 20.76226207 ° N, 155.9797715 ° W, 12 m, 9 August 2013: 1 M 1.6 mm, 1 F 1.7 mm (UF 54370). — Oahu Island. 21.289 ° N, 157.865 ° W, 9 – 12 m, 1 December 2008: 1 M 1.6 mm (UF 15285), 1 ov. F 1.8 mm (UF 15269), 1 ov. F 2.5 mm (UF 15270). — Stn OAH- 420, 21.41212947 ° N, 157.7097973 ° W, 15 m, 23 August 2013: 2 M 1.5 – 1.9 mm, 3 F 1.2 - 1.4 mm, 1 postlarvae 1.0 mm (UF 54388). — Stn OAH- 448, 21.41212947 ° N, 157.7097973 ° W, 15 m, 23 August 2013: 1 F 1.8 mm (UF 54386). — Stn OAH- 516, 21.41212947 ° N, 157.7097973 ° W, 15 m, 23 August 2013: 2 M 1.4 – 1.8 mm, 2 ov. F 2.0 – 2.2 mm, 2 F 1.8 – 2.0 mm (UF 54387). — Stn OAH- 662: 2 M 1.8 – 2.2 mm, 1 F 1.4 mm (UF 54349). — Stn OAH- 738: 1 M 2.2 mm, 1 F 1.4 mm (UF 54350). — Stn OAH- 947: 1 M 1.7 mm (UF 54351). — Pearl and Hermes Atoll. Stn PHR- 1019, 27.785833 ° N, 175.780283 ° W, 14 m, 10 September 2013: 2 F 1.2 – 1.3 mm (UF 54409). — Stn PHR- 1020, 27.785833 ° N, 175.780283 ° W, 14 m, 10 September 2013: 1 M 1.6 mm, 1 ov. F 1.4 mm, 1 F 1.2 mm (UF 54403). — Stn PHR- 1036, 27.78543745 ° N, 175.8235427 ° W, 14 m, 10 September 2013: 2 ov. F 1.4 – 1.5 mm, 2 F 1.3 – 1.4 mm (UF 54406). — Stn PHR- 1079, 27.78543745 ° N, 175.8235427 ° W, 14 m, 10 September 2013: 3 M 1.8 – 2.0 mm (UF 54407). — Stn PHR- 1080, 27.78543745 ° N, 175.8235427 ° W, 14 m, 10 September 2013: 4 M 1.1 – 1.3 mm, 2 F 1.0 – 1.2 mm (UF 54398). — Stn PHR- 1252, 27.785833 ° N, 175.780283 ° W, 14 m, 10 September 2013: 1 F 1.0 mm (UF 54400). — Stn PHR- 754, 27.785833 ° N, 175.780283 ° W, 14 m, 10 September 2013: 2 F 1.4 – 1.5 mm (UF 54402). — Stn PHR- 857, 27.753133 ° N, 175.948767 ° W, 15 m, 11 September 2013: 2 M 1.4 – 1.9 mm, 1 F 1.5 mm (UF 54412). — Stn PHR- 890, 27.785833 ° N, 175.780283 ° W, 14 m, 10 September 2013: 1 M 1.6 mm (UF 54401). — Stn PHR- 892, 27.785833 ° N, 175.780283 ° W, 14 m, 10 September 2013: 4 M 1.6 – 1.8 mm, 5 F 1.3 – 1.6 mm (UF 54404). — Stn PHR- 936, 27.78543745 ° N, 175.8235427 ° W, 14 m, 10 September 2013: 1 F 1.5 mm (UF 54408). — Stn PHR- 937, 27.78543745 ° N, 175.5958235427 ° W, 14 m, 10 September 2013: 1 M 2.0 mm (UF 54405). — Stn PHR- 938, 27.78543745 ° N, 175.8235427 ° W, 14 m, 10 September 2013: 4 M 1.1 – 1.8 mm, 1 ov. F 1.8 mm, 2 F 1.4 – 1.6 mm (UF 54399).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD8E6274F9CFB217445B85E.taxon	diagnosis	Diagnosis (modified from Schnabel & Ahyong 2019). Rostrum margin proximally straight (bottle-shaped); subapical spines present. Carapace ridges clearly distinct, with dense short setae; epigastric region always with 2 spines; parahepatic spines on protogastric region absent; anterior branchial margin with 2 spines; hepatic spine absent; anterior metagastric ridge medially interrupted. Pleonal tergite 3 without posterior transverse ridge. Thoracic sternite 3 anterior margin biconcave, with obtuse median projection. Antennular article 1 with 5 well-developed spines: distomesial spine large; second lateral spine smallest but always distinct. Antennal article 1 distomesial spine overreaching peduncle but not reaching second lateral antennular spine; article 2 distal spines subequal, lateral spine reaching midlength of article 3; article 3 with small mesial spine distally. Flexor margin of Mxp 3 merus with one prominent spine only. P 2 – 4 dactylus extensor margin without upright spines at bases of movable spines. Eggs: Ov. F carried approximately 10 – 25 eggs of 0.4 – 0.5 mm diameter. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD8E6274F9CFB217445B85E.taxon	distribution	Distribution. Hawaiian Islands, between 0 and 32 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD8E6274F9CFB217445B85E.taxon	discussion	Remarks. The species was re-described by Schnabel & Ahyong (2019) who selected a neotype from the Kure Atoll, Hawaii. Phylladiorhynchus integrirostris belongs to the group of species having the epigastric ridge armed with 2 spines and the anterior branchial margin of the carapace armed with two spines only. The species is geographically restricted to the Hawaiian Islands (Schnabel & Ahyong 2019). However, P. integrirostris closely resembles P. lynceus from Chagos Archipelago, Kiribati and Samoa, and P. priasus from Northern Marianas Islands, Guam and Wake Islands. The three species are barely distinguishable morphologically although they are clearly different genetically (see the differences under the Remarks of P. priasus). Most specimens of P. integrirostris and P. priasus present an epigastric scale with plumose setae although, since the setae can be broken and so be unnoticed, this character should be taken cautiously. Furthermore, these three species are also close to P. orpheus (see the differences under the Remarks of P. orpheus). Phylladiorhynchus serrirostris (Melin, 1939), only known from Bonin Islands, has been also considered a synonym of P. integrirostris; however, both are different and valid species (see the differences under the Remarks of P. serrirostris). The specimens of P. integrirostris range from 1.1 to 2.3 mm postorbital carapace length. The 10 and 11 sequences of P. integrirostris from Hawaii diverged 0.3 % for COI and 0.1 % for 16 S, respectively.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFDFE62B4F9CFE39717EB8B2.taxon	description	(Figs. 18, 19, 21 F)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFDFE62B4F9CFE39717EB8B2.taxon	description	Is., Korea, 150 m).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFDFE62B4F9CFE39717EB8B2.taxon	materials_examined	Type material. Lectotype. Japan. Honshu, 110 – 128 m, ov. F 3.6 mm (USNM 26168). Paralectotypes. station data as lectotype, 17 M 3.0 – 4.8 mm, 11 ov. F 2.8 – 3.7 mm (USNM 26168). Other material. Taiwan. Aodi, Taipei County. Stn CP 58, 24.5850 ° N, 122.0966 ° E, 221 – 254 m, 4 August 2000: 3 M 2.7 – 3.2 mm, 1 ov. F 3.5 mm (NTOU). — Stn CP 76, 24.9423 ° N, 122.0251 ° E, 115 – 170 m, 7 May 2001: 1 F 1.8 mm (NTOU). — Stn CP 85, 24.0092 ° N, 122.0090 ° E, 255 – 390 m, 9 May 2001: 6 M 2.2 – 4.0 mm (NTOU). — Stn DW 149, 22.3083 ° N, 121.4895 ° E, 258 m, 20 May 2002: 1 M 2.0 mm (NTOU). — Stn CP 277, 24.3928 ° N, 122.2353 ° E, 1222 – 1261 m, 14 June 2005: 1 M 3.8 mm (NTOU). — Stn OCP 287, 24.9587 ° N, 122.0884 ° E, 259 – 349 m, 8 August 2005: 11 M 2.1 – 2.6 mm, 1 ov. F 2.6 mm, 8 F 1.5 – 2.7 mm (NTOU). — Stn OCP 288, 24.9617 ° N, 122.0891 ° E, 263 – 352 m, 8 August 2005: 27 M 1.1 – 4.2 mm, 2 ov. F 2.4 – 2.6 mm, 13 F 2.1 – 3.0 mm (NTOU). — Stn OCP 293, 24.9728 ° N, 122.0882 ° E, 262 – 232 m, 8 August 2005: 28 M 1.5 – 3.4 mm, 3 ov. F 2.4 – 2.7 mm, 25 F 1.4 – 3.4 mm (NTOU). Indonesia. KARUBAR Stn CP 05, 05.8167 ° S, 132.3000 ° E, 296 – 299 m, 22 October 1991: 1 F 3.1 mm (MNHN- IU- 2016 - 9659). — Stn DW 13, 05.4333 ° S, 132.6333 ° E, 417 – 425 m, 24 October 1991: 1 F 4.8 mm (MNHN-IU- 2014 - 13804), 1 F broken (MNHN-IU- 2014 - 13854). — Stn CP 16, 05.2833 ° S, 132.8333 ° E, 315 – 349 m, 24 October 1991: 1 M 4.2 mm (MNHN-IU- 2014 - 13852), 1 ov. F 3.5 mm (MNHN-IU- 2016 - 426). — Stn CP 25, 05.5000 ° S, 132.8667 ° E, 336 – 346 m, 26 October 1991: 1 F 2.9 mm (MNHN-IU- 2014 - 13856). — Stn CP 36, 06.0833 ° S, 132.7333 ° E, 210 – 268 m, 27 October 1991: 1 ov. F 3.2 mm (MNHN-IU- 2014 - 13853). Chesterfield Islands. EBISCO Stn DW 2603, 19.6294 ° ’ S, 158.7316 ° E, 568 – 570 m, 18 October 2005: 1 ov. F 4.1 mm (MNHN-IU- 2014 - 13855). — Stn DW 2603, 19.6294 ° S, 158.7316 ° E, 568 – 570 m, 18 October 2005: 1 ov. F 3.5 mm (MNHN-IU- 2016 - 425). — KANADEEP Stn DW 4975, 19.7500 ° S, 158.5833 ° E, 386 – 428 m, 9 September 2017: 1 ov. F 3.3 mm (MNHN-IU- 2017 - 2432).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFDFE62B4F9CFE39717EB8B2.taxon	description	Description. Carapace: as long as or slightly broader than long; transverse ridges with dense short setae and few scattered long thick iridescent setae. Gastric region slightly convex, with some transverse ridges: epigastric ridge distinct usually armed with 4 epigastric spines, inner pair longer, often a median scale with thick or plumose setae anterior to epigastric ridge; anterior protogastric ridge not medially interrupted, nearly extending laterally to carapace margin; anterior mesogastric ridge not medially interrupted, laterally interrupted by cervical groove, laterally continuing to first branchial spine; anterior metagastric ridge not medially interrupted, laterally interrupted; posterior epigastric, protogatric, mesogastric, and metagastric ridges scale-like. Mid-transverse ridge uninterrupted, medially slightly depressed, preceded by shallow or indistinct cervical groove, followed by 3 uninterrupted or minutely interrupted ridges, interspersed with 2 short lateral ridge and sometimes few, short scattered scales. Lateral margins slightly convex, with 7 – 8 spines: first anterolateral spine well-developed, reaching anteriorly to level of lateral orbital spine, second spine (hepatic) well-developed, slightly dorsomesially from lateral margin, and followed by 5 – 6 branchial spines (3 anterior and 2 – 3 posterior). Rostrum leaf-like, horizontal, dorsally flattish or slightly concave, [1.6] – 1.9 × as long as broad, length [0.4] – 0.5 and breadth 0.2 – [0.3] that of carapace; lateral margins smooth and convex, with well-developed supraocular basal spines and small subapical spines. Pterygostomian flap ending in blunt tooth, upper margin smooth. Thoracic sternum: As wide as long. Sternite 3 moderately broad, [2.0] – 2.5 × as wide as long, anterior margin with obtuse median projection. Sternite 4 widely contiguous to sternite 3; surface depressed in midline, smooth; greatest width 2.5 – [3.4] × that of sternite 3, 2.6 – [2.8] × as wide as long. Pleon: Elevated ridges with dense short setae and scattered long setae. Tergites 2 – 3 with anterior and posterior transverse elevated ridges; tergite 4 with anterior transverse ridge; tergites 5 – 6 smooth. Eye: Eyestalk length about 0.9 × broader than long, peduncle distally setose, not distinctly expanded proximally, with few short transverse striae on lateral surfaces; maximum corneal diameter 1.2 × rostrum width, as wide as eyestalk (0.95 × maximum peduncle width). Antennule: Article 1 1.3 × longer than wide, with 4 well-developed distal spines: distomesial spine welldeveloped; proximal lateral spine usually absent, minute if present. Antenna: Article 1 with prominent mesial process, distally falling short of lateral antennular spine. Article 2 with minute distomesial spine, distolateral spine larger than distomesial. Article 3 with minute or obsolescent distal mesial spine. Article 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus [0.6] – 0.7 × length of ischium, with well-developed distal spine on extensor and flexor margins. P 1 (lost in lectotype): 3.1 (males), 2.9 – 3.3 (females) × carapace length; subcylindrical, spiny and with scattered long stiff setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus 1.1 – 1.7 length of carapace, 2.1 – 3.2 × as long as carpus. Carpus 1.5 – 2.7 × as long as wide. Palm 1.2 × carpus length, 1.7 – 2.5 × as long as broad. Fingers 0.8 – 0.9 × palm length; fixed finger lateral margin unarmed; movable finger with small to well-developed basal spine. P 2 – 4 (P 2 in lectotype only): Slender, subcylindrical, moderately setose and spinose. Meri successively shorter posteriorly: P 3 merus 0.4 – 0.6 × length of P 2 merus, P 4 merus 0.8 × length of P 3 merus. P 2 merus, 0.6 – [0.9] × carapace length, [4.8] – 6.5 × as long as broad, 1.2 – 1.3 × as long as P 2 propodus; P 3 merus 4.8 – 6.3 × as long as broad, 1.1 – 1.2 × as long as P 3 propodus; P 4 merus 4.4 × as long as broad, as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, distal spine present; flexor margin irregular, with distal spine on P 2 – 3, absent in P 4; P 2 – 3 lateral surface unarmed. Carpi with 2 – 6 spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, smaller or absent on P 4; row granules below extensor margin on lateral surface of P 2 – 4; flexor margin unarmed. Propodi moderately slender, 5.0 – 6.4 × as long as broad; extensor margin irregular usually armed proximally with 3 small spines on P 2 – 3; flexor margin with 3 – 4 slender movable spines in addition to distal pair. Dactyli [0.5] – 0.6 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 6 – 8 movable spines. Eggs. Ov. F carried approximately 30 – 50 eggs of 0.4 – 0.5 mm diameter. Live colour. Carapace and pleon translucent white or diffuse orange red; carapace lateral spines and rostral supraocular basal spines orange red. P 1 diffuse orange-red. P 2 – 4 with diffuse, clear orange-red bandings (after Baba et al. 2009). Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFDFE62B4F9CFE39717EB8B2.taxon	distribution	Distribution. Japan, Korea, Taiwan, Indonesia, Chesterfield Islands, from 110 to 1261 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFDFE62B4F9CFE39717EB8B2.taxon	discussion	Remarks. Phylladiorhynchus integrus was described as Galathea integra by Benedict (1902) from specimens collected in Honshu, Japan. The species was previously collected in other Japanese localities (as G. integra) and from Taiwan, Korea and Eastern China Sea (as P. pusillus) (see above). The examination of the type specimens of the species indicates that it is a valid species, widely distributed in the western Pacific, and differentiated from other species of Phylladiorhynchus by several constant morphological differences. Phylladiorhynchus integrus belongs to the group of the species having 4 spines on the epigastric ridge, 1 well-developed hepatic spine, the anterior metagastric ridge not medially interrupted, the anterior margin of thoracic sternite 3 medially projected and a single spine on the flexor margin of Mxp 3 merus. The species closely resembles P. australis Schnabel & Ahyong, 2019 and P. nui Schnabel & Ahyong, 2019 from Eastern Australia and New Zealand. However, the three species can be distinguished among them by the following characters: - The rostrum lateral margin is straight in P. nui, whereas it is convex in P. australis and P. integrus. - The lateral-most spine of antennular article 1 is very small to indistinct in P. nui and P. integrus, whereas this spine is always distinct in P. australis. and 4. D, left cephalic region, showing antennular and antennal peduncles, ventral view. E, right Mxp 3, lateral view. F, right P 2, lateral view. Scale bar: A, E – H = 1.0 mm; B – D, I = 0.6 mm. - The antennal article 2 is armed with a well-developed distolateral spine, reaching or overreaching the end of article 3 in P. australis, whereas this spine is minute or obsolescent in P. nui and P. integrus. - The P 2 – 4 propodi are more slender in P. nui (6 × as long as broad) than in P. australis and P. integrus (5 × as long as broad). - The flexor margin of the P 2 – 4 dactyli has 9 – 11 movable spines in P. nui, having 6 – 9 spines in P. australis and P. integrus. The specimens of P. integrus range from 1.4 to 4.8 mm postorbital carapace length. Phylladiorhynchus integrus diverged 12 – 13 % (COI) and 5 – 6 % (16 S) from P. nui and P. australis, respectively. The four and three sequences of P. integrus from Indonesia, New Caledonia and Taiwan diverged 0.3 – 4.7 % for COI and 0 – 0 - 8 % for 16 S respectively.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD3E6284F9CFE4074EDBA0A.taxon	description	(Figs. 20, 21 G)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD3E6284F9CFE4074EDBA0A.taxon	materials_examined	Type material. Holotype. French Polynesia. BENTHAUS Stn DW 1894, 27.6688 ° S, 144.3585 ° W, 100 m, 8 November 2002: ov. F 2.3 mm (MNHN- IU- 2019 - 2662). Paratype. Vanuatu, SANTO, no station: 1 M broken (MNHN-IU- 2019 - 2651).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD3E6284F9CFE4074EDBA0A.taxon	etymology	Etymology. From the name Iphiclus, an Argonaut, son of Phylacus and Clymene. The name is considered a substantive in apposition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD3E6284F9CFE4074EDBA0A.taxon	description	Description. Carapace: as long as broad; transverse ridges scattered and indistinct, without setae. Gastric region flattened and smooth, transverse ridges barely distinct: epigastric ridge indistinct, with 2 median spines; 2 lateral scales on anterior protogastric region. Mid-transverse ridge not interrupted, medially depressed, preceded by distinct cervical groove, followed by 1 not medially interrupted ridge and 1 short lateral ridge. Lateral margins slightly convex, with 4 spines: first anterolateral spine well-developed, reaching end of lateral orbital spine, hepatic margin unarmed; anterolateral spine followed by 3 branchial spines (2 anterior and 1 posterior spines). Rostrum triangular, horizontal, dorsally flattish or slightly concave, 1.2 – [1.3] × as long as broad, length 0.3 and breadth 0.3 that of carapace; lateral margins smooth and straight, with well-developed supraocular spines, subapical spines absent. Pterygostomian flap ending in round, upper margin smooth. Thoracic sternum: As wide as long. Sternite 3 moderately broad, [2.2] – 2.5 × as wide as long, anterior margin with obtuse median projection, anterolaterally rounded. Sternite 4 widely contiguous to sternite 3; surface depressed in midline, smooth; greatest width 2.1 – [2.3] × that of sternite 3, 2.8 – [3.2] × as wide as long. Pleon: Transverse ridges with a few scattered short setae. Tergites 2 – 3 with anterior transverse ridge only; tergites 4 – 6 smooth. Eye: Eyestalk as broad as long, peduncle not setose, cornea very expanded distally, maximum corneal diameter 1.3 × rostrum width, [1.2] – 1.3 × maximum peduncle width. Antennule: Article 1 1.3 × longer than wide, with 4 spines: distomesial spine small, distolateral spines with a blunt process or granule (no double or very small distolateral spine); proximal lateral spine small, always present. Antenna: Article 1 with prominent mesial process distally clearly not reaching lateralmost antennular spine. Article 2 with distinct distal spines laterally and mesially. Article 3 with distomesial spine. Article 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus [0.7] × length of ischium, extensor margin unarmed, flexor margin with 3 spines, decreasing in size distally. P 1: 2.8 male, [2.5] (female) × carapace length; subcylindrical, spiny and with scattered long stiff setae and some thick iridescent setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus as long as carapace, 1.5 – [1.7] × as long as carpus. Carpus 1.8 – [2.5] × as long as wide. Palm 0.8 – [0.9] × carpus length, [1.5] – 1.6 × as long as broad. Fingers as long as palm length, fixed finger with distal spine on lateral margin; movable finger with 2 basal spines. P 2 – 4: Slender, subcylindrical, moderately setose and spinose. Meri successively shorter posteriorly: P 3 merus 0.8 – [0.9] × length of P 2 merus, P 4 merus [0.8] – 0.9 × length of P 3 merus. P 2 merus, [0.8] or as long as carapace length, [5.2] – 5.4 × as long as broad, 1.3 – [1.5] × as long as P 2 propodus; P 3 merus 5 – [5.3] × as long as broad, 1.1 – [1.6] × as long as P 3 propodus; P 4 merus 4 – [4.8] × as long as broad, 1.1 – [1.2] × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular but unarmed other than small distal spine; flexor margins of all legs irregular, with distal spine; P 4 lateral surface with median row of 2 small spines, absent in others. Carpi with 2 spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, smaller on P 4; flexor margin unarmed. Propodi slender, [5.0 – 5.5] 5.6 – 7.0 × as long as broad; extensor margin irregular, usually armed with a 1 – 2 proximal spines on P 2 – 4; flexor margin with 3 – 4 slender movable spines in addition to distal pair. Dactyli 0.6 – [0.7] × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 4 – 6 movable spines. Eggs: Ov. F (MNHN- IU- 2019 - 2662) carried 10 eggs of 0.4 mm diameter Live colour. Unknown. Genetic data. No data.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD3E6284F9CFE4074EDBA0A.taxon	distribution	Distribution. French Polynesia and Vanuatu, 100 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD3E6284F9CFE4074EDBA0A.taxon	discussion	Remarks. Phylladiorhynchus iphiclus is easily distiguished from the other species by having the Mxp 3 merus with 3 prominent spines along the flexor margin, the rostrum triangular and 2 spines on the epigastric ridge. Phylladiorhynchus iphiclus resembles to P. boucheti, from Chesterfield Islands, however, they can be distinguished by the following characters: - The epigastric ridge has 2 spines in P. iphiclus, whereas there are 4 spines in P. boucheti. - The gastric ridges are obsolescent or absent in P. iphiclus, whereas the protogastric and mesogastric ridges are distinct in P. boucheti. - The carapace anterior margin has 2 spines in P. iphiclus, whereas there are 3 spines in P. boucheti. - The flexor margin of Mxp 3 merus is armed with 3 spines, whereas there are 2 spines in P. boucheti. Furthermore, the extensor margin is unarmed in P. iphiclus, having a distal spine in P. boucheti.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD0E62F4F9CFCD87460B897.taxon	description	(Figs. 21 H, 22, 54 E) Records requiring verification:	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD0E62F4F9CFCD87460B897.taxon	materials_examined	Type material. Holotype. Madagascar, ATIMO VATAE Stn TB 01, 24.9966 ° S, 47.0950 ° E, 22 m, 30 April 2010: M 2.3 mm (MNHN-IU- 2014 - 13799). Paratypes. Madagascar. ATIMO VATAE Stn CP 3511, 25.2500 ° S, 47.2416 ° E, 97 – 98 m, 29 April 2010: 1 F 2.0 mm (MNHN-IU- 2010 - 2750). — Stn TB 02, 25.0216 ° S, 47.0083 ° E, 18 m, 1 May 2010: 1 M 2.7 mm (MNHN- IU- 2016 - 462). — Stn DW 3530, 24.5983 ° S, 47.5350 ° E, 80 – 86 m, 2 May 2010: 1 M 2.5 mm (MNHN-IU- 2016 - 485). — Stn CP 3546, 25.3783 ° S, 46.7083 ° E, 84 – 85 m, 4 May 2010: 1 M 2.2 mm (MNHN-IU- 2016 - 483), 1 M 1.7 mm (MNHN-IU- 2014 - 13800). — Stn CP 3579, 25.9083 ° S, 45.5533 ° E, 65 – 66 m, 9 May 2010: 1 M 2.1 mm (MNHN- IU- 2016 - 464). — Stn DW 3605, 25.9083 ° S, 44.8500 ° E, 56 – 57 m, 13 May 2010: 2 M 2.3 – 2.5 mm (MNHN-IU- 2016 - 484). — Stn CP 3624, 25.6350 ° S, 45.9500 ° E, 63 – 63 m, 15 May 2010: 1 M 1.8 mm (MNHN-IU- 2016 - 482). Other material. SW Indian Ocean. MD 08 Stn DC 32, 33.8333 ° S, 43.1833 ° E, 40 – 43 m, 15 March 1976: 1 ov. F 3.0 mm (MNHN-IU- 2016 - 467). — Stn CC 31 St 6, 33.1666 ° S, 43.8666 ° E, 36 – 47 m, 15 – 16 March 1976: 1 M 2.5 mm, 1 ov. F 2.4 mm (MNHN-IU- 2016 - 481). — Stn DC 33, 33.8333 ° S, 43.1833 ° E, 25 – 30 m, 16 March 1976: 1 M 2.8 mm (MNHN-IU- 2016 - 466). Reunion Island. MD 32 Stn CP 43, 21.3450 ° S, 55.4483 ° E, 73 – 77 m, 18 August 1982: 5 M 1.5 – 2.3 mm, 5 ov. F 1.8 – 2.3 mm, 4 F 1.5 – 1.8 mm (MNHN-IU- 2019 - 2696). SW Indian Ocean. Walters Shoal-MD 208 Stn WS 03, 33.2033 ° S, 43.8466 ° E, 40 m, 30 April 2017: 1 F 1.4 mm (MNHN-IU- 2016 - 455), 7 M 1.4 – 2.1 mm, 8 F 1.4 – 2.2 mm, 4 postlarvae 1.0 – 1.1 mm (MNHN-IU- 2016 - 459). — Stn WB 05, 33.2516 ° S, 43.9083 ° E, 26 – 30 m, 1 May 2017: 4 M 1.5 – 1.8 mm, 8 F 1.2 – 2.2 mm, 3 postlarvae 1.0 – 1.1 mm (MNHN-IU- 2016 - 456), 6 M 1.7 – 2.7 mm, 1 ov. F 2.5 mm, 4 F 1.4 – 1.6 mm (MNHN-IU- 2016 - 458), 1 M 2.2 mm (MNHN-IU- 2014 - 12606), 1 M 1.9 mm (MNHN-IU- 2014 - 13841). — Stn WS 06, 33.2516 ° S, 43.9083 ° E, 26 m, 1 May 2017: 1 M 1.9 mm, 2 F 1.2 – 1.3 mm (MNHN-IU- 2017 - 3613). — Stn WS 07, 33.2566 ° S, 43.8700 ° E, 30 – 33 m, 2 May 2017: 3 M 1.3 – 1.7 mm, 3 F 1.3 – 1.9 mm (MNHN-IU- 2016 - 487). — Stn WS 08, 33.2283 ° S, 43.9316 ° E, 30 – 33 m, 3 May 2017: 1 M 1.8 mm (MNHN-IU- 2014 - 13840), 2 M 1.3 – 2.2 mm, 5 F 1.3 – 2.3 mm (MNHN-IU- 2017 - 3333), 9 M 1.3 – 2.1 mm, 7 F 1.2 – 2.2 mm, 11 postlarvae 1.0 – 1.1 mm (MNHN-IU- 2016 - 460). — Stn WB 09, 33.2300 ° S, 43.9300 ° E, 27 – 30 m, 4 May 2017: 16 M 1.2 – 3.1 mm, 15 F 1.4 – 2.2 mm, 7 postlarvae 1.0 – 1.1 mm (MNHN-IU- 2014 - 13839), 1 postlarvae 1.0 mm (MNHN-IU- 2016 - 457), 1 F 1.3 mm (MNHN-IU- 2017 - 2960), 2 M 2.1 – 2.4 mm, 4 F 1.5 – 2.1 mm, 10 postlarvae, 0.9 – 1.0 mm (MNHN-IU- 2017 - 3316). — Stn WB 10, 33.1516 ° S, 43.8633 ° E, 30 m, 6 May 2017: 26 M 1.5 – 2.1 mm, 25 F 1.2 – 2.5 mm, 14 postlarvae 1.0 – 1.2 mm (MNHN-IU- 2019 - 2689), 13 M 1.1 – 2.4 mm, 15 F 1.1 – 2.3 mm, 3, postlarvae 1.0 – 1.1 mm (MNHN-IU- 2017 - 3812), 1 M 2.4 mm (MNHN-IU- 2014 - 12639), 2, postlarvae, 1 broken (MNHN-IU- 2017 - 2990).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD0E62F4F9CFCD87460B897.taxon	etymology	Etymology. The new species is named after Janique Etienne, Senior Officer in charge of High Seas projects with Fonds Français pour l’Environnement Mondial (FFEM), in recognition for her support to the Walters Shoals expedition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD0E62F4F9CFCD87460B897.taxon	description	Description. Carapace: [0.9] – 1.1 × as long as broad; transverse ridges with dense short setae and few scattered long and thick iridescent setae. Gastric region with 4 transverse ridges: epigastric ridge distinct with 2 median spines; anterior protogastric ridge not medially interrupted, nearly extending laterally to carapace margin; anterior mesogastric ridge not medially interrupted, laterally continuing to first branchial spine; anterior metagastric ridge scale-like (rarely medially interrupted), sometimes followed by a short small scale. Mid-transverse ridge not interrupted, medially slightly depressed, cervical groove indistinct, followed by 2 not interrupted or minutely interrupted ridges, interspersed with 1 short lateral ridge and few short scales. Lateral margins slightly convex, with 6 – 7 distinct spines: first anterolateral spine well-developed, reaching anteriorly to level of lateral orbital spine, second spine (hepatic) small, slightly dorsomesially from lateral margin, and followed by 4 – 5 branchial spines (3 anterior and 1 – 2 posterior). Rostrum leaf-like, horizontal, dorsally convex, [1.4] – 1.6 × as long as broad, length 0.4 and breadth 0.3 that of carapace; lateral margins smooth or minutely serrated and convex, with well-developed supraocular basal spines and small subapical spines. Pterygostomian flap ending in anterior spine, upper margin smooth. Thoracic sternum: As wide as long. Sternite 3 moderately broad, 1.9 – [3.0] × as wide as long, anterior margin slightly convex or straight. Sternite 4 widely contiguous to sternite 3; anterolaterally smooth, surface depressed in midline, smooth; greatest width 2.0 – [2.5] × that of sternite 3, 2.7 – [3.5] × as wide as long. Pleon: Tergite 2 with anterior and posterior transverse elevated ridges, with short setae and a few scattered long setae; tergites 3 – 4 with anterior transverse ridge only; tergites 5 – 6 smooth. Eye: Eyestalk length about 1.1 × broader than long, peduncle distally setose, not distinctly expanded proximally; cornea expanded distally, maximum corneal diameter 0.9 × rostrum width, as wide as eyestalk. Antennule: Article 1 slightly longer than wide, with 5 distal spines: distomesial spine well-developed; proximal lateral spine small, always present. Antenna: Article 1 with prominent mesial process distally falling well short of lateralmost antennular spine. Article 2 with well-developed distomesial and distolateral spines. Article 3 with distomesial spine and distolateral spines, otherwise only distomesial spine. Article 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus 0.6 – [0.8] × length of ischium, with well-developed distal spine on extensor and flexor margins. P 1: 2.0 – [3.0] (males), 1.7 – 1.9 (females) × carapace length; subcylindrical, spiny and with dense long stiff setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus 0.7 – [1.1] length of carapace, 1.7 – [1.8] × as long as carpus. Carpus [1.5] – 2 × as long as wide. Palm 1.1 × carpus length, [1.4] – 1.7 × as long as broad. Fingers [0.9] 0.8 – 1.0 × palm length; fixed finger with 2 small basal spines; movable finger with 1 – 2 basal spines or unarmed. P 2 – 4: stout, setose and spinose. Meri successively shorter posteriorly: P 3 merus [0.9] – 1.0 × length of P 2 merus, P 4 merus [0.8] – 0.9 × length of P 3 merus. P 2 merus, 0.5 – [0.7] × carapace length, 3.8 – [4.0]. times as long as broad, 1.0 – [1.4] × as long as P 2 propodus; P 3 merus 3.2 – [3.9] × as long as broad, 1.1 – [1.3] × as long as P 3 propodus; P 4 merus [3.3] 2.9 – 3.6 × as long as broad, 1.0 – [1.1] × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, unarmed; flexor margin irregular, with distal spine on P 2 – 3, distal spine absent in P 4, P 4 lateral surface with median row of 2 small spines, absent in P 2 – 3. Carpi with 1 – 3 spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, smaller on P 4; granules below extensor margin on lateral surface of P 2 – 4; flexor margin unarmed. Propodi stout, [4.3 – 4.6] 3.7 – 4.4 × as long as broad; extensor margin irregular, armed with 2 – 4 spines on proximal half, otherwise unarmed; flexor margin with 3 – 4 slender movable spines in addition to distal pair. Dactyli 0.6 – 0.7 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 5 – 6 movable spines. Eggs: Ov. F carried approximately 20 – 35 eggs of 0.4 – 0.5 mm diameter. Live colour. Body uniformly pale orange with subtle pattern of darker orange patches; transverse orange bar across base of rostrum and ocular peduncle. P 1 light orange with scattered reddish spots. P 2 – 4 whitish or translucent, with orange bands. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD0E62F4F9CFCD87460B897.taxon	distribution	Distribution. South West Indian Ocean, Madagascar, Reunion Island and Walter Shoals from 18 to 98 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD0E62F4F9CFCD87460B897.taxon	discussion	Remarks. Phylladiorhynchus janiqueae belongs to the group of species having 2 epigastric spines, 1 hepatic spine, 3 spines on the anterior branchial margin and the pleonal tergite 3 with the anterior ridge only. This species fits with the illustrations from Tirmizi & Javed (1980, 1993) from the Indian Ocean, however we have not examined this material, therefore the identification of these specimens remains dubious. The new species is closely related with P. medea from French Polynesia and New Caledonia (see the differences under the Remarks of this species). The specimens of P. janiqueae range from 1.0 to 3.0 mm postorbital carapace length. The four sequences of P. janiqueae from Madagascar and Walter Shoal diverged 0 % for COI and 0 % for 16 S.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD7E62D4F9CFE7273D7BC66.taxon	description	(Figs. 21 I, 23, 54 F)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD7E62D4F9CFE7273D7BC66.taxon	materials_examined	Type material. Holotype. Papua New Guinea, KAVIENG Stn KS 31, 02.6583 ° S, 150.6283 ° E, 15 m, 12 June 2014: ov. F 2.4 mm (MNHN-IU- 2014 - 13661). Paratypes. Papua New Guinea. KAVIENG Stn KB 06, 02.6866 ° ’ S, 150.6866 ° E, 8 m, 4 June 2014: 1 F 2.0 mm (MNHN-IU- 2014 - 13639). ― Stn KS 15, 02.6866 ° S, 150.6866 ° E, 3 – 5 m, 6 June 2014: 1 ov. F 2.2 mm (MNHN-IU- 2014 - 2176). ― Stn KS 23, 02.6800 ° S, 150.7116 ° E, 4 – 7 m, 8 June 2014: 1 ov. F 1.8 mm (MNHN-IU- 2014 - 13528), 3 M 1.1 – 1.5 mm, 1 ov. F 1.9 mm (MNHN-IU- 2016 - 5835). ― Stn KD 78, 02.7100 ° S, 150.7233 ° E, 6 – 10 m, 26 June 2014, 1 M 2.7 mm (MNHN-IU- 2014 - 13565). Other material. Papua New Guinea. KAVIENG Stn KB 18, 02.6800 ° S, 150.7116 ° E, 4 – 7 m, 8 June 2014: 1 M 2.0 mm (MNHN-IU- 2014 - 13623). ― Stn KD 36, 02.5850 ° S, 150.4833 ° E, 8 m, 16 June 2014: 1 ov. F 2.2 mm (MNHN-IU- 2014 - 13530). ― Stn KS 41, 02.6100 ° S, 150.5483 ° E, 2 – 7 m, 16 June 2014: 2 M 1.7 – 1.9 mm (MNHN- IU- 2014 - 13647). ― Stn KS 43, 02.5866 ° S, 150.4850 ° E, 4 – 12 m, 16 June 2014: 8 M 1.2 – 1.7 mm, 3 ov. F 1.8 – 2.0 mm, 5 F 1.4 – 1.6 mm, 2 postlarvae 1.0 mm (MNHN-IU- 2016 - 5851).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD7E62D4F9CFE7273D7BC66.taxon	etymology	Etymology. Named after Jeff Kinch, formerly Principal of Papua New Guinea National Fisheries Authority’s, National Fisheries College in Kavieng.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD7E62D4F9CFE7273D7BC66.taxon	description	Description. Carapace: Robust or massive, sexually dimorphic (wider on females), 0.9 (males), 0.8 (females) × as long as broad; transverse ridges elevated, upraised dorsally, densely covered with short setae, long and thick setae absent. Gastric region convex (upraised) with some transverse ridges: epigastric ridge distinct, medially interrupted, unarmed; anterior protogastric ridge usually medially and laterally interrupted, with few scales to carapace margin, often followed by few short scales; anterior mesogastric ridge scale-like, laterally interrupted by anterior branch of cervical groove, continuing with few scales, often followed by few short scales; anterior metagastric ridge not medially interrupted, laterally interrupted, followed by scales on posterior metagastric region. Mid-transverse ridge not interrupted, cervical groove distinct, followed by 3 not interrupted or minutely interrupted ridges, interspersed with 1 laterally interrupted or scale-like ridge, and some scattered scales. Lateral margins clearly convex, with 7 – 8 spines: first anterolateral spine well-developed, exceeding lateral orbital spine, second spine (hepatic) small, slightly dorsomesially from lateral margin, and followed by 5 – 6 branchial spines behind distinct anterior cervical groove (3 anterior and 1 – 2 posterior). Rostrum leaf-like, horizontal, dorsally flattish or slightly concave, sexually dimorphic (near bottle-shaped for males) [1.5] 1.4 – 1.7 × as long as broad, length [0.4] – 0.5 and breadth 0.2 – [0.3] that of carapace; lateral margins smooth and convex, with well-developed basal supraocular and subapical spines. Pterygostomian flap ending in blunt tooth; upper margin unarmed. Thoracic sternum: Sternite 3 moderately broad, [2.0] – 2.5 × as wide as long, anterior margin convex, with anterolateral projections. Sternite 4 widely contiguous to sternite 3; surface flattened, smooth; greatest width 3.0 × that of sternite 3, [2.5] – 2.7 × as wide as long. Pleon: Elevated ridges upraised dorsally, with few short setae. Tergites 2 – 4 with anterior and posterior transverse elevated ridges; tergites 5 – 6 smooth. Eye: Eyestalk length about 0.9 × broader than long, peduncle distally setose, not distinctly expanded proximally; maximum corneal diameter 1.0 × rostrum width, about as wide as eyestalk. Antennule: Article 1 longer than wide, with 3 distal spines: distomesial spine absent, pair of distolateral spines present; proximal lateral spine absent. Antenna: Article 1 with prominent mesial process distally clearly not reaching lateral antennular spine. Article 2 unarmed, sometimes with minute distomesial spine. Articles 3 and 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus [0.5] – 0.6 × length of ischium, with 0 – 1 median small spine and well-developed distal spine on extensor margin and 2 spines on flexor margin. P 1: 2.6 – 2.7 × carapace length (male) [1.5] – 1.6 (female); subcylindrical, moderately spiny and with scattered long stiff setae. Merus 0.5 – 0.9 length of carapace, [1.7] 1.6 – 1.8 × as long as carpus, with few scattered spines, mesial spines strongest. Carpus [1.5] – 1.9 × as long as wide with few scattered spines, mesial most prominent. Palm [1.2] – 1.3 × carpus length, 1.3 – 3.0 × as long as broad with scattered small spines on dorsal and ventral surfaces, lateral and mesial margins irregular, with scattered small spines. Fingers unarmed, 0.7 – [0.9] × palm length. P 2 – 4: Moderately stout, setose with few spines. Meri successively shorter posteriorly: P 3 merus 0.7 – 0.9 × length of P 2 merus, P 4 merus 0.8 – 0.9 × length of P 3 merus. P 2 merus, 0.6 × carapace length, 3.8 × as long as broad, 1.1 × as long as P 2 propodus; P 3 merus 3.4 – 3.8 × as long as broad, 1.1 – 1.2 × as long as P 3 propodus; P 4 merus 1.2 × as long as broad, as long as P 4 propodus; extensor margins of P 2 and P 3 with row of few small spines, proximally diminishing, with distal spine; P 4 extensor margin irregular but unarmed; flexor margins irregular, with distal spine on P 2 – 3, absent in P 4. Carpi irregular on extensor margins on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, absent on P 4; row of small spines below extensor margin on lateral surface of P 2 – 3, unarmed on P 4. Propodi stout, 3 – 4 × as long as broad; extensor margins irregular, usually unarmed; flexor margins with 3 – 5 slender movable spines in addition to distal pair. Dactyli 0.7 – 0.8 × length of propodi, ending in incurved, strong, sharp spine; flexor margins with 5 – 6 well-developed dactylar spines, each with 1 spinule. Eggs: Ov. F carried approximately 8 – 20 eggs of 0.3 – 0.5 mm diameter. Colour. Body green-brownish with some whitish and dark brown patches. P 1 – 4 whitish-pale brown, with brownish bands Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD7E62D4F9CFE7273D7BC66.taxon	distribution	Distribution. Papua New Guinea, from 2 to 15 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD7E62D4F9CFE7273D7BC66.taxon	discussion	Remarks. Phylladiorhynchus jeffkinchi belongs to the group of species characterized by having the epigastric ridge unarmed, the carapace and pleon ridges elevated, upraised, and dactylar spines on the flexor margins of the dactyli. The new species resembles P. phanus, from Papua New Guinea and P. marina, from Vanuatu. However, P. jeffkinchi can be distinguished from these species on the basis of the following characters: - The anterior epigastric ridge is always distinct and medially interrupted in P. jeffkinchi, whereas it is often indistinct or scale-like in P. phanus and P. marina. - The anterior protogastric ridge is often medially interrupted in P. jeffkinchi, whereas it is not interrupted in P. phanus and P. marina. - The carapace is densely covered by short setae in P. jeffkinchi, whereas these setae are shorter and less dense in P. phanus and P. marina. - The anterior margin of thoracic sternite 3 is convex and anterolaterally projected in P. jeffkinchi, whereas these anterolateral projections are absent in P. phanus and P. marina. The specimens of P. jeffkinchi range from 1.0 to 2.7 mm postorbital carapace length. The genetic divergences among P. jeffkinchi, P. marina and P. phanus were 7 % (16 S) and larger than 14 % (COI). The three sequences of P. jeffkinchi from Papua New Guinea diverged 0 – 0.3 % for COI and 0 % for 16 S.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD5E6504F9CFA7473D5BAF2.taxon	description	(Figs. 21 K, 24, 54 G – H)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD5E6504F9CFA7473D5BAF2.taxon	materials_examined	Type material: Holotype. Vanuatu, SANTO Stn DB 33, 15.5783 ° S, 167.2300 ° E, 14 – 25 m, 18 September 2006: 1 ov. F 2.1 mm (MNHN-IU- 2014 - 13805). Paratypes. Vanuatu. SANTO Stn ZB 9, 15.6766 ° S, 167.0850 ° E, 5 – 7 m, 02 October 2006: 1 ov. F 1.7 mm (MNHN-IU- 2019 - 2627). — Stn DB 86, 15.6416 ° S, 167.2516 ° E, 13 m, 04 October 2006: 1 ov. F 2.3 mm (MNHN- IU- 2014 - 13809). — Stn DB 1, 15.5516 ° S, 167.2966 ° E, 15 – 25 m, 10 September 2006: 1 M 1.8 mm (MNHN-IU- 2019 - 2628). — Stn FB 72, 15.6016 ° S, 166.9750 ° E, 16 m, 12 October 2006: 1 M 2.2 mm, 1 postlarvae (MNHN- IU- 2014 - 13877). — Stn DB 29, 15.6483 ° S, 167.0850 ° E, 15 m, 17 September 2006: 1 M 1.9 mm, 1 F 1.7 mm (MNHN-IU- 2014 - 13869). — Stn ZB 6, 15.6133 ° S, 167.0216 ° E, 30 m, 28 September 2006: 1 F 1.7 mm (MNHN-IU- 2014 - 13871). New Caledonia Koumac 2.3 Stn KL 39, 20.7511 ° S, 164.23276 ° E, 26 m, 10 November 2019: 1 M 2.1 mm, 2 ov. F 1.9 – 2.5 mm, 3 F 1.7 – 2.0 mm (MNHN-IU- 2014 - 20055). Other material. American Samoa. Olosega Island. OLO- 057, 14.18140174 ° S, 169.6267462 ° W, 14 m, 14 March 2015: 1 ov. F 2.0 mm (UF 54536). — OLO- 211, 14.18140174 ° S, 169.6267462 ° W, 14 m, 14 March 2015: 1 M 2.2 mm (UF 54537). French Polynesia. Society Islands, Moorea Island, Fore reef NE of Tareu Pass, outer reef slope, rubble, 17.4803 ° S, 149.8539 ° W, 35 – 37 m, 2 August 2006: 1 broken (UF 10072). — 17.4826 ° S, 149.8962 ° W, 14 October 2008: 1 ov. F 2.2 mm (UF 15544). — 17.5145 ° S, 149.7616 ° W, 22 m, 23 October 2008: 1 ov. F 1.8 mm (UF 16051). — 17.5303 ° S, 149.7621 ° W, 15 – 32 m, 27 October 2008: 1 ov. F 1.9 mm (UF 16169). Northern Mariana Islands. Saipan Island. Stn SAI- 441, 15.0978 ° N, 145.7434 ° E, 17 m, 11 April 2014, 1 M 1.6 mm, 1 ov. F 2.0 mm (UF 54538). — Stn SAI- 477, 15.0978 ° N, 145.7434 ° E, 17 m, 11 April 2014: 1 M 2.0 mm (UF 54540). — Stn SAI- 584, 15.0978 ° N, 145.7434 ° E, 17 m, 11 April 2014: 1 M 2.0 mm (UF 54540). — Pagan Island. Stn PAG- 694, 18.0706862 ° N, 145.7137933 ° E, 37 m, 20 April 2014: 1 ov. F 2.2 mm, 1 F 1.9 mm (UF 54533). — Stn PAG- 809, 18.0707 ° N, 145.7138 ° E, 37 m, 20 April 2014: 1 M 1.4 mm (UF 54534). Marshall Islands. Wake Island. Stn WAK- 115, 19.2918 ° N, 166.6073 ° E, 14 m, 17 March 2014: 1 F 1.7 mm (UF 54535). Chesterfield Islands, Ebisco, Stn DW 2569, 20.3963 ° S, 158.6699 ° E, 50 m, 14 October 2005: 1 ov. F 1.7 mm (MNHN-IU- 2014 - 13912). New Caledonia. New Caledonia, Lagon Est, Stn 735, 22.0850 ° S, 166.9533 ° E, 15 – 34 m, August 1986: 1 M 2.5 mm (MNHN-IU- 2019 - 2620). — Touho Dive, 7 September 1993: 1 M 1.7 mm, 2 ov. F 2.0, 2.1 mm, 1 F 2.1 mm (MNHN-IU- 2019 - 2614). — Touho, 6 September 1993: 2 M 1.5, 1.6 mm, 2 F 1.7, 1.8 mm (MNHN-IU- 2016 - 498). — Exp. Mont. Komac, 12 m, 7 October 1993: 1 M parasitized 2.5 mm (MNHN-IU- 2019 - 2634). — LIFOU Stn 1430, 20.7916 ° S, 167.1183 ° E, 20 – 25 m, 9 November 2000: 1 ov. F 2.6 mm (MNHN-IU- 2019 - 2622). — Stn 1452, 20.9100 ° S, 167.0350 ° E, 2 – 25 m, 20 November 2000: 1 M 2.2 mm, 1 ov. F 2.0 mm (MNHN-IU- 2019 - 2599). — Stn 1455, 20.9466 ° S, 167.0450 ° E, 15 – 20 m, 25 November 2000: 1 M 2.0 mm, 2 ov. F 2.1 – 2.2 mm (IU- 2019 - 2626). — Koumac 2.3 Stn KB 612, 20.6695 ° S, 164.18941 ° E, 13 m, 01 November 2019: 1 M 1.5 mm, 1 F parasitized 1.6 mm, 1 postlarvae 1.0 mm (MNHN- 2019 - 2580) 1 broken (MNHN-IU- 2014 - 3426). — Stn KB 613, 20.6682 ° S, 164.18962 ° E, 15 m, 01 November 2019: 1 ov. F 1.5 mm (MNHN-IU- 2019 - 1556), 1 M 1.3 mm, 5 ov. F 1.4 – 1.9 mm, 2 F 1.1 – 2.0 mm (MNHN-IU- 2019 - 1573). — Stn KB 615, 20.6943 ° S, 164.24786 ° E, 13 m, 02 November 2019: 1 ov. F 1.5 mm (MNHN-IU- 2019 - 1560), 4 F 1.5 – 1.8 mm (MNHN-IU- 2014 - 20198). — Stn KB 619, 20.5135 ° S, 164.03151 ° E, 12 m, 03 November 2019: 2 M 1.8 – 1.9 mm (MNHN-IU- 2019 - 1562), 1 F 1.8 mm (IU- 2019 - 5518). — Stn KB 622, 20.75144 ° S, 164.23203 ° E, 19 m, 04 November 2019: 2 M 1.4 – 1.5 mm, 2 F parasitized 1.5 – 1.6 mm (MNHN-IU- 2019 - 1572). — Stn KB 623, 20.7518 ° S, 164.2340 ° E, 12 m, 05 November 2019: 2 M 1.5 – 1.6 mm, 3 F 1.1 – 1.8 mm, 1 postlarvae 1.0 mm (MNHN-IU- 2019 - 3418). — Stn KB 624, 20.62347 ° S, 164.14964 ° E, 3 m, 05 November 2019: 1 M 1.3 mm (MNHN-IU- 2019 - 3420), 1 M 1.4 mm (MNHN-IU- 2019 - 3428). — Stn KB 631, 20.52698 ° S, 164.02615 ° E, 3 m, 08 November 2019: 1 M 1.8 mm, 1 postlarvae 1.0 mm (MNHN-IU- 2014 - 20079). — Stn KB 633, 20.59318 ° S, 164.25161 ° E, 41 m, 10 November 2019: 2 M 1.0 – 1.2 mm, 3 ov. F 1.4 – 1.5 mm, 1 F 1.2 mm (MNHN-IU- 2014 - 20048). — Stn KB 636, 20.63948 ° S, 164.18293 ° E, 3 m, 10 November 2019: 1 F 1.9 mm (MNHN-IU- 2014 - 20062), 1 M 1.8 mm, 1 ov. F 2.1 mm, 1 F parasitized 1.7 mm (MNHN-IU- 2014 - 20064), 1 M 1.6 mm (MNHN-IU- 2014 - 20066). — Stn KL 39, 20.7511 ° S, 164.23276 ° E, 26 m, 10 November 2019: 4 M 1.4 – 1.8 mm, 4 F parasitized 1.5 – 1.8 mm (MNHN-IU- 2014 - 20172), 1 F 1.3 mm (MNHN-IU- 2014 - 20046), 1 ov. F 2.3 mm (MNHN-IU- 2014 - 20056). — Stn KL 40, 20.75156 ° S, 164.2343 ° E, 52 m, 10 November 2019: 1 ov. F 2.2 mm (MNHN-IU- 2014 - 20083). — Stn KL 03, 20.67485 ° S, 164.214266 ° E, 50 m, 11 November 2019: 1 F 1.8 mm (MNHN- IU- 2014 - 20069). — Stn KB 638, 20.594533 ° S, 164.10925 ° E, 4 m, 13 November 2019: 1 M 2.0 mm, 1 ov. F 2.6 mm, 1 F parasitized 2.0 mm (MNHN-IU- 2014 - 20119). — Stn KB 639, 20.7511 ° S, 164.23245 ° E, 22 m, 13 November 2019: 1 ov. F 1.7 mm, 3 F 1.6 – 2.2 mm (MNHN-IU- 2014 - 20196). — Stn KB 640, 20.72511 ° S, 164.26738 ° E, 6 m, 13 November 2019: 1 M parasitized 2.0 mm (MNHN-IU- 2014 - 20111), 2 M 1.5 – 2.0 mm, 1 ov. F 2.4 mm (MNHN- IU- 2014 - 20112). — Stn KD 558, 20.72775 ° S, 164.26546 ° E, 6 m, 13 November 2019: 1 F parasitized 1.9 mm (MNHN-IU- 2014 - 20088). — Stn KB 641, 20.7988 ° S, 164.27253 ° E, 36 m, 14 November 2019: 1 broken (MNHN- IU- 2014 - 20102). — Stn KB 643, 20.61416 ° S, 164.13703 ° E, 3 m, 14 November 2019: 2 M 1.4 – 2.5 mm, 6 ov. F 2.4 – 2.6 mm (MNHN-IU- 2014 - 20096), 1 M 2.2 mm (MNHN-IU- 2019 - 5320), 1 ov. F 2.1 mm (MNHN-IU- 2019 - 5319). — Stn KB 645, 20.67573 ° S, 164.2171 ° E, 16 m, 15 November 2019: 1 F 1.3 mm (MNHN-IU- 2014 - 20115), 2 M 1.2 – 1.3 mm, 5 F 1.0 – 1.2 mm (MNHN-IU- 2014 - 20125). — Stn KB 646, 20.644983 ° S, 164.24386 ° E, 12 m, 15 November 2019: 1 F parasitized 2.1 mm (MNHN-IU- 2014 - 20114), 1 M 1.7 mm, 1 postlarvae 0.9 mm (MNHN-IU- 2014 - 20118). — Stn KB 647, 20.664116 ° S, 164.189983 ° E, 38 m, 16 November 2019: 1 ov. F 1.6 mm, 3 F 1.0 – 1.9 mm (MNHN-IU- 2014 - 20139). — Stn KL 27, 20.7878 ° S, 164.27191 ° E, 102 m, 16 November 2019: 1 juv. 1.0 mm (MNHN-IU- 2014 - 20129). — Stn KL 37, 20.8319 ° S, 164.27818 ° E, 81 m, 16 November 2019: 1 M 1.1 mm (MNHN- IU- 2014 - 20132). — Stn KB 649, 20.80583 ° S, 164.26916 ° E, 42 m, 18 November 2019: 5 F 1.7 – 1.9 mm, 5 postlarvae 0.9 – 1.0 mm (MNHN-IU- 2014 - 20154). — Stn KB 651, 20.83145 ° S, 164.280556 ° E, 12 m, 18 November 2019: 1 postlarvae 1.1 mm (MNHN-IU- 2014 - 20143), 1 F 1.0 mm (MNHN-IU- 2014 - 20147). — Stn KB 651, 20.83145 ° S, 164.280556 ° E, 12 m, 18 November 2019: 1 postlarvae 1.1 mm (MNHN-IU- 2014 - 20148), 1 F 1.8 mm (MNHN- IU- 2014 - 20144). — Stn KB 654, 20.44587 ° S, 163.97254 ° E, 15 m, 19 November 2019: 7 F parasitized 1.1 – 2.1 mm (MNHN-IU- 2014 - 20164). — Stn KB 655, 20.68517 ° S, 164.27078 ° E, 6 m, 20 November 2019: 1 M 1.9 mm, 1 F parasitized 2.2 mm (MNHN-IU- 2014 - 20193). — Stn KB 659, 20.66049 ° S, 164.26852 ° E, 15 m, 21 November 2019: 1 M 1.9 mm (MNHN-IU- 2014 - 20176). — Stn KB 664, 20.66925 ° S, 164.230805 ° E, 3 m, 23 November 2019: 1 F parasitized 1.8 mm (MNHN-IU- 2014 - 20184). — Stn ARMS 2 B, November 2019: 1 M 1.7 mm (MNHN-IU- 2014 - 20074). — Stn ARMS 8 B, November 2019: 1 M 1.8 mm, 2 ov. F 2.2. – 2.3 mm (MNHN-IU- 2014 - 20162).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD5E6504F9CFA7473D5BAF2.taxon	etymology	Etymology. The new species is named after Pascale Joannot, Head of the MNHN Expeditions programme, who has been instrumental in raising support for the Our Planet Reviewed New Caledonia expeditions.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD5E6504F9CFA7473D5BAF2.taxon	description	Description. Carapace: as long as or slightly broader than long, sexually dimorphic (wider on females) 0.9 – 1.0 (males), [0.8] – 0.9 (females); transverse ridges with dense short setae and few scattered thick and iridescent setae. Gastric region slightly convex with 4 transverse ridges: epigastric ridge distinct with 2 median spines and some lateral short scales, often followed by small scattered scales; anterior protogastric ridge not medially interrupted, nearly extending laterally to carapace margin, often followed by small scattered scales; anterior mesogastric ridge not medially interrupted, continuing to first branchial spine; anterior metagastric not medially interrupted, sometimes followed by short scattered scales on posterior metagastric region. Mid-transverse ridge not interrupted, medially slightly depressed, cervical groove indistinct, followed by 2 not interrupted or minutely interrupted ridges, interspersed with 2 short lateral ridges and few short scales. Lateral margins convex, with 6 distinct spines: first anterolateral spine well-developed, reaching anteriorly to level of lateral orbital spine, second spine (hepatic) welldeveloped, slightly dorsomesially from lateral margin, and followed by 4 – 5 branchial spines (3 anterior and 1 – 2 posterior). Rostrum leaf-like, horizontal, dorsally flattish or slightly concave, sexually dimorphic (shorter and wider on females) 1.5 – 1.8 (males), [1.1] – 1.6 × as long as broad, length [0.3] – 0.4 and breadth 0.2 – [0.3] that of carapace; lateral margins smooth and convex, with well-developed supraocular basal spines and small subapical spines. Pterygostomian flap ending in blunt tooth, upper margin smooth. Thoracic sternum: Slightly wider than long. Sternite 3 broad, [2.8] – 3.0 × as wide as long, anterior margin convex, anterolaterally rounded. Sternite 4 widely contiguous to sternite 3; anterolaterally rounded, surface depressed in midline, smooth; greatest width [2.9] – 3.0 × that of sternite 3, 3.0 – [3.5] × as wide as long. Pleon: Elevated ridges with short setae and a few scattered long setae. Tergites 2 – 3 with anterior and posterior transverse elevated ridges; tergite 4 with anterior transverse ridge, posterior transverse ridge absent; tergites 5 – 6 smooth. Eye: Eyestalk length about 0.9 – [1.1] × broader than long, peduncle distally setose, not distinctly expanded proximally; cornea expanded distally, maximum corneal diameter [0.9] × rostrum width, as wide as eyestalk. Antennule: Article 1 slightly longer than wide, with 5 distal spines: distomesial spine well-developed; proximal lateral spine small, always present. Antenna: Article 1 with prominent mesial process, distally falling well short of lateralmost antennular spine. Article 2 with well-developed distomesial and distolateral spines. Article 3 with distomesial spine. Article 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus 0.8 × length of ischium, with well-developed distal spine on extensor and flexor margins. P 1: [3.3] 3.0 – 3.5 (males), 1.8 – 2.2 (females) × carapace length; subcylindrical, spiny and with long stiff setae and dense thick iridescent and plumose setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus 0.7 – [1.2] length of carapace, [1.6] – 2.2 × as long as carpus. Carpus 1.5 – [2.0] × as long as wide. Palm 1.2 × carpus length, 1.9 – [2.2] × as long as broad. Fingers [0.7] – 1.0 × palm length; fixed finger unarmed; movable finger with 1 – 2 basal spines. P 2 – 4: Setose and spinose. Meri successively shorter posteriorly: P 3 merus 0.9 × length of P 2 merus, P 4 merus 0.8 – 0.9 × length of P 3 merus. P 2 merus, 0.5 × carapace length, 3.5 – 3.6 × as long as broad, 1.0 – 1.1 × as long as P 2 propodus; P 3 merus 3.0 – 4.3 × as long as broad, 1.1 – 1.3 × as long as P 3 propodus; P 4 merus 2.6 – 3.0 × as long as broad, as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, unarmed; flexor margin irregular, with distal spine on P 2 – 3, absent on P 4. Carpi with 1 – 3 spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, absent on P 4; granules below extensor margin on lateral surface of P 2 – 4; flexor margin unarmed. Propodi stout, [4.0 – 5.0] 3.7 – 4.8 × as long as broad; extensor margin irregular; flexor margin with 3 slender movable spines in addition to distal pair. Dactyli [0.6 – 0.7] × length of propodi, ending in incurved, strong, sharp spine; flexor margin with cuticular spines at basis of 5 – 6 movable spines. Eggs: Ov. F carried approximately 20 – 25 eggs of 0.3 mm diameter. Live colour. Variable; base colour white or pale beige. Carapace with fine red to brown markings (Fig 54 H) or with dense patches of red to dark brown chromatophores (Fig 54 G). Rostrum with dark transverse bar at least across base. P 1 – 4 with dark brown bars and scattered white spots. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD5E6504F9CFA7473D5BAF2.taxon	distribution	Distribution. French Polynesia, Vanuatu, Papua New Guinea, Guam Island, Northern Mariana Islands, American Samoa and New Caledonia, from 5 to 102 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFD5E6504F9CFA7473D5BAF2.taxon	discussion	Remarks. Phylladiorhynchus joannotae belongs to the group of species having 2 epigastric spines, 1 hepatic spine, 3 spines on the anterior branchial margin of the carapace and cuticular spines along the flexor margin of the P 2 – 4 dactyli. The morphologically and genetically closest species is P. amphion, from Western Australia. However, these species can be distinguished in basis of the following characters: - The metagastric ridge is usually interrupted in P. amphion, whereas it is not medially interrupted in P. joannotae. - P. joannotae has iridescent setae in the carapace and pleon, whereas these setae are absent in P. amphion. The specimens of P. joannotae range from 1.0 to 2.6 mm postorbital carapace length. P. joannotae showed divergences of 9 % (COI) and 6 % (16 S) with the closest relative (P. amphion). The 11 sequences of P. joannotae from New Caledonia and Vanuatu diverged 0 – 2 % for COI and 0 – 1 % for 16 S.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFA8E6504F9CFB8D7101BFBC.taxon	diagnosis	Diagnosis (modified from Schnabel & Ahyong 2019). Rostrum without subapical spines; lateral margins slightly concave. Carapace with epigastric ridge with 5 spines; anterior branchial margin with 2 spines; anterior protogastric ridge not interrupted medially, without scale behind median epigastric spine; anterior mesogastric ridge not medially interrupted or with small medial gap; anterior metagastric ridge medially interrupted. Thoracic sternite 3 anterior margin sinuous, with shallow median concavity. Antennular article 1 with 5 well-developed spines; distomesial spine small, similar to lower distolateral spine. Antennal article 1 mesial process distally falling well short of second lateral antennular spine; article 2 distomesial spine longer than distolateral; article 3 with [small] to distinct distomesial spine. Flexor margin of Mxp 3 merus with 2 prominent spines. P 2 – 4 dactylus flexor margin without upright spines at bases of movable spines.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFA8E6504F9CFB8D7101BFBC.taxon	distribution	Distribution. New Zealand, Kermadec Islands, 195 – 287 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFA8E6504F9CFB8D7101BFBC.taxon	discussion	Remarks. As Schnabel & Ahyong (2019) pointed out, the species is characterized by the presence of 5 epigastric spines and 2 spines on the anterior branchial margin. The closest species is P. punctatus, from New Caledonia. Both species can be differentiated by the interrumptions in the gastric ridges (see the differences under the Remarks of this species).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFA8E65B4F9CF95773EBB8CF.taxon	description	(Figs. 21 L, 25, 54 I)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFA8E65B4F9CF95773EBB8CF.taxon	materials_examined	Type material. Holotype. New Caledonia, Koumac 2.3 Stn KB 629, 20.59121 ° S, 164.21503 ° E, 10 m, 7 November 2019: F 1.6 mm (MNHN-IU- 2019 - 3427). Paratypes. New Caledonia. LIFOU Stn 1436, 20.9250 ° S, 167.0700 ° E, 10 – 20 m, 10 November 2000: 1 F 1.8 mm (MNHN-IU- 2014 - 13847). New Caledonia. Koumac 2.3 Stn KL 07, 20.64605 ° S, 164.18498 ° E, 82 m, 15 November 2019: 1 M 1.5 mm (MNHN-IU- 2014 - 20121). — Stn KL 22, 20.4459 ° S, 163.97176 ° E, 47 m, 19 November 2019: 1 F 1.4 mm (MNHN- IU- 2014 - 20170) parasitized. — Stn KL 25, 20.75421 ° S, 164.22986 ° E, 65 m, 10 November 2019: 1 F 1.8 mm (MNHN-IU- 2014 - 20035). — Stn KL 03, 20.67485 ° S, 164.214266 ° E, 50 m, 11 November 2019: 1 F 1.8 mm (MNHN- IU- 2014 - 20069).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFA8E65B4F9CF95773EBB8CF.taxon	etymology	Etymology. Named after the expedition KOUMAC aimed to catalogue the marine biodiversity of the lagoons of Koumac, New Caledonia. The name is considered as a substantive in apposition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFA8E65B4F9CF95773EBB8CF.taxon	description	Description. Carapace: Robust or massive, [0.9] – 1.1 × as long as broad; transverse ridges elevated, serrated, with few short setae, and few scattered long thick iridescent setae. Gastric region convex (upraised dorsally), with some transverse ridges: epigastric ridge indistinct, with 5 spines (1 median and 2 pairs of small spines laterally); anterior protogastric ridge not medially interrupted, laterally with some scales, posterior protogastric region often with a few scales; anterior mesogastric ridge scale-like, laterally interrupted by anterior branch of cervical groove; anterior metagastric ridge scale-like, followed by few scales on posterior metagastric area. Mid-transverse ridge uninterrupted or minutely interruped, medially depressed, preceded by distinct cervical groove, followed by 1 – 2 uninterrupted or minutely interrupted ridges, interspersed with 2 short lateral ridges and scattered short scales. Lateral margins clearly convex, with 7 spines: first anterolateral spine well-developed, reaching end of lateral orbital spine, second spine (hepatic) well-developed, slightly dorsomesially from lateral margin, and followed by 5 branchial spines (3 anterior and 2 posterior). Rostrum leaf-like, horizontal, dorsally flattish or slightly concave, 1.3 – [1.4] × as long as broad, length 0.4 and breadth 0.3 that of carapace; lateral margins serrated and convex, with well-developed supraocular spines, subapical spines well-developed. Pterygostomian flap ending in blunt tooth, upper margin serrated, with series of uprised striae. Thoracic sternum: As wide as or slightly wider than long, lateral margins of posterior half slightly divergent. Sternite 3 moderately broad, 2.5 – [2.8] × as wide as long, anterior margin concave. Sternite 4 widely contiguous to sternite 3; surface flattened, smooth; greatest width 2.7 – [2.9] × that of sternite 3, 2.6 × as wide as long. Pleon: Elevated ridges upraised dorsally, with a few scattered short setae. Tergites 2 – 3 with anterior and posterior transverse elevated ridges; tergite 4 with anterior transverse ridge; tergites 5 – 6 smooth. Eye: Eyestalk length about 1.4 – [1.6] × broader than long, peduncle distally setose, not distinctly expanded proximally; maximum corneal diameter 1.1 × rostrum width, as wide as eyestalk. Antennule: Article 1 slightly longer than wide, with 4 spines: distomesial spine well-developed; proximal lateral spine absent. Antenna: Article 1 with prominent mesial process distally clearly not reaching lateral antennular spine. Article 2 with small distal spines laterally and mesially. Article 3 often with a minute mesial spine. Article 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus 0.6 × length of ischium at midlength, with 0 – 1 median and 1 well-developed distal spines on extensor margin and 1 strong spine on flexor margin. P 1: [2.5] – 3.0 (males) × carapace length, not known in females, subcylindrical, spiny and with scattered long stiff setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus [1.0] – 1.1 length of carapace, [2.1] – 2.7 × as long as carpus. Carpus [2.2] × as long as wide. Palm [1.3] – 1.4 × carpus length, 2.1 – [2.8] × as long as broad. Fingers unarmed, [0.7] – 0.8 × palm length. P 2 – 4 (lost in most specimens): Stout, subcylindrical, moderately setose and spinose, with few scattered plumose setae. Meri successively shorter posteriorly: P 3 merus [0.6] × length of P 2 merus, P 4 merus [0.9] × length of P 3 merus; P 2 merus, [0.6] × carapace length, [5] × as long as broad, [1.1] × as long as P 2 propodus; P 3 merus [4.4] × as long as broad, [0.9] × as long as P 3 propodus; P 4 merus [3.3] × as long as broad, [0.8] × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, with small distal spine; flexor margins of P 2 – 4 irregular, each with distal spine. Carpi with 1 or 2 spines on extensor margin on P 2 – 3, unarmed on P 4; row of small spines below extensor margin on lateral surface of P 2 – 3, unarmed on P 4; flexor margins with distal spine. Propodi moderately slender, 6 – 7 × as long as broad; extensor margin irregular, usually armed with 1 – 3 well-developed spines on P 2 – 4; flexor margin with 2 – 5 slender movable spines in addition to distal pair. Dactyli 0.7 – 0.8 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 5 – 6 movable spines. Eggs. No data. Live colour. Body grey or red, with median pale band. P 2 – 4 whitish, with brownish-grey bands. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFA8E65B4F9CF95773EBB8CF.taxon	distribution	Distribution. New Caledonia, Koumac, between 10 and 82 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFA8E65B4F9CF95773EBB8CF.taxon	discussion	Remarks. Phylladiorhynchus koumac belongs to the species group that has 5 epigastric spines, 3 spines on the anterior branchial margin, gastric region strongly convex, the leaf-like rostrum (margins clearly convex and subapical spines present), and elevated ridges upraised dorsally. The new species closely resembles P. pulchrus, from French Polynesia, the Philippines, and Vanuatu (see the differences under the Remarks of this species). The specimens of P. koumac range from 1.4 to 1.8 mm postorbital carapace length. The four sequences of P. koumac from New Caledonia diverged 0 – 0.5 % for COI and 0 % for 16 S.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFA3E65E4F9CFD9C7348BDA9.taxon	description	(Figs. 26, 27 C – D, 33 A)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFA3E65E4F9CFD9C7348BDA9.taxon	materials_examined	Type material. Holotype. New Caledonia, LIFOU Stn 1457 20.7800 ° S, 167.0458 ° E, 5 – 10 m, 27 November 2000: ov. F 2.4 mm (MNHN-IU- 2014 - 13739). Paratypes. New Caledonia. Exp. Mont. Komac 12 m, 7 October 1993: 1 F 1.7 mm (IU- 2019 - 2635). — LIFOU Stn 1435, 20.9200 ° S, 167.0116 ° E, 5 – 30 m, 8 November 2000: 1 ov. F 2.0 mm (MNHN-IU- 2019 - 2608). — Stn 1430, Stn 1430, 20.7916 ° S, 167.1183 ° E, 20 – 25 m, 9 November 2000: 2 ov. F 2.2 – 2.4 mm (MNHN-IU- 2019 - 2632). — Stn 1457, 20.7800 ° S, 167.0458 ° E, 5 – 10 m, 27 November 2000: 1 ov. F 2.4 mm (MNHN-IU- 2014 - 13739). — Koumac 2.3 Stn KB 631, 20.52698 ° S, 164.02615 ° E, 3 m, 08 November 2019: 1 F parasitized 1.7 mm (MNHN-IU- 2019 - 2581). — Stn KL 39, 20.7511 ° S, 164.23276 ° E, 26 m, 10 November 2019: 2 F parasitized 1.8 – 2.0 mm (MNHN-IU- 2019 - 2582), 1 F 1.8 mm (MNHN-IU- 2014 - 20123). — Stn ARMS 9 C: 1 ov. F 1.1 mm (MNHN-IU- 2014 - 20145). — Stn KB 659, 20.66049 ° S, 164.26852 ° E, 15 m, 21 November 2019: 1 M 2.3 mm (MNHN-IU- 2014 - 20175). Mariana Islands. Guam Island. Stn GUA- 645, 13.57847 ° N, 144.82831 ° E, 10 m, 25 March 2014: 1 ov. F 2.0 mm (UF 54543). Other material. Japan. Okinawa. Iriomote Is. Nakano Beach, 24.4323 ° N, 123.7916 ° E, 19 m, 9 July 2010: 1 M 2.0 mm (UF 26910). Papua New Guinea. KAVIENG Stn KS 33, 02.6366 ° S, 150.6450 ° E, 8 – 10 m, 13 June 2014: 1 ov. F 2.8 mm (MNHN-IU- 2014 - 13559), 1 M 2.0 mm (MNHN-IU- 2014 - 13645). Vanuatu. SANTO Stn DB 1, 15.5516 ° S, 167.2966 ° E, 15 – 25 m, 10 September 2006: 1 M 2.0 mm, 1 ov. F 1.9 mm (MNHN-IU- 2019 - 2629). — Stn DR 21, 15.6083 ° S, 167.0233 ° E, 22 – 25 m, 15 September 2006: 1 F 1.3 mm (MNHN-IU- 2014 - 13884). — Stn DB 29, 15.6483 ° S, 167.0850 ° E, 15 m, 17 September 2006: 1 M 1.5 mm (MNHN- IU- 2019 - 2624). — Stn DS 49, 15.6450 ° S, 167.0866 ° E, 10 – 17 m, 21 September 2006: 1 M 2.4 mm (MNHN-IU- 2019 - 2631). — Stn DB 63,15.4483 ° S, 167.2633 ° E, 21 m, 25 September 2006: 2 ov. F 2.0 – 2.1 mm (MNHN-IU- 2014 - 13872). — Stn DB 65, 15.4300 ° S, 167.2166 ° E, 13 m, 26 September 2006: 2 M 2.0 – 2.2 mm, 4 ov. F 1.7 – 2.1 mm, 3 F 1.8 – 2.3 mm (MNHN-IU- 2014 - 13870). — Stn DB 69, 15.4066 ° S, 167.2166 ° E, 38 m, 27 September 2006: 1 M 2.0 mm (MNHN-IU- 2029 - 2625). — Stn DB 75, 15.3816 ° S, 167.1983 ° E, 20 m, 28 September 2006: 1 M 2.0 mm (MNHN-IU- 2014 - 13885). — Stn FB 43, 15.4733 ° S, 167.2483 ° E, 19 m, 30 September 2006: 1 M 2.0 mm, 2 ov. F 1.7 – 1.8 mm (MNHN-IU- 2019 - 2590). — Stn FB 64, 15.5900 ° S, 166.9866 ° E, intertidal, 10 October 2006: 1 M 3.0 mm (MNHN-IU- 2014 - 13807). — no Stn number: 1 M 1.9 mm (MNHN-IU- 2019 - 2623). Mariana Islands. Guam Island. Mid-western coast, at Orote northern tip, dead coral, 25 m, 20 June 2002: 2 M 1.1 – 1.7 mm, 1 ov. F 1.6 (UF 3122). — Mariana Islands. Guam Island. Stn GUA- 597, 13.57847 ° N, 144.82831 ° E, 11 m, 25 March 2014: 3 M 1.4 – 1.7 mm, 6 ov. F 1.5 – 2.4 mm (UF 54509). — Stn GUA- 693, 13.57847 ° N, 144.82831 ° E, 11 m, 25 March 2014: 1 M 2.0 mm (UF 54510). — Stn GUA- 698, 13.57847 ° N, 144.82831 ° E, 11 m, 25 March 2014: 1 ov. F 1.8 mm (UF 54511). — Stn GUA- 880, 13.30553 ° N, 144.65257 ° E, 12 m, 27 March 2014: 1 ov. F 1.9 mm (UF 54503). — Stn GUA- 877, 13.30553 ° N, 144.65257 ° E, 12 m, 27 March 2014: 1 M 1.4 mm (UF 54504). — Stn GUA- 756, 13.30553 ° N, 144.65257 ° E, 12 m, 27 March 2014: 1 M 1.9 mm, 1 ov. F 2.0 mm, 1 F 2.1 mm (UF 54505). — Stn GUA- 805, 13.30553 ° N, 144.65257 ° E, 12 m, 27 March 2014: 2 ov. F 2.0 – 2.1 mm (UF 54506). — Stn GUA- 810, 13.30553 ° N, 144.65257 ° E, 12 m, 27 March 2014: 1 F parasitized, 2.0 mm (UF 54507). — Stn GUA- 878, 13.30553 ° N, 144.65257 ° E, 12 m, 27 March 2014: 1 M 1.8 mm (UF 54508). — Stn GUA- 947, 13.48871 ° N, 144.87796 ° E, 14 m, 28 March 2014: 1 M 1.8 mm (UF 54512). — Stn GUA- 1216, 13.44878 ° N, 144.62596 ° E, 14 m, 31 March 2014: 2 M 1.6 – 2.0 mm, 1 ov. F 1.9 mm (UF 54513). — Stn GUA- 1201, 13.44878 ° N, 144.62596 ° E, 14 m, 31 March 2014: 1 ov. F 1.5 mm (UF 54514). — Stn GUA- 1204, 13.44878 ° N, 144.62596 ° E, 14 m, 31 March 2014: 1 M 2.0 mm (UF 54515). Northern Mariana Islands. Saipan Island. Stn SAI- 442, 15.0978 ° N, 145.7434 ° E, 17 m, 11 April 2014: 4 M 1.2 – 2.0 mm, 5 ov. F 1.7 – 2.1 mm (UF). — Stn SAI- 543, 15.0978 ° N, 145.7434 ° E, 17 m, 11 April 2014: 4 M 1.1 – 1.9 mm, 2 ov. F 1.7 – 1.9 mm (UF 54517). — Stn SAI- 544, 15.09782849 ° N, 145.7434357 ° E, 17 m, 11 April 2014: 1 M 2.0 mm, 2 ov. F 2.1 – 2.3 mm (UF 54518). — Stn SAI- 641, 15.15620 ° N, 145.6899 ° E, 11 m, 17 April 2014: 4 M 1.2 – 2.1 mm, 8 ov. F 1.8 – 2.5 mm (UF 54519). — Stn SAI- 586, 15.0978 ° N, 145.7434 ° E, 17 m, 11 April 2014: 2 M 2.1 – 2.4 mm, 3 ov. F 1.5 – 2.1 mm (UF 54520). — Stn SAI- 437, 15.0978 ° N, 145.7434 ° E, 17 m, 11 April 2014: 3 M 1.9 – 2.3 mm, 1 ov. F 1.8 mm (UF 54521). — Stn SAI- 708, 15.1562 ° N, 145.6899 ° E, 11 m, 17 April 2014: 2 M 1.8 – 1.9 mm, 4 ov. F 1.8 – 2.0 mm, 1 F 1.2 mm (UF 54522). — Stn SAI- 756, 15.1562 ° N, 145.6899 ° E, 11 m, 17 April 2014: 3 ov. F 1.7 – 2.1 mm (UF 54523). — Stn SAI- 789, 15.1562 ° N, 145.6899 ° E, 11 m, 17 April 2014: 3 M 1.9 – 2.3 mm, 4 ov. F 1.7 – 2.2 mm, 1 F 1.2 mm (UF 54524). — Stn SAI- 878, 15.27406 ° N, 145.79106 ° E, 9 m, 18 April 2014: 1 F 1.8 mm (UF 54525). — Stn SAI- 889, 15.27406 ° N, 145.79106 ° E, 9 m, 18 April 2014: 11 M 1.5 – 2.3 mm, 8 ov. F 1.4 – 2.0 mm (UF 54526). — Stn SAI- 957, 15.27406 ° N, 145.79106 ° E, 9 m, 18 April 2014: 3 M 1.5 – 2.0 mm, 5 ov. F 1.6 – 2.2 mm, 1 F 1.8 mm (UF 54527). — Stn SAI- 786, 15.15620 ° N, 145.68996 ° E, 10 m, 17 April 2014: 1 postlarvae (UF 54528). American Samoa. Olosega Island. Stn OLO- 186, 14.18140174 ° S, 169.6267462 ° W, 14 m, 14 March 2015: 1 ov. F 2.0 mm (UF 54529). — Tutuila Island. Stn TUT- 117, 14.36046 ° S, 170.75024 ° W, 17 m, 26 February 2015: 1 M 1.7 mm, 1 ov. F 1.9 mm (UF 54530). — Stn TUT- 374, 14.36613333 ° S, 170.7628833 ° W, 12 m, 5 March 2015: 1 M 1.9 mm, 1 ov. F 2.0 mm (UF 54531). — Rose Atoll. Stn ROS- 689, 14.55965 ° S, 168.1601167 ° W, 14 m, 18 March 2015: 1 M 1.9 mm (UF 54532).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFA3E65E4F9CFD9C7348BDA9.taxon	etymology	Etymology. Named after our colleague and host Laure Corbari, curator of the Crustacean Collection of the Muséum national d’Histoire naturelle, Paris.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFA3E65E4F9CFD9C7348BDA9.taxon	description	Description. Carapace: Slightly broader than long; transverse ridges with dense short setae and long and thick iridescent setae. Gastric region slightly convex with 4 transverse ridges: epigastric ridge distinct with 2 median spines, often with short scales laterally; anterior protogastric ridge not medially interrupted, extending laterally to carapace margin, sometimes followed by small scales on posterior protogatric ridge; anterior mesogastric ridge not medially interrupted, laterally continuing to first branchial spine, followed by some lateral scales; anterior metagastric ridge medially interrupted or not medially interrupted, followed always by one median scale and often other small scales in posterior metagastric region. Mid-transverse ridge not interrupted, medially depressed, cervical groove indistinct, followed by 2 not interrupted or minutely interrupted ridges, interspersed with 1 – 2 short lateral ridges and few short scales. Lateral margins slightly convex, with 6 – 7 distinct spines: first anterolateral spine well-developed, exceeding level of lateral orbital spine, second spine (hepatic) small, slightly dorsomesially from lateral margin, and followed by 5 branchial spines (3 anterior and 1 – 2 posterior). Rostrum leaf-like, horizontal, dorsally concave, [1.1] – 1.4 × as long as broad, length [0.3] – 0.4 and breadth 0.3 that of carapace; lateral margins serrated and strongly convex, with well-developed supraocular basal spines and strong subapical spines (tridentiform). Pterygostomian flap ending in sharp tooth, upper margin smooth, with series of elevated striae. Thoracic sternum: As wide as long. Sternite 3 quadrangular, [1.5] 1.0 – 1.8 × as wide as long, anterior margin serrated, with a median projection, anterolaterally rounded. Sternite 4 widely contiguous to sternite 3; anterolaterally serrated, surface depressed in midline, smooth; greatest width 2.6 – [3.0] × that of sternite 3, 3.0 × as wide as long. Pleon: Elevated ridges with short setae and a few scattered long setae. Tergites 2 – 3 with anterior and posterior transverse elevated ridges; tergite 4 with anterior transverse ridge only; tergites 5 – 6 smooth. Eye: Eyestalk length about [1.1] – 1.2 × broader than long, peduncle distally setose, not distinctly expanded proximally; cornea expanded distally, maximum corneal diameter 0.8 – [0.9] × rostrum width, narrower than eyestalk (0.8 × maximum peduncle width). Antennule: Article 1 slightly longer than wide, with 5 distal spines: distomesial spine well-developed; proximal lateral spine small, always present. Antenna: Article 1 with prominent mesial process, distally falling well short or overreaching lateralmost antennular spine. Article 2 with well-developed distomesial and distolateral spines. Article 3 with distomesial spine. Article 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus 0.8 × length of ischium, with well-developed distal spine on extensor and flexor margins. P 1: 2.3 – 3.1 (males), [1.4] – 2.1 (females) × carapace length; subcylindrical, spiny and with long stiff setae and iridescent setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus [0.5] – 0.9 length of carapace, 1.4 – [1.6] × as long as carpus. Carpus [1.6] – 2.3 × as long as wide. Palm [1.1] 1.0 – 1.2 × carpus length, [1.7] 1.6 – 2.2 × as long as broad. Fingers [0.7] – 0.8 × palm length; fixed finger with small basal spine; movable finger unarmed. P 2 – 4 (P 4 lost in the holotype): Stout, setose, with iridiscent setae, rugose and spinose. Meri successively shorter posteriorly: P 3 merus [0.9] – 1.0 × length of P 2 merus, P 4 merus 0.8 – 0.9 × length of P 3 merus. P 2 merus, 0.5 – [0.6] × carapace length, [3.2] – 3.7 × as long as broad, [1.3] 1.2 – 1.5 × as long as P 2 propodus; P 3 merus 2.5 – [3.2] × as long as broad, [1.1] – 1.4 × as long as P 3 propodus; P 4 merus 2.3 – 2.8 × as long as broad, 0.9 – 1.4 × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, unarmed; flexor margin irregular, with distal spine on P 2 – 3, distal spine absent in P 4. Carpi with 1 – 3 spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, smaller on P 4; granules below extensor margin on lateral surface of P 2 – 4; flexor margin unarmed. Propodi stout, [4.2 – 4.7] 3.3 – 5.0 × as long as broad; extensor margin irregular, armed with 2 – 4 spines; flexor margin with 3 – 6 slender movable spines in addition to distal pair. Dactyli [0.6] – 0.8 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 5 – 6 movable spines. Eggs: Ov. F carried approximately 7 – 35 eggs of 0.3 – 0.5 mm diameter. Live colour. Body brownish or dark green with small dark brown spots. P 1 – 4 whitish, with dark brown spots and stripes. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFA3E65E4F9CFD9C7348BDA9.taxon	distribution	Distribution. Japan, Papua New Guinea, Vanuatu, New Caledonia, Guam Island, Northern Mariana Islands and American Samoa. Intertidal to 38 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFA3E65E4F9CFD9C7348BDA9.taxon	discussion	Remarks. Phylladiorhynchus laureae belongs to the group of species having 2 epigastric spines, 1 hepatic spine and 3 spines on anterior branchial margin. Phylladiorhynchus laureae closely resembles to P. gustavi from French Polynesia. Both species are very similar morphologically, although very different genetically. Both species can be distinguished by the following subtle characters: - The anterior metagastric ridge is often scale-like in P. gustavi, whereas it is continuous (holotype) or medially interrupted in P. laureae. - The metagastric ridge is followed by a large scale in P. laureae, whereas this scale is very small or absent in P. gustavi. - The anterior margin of thoracic sternite 3 is medially projected in P. laureae, whereas it is usually convex in P. gustavi. Specimens of P. laureae range from 1.1 to 2.5 mm postorbital carapace length. The sequences of P. laureae (COI) were 8 % divergent from P. gustavi. The five sequences of P. laureae from Guam, Japan, New Caledonia and Papua New Guinea diverged 1.8 %% for COI and 0.6 % for 16 S.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFA6E65C4F9CFB4373B6BE52.taxon	description	(Fig. 28)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFA6E65C4F9CFB4373B6BE52.taxon	materials_examined	Type material. Lectotype. Chile, Corral, Valdivia: ov. F 2.7 mm (USNM 32261). Paralectotype. Chile, Corral, Valdivia: 1 ov. F 2.6 mm (USNM 32261).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFA6E65C4F9CFB4373B6BE52.taxon	description	Description. Carapace: Slightly wider than long. Gastric region slightly convex with 4 transverse ridges: epigastric ridge distinct with 2 median pairs of spines symmetrically distant of median area; anterior protogastric ridge not medially interrupted, nearly extending laterally to carapace margin; anterior mesogastric ridge not medially interrupted, laterally interrupted by cervical groove, laterally continuing uninterrupted to first branchial spine; anterior metagastric ridge scale-like. Mid-transverse ridge uninterrupted, preceded by shallow or undistinct cervical groove, followed by 2 uninterrupted or minutely interrupted ridges, interspersed with 1 short lateral ridge. Lateral margins convex, with 7 spines: first anterolateral spine well-developed, reaching anteriorly to level of lateral orbital spine, second spine (hepatic) small, slightly dorsomesially from lateral margin, and followed by 5 branchial spines behind distinct anterior cervical groove (3 anterior and 2 posterior). Rostrum leaf-like, horizontal, dorsally flattish or slightly concave, 1.3 × as long as broad, length 0.3 and breadth 0.2 that of carapace; lateral margins smooth and convex, with well-developed supraocular basal spines and small subapical spines. Pterygostomian flap ending in blunt tooth, upper margin smooth. Thoracic sternum: As wide as long. Sternite 3 moderately broad, twice as wide as long, anterior margin convex, anterolaterally convex. Sternite 4 widely contiguous to sternite 3; surface depressed in midline, smooth; greatest width 3 × that of sternite 3, 3.2 × as wide as long. Pleon: Tergite 2 with anterior and posterior transverse elevated ridges; tergites 3 – 4 with anterior transverse ridge only; tergites 5 – 6 smooth. Eye: Eyestalk length about 1.2 × broader than long, peduncle distally setose, not distinctly expanded proximally; maximum corneal diameter 1.2 × rostrum width, as wide as eyestalk (as wide as maximum peduncle width). Antennule: Article 1 slightly longer than wide, with 4 well-developed distal spines: distomesial spine welldeveloped; proximal lateral spine absent or present as a granule. Antenna: Article 1 with prominent mesial process distally falling well short of lateral antennular spine. Article 2 and 3 with well-developed distomesial and distolateral spines. Article 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus 0.7 – 0.8 × length of ischium, with well-developed distal spine on extensor margin and 1 much larger spine at flexor margin. P 1: 3 × carapace length; subcylindrical, spiny; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus 1.1 length of carapace, 1.8 × as long as carpus. Carpus 1.8 × as long as wide. Palm 1.2 × carpus length, 1.8 × as long as broad. Fingers 0.8 × palm length; some marginal spines along proximal half of movable and fixed fingers. P 2 (presumably, other walking legs lost): Stout, moderately setose and spinose. Merus, 0.7 × carapace length, 3.7 × as long as broad, 1.5 × as long as propodus; extensor margin with row of spines, proximally diminishing, with prominent distal spine; flexor margin irregular, with distal spine; lateral surface with scales. Carpus with 3 spines on extensor margin; row of small acute granules below extensor margin on lateral surface; flexor margin unarmed other than distal spine. Propodi stout, 3.8 × as long as broad; extensor margin irregular unarmed; flexor margin with 3 slender movable spines in addition to distal pair. Dactyli 0.6 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 4 movable spines. Live colour. Unknown. Genetic data. No data.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFA6E65C4F9CFB4373B6BE52.taxon	distribution	Distribution. Only known from Corral, Valdivia (Chile), unknown depth.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFA6E65C4F9CFB4373B6BE52.taxon	discussion	Remarks. Phylladiorhynchus lenzi was described by Rathbun (1907) from 4 specimens collected by C. E. Porter in waters of Corral, Valdivia Province (Chile), with a very short diagnosis and one photo. The species was later reported in other localities along the coast (e. g. Concepcion area), as well as in the oceanic islands (e. g. Juan Fernandez, Salas y Gomez) (see above). However, until now, no complete description of the species existed, avoiding a comparison among the specimens from different localities as well as with other species. Furthermore, when examining specimens from different Chilean localities, e. g. Corral (type specimens) and Valparaiso (coll. Porter, 1899, deposited in the MNHN of Paris) we found that they belong to two different species (P. lenzi and P. porteri). Unfortunately, we failed to examine specimens from Chilean oceanic islands, so the identity of the specimens of Phylladiorhynchus reported from these islands remains to be confirmed. Phylladiorhynchus lenzi has been considered a synonym of P. pusillus (Henderson, 1885) (Baba et al. 2008; Schnabel & Ahyong 2019). Phylladiorhynchus lenzi belongs to the group of species having usually 4 spines on the epigastric ridge, the anterior metagastric ridge scale-like, a very small hepatic spine, the anterior margin of the thoracic sternite 3 convex and one spine on the flexor margin of the Mxp 3 merus. This group contains four species: P. pusillus from New Zealand and Australia, P. lenzi, from Chile, P. porteri, from Chile, and P. poeas from French Polynesia. Differences between P lenzi and P. porteri are given under the Remarks of the latter species. Phylladiorhynchus lenzi can be distinguished from P. pusillus and P. poeas by subtle but constant differences: - The proximal lateral spine of the antennular article is always distinct in P. pusillus, whereas this spine is very small to indistinct in P. lenzi and P. poeas. - The antennal article 3 is armed with a small distomesial spine in P. pusillus, whereas this article has welldeveloped distomesial and distolateral spines in P. lenzi and P. poeas.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFA5E6404F9CFF2971BEB896.taxon	description	(Figs. 29, 30 J – L, S – T, 33 B, 55 A, 56 D – I)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFA5E6404F9CFF2971BEB896.taxon	materials_examined	Type material. Holotype. Taiwan, Kenting County, Houbihu, 21.936361 ° N, 120.746806 ° E, 7 – 10 m, 2 June 2019: F 2.2 mm (NMMBCD 5596). Paratypes. Taiwan. Kenting County. Small fishing port S of Wanlitung. 21.9706 ° N 120.7156 ° E, 5 – 8 m, 30 June 2007: 1 ov. F 2.6 (UF 11812). — Kenting County, National Museum of Marine Biology & Aquarium, Water outlet, 21.9319 ° N, 120.7344 ° E, 8 – 10 m, 23 May 2011: 1 M 3.4 mm (NMMBCD). — 21.9319 ° N, 120.7344 ° E, 8 – 10 m, 24 May 2011: 1 ov. F 3.0 mm, 1 F 3.2 mm (NMMBCD). — 21.9230 ° N, 120.8319 ° E, 7 – 10 m, 30 May 2011: 9 M 2.6 – 3.7 mm, 2 F 2.9 – 3.2 mm, 9 ov. F 1.7 – 3.1 mm (NMMBCD). — 21.9319 ° N, 120.7447 ° E, 8 – 10 m, 25 July 2011: 2 M 3.0 – 3.1 mm, 1 ov. F 2.1 mm (NMMBCD). — 21.9319 ° N, 120.7447 ° E, 8 – 10 m, 5 October 2011: 2 F 2.5 – 3.1 mm (NMMBCD). — Houbihu, 21.9361 ° N, 120.7466 ° E, 7 – 10 m, 2 June 2019: 1 M 1.7 mm, 1 ov. F 2.5 mm, 1 F 1.4 mm (MNHN- IU- 2016 - 1490). Other material. Kenting County, National Museum of Marine Biology & Aquarium, Water outlet, 21.9319 ° N, 120.7447 ° E, 8 – 10 m, 13 October 2011: 1 M 2.2 mm (NMMBCD). — 21.9319 ° N, 120.7447 ° E, 8 – 10 m, 19 November 2011: 1 F 2.6 mm (NMMBCD). — 21.9319 ° N, 120.7447 ° E, 8 – 10 m, 3 May 2012: 1 ov. F 3.0 mm (NMMBCD). — Houbihu, 21.9361 ° N, 120.7466 ° E, 7 – 10 m, 14 May 2012: 1 ov. F 2.9 mm (NMMBCD). — Houbihu, 21.9361 ° N, 120.7466 ° E, 7 – 10 m, 29 June 2013: 1 ov. F 2.5 mm (NMMBCD). — Hojie, 21.9553 ° N, 120.7116 ° E, 24 – 26 m, 26 February 2013: 1 ov. F 2.7 mm (NMMBCD).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFA5E6404F9CFF2971BEB896.taxon	etymology	Etymology. The species is named after our colleague Chia-Wei Lin, from the National museum of marine biology & aquarium, Pingtung, Taiwan, who collected most of the specimens.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFA5E6404F9CFF2971BEB896.taxon	description	Description. Carapace: As long as or slightly longer than broad; transverse ridges with dense short setae and some thick setae. Gastric region slightly convex with some transverse ridges: epigastric ridge distinct with 2 median spines and some lateral short scales, followed by small short scales on posterior epigastric region; anterior protogastric ridge not medially interrupted, nearly extending laterally to carapace margin, often followed by uninterrupted posterior protogastric ridge or short scales; anterior mesogastric ridge not medially interrupted, laterally interrupted by anterior branch of cervical groove, and continuing uninterrupted to first branchial spine; anterior metagastric ridge not medially interrupted, followed by posterior scale-like metagastric ridge. Mid-transverse ridge not interrupted, medially depressed, followed by shallow or indistinct cervical groove, followed by 2 – 3 not interrupted or minutely interrupted ridges, interspersed with 2 interrupted riges and few short scales. Lateral margins slightly convex, with 6 distinct spines: first anterolateral spine well-developed, reaching anteriorly to level of lateral orbital spine, second spine (hepatic) well-developed, slightly dorsomesially from lateral margin, and followed by 4 branchial spines (3 anterior and 1 posterior). Rostrum dagger-like, horizontal dorsally flattish or slighly concave [1.5] – 1.8 × as long as broad, length [0.4] – 0.5 and breadth 0.2 – [0.3] that of carapace; lateral margins smooth and straight or nearly straight, with well-developed supraocular basal spines and subapical spines. Pterygostomian flap with anterior spine, upper margin smooth. Thoracic sternum: As wide as long. Sternite 3 moderately broad, [1.9] – 2.3 × as wide as long, anterior margin convex, with a blunted median projection, anterolaterally rounded. Sternite 4 widely contiguous to sternite 3; anterolaterally smooth, surface depressed in midline, smooth; greatest width [2.5] – 2.9 × that of sternite 3, 2.7 – [3.5] × as wide as long. Pleon: Elevated ridges with short setae and a few scattered long setae. Tergite 2 – 4 with anterior and posterior transverse elevated ridges; tergites 5 – 6 smooth. Eye: Eyestalk length about 0.9 – [1.1] × broader than long, peduncle distally setose, not distinctly expanded proximally, with few short transverse striae on lateral surfaces; cornea expanded distally, maximum corneal diameter [0.9] – 1.0 × rostrum width, as wide as eyestalk. Antennule: Article 1 longer than wide, with 5 distal spines: distomesial spine well-developed; proximal lateral spine small, always present. Antenna: Article 1 with prominent mesial process, distally falling well short of lateralmost antennular spine. Article 2 with well-developed distomesial and distolateral spines. Article 3 unarmed or with small distomesial spine. Article 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus [0.6] – 0.7 × length of ischium, with well-developed distal spine on extensor and flexor margins. P 1: Slender, 3.0 (males), 2.0 – [2.3] (females) × carapace length; subcylindrical, spiny and with long stiff setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus 0.7 – [0.9] length of carapace, 1.5 – [1.8] × as long as carpus. Carpus [2.5] – 3.1 × as long as wide. Palm 1.1 × carpus length, 2.3 – [2.7] × as long as broad. Fingers 0.7 × palm length; fixed finger with 0 – 1 basal spines; movable finger with 1 basal spine. P 2 – 4: setose and spinose. Meri successively shorter posteriorly: P 3 merus 0.9 × length of P 2 merus, P 4 merus [0.8] – 0.9 × length of P 3 merus. P 2 merus, 1.5 – [1.8] × carapace length, [3.4] – 4.4 × as long as broad, [1.1] 1.1 × as long as P 2 propodus; P 3 merus [3.6] – 3.7 × as long as broad, 1.0 – [1.2] × as long as P 3 propodus; P 4 merus [3.2] – 3.7 × as long as broad, 1.0 – [1.1] × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, unarmed; flexor margin irregular, with distal spine on P 2 – 3, absent or small on P 4. Carpi with 2 – 4 small spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, absent on P 4; row of granules below extensor margin on lateral surface of P 2 – 4; flexor margin unarmed. Propodi stout, 4.0 – 5.0 × as long as broad; extensor margin irregular; flexor margin with 3 – 6 slender movable spines in addition to distal pair. Dactyli [0.6 – 0.7] 0.5 – 0.7 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with cuticular spines at basis of 5 – 6 movable spines. Eggs: Ov. F carried approximately 4 – 15 eggs of 0.5 mm diameter. Live colour. Variable colour pattern. Body dark to light brown, sometimes orange or with large whitish spots and bands. P 1 – 4 with dusky and whitish bands. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFA5E6404F9CFF2971BEB896.taxon	distribution	Distribution. Taiwan, Kenting, between 5 and 26 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFA5E6404F9CFF2971BEB896.taxon	discussion	Remarks. Phylladiorhynchus lini belongs to the group of species having 2 epigastric spines, 1 hepatic spine, 3 spines on the anterior branchial margin, rostrum dagger-like (margin straights or nearly straight) and dactylar spines along the flexor margin of the P 2 – 4 dactyli. The new species is closely related to P. spinosus Schnabel & Ahyong, 2019, from New Zealand and Australia (see the differences under the Remarks of P. spinosus). The specimens of P. lini range from 1.4 to 3.4 mm postorbital carapace length. The two sequences of P. lini from Taiwan diverged 0.6 % (COI) and 0.2 % (16 S).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB8E6464F9CFE64710DBF4E.taxon	description	(Figs. 31, 33 C, 37 E – F)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB8E6464F9CFE64710DBF4E.taxon	materials_examined	Type material. Holotype. Chagos Archipelago, Great Chagos Bank, Brothers Island. Stn CH 0614 dead branching coral heads. Outer Reef Slope, 8 – 12 m, February 2012: ov. F 2.0 mm (MNHN-IU- 2019 - 2593) Paratypes. Chagos Archipelago, Great Chagos Bank, Eagle Island. Stn CH 1364 dead branching coral heads. Outer Reef Slope, 8 – 12 m, February 2012: 1 ov. F 1.4 mm, 1 F 1.3 mm (MNHN-IU- 2019 - 2594). — Stn CH 0447, 1 M 1.7 mm (OUMNH. ZC. 2014 - 09 - 064). — Stn CH 0230, 1 M 1.2 mm (OUMNH. ZC. 2014 - 09 - 065). Kiribati. Line Islands. 9.91 ° S, 150.21 ° W, 11 m, 8 November 2013: 1 M 1.9 mm (UF 41803). — Millenium island. 9.91 ° S, 150.21 ° W, 12 m, 5 November 2013: 1 M 1.8 mm 1 ov. F 1.3 mm (UF 40895). Other material. American Samoa, Olosega Island. Stn OLO- 187, 14.18140174 ° S, 169.6267462 ° W, 14 m, 14 March 2015: 1 M 1.8 mm (UF 54480). — Stn OLO- 094, 14.18140174 ° S, 169.6267462 ° W, 14 m, 14 March 2015: 1 ov. F 2.1 mm (UF 54481). — Rose Atoll. Stn ROS- 610, 14.54895 ° S, 168.13792 ° W, 15 m, 17 March 2015: 8 M 1.4 – 2.1 mm, 8 ov. F 1.5 – 2.3 mm (UF 54482). — Stn ROS- 527, 14.54895 ° S, 168.13792 ° W, 15 m, 17 March 2015: 1 M 1.9 mm, 1 ov. F 1.8 mm (UF 54483). — Stn ROS- 562, 14.54895 ° S, 168.13792 ° W, 15 m, 17 March 2015: 10 M 1.4 – 2.1 mm, 13 ov. F 1.4 – 2.1 mm, 2 F 1.5 – 1.8 mm (UF 54484). — Ofu Island. Stn OFU- 121, 14.17765914 ° S, 169.649504 ° W, 14 m, 21 March 2015: 1 ov. F 2.3 mm (UF 54477). — Stn OFU- 168, 14.17765914 ° S, 169.649504 ° W, 14 m, 21 March 2015: 1 M 1.9 mm, 1 ov. F, 2.2 mm (UF 54478). — Stn OFU- 280, 14.18628 ° S, 169.6599 ° W, 14 m, 26 March 2015: 1 M 1.8 mm, 1 ov. F 1.9 mm (UF 54479). Kiribati. Phoenix islands, Orona island. 4.519 ° S 172.227 ° W, 12 m, 17 September 2015: 1 ov. F 1.7 mm (UF 51296). — 4.520 ° S, 172.230 ° W, 15 m, 18 September 2015: 1 M 1.5 mm (UF 51370), 2 M 1.8 – 1.9 mm 1 ov. F 1.8 mm (UF 51363). — Nikumaroro Island. 4.656 ° S, 174.545 ° W, 14 m, 23 September 2015: 1 ov. F 1.7 mm (UF 51559). — 4.694 ° S 174.490 ° E, 7.5 – 9 m, 25 September 2015: 1 M 1.3 mm (UF 51644). Western Australia. Hibernia Reef. Stn 142 / K 13, 11 ° 59.292 ’ S, 123 ° 21.154 ’ E, no depth, 4 October 2013: 1 M 2.1 mm (WAM C 55689). — Rowley Shoals. Mermaid Reef. Stn 178 / K 14 - T 1, 17 ° 09.69 ’ S, 119 ° 38.826 ’ E, 12 – 18 m, 13 October 2014: 1 ov. F 1.6 mm (WAM C 53889).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB8E6464F9CFE64710DBF4E.taxon	etymology	Etymology. From the name Lynceus, an Argonaut, son of Aphareus and Arene. The name is considered a substantive in apposition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB8E6464F9CFE64710DBF4E.taxon	description	Description. Carapace: As long as or slightly longer than broad; transverse ridges with dense short setae and few scattered long and thick iridiscent setae. Gastric region slightly convex with 4 transverse ridges: epigastric ridge distinct with 2 spines (Samoa and Kiribati specimens often with 1 median produced scale usually with thick plumose setae before epigastric ridge), short scales laterally; anterior protogastric ridge not medially interrupted, nearly extending laterally to carapace margin; anterior mesogastric ridge not medially interrupted, laterally continuing to first branchial spine; sometimes followed by some short small scales; anterior metagastric ridge medially interrupted, laterally continuing to second branchial spine. Mid-transverse ridge not interrupted, cervical groove indistinct, followed by 2 not interrupted or minutely interrupted ridges, interspersed with 1 short lateral ridge and sometimes few, short scattered scales. Lateral margins slightly convex, with 4 distinct spines: first anterolateral spine welldeveloped, reaching anteriorly to level of lateral orbital spine, hepatic margin unarmed; anterolateral spine followed by 3 branchial spines (2 anterior and 1 posterior spines). Rostrum leaf-like, horizontal, dorsally concave, [1.5] – 1.6 × as long as broad, length [0.3] – 0.4 and breadth 0.2 – [0.3] that of carapace; lateral margins serrated and convex, with well-developed supraocular basal spines and small subapical spines. Pterygostomian flap ending in blunt tooth, upper margin smooth. Thoracic sternum: As wide as long. Sternite 3 quadrangular, slightly wider than long (1.5 × as wide as long), anterior margin with median blunted projection, lateral margins rounded. Sternite 4 widely contiguous to sternite 3; surface depressed in midline, smooth; greatest width 3.0 × that of sternite 3, [2.7] 2.5 – 3.0 × as wide as long. Pleon: Elevated ridges with short setae and a few scattered long setae. Tergite 2 with anterior and posterior transverse elevated ridges; tergites 3 – 4 with anterior transverse ridge; tergites 5 – 6 smooth. Eye: Eyestalk length about 0.9 – [1.0] × broader than long, peduncle distally setose, slightly expanded proximally; maximum corneal diameter 0.8 – [0.9] × rostrum width, narrower than eyestalk (0.8 × maximum peduncle width). Antennule: Article 1 slightly longer than wide, with 5 distal spines: distomesial spine small; proximal lateral spine small, always present. Antenna: Article 1 with prominent mesial process, distally falling well short of lateralmost antennular spine. Article 2 with well-developed distomesial and distolateral spines. Article 3 with distomesial spine. Article 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus 0.6 – [0.8] × length of ischium, with distal spine on extensor and flexor margins. P 1: 1.7 – 2.0 (males), 2.0 (females) × carapace length; subcylindrical, spiny and with dense long stiff setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus 0.7 – 0.8 length of carapace, 2.0 – 2.1 × as long as carpus. Carpus 1.2 – 1.3 × as long as wide. Palm 1.2 – 1.3 × carpus length, 1.5 – 1.7 × as long as broad. Fingers unarmed, 0.6 – 0.9 × palm length. P 2 – 4: Stout, setose and spinose. Meri successively shorter posteriorly: P 3 merus 0.7 – [0.8] × length of P 2 merus, P 4 merus 0.8 × length of P 3 merus. P 2 merus, 0.7 – [0.8] × carapace length, [3.6] – 3.8 × as long as broad, [1.2] – 1.3 × as long as P 2 propodus; P 3 merus 3.7 × as long as broad, [1.0] – 1.1 × as long as P 3 propodus; P 4 merus [2.8] 2.7 – 3.2 × as long as broad, [1.0] – 1.1 × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, unarmed; flexor margin irregular, with distal spine on P 2 – 3, unarmed on P 4. Carpi with 1 – 3 prominent spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, smaller on P 4; row of small spines or granules below extensor margin on lateral surface of P 2 – 3, unarmed on P 4; flexor margin unarmed. Propodi stout, [3.8] 3.3 – 4.3 × as long as broad; extensor margin irregular, armed with 1 – 3 spines on proximal half or unarmed; flexor margin with 3 – 4 slender movable spines in addition to distal pair. Dactyli 0.7 – [0.8] × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 4 – 6 movable spines. Eggs: Ov. F carried approximately 5 – 25 eggs of 0.3 – 0.4 mm diameter. Live colour. Unknown. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB8E6464F9CFE64710DBF4E.taxon	distribution	Distribution. Chagos Archipelago, Western Australia, Kiribati, American Samoa, between 7.5 and 18 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB8E6464F9CFE64710DBF4E.taxon	discussion	Remarks. Phylladiorhynchus lynceus belongs to the group of species with 2 epigastric spines and 2 spines on the anterior branchial margin. The closest species are: P. integrirostris, from Hawaii and P. priasus from the Mariana and Marshall Islands. The three species are barely distinguishable morphologically, although they are genetically very different (see the differences under the Remarks of P. priasus). Furthermore, these three species are also close to P. orpheus (see the differences under the Remarks of P. orpheus). The specimens of P. lynceus range from 1.2 to 2.3 mm postorbital carapace length. The four and seven sequences of P. lynceus from American Samoa and Chagos and Kiribati diverged 1.0 – 2.3 % (COI) and 0 – 1 % (16 S), respectively.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFBEE6454F9CF91C7342BB9A.taxon	description	(Fig. 32) Phylladiorhynchus ikedai. — Baba, 1991, 485 (in part, only some specimens from CALSUB Pl 16).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFBEE6454F9CF91C7342BB9A.taxon	materials_examined	Type material. Holotype. Chesterfield Islands, KANADEEP Stn DW 4962, 23.0333 ° S, 159.4666 ° E, 315 – 1260 m, 6 September 2017: M 2.3 mm (MNHN-IU- 2017 - 2736) Paratypes. New Caledonia. CALSUB Pl 16, 20.6300 ° S, 167.0450 ° E, 500 m, 7 March 1989: 1 M broken (MNHN- IU- 2013 - 19942), 1 M 1.8 mm (MNHN-IU- 2013 - 19943). — KANACONO Stn CP 4684, 22.4000 ° S, 167.4000 ° E, 493 – 694 m, 14 August 2016: 1 M 1.7 mm (MNHN-IU- 2017 - 11683). — Stn DW 4749, 23.6333 ° S, 167.7333 ° E, 440 – 457 m, 24 August 2016: 1 ov. F 3.1 mm (MNHN-IU- 2017 - 11662).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFBEE6454F9CF91C7342BB9A.taxon	etymology	Etymology. Named after Philippe Maestrati, curatorial assistant of the general mollusc collection of the Muséum national d’Histoire naturelle, Paris.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFBEE6454F9CF91C7342BB9A.taxon	description	Description. Carapace: 1.1 – [1.2] × as long as broad; transverse ridges with dense short setae and scattered long setae. Gastric region flattened with 4 transverse ridges: epigastric ridge indistinct, with 4 spines (2 pairs of spines laterally, sometimes with median granule); anterior protogastric ridge not medially interrupted, nearly extending laterally to carapace margin; anterior mesogastric ridge not medially interrupted, or minutely interrupted, laterally interrupted by anterior branch of cervical groove; anterior metagastric ridge scale-like. Mid-transverse ridge not interrupted, preceded by shallow cervical groove, followed by 2 uninterrupted or minutely interrupted ridges, interspersed with 1 – 2 short lateral ridges and occasionally with few, short scattered scales (absent in holotype). Lateral margins straight or slightly convex, with 7 spines: first anterolateral spine well-developed, reaching or slightly exceeding level of lateral orbital spine, second spine (hepatic) small, slightly dorsomesially from lateral margin, and followed by 5 branchial spines (3 anterior and 2 posterior). Rostrum dagger-like, horizontal, dorsally slightly or deeply concave, 2.0 – [2.1] × as long as broad, length 0.4 – [0.5] and breadth 0.3 that of carapace; lateral margins smooth, convex, with well-developed supraocular spines, subapical spines distinct. Pterygostomian flap ending in blunt tooth, upper margin smooth. Thoracic sternum: As wide as long. Sternite 3 moderately broad 2.0 – [2.3] × as wide as long anterior margin convex. Sternite 4 narrowly contiguous to sternite 3; surface depressed in midline, smooth; greatest width 3 × that of sternite 3, 2.5 – [3.0] × as wide as long. Pleon: Elevated ridges with short setae and a few scattered long setae Tergite 2 with anterior and posterior transverse elevated ridges; tergites 3 – 4 with anterior transverse ridges only; tergites 5 – 6 smooth. Eye: Eyestalk as broad as long, peduncle distally setose, not distinctly expanded proximally, maximum corneal diameter 0.9 – [1.0] × rostrum width, as wide as eyestalk. Antennule: Article 1 1.3 × longer than wide, with 5 distal spines, distomesial spine well-developed; proximal lateral spine small. Antenna: Article 1 with prominent mesial process distally clearly not reaching lateralmost antennular spine. Article 2 unarmed, sometimes with minute distolateral spine. Articles 3 and 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus half length of ischium, extensor margin with distal spine, flexor margin with 2 strong spines. P 1: [2.9] × carapace length (males), 2.8 (female), subcylindrical, spiny and with scattered long stiff setae and dense short thick setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, mesial spines usually stronger than others. Merus [1.1] 1.1 length of carapace, [2.1] – 2.3 × as long as carpus. Carpus [2.4] – 2.5 × as long as wide. Palm 1.4 × carpus length, [2.9] – 3.4 × as long as broad. Fingers unarmed, 0.7 × palm length. P 2 – 4 (lost in holotype): Slender, subcylindrical, moderately setose and spinose. Meri successively shorter posteriorly: P 3 merus 0.7 × length of P 2 merus, P 4 merus 0.9 × length of P 3 merus. P 2 merus, 0.8 × carapace length, 9 × as long as broad, 1.3 × as long as P 2 propodus; P 3 merus 5 × as long as broad, 1.3 × as long as P 3 propodus; P 4 merus 4 × as long as broad, 1.2 × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, unarmed; flexor margins of P 2 – 4 irregular; P 4 lateral surface with 4 small spines. Carpi extensor margin with 8 – 9 small spines on extensor margin on P 2 – 3; distal spine prominent on P 2, smaller or absent in P 3 – 4; flexor margin unarmed. Propodi moderately slender, 6.3 – 7.0 × as long as broad; extensor margin irregular, usually serrated, with 7 – 10 small spines; flexor margins with 2 – 3 slender movable spines in addition to distal pair. Dactyli 0.5 – 0.6 × length of propodi, ending in incurved, strong, sharp spine; flexor margins with 8 – 10 movable spines. Eggs: no data. Live colour. Unknown. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFBEE6454F9CF91C7342BB9A.taxon	distribution	Distribution. New Caledonia, Chesterfield Islands, from 315 to 1260 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFBEE6454F9CF91C7342BB9A.taxon	discussion	Remarks. Phylladiorhynchus maestratii belongs to the group of species having the epigastric ridge with 4 spines and often one median process or scale with thick plumose setae (rarely with 5 spines), 3 spines on the anterior branchial margin of the carapace and the Mxp 3 merus with two prominent spines along the flexor margin. Phylladiorhynchus maestratii is close to P. cepheus from French Polynesia and P. butes, from New Caledonia, Chesterfield Islands, Indonesia and French Polynesia, however these species can be distinguished on basis of the following characters: - The median epigastric process, spine or scale with thick setae is always present in P. cepheus and P. butes, whereas this process is absent or is present as a simple granule in P. maestratii. - The rostrum is less than twice as long as wide in P. cepheus and P. butes, whereas it is more than twice as long as wide in P. maestratii. - Thoracic sternite 4 is widely contiguous to sternite 3 in P. cepheus and P. butes, whereas it is narrowly contiguous in P. maestratii. - The P 2 – 4 dactyli flexor margins have 8 – 10 movable spines in P. maestratii, whereas these margins have 6 – 7 movable spines in P. cepheus and 5 – 8 in P. butes. - Antennal article 2 is armed distally with well-developed mesial and lateral spines in P. butes rather than unarmed, at most with small distomesial spine or granule in P. cepheus and P. maestratii. The genetic divergences among these species were larger than 7 % (COI) and 5 % (16 S). The specimens of P. maestratii range from 1.6 to 3.1 mm postorbital carapace length.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFBDE64A4F9CFD68734BBE52.taxon	description	(Figs. 33 D, 34, 55 B)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFBDE64A4F9CFD68734BBE52.taxon	materials_examined	Type material. Holotype. Vanuatu, SANTO Stn DB 14, 15.5150 ° S, 167.18333 ° E, 10 – 14 m, 13 October 2006: ov. F 2.0 mm (MNHN-IU- 2014 - 13887). Paratypes. Vanuatu. SANTO Stn DS 6 light, 15.5150 ° S, 167.18333 ° E, 8 – 15 m, 11 September 2006: 1 M 1.4 mm, 3 ov. F 1.7 – 1.8 mm (MNHN-IU- 2014 - 13893). ― Stn FR 2, 15.5550 ° S, 167.1466 ° E, 1 – 25 m, 11 September 2006: 1 F 1.8 mm (MNHN-IU- 2014 - 13889). ― Stn DB 16, 15.591667 ° S, 167.2633 ° E, 32 – 40 m, 14 September 2006: 1 M 1.5 mm (MNHN-IU- 2014 - 13891). ― Stn DB 40, 15.4966 ° S, 167.2516 ° E, 5 m, 19 September 2006: 3 M 1.5 – 1.6 mm (MNHN-IU- 2019 - 2646), 1 ov. F 1.5 mm (MNHN-IU- 2016 - 453), 1 F 1.6 mm (MNHN-IU- 2014 - 13862). Other material. Vanuatu. SANTO Stn DB 46, 15.4800 ° S, 167.2533 ° E, 2 – 3 m, 20 September 2006: 1 F 1.7 mm (MNHN-IU- 2019 - 2644). ― Stn DB 53, 15.4800 ° S, 167.2533 ° E, 5 m, 22 September 2006: 5 M 1.4 – 1.8 mm, 11 ov. F 1.2 – 1.8 mm, 5 F 1.3 – 2.1 mm (MNHN-IU- 2019 - 2652), 8 M 1.2 – 1.5 mm, 3 ov. F 1.7 – 2.0 mm, 1 F 1.4 mm (MNHN-IU- 2019 - 2645). ― Stn AT 42, 15.6250 ° S, 167.0383 ° E, 112 - 148 m, 28 September 2006: 1 M 1.4 mm (MNHN-IU- 2014 - 13895). ― Stn NS 36, 15.5283 ° S, 167.1583 ° E, 2 – 3 m, 2 October 2006: 1 ov. F 1.5 mm (MNHN-IU- 2014 - 13806). ― Stn FB 40, 15.3816 ° S, 167.1950 ° E, 9 m, 29 September 2006: 1 postlarvae 1.6 mm (MNHN-IU- 2014 - 13876). ― Stn DB 83, 15.7233 ° S, 167.2500 ° E, 6 m, 3 October 2006: 1 M 1.4 mm (MNHN- IU- 2014 - 13892). ― Stn DS 91, 15.5616 ° S, 167.1400 ° E, 7 m, 6 October 2006: 2 M 1.4 – 1.5 mm, 1 ov. F 2.3 mm (MNHN-IU- 2014 - 13888). ― Stn FB 56, 15.5866 ° S, 167.0350 ° E, 3 – 18 m, 7 October 2006: 1 ov. F 1.6 mm (MNHN- IU- 2014 - 13896). ― Stn EP 34, 15.5550 ° S, 167.2150 ° E, 40 – 60 m, 14 October 2006: 1 ov. F 1.5 mm (MNHN-IU- 2019 - 2647). ― Stn EP 36, 15.5550 ° S, 167.3226 ° E, 20 – 60 m, 14 October 2006: 2 M 1.4 – 2.2 mm, 1 ov. F 1.9 mm (MNHN-IU- 2019 - 2650), 4 M 1.4 – 1.8 mm, 3 ov. F 1.9 – 2.2 mm, 1 F 1.4 mm (MNHN-IU- 2019 - 2649), 1 M 1.5 mm, 1 ov. F 2.0 mm, 4 F 1.6 – 2.3 mm (MNHN-IU- 2019 - 2653). ― Stn LD 31, 15.5716 ° S, 167.1983 ° E, 1 – 3 m, 14 October 2006: 2 ov. F 2.4 – 2.6 mm (MNHN-IU- 2014 - 13890). ― Stn FB 80, 15.5516 ° S, 167.1600 ° E, 2 m, 14 October 2006: 1 F 1.7 mm (MNHN-IU- 2014 - 13894). ― Stn LD 35, 15.5466 ° S, 167.1933 ° E, 3 – 8 m, 16 October 2006: 2 M 1.4 - 2.3 mm, 3 ov. F 1.3 – 2.1 mm, 1 F 2.2 mm (MNHN-IU- 2014 - 13897). ― SE corner of Espiritu Santo Island (mixed illegible or confused labels) Sep / Oct 2006: 5 F 1.7 – 2.1 mm (MNHN-IU- 2019 - 2648).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFBDE64A4F9CFD68734BBE52.taxon	etymology	Etymology. Named in memorian of Marina Alcobendas, from the Museo Nacional de Ciencias Naturales, Madrid, evolutionary biologist and dear colleague. The name is considered as a substantive in apposition	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFBDE64A4F9CFD68734BBE52.taxon	description	Description. Carapace: Robust or massive, sexually dimorphic (wider on females), 0.9 – 1.2 (males), [0.8] – 0.9 (females) × as long as broad; transverse ridges elevated, upraised dorsally, with few short setae and scattered long thick plumose setae. Gastric region convex (upraised) with some transverse ridges: epigastric ridge unarmed, scalelike, often undistinct; anterior protogastric ridge not medially interrupted, laterally interrupted with few scales, often followed by few short scales; anterior mesogastric ridge scale-like, laterally interrupted by anterior branch of cervical groove, continuing with few scales, often followed by few short scales; anterior metagastric ridge medially uninterrupted, followed by few scales on posterior metagastric region. Mid-transverse ridge not interrupted, cervical groove distinct, followed by 2 not interrupted or minutely interrupted ridges, interspersed with 1 laterally interrupted or scale-like ridge and 1 – 2 short lateral ridges. Lateral margins clearly convex, with 7 – 8 spines: first anterolateral spine well-developed, exceeding lateral orbital spine, second spine (hepatic) minute, often obsolescent in males, slightly dorsomesially from lateral margin, and followed by 5 – 6 branchial spines (3 anterior and 2 – 3 posterior). Rostrum leaf-like, horizontal, dorsally slightly or deeply concave, [1.5] 1.6 – 1.9 × as long as broad, length [0.4] 0.3 – 0.5 and breadth [0.2] – 0.3 that of carapace; lateral margins minutely serrated and convex, with well-developed supraocular and subapical spines (trispinose). Pterygostomian flap ending in blunt tooth; upper margin unarmed. Thoracic sternum: Sternite 3 moderately broad, [1.8] – 1.9 × as wide as long, anterior margin convex, anterolaterally rounded. Sternite 4 widely contiguous to sternite 3; surface flattened, smooth; greatest width [2.5] 3.0 – 3.1 × that of sternite 3, 2.0 – [2.1] × as wide as long. Pleon: Elevated ridges upraised dorsally, with few short setae. Tergites 2 – 4 with anterior and posterior transverse elevated ridges; tergites 5 – 6 smooth. Eye: Eyestalk length about 1.0 × broader than long, peduncle distally setose, not distinctly expanded proximally; maximum corneal diameter 0.9 – [1.0] × rostrum width, as wide as eyestalk. Antennule: Article 1 longer than wide, with 3 – 4 distal spines: distomesial spine small or obsolescent; proximal lateral spine absent. Antenna: Article 1 with prominent mesial process, distally not reaching lateral antennular spine. Article 2 often with minute distal spines laterally and mesially. Articles 3 and 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus [0.6] × length of ischium, with 1 well-developed distal spine on extensor margin and 2 spines on flexor margin, proximal spine slightly larger than distal. P 1: 2.0 × carapace length (males), [1.3] – 1.6 (females), subcylindrical, with scattered spines and long stiff setae; merus and carpus with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus [0.5] – 1.0 length of carapace, [1.5] – 2.1 × as long as carpus. Carpus 1.5 – [2.0] × as long as wide. Palm [0.7] – 0.9 × carpus length, [1.4] 1.2 – 1.5 × as long as broad with scattered small spines on dorsal and ventral surfaces, lateral and mesial margins irregular, with 1 small spine on distomesial margin. Fingers unarmed, 1.0 – [1.1] × palm length. P 2 – 4: Moderately stout, setose and moderately spinose. Meri successively shorter posteriorly: P 3 merus 0.8 – [0.9] × length of P 2 merus, P 4 merus 0.7 – [0.9] × length of P 3 merus. P 2 merus [1.5] 1.7 – 2.4 × carapace length, [4.0] – 4.2 × as long as broad, 1.3 × as long as P 2 propodus; P 3 merus [3.5] – 3.6 × as long as broad, [1.1] – 1.2 × as long as P 3 propodus; P 4 merus [2.5] 3.0 – 3.3 × as long as broad, as long as P 4 propodus; extensor margins of P 2 and P 3 with row of few small spines, proximally diminishing in size, with well-developed distal spine; P 4 extensor margin irregular but unarmed; flexor margins irregular, with distal spine on P 2 – 3, absent in P 4. Carpi with 1 spine on extensor margins of P 2 and often in P 3, unarmed on P 4; distal spine prominent on P 2 – 3, absent on P 4; row of small spines below extensor margin on lateral surface of P 2 and may be present on P 3?, unarmed on P 4. Propodi stout, [4.5 – 5.0] 4.4 – 4.5 × as long as broad; extensor margins irregular, usually unarmed; flexor margins with 3 – 4 slender movable spines in addition to distal pair. Dactyli [0.8] – 0.9 × length of propodi, ending in incurved, strong, sharp spine; flexor margins with 5 – 6 well-developed dactylar spines, each with 1 spinule. Eggs: Ov. F carried approximately 7 – 25 eggs of 0.3 – 0.4 mm diameter. Live colour. Body brownish, with brown patches and spots. P 1 – 4 pale brown, with dark spots and bands. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFBDE64A4F9CFD68734BBE52.taxon	distribution	Distribution. Vanuatu, between 2 and 148 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFBDE64A4F9CFD68734BBE52.taxon	discussion	Remarks. Phylladiorhynchus marina belongs to the group of species having the epigastric ridge unarmed, the carapace and pleon ridges elevated, upraised, and dactylar spines on the flexor margins of the dactyli. The new species is closely related to P. phanus from Papua New Guinea (see the differences under the Remarks of this species). The specimen of P. marina range from 1.2 to 2.6 mm size postorbital carapace length. The two sequences of P. marina from Vanuatu diverged 0.8 % for COI and 0.8 % for 16 S.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB1E64F4F9CFF297398BEC1.taxon	description	(Figs. 33 E, 35, 55 C)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB1E64F4F9CFF297398BEC1.taxon	materials_examined	Type material. Holotype. French Polynesia. Rapa, Stn 30, 27.6366 ° S, 144.3033 ° W, 16 – 20 m, 16 / 18 November 2002: M 2.2 mm (MNHN-IU- 2016 - 493). Paratypes. French Polynesia. Rapa, Stn 30, 27.6366 ° S, 144.3033 ° W, 16 – 20 m, 16 / 18 November 2002: 1 M 2.2 mm (MNHN-IU- 2016 - 493), 6 M 2.0 – 2.5 mm (MNHN-IU- 2016 - 509), 1 M broken (MNHN-IU- 2016 - 494), 1 M 2.0 mm, 5 ov. F 2.1 – 2.7 mm (MNHN-IU- 2014 - 13905). — Stn 32, 27.5966 ° S, 144.3833 ° W, 15 – 20 m, 18 / 23 November 2002: 4 M 2.0 – 2.5 mm, 8 ov. F 2.2 – 2.7 mm (MNHN-IU- 2019 - 2603), 1 M 2.8 mm, 2 ov. F 1.8 – 2.1 mm, 1 F 2.0 mm (MNHN-IU- 2016 - 1493). — Stn 33, 27.5800 ° S, 144.3466 ° W, 30 m, 19 November 2002: 4 M 2.0 – 3.0 mm, 2 ov. F 2.1 – 2.5 mm, 1 F 2.0 mm (MNHN-IU- 2019 - 2607). — Stn 36, 27.5583 ° S, 144.3466 ° W, 27 m, 21 November 2002: 1 M, 2.5 mm, 4 ov. F 2.0 – 2.6 mm (MNHN-IU- 2019 - 2606). New Caledonia. Lagon Nord. Stn DW 1097, 19.8616 ° S, 163.7083 ° E, 33 – 34 m, 24 October 1989: 1 M 2.1 mm (MNHN-IU- 2014 - 13795). Other material. South China Sea. Macclesfield Bank. Stn 24, 15.4416 ° N, 114.2333 ° E, 24 – 63 m, May 1892: 1 ov. F 2.4 mm (MNHN-IU- 2016 - 9639). Chesterfield Islands. CORAIL 2 Stn CP 25, 20.4166 ° S, 161.0833 ° E, 67 – 70 m, 22 July 1988: 1 M 1.6 mm (MNHN-IU- 2016 - 472 (Ga- 2050 )). New Caledonia. Touho. September 1993: 1 ov. F 2.4 mm (MNHN-IU- 2019 - 2616). — Koumac 2.3 Stn KL 25, 20.75421 ° S, 164.22986 ° E, 65 m, 10 November 2019: 1 M 1.4 mm (MNHN-IU- 2014 - 20045). — Stn KL 03, 20.67485 ° S, 164.214266 ° E, 50 m, 11 November 2019: 2 ov. F 2.0 – 2.3 mm (MNHN-IU- 2014 - 20052), 1 F parasitized 1.9 mm (MNHN-IU- 2014 - 20063). — Stn KB 642, 20.7093 ° S, 164.25715 ° E, 28 m, 14 November 2019: 2 M 1.7 – broken (MNHN-IU- 2014 - 20109). — Stn KB 647, 20.664116 ° S, 164.189983 ° E, 38 m, 16 November 2019: 2 M 1.5 – 2.0 mm (MNHN-IU- 2014 - 20140). — Stn KL 22, 20.4459 ° S, 163.97176 ° E, 47 m, 19 November 2019: 2 M 1.9 – 2.0 mm (MNHN-IU- 2014 - 20158). 2 M 1.5 – 1.7 mm, 2 ov. F 1.9 – 2.0 mm, 3 postlarvae 0.9 – 1.0 mm (MNHN- IU- 2014 - 20169). French Polynesia. BENTHAUS Stn CP 1918, 27.0500 ° S, 146.0666 ° W, 130 – 140 m, 12 November 2002: 3 M 2.5 mm, 5 ov. F 2.5 – 3.4 mm (MNHN-IU- 2019 - 2591). — Stn CP 1922, 27.0667 ° S, 146.0667 ° W, 150 – 163 m, 12 November 2002: 1 ov. F 2.1 mm (MNHN-IU- 2019 - 2597). — Rapa. Stn 4, 27.5716 ° S, 144.3683 ° W, 18 m, 4 November 2002: 7 M 2.0 – 3.4 mm, 3 ov. F 2.6 – 3.0 mm, 5 F 1.5 – 2.0 mm (MNHN-IU- 2019 - 2585, MNHN-IU- 2014 - 13908). — Stn 5, 27.0933 ° S, 144.3083 ° W, 8 m, 4 November 2002: 4 M 2.6 – 3.4 mm, 3 ov. F 2.5 – 3.6 mm (MNHN- IU- 2019 - 2587). — Stn 9, 27.6216 ° S, 144.3700 ° W, 3 – 24 m, 6 November 2002: 1 M 2.2 mm (MNHN-IU- 2014 - 13907), 1 ov. F 2.2 mm (MNHN-IU- 2014 - 13901). — Stn 10, 27.5800 ° S, 144.3800 ° W, 16 – 18 m, 7 November 2002: 1 ov. F 2.5 mm (MNHN-IU- 2019 - 2584). — Stn 11, 27.6200 ° S, 144.3033 ° W, 2 m, 7 November 2002: 1 ov. F 2.2 mm (MNHN-IU- 2019 - 2605). — Stn 14, 27.5966 ° S, 144.2266 ° W, 2 m, 8 November 2002: 1 M 2.1 mm, 1 ov. F 2.3 mm (MNHN-IU- 2019 - 2586). — Stn 16, 27.6050 ° S, 144.3066 ° W, 5 m, 8 November 2002: 1 ov. F 2.1 mm (MNHN-IU- 2014 - 13906). — Stn 20, 27.5900 ° S, 144.3883 ° W, 5 m, 12 November 2002: 4 ov. F 2.2 – 2.8 mm (MNHN-IU- 2019 - 2588). — Stn 25, 27.6400 ° S, 144.3150 ° W, 3 m, 13 November 2002: 1, ov. F, 1.8 mm (MNHN-IU- 2014 - 13902). — Stn 27, 27.6450 ° S, 144.3116 ° W, 6 m, 14 November 2002: 1 M 2.3 mm (MNHN-IU- 2014 - 13910). — Stn 29, 27.5716 ° S, 144.3500 ° W, 2 – 4 m, 15 November 2002: 1 M 2.7 mm, 4 ov. F, 2.0 – 2.6 mm (MNHN-IU- 2019 - 2636), 1 ov. F 2.2 mm (MNHN-IU- 2014 - 13903).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB1E64F4F9CFF297398BEC1.taxon	etymology	Etymology. From the name Medea, daughter of Aeetes, joined the Argo when the Fleece was recovered. The name is considered a substantive in apposition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB1E64F4F9CFF297398BEC1.taxon	description	Description. Carapace: as long as or slightly longer than broad; transverse ridges with dense short setae and few scattered iridescent setae. Gastric region slightly convex with 4 transverse ridges: epigastric ridge distinct with 2 median spines and some lateral short scales, often followed by some short scales; anterior protogastric ridge not medially interrupted, extending laterally to carapace margin followed by some short scales posteriorly; anterior mesogastric ridge not medially interrupted, laterally interrupted by cervical groove; anterior metagastric scale-like, often followed by some short scales. Mid-transverse ridge uninterrupted, medially slightly depressed, cervical groove distinct, followed by 2 not interrupted or minutely interrupted ridges, interspersed with 1 short lateral ridge and some short scales. Lateral margins nearly straight, with 6 distinct spines: first anterolateral spine well-developed, reaching anteriorly to level of lateral orbital spine, second spine (hepatic) small, slightly dorsomesially from lateral margin, and followed by 4 branchial spines (3 anterior and 1 posterior). Rostrum leaf-like, horizontal, dorsally convex, 1.3 – [1.5] × as long as broad, length 0.3 – [0.4] and breadth [0.2] – 0.3 that of carapace; lateral margins serrated and convex, with well-developed basal supraocular spines, subapical spines absent or obsolescent. Pterygostomian flap with anterior spine, upper margin serrated. Thoracic sternum: As wide as long. Sternite 3 moderately broad, [1.5] – 2.9 × as wide as long, anterior margin convex, anterolaterally rounded. Sternite 4 widely contiguous to sternite 3; anterolaterally serrated, surface depressed in midline, smooth; greatest width 2.2 – [3.5] × that of sternite 3, 2.3 – [3.0] × as wide as long. Pleon: Elevated ridges with short setae and a few scattered long setae. Tergite 2 with anterior and posterior transverse elevated ridges; tergites 3 – 4 with anterior transverse ridge; tergites 5 – 6 smooth. Eye: Eyestalk length about 1.0 – [1.1] × broader than long, peduncle distally setose, not distinctly expanded proximally, with few short transverse striae on lateral surfaces; cornea expanded distally, maximum corneal diameter 0.8 – [0.9] × rostrum width, as wide as eyestalk. Antennule: Article 1 slightly longer than wide, with 5 distal spines: distomesial spine well-developed; proximal lateral spine small, always present. Antenna: Article 1 with prominent mesial process, distally not reaching lateralmost antennular spine. Article 2 with well-developed distomesial and distolateral spines. Article 3 and 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus 0.7 – [0.8] × length of ischium, with well-developed distal spine on extensor and flexor margins. P 1 (lost in holotype): 2.5 – 3.8 (males), 2.6 (females) × carapace length, subcylindrical, spiny and with long stiff setae; merus, carpus and pam with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus 0.8 length of carapace, 2.2 × as long as carpus. Carpus 1.8 × as long as wide. Palm 1.6 × carpus length, 3.4 × as long as broad. Fingers 0.5 × palm length, fixed finger with 2 basal spines; movable finger with 1 – 2 basal spines. P 2 – 4: stout, setose and spinose. Meri successively shorter posteriorly: P 3 merus 0.9 × length of P 2 merus, P 4 merus [0.8] – 0.9 × length of P 3 merus. P 2 merus, 0.6 – [0.7] × carapace length, [4.0] – 4.5 × as long as broad, [1.2] – 1.3 × as long as P 2 propodus; P 3 merus [3.5] – 4.1 × as long as broad, 1.1 – [1.2] × as long as P 3 propodus; P 4 merus 3.6 – [3.7] × as long as broad, 0.9 – [1.0] × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, unarmed; flexor margin irregular, with distal spine on P 2 – 3, absent on P 4. Carpi with 2 spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, absent on P 4; granules below extensor margin on lateral surface of P 2 – 4; flexor margin unarmed. Propodi moderately stout, [4.5 – 5.5] 4.5 – 6.3 × as long as broad; extensor margin irregular; flexor margin with 3 – 6 slender movable spines in addition to distal pair. Dactyli 0.5 – 0.7 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 5 – 6 movable spines. Eggs: Ov. F carried approximately 10 – 25 eggs of 0.3 – 0.4 mm diameter. Live colour. Base colour of body light orange-yellow, with small dark orange spots. P 1 whitish, distal tip finger orange. P 2 – 4 light whitish-translucent, covered by whitish spots and dark orange stripes. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB1E64F4F9CFF297398BEC1.taxon	distribution	Distribution. South China Sea (Macclesfield Bank), New Caledonia and French Polynesia, between 2 and 163 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB1E64F4F9CFF297398BEC1.taxon	discussion	Remarks. Phylladiorhynchus medea belongs to the group of species having 2 epigastric spines, 1 hepatic spine, 3 spines on the anterior branchial margin and the pleonal tergite 3 with the anterior ridge only. The new species closely resembles P. janiqueae, but both species can be distinguished by the following characters: - The rostrum has small subapical spines in P. janiqueae, whereas these spines are absent or obsolescent in P. medea. - The anterior upper margin of the pterygostomian flap is smooth in P. janiqueae, whereas this margin is usually serrated in P. medea. - The antennal article 3 has a small distomesial spine in P. janiqueae, whereas this spine is absent in P. medea. Phylladiorhynchus medea is also close to P. zetes from the French Polynesia (see the differences under the Remarks of this species). The specimens of P. medea range from 0.9 to 3.4 mm postorbital carapace length. The genetic divergences between P. medea and P. janiqueae were 11 % (COI) and 6 % (16 S). The six sequences of P. medea from New Caledonia and French Polynesia diverged 0.6 % (COI) and 0.3 % (16 S).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB4E64C4F9CFF297325BC71.taxon	description	(Fig. 33 F)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB4E64C4F9CFF297325BC71.taxon	materials_examined	Type material. Paratype. Western Australia. Dampier Archipelago. Stn DA 3 / 99 / 61, 20.5776 ° S, 116.6620 ° E, 3 – 5 m, 14 September 1999: 1 M 2.7 mm (WAM C 25991). Other material. Western Australia. Montgomery Island. Stn 22 / K 09 - adhoc, 15.944317 ° S, 124.27330 ° E, no depth, 22 October 2009: 1 ov. F 2.2 mm (WAM C 44013). — Stn 24 / K 09 - T 2, 16.014417 ° S, 124.173100 ° E, no depth, in sponge, 23 October 2009: 2 M 1.2 – 1.9 mm (WAM C 44014). — Stn 27 / K 09 - Q 3, 15.0000 ° S, 124.0000 ° E, no depth, 24 October 2009: 1 F 2.2 mm (WAM C 44038). — Long Reef. Stn 50 / K 10 - Q 2, 13.91545 ° S, 125.774317 ° E, 2 m, 22 October 2010: 1 M 2.3 mm, 1 ov. F 2.4 mm (WAM C 46587), 1 F 2.2 mm (WAM C 46583). — Adele Island. Stn 07 / K 09 - Q 3, 15.49123 ° S, 123.16330 ° E, 0 m, 5 October 2009: 2 F 1.4 – 1.8 mm (WAM C 44006). — Stn 11 / K 09 - T 1, 15.581583 ° S, 123.163200 ° E, 0 – 3.5 m, 17 October 2009: 1 ov. F 2.3 mm (WAM C 44003). — Ningaloo Reef. 22.7473 ° S, 113.0752 ° E, 11 m, 1 May 2009: 1 M 2.5 mm (UF 22848).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB4E64C4F9CFF297325BC71.taxon	diagnosis	Diagnosis. Rostrum with subapical spines; lateral margins straight. Epigastric ridge of carapace unarmed, medially interrupted; anterior branchial margin of carapace with 2 spines; anterior protogastric and mesogastric ridges not interrupted medially; anterior metagastric ridge medially interrupted. Pterygostomian flap with small spine on upper margin. Anterior margin of thoracic sternite 3 slightly convex, with lateral projections Antennular article 1 with 5 spines; distomesial spine small, similar to lower distolateral spine. Antennal article 1 mesial process distally falling well short of second lateral antennular spine; article 2 with distomesial and distolateral spines subequal; article 3 with small distolateral spine. Flexor margin of Mxp 3 merus with 1 prominent spine. P 2 – 4 dactylus flexor margins with 5 – 6 well-developed dactylar spines, each with 1 spinule. Eggs: Ov. F carried approximately 30 – 40 eggs of 0.4 – 0.5 mm diameter. Live colour. Unknown. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB4E64C4F9CFF297325BC71.taxon	distribution	Distribution. The species was described from the Dampier Archipelago (northwestern Australia) (Macpherson, 2008). The new records extend its distribution range along the Western Australia, between 0 and 11 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB4E64C4F9CFF297325BC71.taxon	discussion	Remarks. Phylladiorhynchus nudus belongs to group of species having the epigastric ridge unarmed, the carapace and the pleon ridges elevated, upraised, and dactylar spines on the flexor margin of the P 2 – 4 dactyli. The group includes P. marina, P. jeffkinchi, P. nudus and phanus. However, P. nudus can be easily distinguished from the rest of the species of this group by the presence of a spine on the upper margin of the pterygostomiam flap, whereas this margin is unarmed in the other species. The genetic divergences between P. nudus and other species were always higher than 14 % for COI and 13 % for 16 S.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB4E64D4F9CFA7E71F8BB07.taxon	description	(Fig. 33 G)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB4E64D4F9CFA7E71F8BB07.taxon	materials_examined	Material examined. New Zealand. Iceberg Seamount, Stn. TAN 0905 / 119, 44.158 – 44.162 ° S, 174.555 – 174.552 ° W, 487 – 616 m, 28 Jun 2009: 11 M 5.0 – 9.0 mm, 18 F 4.1 – 8.2 mm (MNHN-IU- 2019 - 2595, ex NIWA 54729).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB4E64D4F9CFA7E71F8BB07.taxon	diagnosis	Diagnosis (modified from Schnabel & Ahyong 2019). Rostrum lateral margins straight or very slightly convex (in small specimens); subapical spines present. Epigastric region typically with 4 spines, small specimens can have lateral pair much smaller (sometimes granular) than mesial pair; anterior branchial margin of carapace with 3 spines; anterior metagastric ridge continuous, not medially interrupted. Pleonal tergite 3 with anterior and posterior transverse ridge. Anterior margin of thoracic sternite 3 biconcave, with obtuse median projection. Antennular article 1 with 4 well-developed spines and minute second lateral spine: distomesial spine long; second (proximal) lateral spine minute. Antennal article 1 mesial process distally reaching at least to level of second lateral antennular spine; article 2 distolateral spine distinctly larger than minute distomesial spine (if present), article 3 distolateral spine minute or absent. Flexor margin of Mxp 3 merus with one prominent spine only. P 2 – 4 dactylus extensor margin without upright spines at bases of movable spines. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB4E64D4F9CFA7E71F8BB07.taxon	distribution	Distribution. Southeastern Australia, Tasmania, New Zealand, between 46 and 1246 m (from Schnabel & Ahyong 2019).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB4E64D4F9CFA7E71F8BB07.taxon	discussion	Remarks. The species closely resembles P. australis Schnabel & Ahyong, 2019, from New Zealand and Southern Australia, and P. integrus (Benedict, 1902) from Japan to Chesterfield Islands (see the differences under the Remarks of P. integrus).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB5E6704F9CFDD4755FBE8A.taxon	description	(Figs. 33 H, 36, 55 D)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB5E6704F9CFDD4755FBE8A.taxon	materials_examined	Type material. Holotype. French Polynesia, Society Islands, Moorea Island, 17.4758 ° S, 149.8322 ° W, 20 m, 25 July 2006: ov. F 2.0 mm (UF 9732). Paratypes. French Polynesia. Marquesas Islands. PAKAIHI I TE MOANA Stn MQ 2 - GR-B, 8 ° 56.231 ’ S, 140 ° 07.240 ’ W, 20 – 23 m, January 2012: 1 M broken (MHNH-IU- 2014 - 13730). French Polynesia. Society Islands. Moorea Island. 17.5145 ° S, 149.7616 ° W, 20 m, 23 October 2008: 1 M 2.1 mm (UF 36172). — 17.4764 ° S, 149.8316 ° W, 13 m, 31 January 2012: 1 M 2.0 mm (UF 33772), 1 M 2.7 mm (UF 33773). — 17.4759 ° S, 149.8419 ° W, 13 m, 3 February 2012: 1 F 1.9 mm (UF 33866), 1 M 1.6 mm, 1 F 2.4 mm (UF 33867), 1 ov. F 2.2 mm (UF 33961), 4 M 1.1 – 1.9 mm, 6 ov. F 1.8 – 2.0 mm (UF 33962). Other material. French Polynesia. Society Islands. Moorea Island. 17.4764 ° S, 149.8316 ° W, 13 m, 6 February 2012: 6 M 1.4 – 2.0 mm, 5 ov. F 1.6 – 2.0 mm, 1 F 2.2 mm (UF 34088). — 17.4785 ° S, 149.8477 ° W, 13 m, 7 February 2012: 1 M 2.0 mm, 4 ov. F 1.6 – 2.3 mm, 1 F 1.7 mm (UF 34158). — 17.4785 ° S, 149.8477 ° W, 13 m, 8 February 2012: 1 ov. F 2.6 mm (UF 34209). — BENTHAUS Stn DW 1968, 23 ° 23 ’ S, 150 ° 44 ’ W, 100 – 120 m, 20 November 2002: 1 M 2.3 mm (MNHN-IU- 2019 - 2633). Northern Mariana Islands. Saipan Island. Stn SAI- 435, 15.0978 ° N, 145.7434 ° E, 17 m, 11 April 2014: 1 ov. F 2.0 mm (UF 54495). — Stn SAI- 640, 15.15620 ° N, 145.6899 ° E, 11 m, 17 April 2014: 4 M 1.3 – 2.0 mm, 1 F 2.3 mm (UF 54489). — Stn SAI- 788, 15.15620417 ° N, 145.6899585 ° E, 11 m, 17 April 2014: 1 M 1.4 mM 1 F 2.0 mm (UF 54502). — Stn SAI- 947, 15.27406 ° N, 145.79106 ° E, 9 m, 18 April 2014: 2 M 1.6 – 2.0 mm, 1 ov. F 1.8 mm (UF 54498). — Pagan Island. Stn PAG- 404, 18.10734 ° N, 145.78587 ° E, 11 m, 20 April 2014: 2 M 1.3 – 2.3 mm (UF 54491). — Stn PAG- 409, 18.10734 ° N, 145.78587 ° E, 11 m, 20 April 2014: 1 F 1.9 mm (UF 54488). — Stn PAG- 478, 18.1073 ° N, 145.7858 ° E, 11 m, 20 April 2014: 1 M 1.5 mm, 1 ov. F 2.0 mm (UF 54486). — Stn PAG- 695, 18.0706 ° N, 145.7137 ° E, 11 m, 20 April 2014: 1 M 2.0 mm, 1 ov. F 1.9 mm (UF 54499). — Stn PAG- 696, 18.0706 ° N, 145.7137 ° E, 11 m, 20 April 2014: 2 ov. F 2.1 – 2.4 mm (UF 54500). — Stn PAG- 1149, 18.0706 ° N, 145.7137 ° E, 11 m, 20 April 2014: 1 M 1.9 mm (UF 54497). — Stn PAG- 839, 18.11947 ° N, 145.7555 ° E, 12 m, 23 April 2014: 1 M 1.5 mm, 2 ov. F 1.6 – 1.8 mm (UF 54487). — Stn PAG- 898, 18.11947 ° N, 145.7555 ° E, 12 m, 23 April 2014: 3 M 1.5 – 2.4 mm 2 ov. F 2.0 – 2.3 mm (UF 54485). — Maug Islands. Stn MAU- 475, 20.0141 ° N, 145.2342 ° E, 15 m, 03 May 2014: 1 M 1.6 mm (UF 54490). Mariana Islands. Guam Island. Stn GUA- 645, 13.57847 ° N, 144.82831 ° E, 11 m, 25 March 2014: 1 M 1.3 mm, 1 ov. F 1.8 mm (UF 54542). — Stn GUA- 688, 13.57847 ° N, 144.82831 ° E, 11 m, 25 March 2014: 1 ov. F 2.4 mm (UF 54496). American Samoa. Rose Atoll. Stn ROS- 777, 14.5293 ° S, 168.1534 ° W, 10 m, 16 March 2015: 1 F 1.2 mm (UF 54493). — Stn ROS- 689, 14.5596 ° S, 168.1601 ° W, 45 m, 18 March 2015: 1 M 1.2 mm (UF 54492). — Tutuila Island. Stn TUT 138, 14.3604 ° S, 170.7502 ° W, 17 m, 26 February 2015: 1 F 2.1 mm (UF 54494). Kiribati. Line islands, Millenium Island. 9.91 ° S, 150.21 ° W, 12 m, 5 November 2013: 1 ov. F 2.0 mm, in dead Pocillopora (UF 39269). — Jarvis Island. Stn JAR- 653, 0.36900 ° S, 160.0081 ° W, 16 m, 10 April 2015: 1 M 1.8 mm (UF 54501). Papua New Guinea. PAPUA NIUGINI Stn PB 07, 05.1800 ° S, 145.8300 ° E, 22 m, 30 December 2012: 1 M 2.0 mm (MNHN-IU- 2013 - 397). — KAVIENG Stn KB 28, 02.7283 ° S, 150.6400 ° E, 15 – 26 m, 11 June 2014: 1 M 1.7 mm (MNHN-IU- 2014 - 13657). New Caledonia. LIFOU Stn 1429, 20.7916 ° S, 167.1183 ° E, 8 – 18 m, 24 November 2000: 2 ov. F 2.1 – 2.4 mm (MNHN-IU- 2019 - 2601).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB5E6704F9CFDD4755FBE8A.taxon	etymology	Etymology. From the name Orpheus, an Argonaut, son of Calliope and Oeagrus. The name is considered a substantive in apposition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB5E6704F9CFDD4755FBE8A.taxon	description	Description. Carapace: [1.1] – 1.2 longer than broad; transverse ridges with a few short setae. Gastric region flattened with 4 transverse ridges (usually barely distinct): epigastric ridge with 2 median spines; anterior protogastric ridge not medially interrupted, nearly extending laterally to carapace margin; anterior mesogastric ridge not medially interrupted, laterally continuing to first branchial spine, followed by some short lateral scales; anterior metagastric ridge scale-like, often followed by a short scale. Mid-transverse ridge not interrupted, medially depressed, preceded by cervical groove distinct, followed by 2 not interrupted or minutely interrupted ridges, interspersed with 1 short lateral ridge and sometimes few, short scattered scales. Lateral margins straight or slightly convex, with 4 distinct spines: first anterolateral spine well-developed, reaching anteriorly to level of lateral orbital spine, hepatic margin unarmed; anterolateral spine followed by 3 branchial spines (2 anterior and 1 posterior). Rostrum leaf-like, horizontal, dorsally concave, [1.5] 1.4 – 1.6 × as long as broad, length 0.3 – [0.4] and breadth 0.2 – [0.3] that of carapace; lateral margins serrated and convex, with well-developed supraocular spines, and small subapical spines. Pterygostomian flap ending in acute tooth, upper margin smooth. Thoracic sternum: As wide as long. Sternite 3 moderately broad, [1.8] – 2.0 × as wide as long, anterior margin convex, with blunted median projection, lateral margins rounded. Sternite 4 widely contiguous to sternite 3; surface depressed in midline, smooth; greatest width 3.0 × that of sternite 3, 2.6 – [2.7] × as wide as long. Pleon: Elevated ridges with a few scattered short setae. Tergite 2 with anterior and posterior transverse elevated ridges with short setae and a few scattered long setae; tergites 3 – 4 with anterior transverse ridge; tergites 5 – 6 smooth. Eye: Eyestalk length about [1.0] – 1.1 × broader than long, peduncle distally setose, slightly expanded proximally; maximum corneal diameter 0.8 – [0.9] × rostrum width, as wide as eyestalk. Antennule: Article 1 slightly longer than wide, with 5 distal spines: distomesial spine well-developed; proximal lateral spine small, always present. Antenna: Article 1 with prominent mesial process distally not reaching lateralmost antennular spine. Article 2 with well-developed distomesial and distolateral spines. Article 3 with distomesial spine. Article 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus [0.7] 0.7 × length of ischium, with well-developed distal spine on extensor and flexor margins. P 1: 1.9 – 2.3 (males), 1.8 – [1.9] (females) × carapace length; subcylindrical, spiny and with dense long stiff setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus 0.6 – 0.7 length of carapace, 1.3 – 1.9 × as long as carpus. Carpus 1.3 – 1.7 × as long as wide. Palm 1.1 × carpus length, 1.1 – 1.4 × as long as broad. Fingers [0.9] – 1.1 × palm length; fixed finger unarmed; movable finger with basal spine. P 2 – 4: Stout, moderately setose and spinose. Meri successively shorter posteriorly: P 3 merus 0.9 × length of P 2 merus, P 4 merus [0.8] – 0.9 × length of P 3 merus. P 2 merus, [0.5] – 0.6 × carapace length, 4.6 – [4.7] × as long as broad, [1.1] – 1.2 × as long as P 2 propodus; P 3 merus [4.5] 4.4 – 4.7 × as long as broad, [1.1] – 1.3 × as long as P 3 propodus; P 4 merus [3.4] 3.2 – 3.6 × as long as broad, [1.0] – 1.1 × as long as P 4 propodus; extensor margin of P 2 – 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, unarmed; flexor margin irregular, with distal spine on P 2 – 4. Carpi extensor margin unarmed on P 2 – 4; distal spine prominent on P 2 – 3, smaller on P 4; extensor margin on lateral surface of P 2 – 4 unarmed; flexor margin unarmed. Propodi stout, [4.7] 4.5 – 5.0 × as long as broad; extensor margin unarmed; flexor margin with 3 – 4 slender movable spines in addition to distal pair. Dactyli [0.6] – 0.8 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 4 – 5 movable spines. Eggs: Ov. F carried approximately 5 – 20 eggs of 0.3 – 0.5 mm diameter. Live colour. Body orange, with dark orange spots and often with blue-purple patches and bands. P 1 whitish, with brownish stripes. P 2 – 4 whitish, with brown transverse stripes. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB5E6704F9CFDD4755FBE8A.taxon	distribution	Distribution. Papua New Guinea, Northern Mariana Islands, Guam Island, American Samoa (Rose Atoll), Kiribati, New Caledonia and French Polynesia from 8 to 120 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FFB5E6704F9CFDD4755FBE8A.taxon	discussion	Remarks. Phylladiorhynchus orpheus belongs to the group of species with 2 median epigastric spies, without hepatic spine and 2 spines on the anterior branchial margin. Phylladiorhynchus orpheus is closely related to P. lynceus, from Chagos, Western Australia, Kiribati and Samoa, P. priasus, from Mariana and Marshall Islands, and P. integrirostris from Hawaii. However, the new species can be easily distinguished from these species by the following differences: - The rostral margins are usually smooth in P. orpheus, whereas these margins are always serrated in the other species. - The carapace ridges are flattened and often indistinct in P. orpheus, whereas these ridges are elevated and distinct in the other species. - The metagastric ridge is scale-like in P. orpheus, whereas this ridge is medially interrupted in the other species. - The extensor margins of P 2 – 4 propodi are unarmed in P. orpheus, whereas these margins are armed with 1 – 3 proximal spines in the other species. The specimens of P. orpheus from Kiribati, Phoenix Islands, and from Papua New Guinea have the thoracic sternite 3 with a different shape. Furthermore, they have a different colour pattern. A revision of additional material from both localities, including genetic data, will be necessary to clarify if these small differences are intraspecific variations. The genetic divergences between P. orpheus and the other closely related species were very high: 19 – 22 % (COI) and 13 – 16 % (16 S). The six sequences of P. orpheus from French Polynesia and Northern Mariana Islands diverged up to 2 % for COI. The specimens of P. orpheus range from 1.1 to 2.7 mm postorbital carapace length.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF89E6774F9CFF297557BD62.taxon	description	(Figs. 33 P, 38, 55 E)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF89E6774F9CFF297557BD62.taxon	materials_examined	Type material. Holotype. Mayotte and Comores Islands, BIOMAGLO Stn DW 4800, 11.4500 ° S, 47.3166 ° E, 240 – 255 m, 24 January 2017: M 3.7 mm (MNHN-IU- 2016 - 7002). Paratypes. Reunion Island. MD 32 Stn DC 2, 21.2066 ° S, 55.8233 ° E, 160 – 190 m, 12 August 1982: 1 M 2.7 mm (MNHN-IU- 2019 - 2691). — Stn CP 177, no data: 1 M 2.3 mm (MNHN-IU- 2019 - 2699). Mayotte and Comores Islands. BIOMAGLO Stn DW 4800, 11.4500 ° S, 47.3166 ° E, 240 – 255 m, 24 January 2017: 1 F 3.3 mm (MNHN-IU- 2016 - 7003). — Stn DW 4838, 11.9833 ° S, 43.5166 ° E, 185 – 267 m, 29 January 2017: 1 ov. F 3.4 mm (MNHN-IU- 2017 - 428).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF89E6774F9CFF297557BD62.taxon	etymology	Etymology. Named after our colleague Paula Martin-Lefèvre, Collection Manager in the Muséum national d’Histoire naturelle, Paris. The name is considered a substantive in apposition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF89E6774F9CFF297557BD62.taxon	description	Description. Carapace: as long as or slightly longer than broad; transverse ridges with few short setae. Gastric region flattened with 4 transverse ridges: epigastric ridge indistinct, with 5 spines (1 median and 2 pairs of spines laterally); anterior protogastric ridge not medially interrupted or minutely medially interrupted, nearly extending laterally to carapace margin; anterior mesogastric ridge not medially interrupted, laterally interrupted by anterior branch of cervical groove, continuing to first branchial spine; anterior metagastric ridge scale-like. Mid-transverse ridge not interrupted, medially depressed, preceded by shallow to distinct cervical groove, followed by 2 not interrupted or minutely interrupted ridges, interspersed with 1 – 2 short lateral ridges and sometimes few, short scattered scales. Lateral margins slightly convex, with 7 spines: first anterolateral spine well-developed, reaching anteriorly to level of lateral orbital spine, second spine (hepatic) small, slightly dorsomesially from lateral margin, and followed by 5 branchial spines (3 anterior and 2 posterior). Rostrum triangular, horizontal, dorsally flattish or slightly concave, [1.2] – 1.8 × as long as broad, length 0.3 – [0.4] and breadth 0.2 – [0.3] that of carapace; lateral margins smooth and straight or slightly concave, with well-developed supraocular spines, subapical spines absent. Pterygostomian flap ending in blunt tooth, upper margin smooth. Thoracic sternum: 1.2 as wide as long. Sternite 3 moderately broad, [2.3] 2.0 – 3.0 × as wide as long, anterior margin straight or with a median feeble excavation, moderately produced anterolaterally. Sternite 4 widely contiguous to sternite 3; surface not depressed in midline, smooth; greatest width 2.0 – [3.0] × that of sternite 3, [2.5] 2.0 – 3.0 × as wide as long. Pleon: Transverse elevated ridges with short setae. Tergite 2 with anterior and posterior transverse elevated ridges; tergites 3 and 4 with anterior transverse ridge; tergites 5 – 6 smooth. Eye: Eyestalk length about 1.0 – [1.2] × broader than long, peduncle distally setose, not distinctly expanded proximally; maximum corneal diameter 1.0 × rostrum width, as wide as eyestalk. Antennule: Article 1 1.5 × longer than wide, with 5 spines: distomesial spine small; proximal lateral spine small, always present. Antenna: Article 1 with prominent mesial process, distally falling well short of lateralmost antennular spine. Article 2 with well-developed distal spines, distomesial larger than distolateral. Article 3 with often with a small to distinct distomesial spine. Article 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus [0.6] 0.6 × length of ischium, with well-developed distal spine on extensor margin and 2 strong spines on flexor margin. P 1: 2.8 – [3.5] (males), 2.8 – 3.0 (females) × carapace length; subcylindrical, spiny and with scattered long stiff setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus 1.0 – [1.3] length of carapace, [1.8] – 2.5 × as long as carpus. Carpus [2.0] – 2.4 × as long as wide. Palm [1.1] – 1.4 × carpus length, [1.5] – 2 × as long as broad. Fingers [0.9] – 1.0 × longer than palm; fixed finger unarmed or with basal spine; movable finger with well-developed basal spine. P 2 – 4: Slender, subcylindrical, moderately setose and spinose. Meri successively shorter posteriorly: P 3 merus [0.7] – 0.8 × length of P 2 merus, P 4 merus [0.8] – 0.9 × length of P 3 merus. P 2 merus, [0.8] – 0.9 × carapace length, 5.6 – [6.0] × as long as broad, 1.4 × as long as P 2 propodus; P 3 merus [6.5] – 7.1 × as long as broad, [1.2] – 1.4 × as long as P 3 propodus; P 4 merus [5.0] – 5.4 × as long as broad, 1.2 × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, withdistal spine; flexor margins of P 2 – 4 irregular, each with distal spine; P 4 lateral surface with median row 4 small spines, absent in others. Carpi with 3 or 4 spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, minute on P 4; row of small spines below extensor margin on lateral surface of P 2 – 3, unarmed on P 4; flexor margin unarmed. Propodi slender, [7.2 – 7.3] 7.0 – 7.5 × as long as broad; extensor margin irregular, usually unarmed; flexor margin with 3 – 4 movable spines in addition to distal pair. Dactyli 0.5 – [0.6] × length of propodi, ending in incurved, strong, sharp spine; flexor margin with [7 – 8] 6 – 8 movable spines. Eggs: Ov. F (MNHN-IU- 2017 - 428) carried 18 eggs of 0.3 – 0.4 mm diameter. Live colour. Body pale orange, with golden-yellow spots. P 1 orange, with whitish. P 2 – 4 light orange, with dark bands. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF89E6774F9CFF297557BD62.taxon	distribution	Distribution. Mayotte and Comores Islands, Reunion Island, between 160 and 267 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF89E6774F9CFF297557BD62.taxon	discussion	Remarks. Phylladiorhynchus paula belongs to the group of species that present 5 epigastric spines, the rostrum margin straight, the subapical spines of the rostrum absent, 3 spines on the anterior branchial margin, and the Mxp 3 merus with two prominent spines along the flexor margin. Phylladiorhynchus paula closely resembles to P. acastus, from the Philippines, Papua New Guinea, Vanuatu, Chesterfield Islands and New Caledonia, and P. argus, from French Polynesia, New Caledonia and Chesterfield Islands, however, they can be distinguished by the following characters: - The dorsal carapace ridges have scattered long thick iridescent setae in P. acastus, whereas these setae are absent in P. paula and they are short in P. argus. - The anterior mesogastric ridge is medially interrupted in P. argus, whereas it is not medially interrupted in P. acastus and P. paula. - Pleonal tergite 3 has 2 ridges (anterior and posterior) in P. acastus, whereas there is only one anterior ridge in P. argus and P. paula. - The shape of thoracic sternite 3 is broad in P. argus, being more than 4 × as wide as long, compared to moderately broad in P. acastus and P. paula (2 – 3 × as wide as long). - Antennal article 3 has a small to well-developed distomesial spine in P. paula and P. acastus, whereas this article is unarmed in P. argus. The genetic distances between these species were always quite large: P. paula diverged 10 – 12 % (COI) and 3 – 5 % (16 S) from P. argus and P. acastus. The two sequences of P. paula from Mayotte and Comores Islands diverged 0.6 % for COI and 0 % for 16 S. The specimens of P. paula range from 2.3 to 3.7 mm postorbital carapace length.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF8FE6754F9CFB707438BEC2.taxon	description	(Figs. 39, 55 F)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF8FE6754F9CFB707438BEC2.taxon	materials_examined	Type material. Holotype. French Polynesia, Rapa. Stn 28, 27.6400 ° S, 144.3433 ° W, 30 m, 15 November 2002: ov. F 2.8 mm (MNHN-IU- 2014 - 13734). Paratypes. French Polynesia. Society Islands, Moorea. Off E Opunohu pass, at Shark Feeding buoy, outer reef slope from within rubble, 17.4817 ° S, 149.8558 ° W, 17 – 18 m, 16 October 2008: 1 F 3.0 mm (UF 15626). — Between Temae and Afarealtu. Outer reef slope, 17.5145 ° S, 149.761 ° W, 20 m, 23 October 2008: 1 ov. F 2.2 mm (UF 36172), 1, ov. F 2.4 mm (UF 16035), 1 M 2.0 mm (UF 16078). Mariana Islands. Guam Island. Glass Breakwater. Near mouth of Apra harbour, among rocks, 3 – 6 m, 17 October 2001: 1 M 3.3 mm (UF 1224). Other material. New Caledonia. Exp. Mont. Komac, 12 m, 7 October 1993: 1 F 2.0 mm (MNHN- IU- 2016 - 468). French Polynesia. Between Temae and Afarealtu. Outer reef slope, 17.4759 ° S, 149.8419 ° W, 11 m, 30 January 2012: 1 ov. F 2.1 mm (UF 34661).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF8FE6754F9CFB707438BEC2.taxon	etymology	Etymology. From the name Peneleos, an Argonaut, son of Hippalmus and Asterope. The name is considered a substantive in apposition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF8FE6754F9CFB707438BEC2.taxon	description	Description. Carapace: As long as or slightly longer than broad; transverse ridges with dense short setae. Gastric region with 4 transverse ridges: epigastric ridge distinct, with 2 median spines (rarely with some outer small spine or granules), short scales laterally; anterior protogastric ridge not medially interrupted, nearly extending laterally to carapace margin, sometimes followed by a posterior protogatric ridge; anterior mesogastric ridge not medially interrupted, laterally continuing to first branchial spine, followed by some short lateral scales; anterior metagastric ridge not medially interrupted, followed by some short scales. Mid-transverse ridge not interrupted, medially depressed, cervical groove indistinct, followed by 2 – 3 not interrupted or minutely interrupted ridges, interspersed with 1 short lateral ridge and few short scales. Lateral margins nearly straight or slightly convex, with 5 distinct spines: first anterolateral spine well-developed, reaching anteriorly to level of lateral orbital spine, second spine (hepatic) small, slightly dorsomesially from lateral margin, and followed by 3 branchial spines (2 anterior and 1 posterior). Rostrum leaf-like, horizontal, dorsally flattish, 1.6 – [1.7] × as long as broad, length 0.3 – [0.4] and breadth 0.2 that of carapace; lateral margins serrated and convex, with well-developed supraocular spines, subapical spines absent. Pterygostomian flap ending in sharp tooth, upper margin smooth. Thoracic sternum: As wide as long. Sternite 3 broad, 3.4 – [4.0] × as wide as long, anterior margin serrated and convex, with a median projection, produced anterolaterally. Sternite 4 widely contiguous to sternite 3; anterolaterally serrated, surface depressed in midline, smooth; greatest width 1.3 – [1.4] × that of sternite 3, [3.1] – 3.2 × as wide as long. Pleon: Elevated ridges, with short setae and a few scattered long setae. Tergite 2 with anterior and posterior transverse; tergites 3 – 4 with anterior transverse ridge; tergites 5 – 6 smooth. Eye: Eyestalk length about [0.7] – 0.8 × broader than long, peduncle distally setose, not distinctly expanded proximally; cornea expanded distally, maximum corneal diameter [1.0] – 1.1 × rostrum width, 0.7 – 0.8 × maximum peduncle width. Antennule: Article 1 slightly longer than wide, with 5 distal spines: distomesial spine well-developed; proximal lateral spine small, always present. Antenna: Article 1 with prominent mesial process, nearly reaching end of proximal lateral antennular spine. Article 2 with well-developed distomesial and distolateral spines. Article 3 and 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus 0.6 – [0.7] × length of ischium, with well-developed distal spine on extensor and flexor margins. P 1 (lost in holotype): 2.8 – 3.8 (males), 2.3 – 2.4 (females) × carapace length; subcylindrical, spiny and with long stiff setae and dense thick iridescent and plumose setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus 0.9 – 1.4 length of carapace. Carpus 1.7 – 1.9 × as long as wide. Palm 1.0 – 1.4 × carpus length, 2.0 – 2.5 × as long as broad. Fingers 0.6 – 1.0 × palm length; fixed finger with small basal spine; movable finger unarmed (large basal spine in 39 F) fingers also extremely gaping. P 2 – 4: Slender, densely setose and spinose. Meri successively shorter posteriorly: P 3 merus 0.7 – [0.8] × length of P 2 merus, P 4 merus [0.8] – 0.9 × length of P 3 merus. P 2 merus, 0.7 – [0.8] × carapace length, [4.0] – 5.0 × as long as broad, [1.2] 1.3 × as long as P 2 propodus; P 3 merus 3.8 – 4 × as long as broad, [1.0] – 1.1 × as long as P 3 propodus; P 4 merus 3.6 – [3.6] × as long as broad, [1.0] – 1.1 × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, unarmed; flexor margin irregular, with distal spine on P 2 – 3, distal spine absent in P 4. Carpi with 1 – 3 spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, smaller on P 4; granules below extensor margin on lateral surface of P 2 – 4; flexor margin unarmed. Propodi moderately slender, [5.0 – 5.8] 4.9 – 6.0 × as long as broad; extensor margin irregular, armed with 2 – 4 spines on proximal half; flexor margin with 3 – 4 slender movable spines in addition to distal pair. Dactyli 0.5 – 0.6 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 5 – 6 movable spines. Eggs: Ov. F carried approximately 6 – 20 eggs of 0.3 – 0.4 mm diameter. Live colour. Body light orange, with dark orange-brownish small spots. P 1 light orange, tip of fingers dark orange-reddish. P 2 – 4 whitish, with dark orange-brownish bands. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF8FE6754F9CFB707438BEC2.taxon	distribution	Distribution. Guam Island, French Polynesia and New Caledonia, between 3 and 30 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF8FE6754F9CFB707438BEC2.taxon	discussion	Remarks. Phylladiorhynchus peneleos belongs to the group of species having 2 median epigastric spines, the hepatic margin armed with 1 spine, and 2 spines on the anterior branchial margin. The new species is morphologically close to P. bahamut, from the Red Sea. However, they can be distinguished by the following characters: - The gastric region has several secondary ridges in P. peneleos, whereas these ridges are absent in P. bahamut. - The carapace ridges have some iridescent thick setae in P. bahamut, whereas these setae are in P. peneleos. - Thoracic sternite 3 is broad, more than 3 × wider than long, in P. peneleos, whereas the sternite 3 is less than twice wider than long in P. bahamut. - The cornea is wider than the stalk in P. peneleos, whereas it is 0.7 – 0.8 × width of the eyestalk in P. bahamut. - The P 1 has plumose setae in P. peneleos, whereas these setae are absent in P. bahamut. The genetic divergences between P. peneleos and P. bahamut were very high 32 % (COI) and 14 % (16 S). The four-six sequences of P. peneleos from French Polynesia diverged 0.15 % for COI and 0 % for 16 S respectively. The specimens of P. peneleos range from 2.0 to 3.3 mm postorbital carapace length.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF82E6784F9CFF29726FBAF2.taxon	description	(Figs. 33 K, 40, 55 G)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF82E6784F9CFF29726FBAF2.taxon	materials_examined	Type material. Holotype. Madagascar. ATIMO VATAE Stn TB 02, 25.0216 ° S, 47.0083 ° E, 18 m, 1 May 2010: M 3.2 mm (MNHN-IU- 2016 - 486). Paratypes. Madagascar. ATIMO VATAE Stn TP 02, 25.0250 ° S, 47.0233 ° E, 25 – 30 m, 29 April 2010: 1 ov. F 2.7 mm (MNHN-IU 2010 - 2730). — Stn TB 02 - TB 03, 25.0216 ° S, 47.0083 ° E, 18 m, 1 May 2010: 2 M 2.5 – 2.6 mm (MNHN-IU 2016 - 463). — Stn TB 02, 25.0216 ° S, 47.0083 ° E, 18 m, 1 May 2010: 1 ov. F 2.5 mm (MNHN-IU 2016 - 461). — Stn TB 03, 25.0216 ° S, 47.0083 ° E, 18 m, 1 May 2010: 2 ov. F 2.8 – 3.2 mm (MNHN-IU 2016 - 465). Other material. Madagascar. ATIMO VATAE Stn TB 05, 25.0366 ° S, 47.0066 ° E, 23 m, 1 May 2010: 2 ov. F 2.7 – 2.9 mm, 2 F 2.5 – 2.6 mm (MNHN-IU 2010 - 2742).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF82E6784F9CFF29726FBAF2.taxon	etymology	Etymology. Named after biologist Pepe Fernández, from the Museo Nacional de Ciencias Naturales, Madrid, and especially interested in registering new species; in gratitude for his enthusiasm and support.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF82E6784F9CFF29726FBAF2.taxon	description	Description. Carapace: As long as or slightly longer than broad, transverse ridges with dense short setae and few scattered long and thick iridescent setae. Gastric region slightly convex with 4 transverse ridges: epigastric ridge distinct with 2 median spines, often followed by short scales; anterior protogastric ridge not medially interrupted, nearly extending laterally to carapace margin, often followed by uninterrupted (in large specimens) or scale-like posterior protogastric ridge; anterior mesogastric ridge not medially interrupted, laterally continuing to first branchial spine, followed by some short lateral scales; anterior metagastric ridge not medially interrupted, followed by short scales. Mid-transverse ridge not interrupted, medially depressed, preceeded by a shallow or indistinct cervical groove, followed by 2 not interrupted or minutely interrupted ridges, interspersed with 2 short lateral ridges and sometimes few, short scattered scales. Lateral margins slightly convex, with 4 distinct spines: first anterolateral spine welldeveloped, reaching anteriorly to level of lateral orbital spine, hepatic margin unarmed; anterolateral spine followed by 3 – 4 branchial spines (2 anterior and 1 – 2 posterior). Rostrum leaf-like, horizontal, dorsally flattened, 1.3 – [1.5] × as long as broad, length 0.3 – [0.4] and breadth [0.2] – 0.3 that of carapace; lateral margins smooth and convex, with well-developed supraocular spines, subapical spines absent or obsolescent. Pterygostomian flap ending in acute tooth, upper margin smooth. Thoracic sternum: As wide as long. Sternite 3 moderately broad, [1.8] 1.5 – 2.0 × as wide as long, anterior margin convex, anterolaterally rounded. Sternite 4 widely contiguous to sternite 3, surface depressed in midline, smooth; greatest width [2.8] – 3.0 × that of sternite 3, [2.8] – 3.0 × as wide as long. Pleon: Elevated ridges with short setae and a few scattered long setae. Tergite 2 with anterior and posterior transverse elevated ridges; tergites 3 – 4 with anterior transverse ridge; tergites 5 – 6 smooth. Eye: Eyestalk length about 0.9 × broader than long, peduncle distally setose, not distinctly expanded proximally; maximum corneal diameter 0.7 – [0.9] × rostrum width, as wide as eyestalk. Antennule: Article 1 slightly longer than wide, with 5 distal spines: distomesial spine well-developed; proximal lateral spine small, always present. Antenna: Article 1 with prominent mesial process nearly reaching end of lateralmost antennular spine. Article 2 with well-developed distomesial and distolateral spines. Article 3 and 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus [0.6] – 0.6 × length of ischium, with well-developed distal spine on extensor and flexor margins. P 1: 2.9 – [3.3] (males), 2.2 – 2.8 (females) × carapace length; subcylindrical, spiny and with long stiff setae and dense thick and plumose setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus 1.1 – [1.3] length of carapace, [1.9] 1.6 – 2.2 × as long as carpus. Carpus [2.6] 1.9 – 2.7 × as long as wide. Palm 0.9 – [1.2] × carpus length, 1.5 – [2.3] × as long as broad. Fingers [0.7] – 1.0 × palm length; fixed finger unarmed; movable finger with a basal spine. P 2 – 4: Moderately stout, densely setose and spinose. Meri successively shorter posteriorly: P 3 merus 0.6 × length of P 2 merus, P 4 merus 0.8 – 0.9 × length of P 3 merus. P 2 merus, 1.8 – [2.1] × carapace length, 3.8 – [5.0] × as long as broad, [1.2] – 1.4 × as long as P 2 propodus; P 3 merus [3.9] 3.7 – 4.0 × as long as broad, 0.9 – [1.1] × as long as P 3 propodus; P 4 merus 4 × as long as broad, as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, unarmed; flexor margin irregular, with distal spine on P 2 – 4, P 4 lateral surface with median row of 2 spines. Carpi with 1 – 4 minute spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, smaller on P 4; row of small spines below extensor margin on lateral surface of P 2 – 3; flexor margin unarmed. Propodi moderately stout, [5.0 – 5.6] 4.0 – 5.8 × as long as broad; extensor margin irregular, armed with 3 spines on proximal half; flexor margin with 3 – 4 slender movable spines in addition to distal pair. Dactyli [0.5 – 0.6] 0.5 – 0.7 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 5 – 6 movable spines. Eggs: Ov. F carried approximately 20 – 30 eggs of 0.3 – 0.6 mm diameter. Live colour. Body green-orange, with red and orange spots and patches. P 1 green, tip of fingers dark orangereddish. P 2 – 4 light green-orange, with dark green-brownish bands. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF82E6784F9CFF29726FBAF2.taxon	distribution	Distribution. Madagascar, between 18 and 30 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF82E6784F9CFF29726FBAF2.taxon	discussion	Remarks. Phylladiorhynchus pepei belongs to the group of species having 2 median epigastric spines, the hepatic margin unarmed and 2 spines on the anterior branchial margin. This species is easily characterized by the absence of subapical spines. The new species closely resembles P. barbeae, from Papua New Guinea, Vanuatu and New Caledonia. However, both species can be distinguished on the basis of the following characters: - The anterior margin of thoracic sternite 3 is straight or slightly convex, and produced anterolaterally in P. barbeae, whereas this margin is medially produced, and anterolaterally rounded in P. pepei. - Antennal article 3 is often armed with a distomesial spine in P. barbeae, whereas this article is unarmed in P. pepei. - The P 2 – 4 propodi are slender in P. barbeae (5 – 7 × as long as wide), whereas they are stout (<5.8 × as long as wide) in P. pepei. - The colour pattern of the carapace and pleon are whitish, with reddish patches, in P. barbeae, whereas the colour pattern is green or light orange, with no reddish patches, in P. pepei. The genetic divergences between P. pepei and P. barbeae were 7 % (COI) and 3 % (16 S). The two sequences of P. pepei from Madagascar were divergent by 0.6 % for 16 S. The specimens of P. pepei range in size range from 2.5 to 3.2 mm postorbital carapace length.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF80E67F4F9CFB80736BBAD6.taxon	description	(Figs. 33 L, 41, 55 H)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF80E67F4F9CFB80736BBAD6.taxon	materials_examined	Type material. Holotype. Papua New Guinea. PAPUA NIUGINI Stn PB 12, 05.1966 ° S, 145.8133 ° E, 7 – 15 m, 30 December 2012: M 1.6 mm (MNHN-IU- 2014 - 13812). Paratypes. Papua New Guinea. PAPUA NIUGINI Stn PS 02, 05.2016 ° S, 145.8216 ° E, 15 – 17 m, 6 November 2012: 2 F 1.1 – 1.4 mm (MNHN-IU- 2019 - 2669). — Stn PD 19, 05.0900 ° S, 145.8083 ° E, 3 – 10 m, 13 November 2012: 2 M 1.3 – 1.7 mm, 1 ov. F 1.8 mm (MNHN-IU- 2019 - 2665). — Stn PD 23, 05.1000 ° S, 145.8200 ° E, 3 – 7 m, 14 November 2012: 1 M 2.0 mm, 2 ov. F 1.8 – 2.2 mm (MNHN-IU- 2014 - 13900). — Stn PD 24, 05.0883 ° S, 145.8100 ° E, 3 – 6 m, 14 November 2012: 2 ov. F 2.0 – 2.2 mm, 2 F 1.4 – 2.0 mm (MNHN-IU- 2016 - 450), 1 M 1.6 mm (MNHN- IU- 2016 - 451). — Stn PB 08, 05.1833 ° S, 145.8066 ° E, 4 – 5 m, 30 December 2012: M 2.1 mm, 3 ov. F 1.8 – 2.2 mm (MNHN-IU- 2019 - 2687). Other material. Papua New Guinea. PAPUA NIUGINI Stn PD 25, 05.0833 ° S, 145.8183 ° E, 3 – 5 m, 14 November 2012: 1 M postlarva 0.9 mm, 1 F 1.5 mm (MNHN-IU- 2019 - 2654). — Stn PB 19, 05.0850 ° S, 145.8100 ° E, 10 m, 16 November 2012: 3 M 1.0 – 1.5 mm (MNHN-IU- 2019 - 2673). — Stn PR 69, 05.0266 ° S, 145.8016 ° E, 2 – 15 m, 20 November 2012: 1 M 1.8 mm (MNHN-IU- 2014 - 13811). — Stn PD 56, 05.1616 ° S, 145.8050 ° E, 2 – 5 m, 29 November 2012: 1 ov. F 1.6 mm (MNHN-IU- 2019 - 2658). — Stn PR 129, 05.1883 ° S, 145.8250 ° E, 1 – 24 m, 29 November 2012: 1 ov. F 1.7 mm (MNHN-IU- 2014 - 13899). — Stn PR 142, 05.1966 ° S, 145.8200 ° E, 2 – 18 m, 30 November 2012: 1 M 1.2 mm, 1 F 1.3 mm (MNHN-IU- 2013 - 7074). — Stn PD 62, 05.1633 ° S, 145.8066 ° E, 1 – 3 m, 1 December 2012: 1 M 1.8 – 2.0 mm, 1 ov. F 1.8 mm (MNHN-IU- 2014 - 13879). — Stn PD 79, 05.1166 ° S, 145.8083 ° E, 20 m, 10 December 2012: 1 M 1.5 mm, 1 ov. F 1.8 mm (MNHN-IU- 2019 - 2671). — Stn PD 31, 05.0883 ° S, 145.8016 ° E, 1 – 6 m, 12 – 13 December 2012: 1 M 1.8 mm (MNHN-IU- 2014 - 13878). — Stn PD 19, 05.0900 ° S, 145.8083 ° E, 3 – 10 m, 30 December 2012: 1 M 1.5 mm (MNHN-IU- 2017 - 1343). — Stn PB 15, 05.0783 ° S, 145.8150 ° E, 5 m, 30 December 2012: 2 ov. F 1.5 – 1.6 mm, 1 F 1.8 mm (MNHN-IU- 2019 - 2659). — Stn PB 08, 05.1833 ° S, 145.8066 ° E, 4 – 5 m, 30 December 2012: 1 ov. F 1.6 mm (MNHN-IU- 2013 - 355), 1 M 1.4 mm, 6 ov. F 1.8 – 2.2 mm, 1 F 1.7 mm (MNHN- IU- 2016 - 454), 1 M 1.4 mm (MNHN-IU- 2014 - 13898). — Stn PS 31, 05.1361 ° S, 145.8236 ° E, 10 – 37 m, 30 December 2012: 1 M 1.3 mm (MNHN-IU- 2013 - 861). — Stn PS 15, 05.0965 ° S, 145.8032 ° E, 12 m, 30 December 2012: 1 M 1.7 mm, 1 ov. F 1.9 mm (MNHN-IU- 2019 - 2672). — Stn PB 37, 05.2650 ° S, 145.7850 ° E, 10 m, 30 December 2012: 1 ov. F 2.3 mm (MNHN-IU- 2016 - 1483). — Stn PS 08, 05.1839 ° S, 145.8071 ° E, 8 m, 30 December 2012: 1 M 1.4 mm, 1 ov. F 1.4 mm, 7 F 1.0 – 1.4 mm (MNHN-IU- 2014 - 13864). — Stn PB 19, 05.0850 ° S, 145.8100 ° E, 10 m, 30 December 2012: 2 M, 1.4 – 1.8 mm (MNHN-IU- 2014 - 13880). — Stn PS 23, 05.0762 ° S, 145.8201 ° E, 21 m, 30 December 2012: 1 M 1.7 mm (MNHN-IU- 2019 - 2657). — Stn PB 24, 04.9850 ° S, 145.7933 ° E, 1 m, 30 December 2012: 1 M 1.3 mm (MNHN-IU- 2019 - 2656). — Stn PB 53, 05.1350 ° S, 145.8033 ° E, 3 m, 30 December 2012: 1 F 1.5 mm (MNHN-IU- 2019 - 2655). — KAVIENG Stn KB 54, 02.5316 ° S, 150.4583 ° E, 11 m, 20 June 2014: 1 ov. F 2.5 mm (MNHN- IU- 2014 - 2893).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF80E67F4F9CFB80736BBAD6.taxon	etymology	Etymology. From the name Phanus, an Argonaut, son of Dionysus and Ariadne. The name is considered a substantive in apposition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF80E67F4F9CFB80736BBAD6.taxon	description	Description. Carapace: Robust or massive, sexually dimorphic (wider on females) 0.9 – [1.0] (males), 0.7 – 0.9 (females) × as long as broad; transverse ridges elevated, upraised dorsally, with short setae and scattered thick setae. Gastric region convex (upraised) with some transverse ridges: epigastric ridge unarmed, scale-like, continuing laterally with few small scales, often indistinct; anterior protogastric ridge not medially interrupted, laterally interrupted with some scales, often followed by some short scales; anterior mesogastric ridge scale-like, laterally interrupted by anterior branch of cervical groove, continuing with few scales, often followed by some short scales; anterior metagastric ridge medially uninterrupted, laterally interrupted, followed by some scales on posterior metagastric region. Mid-transverse ridge not interrupted, cervical groove distinct, followed by 2 uninterrupted or minutely interrupted ridges, interspersed with 1 laterally interrupted or scale-like ridge and 1 – 2 short lateral ridges. Lateral margins clearly convex, with 7 – 8 spines: first anterolateral spine well-developed, exceeding lateral orbital spine, second spine (hepatic) minute, often obsolescent in males, slightly dorsomesially from lateral margin, and followed by 5 – 6 branchial spines behind distinct anterior cervical groove (3 anterior and 2 – 3 posterior). Rostrum leaf-like, horizontal, dorsally slightly or deeply concave, [1.5] 1.6 – 1.9 × as long as broad, length 0.3 – [0.5] and breadth 0.2 – [0.3] that of carapace; lateral margins minutely serrated and convex, with well-developed supraocular and subapical spines. Pterygostomian flap ending in acute tooth; surface unarmed. Thoracic sternum: As wide as long or slightly wider than long, lateral margins of posterior half slightly divergent. Sternite 3 moderately broad, [1.9] – 2.4 × as wide as long, anterior margin convex. Sternite 4 widely contiguous to sternite 3; surface flattened, smooth; greatest width [2.8] – 3.1 × that of sternite 3, 2.0 – [2.3] × as wide as long. Pleon: Elevated ridges upraised dorsally, with few short setae. Tergites 2 – 4 with anterior and posterior transverse elevated ridges; tergites 5 – 6 smooth. Eye: Eyestalk length about 1.0 × broader than long, peduncle distally setose, not distinctly expanded proximally, with few short transverse striae on lateral surfaces; maximum corneal diameter 0.9 – [1.0] × rostrum width, as wide as eyestalk. Antennule: Article 1 longer than wide, with 3 – 4 distal spines: distomesial spine small or obsolescent; proximal lateral spine absent. Antenna: Article 1 with prominent mesial process distally not reaching lateral antennular spine. Article 2 often with minute distal spines laterally and mesially. Articles 3 and 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margin. Merus 0.6 × length of ischium, with 1 well-developed distal spine on extensor margin and 2 spines, subequal in size, on flexor margin. P 1: [2.4] 2.0 – 2.7 × carapace length (males), 1.3 (females), subcylindrical, with scattered spines and long stiff setae; merus and carpus with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus [0.9] 0.5 – 1.0 length of carapace, [1.6] 1.5 – 2.1 × as long as carpus. Carpus 1.1 – [1.9] × as long as wide. Palm [0.9] 0.7 – 1.0 × carpus length, 1.1 – [1.4] × as long as broad with scattered small spines on dorsal and ventral surfaces, lateral and mesial margin irregular, with 1 small spine on distal mesial margin. Fingers unarmed, [0.9] – 1.3 × palm length. P 2 – 4 (P 2 – 3 absent from holotype): Moderately stout, setose and few spinose. Meri successively shorter posteriorly: P 3 merus 0.9 × length of P 2 merus, P 4 merus 0.7 – 0.8 × length of P 3 merus. P 2 merus, 1.7 – 2.4 × carapace length, 3.8 – 4.0 × as long as broad, 1.2 × as long as P 2 propodus; P 3 merus 3.6 – 4.0 × as long as broad, 1.1 – 1.2 × as long as P 3 propodus; P 4 merus 2.7 – 2.8 × as long as broad, as long as P 4 propodus; extensor margins of P 2 and P 3 with row of few small spines, proximally diminishing, with well-developed distal spine; P 4 extensor margin irregular but unarmed, distal spine absent; flexor margins of all legs irregular, with distal spine on P 2 – 3, distal spine absent in P 4. Carpi with 1 spine on extensor margins on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, absent on P 4; row of small spines below extensor margins on lateral surface of P 2 – 3, unarmed on P 4. Propodi stout, 4.0 – 5.0 × as long as broad; extensor margins irregular, usually unarmed; flexor margins with 5 – 6 slender movable spines in addition to distal pair. Dactyli 0.7 – 0.8 × length of propodi, ending in incurved, strong, sharp spine; flexor margins with 5 – 6 well-developed dactylar spines, each with 1 spinule. Eggs: Ov. F carried approximately 8 – 30 eggs of 0.3 – 0.4 mm diameter. Live colour. Body orange, with dark orange-reddish patches. P 1 pale yellow, with dark orange-brownish bands. P 2 – 4 pale yellow, with dark orange bands. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF80E67F4F9CFB80736BBAD6.taxon	distribution	Distribution. Papua New Guinea, between 1 and 37 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF80E67F4F9CFB80736BBAD6.taxon	discussion	Remarks. Phylladiorhynchus phanus belongs to the group of species having the epigastric ridge unarmed, the carapace and pleon ridges elevated, upraised, and dactylar spines on the flexor margins of the P 2 - 4 dactyli. The new species is closely related to P. marina from Vanuatu. Both species are highly similar, and very difficult to distinguish using morphological characters only, therefore they are considered cryptic species. Both species might be distinguished on the basis of their different coloration (the base color of the carapace is orange in P. phanus, whereas the overall colour pattern is brownish in P. marina, compare Figures 55 B and 55 H), however, we can’t discount intraspecific variability and this character might not be reliable. There are some scattered long plumose setae on the carapace ridges of P. marina, whereas these setae are always absent in P. phanus. On the other hand, the setae can be missing in preserved specimens. P. phanus is also close to P. phlias, from Papua New Guinea (see the differences under the Remarks of this species). The genetic divergences between P. phanus and P. marina were 13 % (COI) and 4 % (16 S). The two and three sequences of P. phanus from Papua New Guinea diverged 0.7 % (COI) and 0.1 % (16 S). The specimens of P. phanus range in size from 0.9 to 2.5 mm postorbital carapace length.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF87E67D4F9CFBB17280B9CA.taxon	description	(Figs. 42, 50 A, 55 I)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF87E67D4F9CFBB17280B9CA.taxon	materials_examined	Type material. Holotype. Papua New Guinea. KAVIENG Stn KZ 16, 02.5783 ° S, 150.7916 ° E, 1 – 2 m, 23 June 2014: ov. F 2.2 mm (MNHN-IU- 2014 - 13535). Paratypes. Papua New Guinea. PAPUA NIUGINI Stn PS 09, 05.2050 ° S, 145.8133 ° E, 8 – 10 m, 9 November 2012: 1 M 1.2 mm, 1 ov. F 1.8 mm (MNHN-IU- 2014 - 13886). — Stn PD 19, 05.0900 ° S, 145.8083 ° E, 10 m, 13 November 2012: 3 M 1.2 – 1.8 mm, 2 ov. F 1.5 – 1.7 mm, 2 F broken (MNHN-IU- 2019 - 2674), 1 ov. F 1.3 mm (MNHN-IU- 2013 - 703). — Stn PS 13, 05.0984 ° S, 145.8210 ° E, 8 m, 30 December 2012: 7 M 1.1 – 1.4 mm, 3 ov. F 1.4 – 1.6 mm, 1 F 1.0 mm (MNHN-IU- 2014 - 13881). Other material. Papua New Guinea. PAPUA NIUGINI Stn PD 24, 05.0883 ° S, 145.8100 ° E, 3 – 6 m, 14 November 2012: 2 ov. F 1.5 – 1.9 mm 2 F 1.9 – 2.0 mm (MNHN-IU- 2019 - 2660), 1 M 1.4 mm (MNHN-IU- 2016 - 452). — Stn PD 65, 05.1416 ° S, 145.8083 ° E, 1 – 4 m, 1 December 2012: 1 M 1.3 mm (MNHN-IU- 2014 - 139139). — Stn PB 50, 05.0783 ° S, 145. 05 ° 04.7 ’ S, 145 ° 48.9 ’ E 8150 ° E, 3 m, 30 December 2012: 1 M 1.7 mm (MNHN-IU- 2016 - 1492). — Stn PS 12, 05.0888 ° S, 145.8094 ° E, 6 m, 30 December 2012: 1 M 1.4 mm, 2 ov. F 1.3 – 1.5 mm, 2 F 1.1 – 1.4 mm (MNHN-IU- 2014 - 13866). — Stn PS 18, 05.0266 ° S, 145.8009 ° E, 16 m, 30 December 2012: 1 ov. F 1.5 mm (MNHN- IU- 2019 - 2666). — KAVIENG Stn KS 07, 02.6866 ° S, 150.6866 ° E, 8 m, 4 June 2014: 1 F 1.4 mm (MNHN-IU- 2016 - 5832). — Stn KD 36, 02.5850 ° S, 150.4833 ° E, 8 m, 16 June 2014: 1 F 1.4 mm (MNHN-IU- 2014 - 13695).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF87E67D4F9CFBB17280B9CA.taxon	etymology	Etymology. From the name Phlias, an Argonaut, son of Dionysus and Ariadne. The name is considered a substantive in apposition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF87E67D4F9CFBB17280B9CA.taxon	description	Description. Carapace: Robust or massive, sexually dimorphic (wider on females) 0.9 – 1.0 (males), [0.7] – 0.8 (females) × as long as broad; transverse ridges elevated, upraised dorsally, with very few short setae. Gastric region convex (upraised) with some transverse ridges: epigastric ridge indistinct, unarmed, with some scales, anterior protogastric ridge not medially interrupted, laterally interrupted with some scales to carapace margin, sometimes followed by a few some short scales posteriorly; anterior mesogastric ridge scale-like, otherwise indistinct, laterally interrupted by anterior branch of cervical groove, laterally continuing with few scales; often followed by short scales; anterior metagastric ridge scale-like, followed by few scales on posterior metagastric region. Mid- transverse ridge not interrupted, preceded by shallow or indistinct cervical groove, followed by 2 uninterrupted or minutely interrupted ridges, interspersed with 1 laterally interrupted, complete or scale-like ridge and 1 short lateral ridge. Lateral margins clearly convex, with 4 – 5 spines: first anterolateral spine well-developed, reaching end of lateral orbital spine, hepatic spine absent, followed by 3 – 4 spines (2 anterior well-developed branchial spines subequal in size, second slightly dorsomesially from lateral margin, 1 – 2 posterior branchial spines very small). Rostrum bottle-shaped (proximally straight and distally convex), horizontal, dorsally flattish or slightly concave, [1.1] – 1.3 (females), 1.8 (males) × as long as broad, length [0.3] – 0.5 and breadth 0.2 that of carapace; lateral margins finely serrated, with well-developed supraocular spines, subapical spines small. Pterygostomian flap ending in blunt tooth; upper margin smooth. Thoracic sternum: As wide as long or slightly wider than long, lateral margins of posterior half slightly divergent. Sternite 3 moderately broad, [2.6] 1.7 – 2.7 × as wide as long, anterior margin straight or slightly convex. Sternite 4 widely contiguous to and wider than sternite 3; surface flattened, smooth; greatest width 3.3 – [3.4] × that of sternite 3, 2.4 – [2.5] × as wide as long. Pleon: Elevated ridges upraised dorsally, with few short setae. Tergites 2 – 3 with anterior and posterior transverse elevated ridges; tergite 4 with anterior transverse ridge only; tergites 5 – 6 smooth. Eye: Eyestalk length about 0.9 × broader than long, peduncle distally setose, not distinctly expanded proximally; maximum corneal diameter 0.9 × rostrum width, as wide as eyestalk. Antennule: Article 1 slightly longer than wide, with 3 – 4 distal spines: distomesial spine minute or absent; proximal lateral spine absent. Antenna: Article 1 with prominent mesial process distally not reaching lateral antennular spine. Article 2 unarmed, sometimes with minute distomesial spine. Articles 3 and 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus [0.4] 0.4 × length of ischium, with 0 – 1 median small spine and well-developed distal spine on extensor margin and 2 smaller spines, subequal in size, on flexor margin. P 1 (lost in holotype): 2.2 – 2.3 × carapace length (males), 1.4 (females); subcylindrical, with scattered spines and long stiff setae, with some thick short setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus 0.5 – 0.8 length of carapace, 1.7 – 2.0 × as long as carpus. Carpus 1.3 – 1.7 × as long as wide. Palm 0.8 – 1.0 × carpus length, 1.2 – 1.3 × as long as broad irregular and unarmed in all the surfaces, otherwise a small spine on basal mesial margin. Fingers unarmed, 1.0 – 1.3 × palm length. P 2 – 4 (lost in most specimens): Stout, moderately setose and spinose. Meri successively shorter posteriorly: P 3 merus 0.9 × length of P 2 merus, P 4 merus 0.7 – 0.9 × length of P 3 merus. P 2 merus, 0.4 – 0.5 × carapace length, 3.2 × as long as broad, 1.2 – 1.3 × as long as P 2 propodus; P 3 merus 3.2 – 3.5 × as long as broad, 1.0 – 1.2 × as long as P 3 propodus; P 4 merus 2.8 – 3.2 × as long as broad, as long as P 4 propodus; extensor margins of P 2 – 3 irregular, unarmed, with small distal spine; P 4 extensor margin irregular, unarmed, distal spine absent; flexor margins of all legs irregular, with distal spine on P 2 – 3, distal spine absent in P 4. Carpi extensor margins irregular or granulated, unarmed on P 2 – 4. Propodi stout, 3.5 – 4.5 × as long as broad; extensor margins irregular, usually unarmed; flexor margins with 2 – 4 slender movable spines in addition to distal pair. Dactyli 0.7 – 0.9 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 4 – 6 well-developed dactylar spines, each with movable spinule. Eggs: Ov. F carried approximately 5 – 15 eggs of 0.4 – 0.5 mm diameter. Live colour. Body whitish- pale yellow, with small brownish spots. P 1 whitish-pale yellow. P 2 – 4 whitish, with translucid bands. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF87E67D4F9CFBB17280B9CA.taxon	distribution	Distribution. Papua New Guinea, between 1 and 16 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF87E67D4F9CFBB17280B9CA.taxon	discussion	Remarks. Phylladiorhynchus phlias belongs to the group of species characterized by having the epigastric ridge unarmed, the carapace and pleon ridges elevated, upraised, and dactylar spines on the flexor margins of the P 2 - 4 dactyli. The closest species are P. phanus, from Papua New Guinea, and P. marina, from Vanuatu, however these species can be distinguished on the basis of the following morphological characters: - The rostrum is bottle-shaped in P. phlias, whereas it is leaf-like in the other species. - The hepatic spine is absent in P. phlias, whereas it is present in P. phanus and P. marina. - The anterior branchial margin has 2 branchial spines in P. phlias, whereas this margin has 3 spines in in the other species. The genetic divergences between P. phlias, and these species exceeded 15 % (COI) and 12 % (16 S), respectively. The two sequences of P. phlias from Papua New Guinea diverged 0.5 % for COI. The specimens of P. phlias range in size 1.1 – 2.2 mm postorbital carapace length.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF9AE6604F9CFF2970CCBF4E.taxon	description	(Figs. 43, 50 B, 56 A)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF9AE6604F9CFF2970CCBF4E.taxon	materials_examined	Type material. Holotype. French Polynesia. BENTHAUS Stn DW 1866, 28.9833 ° S, 140.2475 ° W, 50 – 100 m, 4 November 2002: M 2.8 mm (IU- 2014 - 13736). Paratypes. French Polynesia. BENTHAUS Stn DW 1866, 28.9833 ° S, 140.2475 ° W, 50 – 120 m, 4 November 2002: 5 M 2.2 – 3.5 mm, 5; ov. F 2.1 – 3.4 mm 2 F 3.0 – 3.4 mm, 1 M 3.4 mm (MNHN-IU- 2016 - 9658), 1 M 2.5 mm (MNHN-IU- 2014 - 13859). — Stn DW 1867, 28.9833 ° S, 140.2333 ° W, 127 – 170 m, 4 November 2002: 2 M 2.0 – 2.2 mm (MNHN-IU- 2019 - 2681). — Stn DW 1876, 28.9833 ° S, 140.2500 ° W, 150 – 160 m, 4 November 2002: 2 M 2.1 – 2.8 mm (MNHN-IU- 2019 - 2693). — Stn DW 1898, 27.5666 ° S, 144.4500 ° W, 580 – 820 m, 8 November 2002: 1 M 3.3 mm (MNHN-IU- 2014 - 13737). Other material. French Polynesia. BENTHAUS Stn DW 1877, 28.9833 ° S, 140.2500 ° W, 59 – 150 m, 4 November 2002: 1 M 2.8 mm (MNHN-IU- 2014 - 13904), 8 M 2.3 – 3.7 mm, 5 ov. F 2.5 – 3.4 mm, 3 F 2.5 – 2.8 mm (MNHN-IU- 2014 - 13738), 1 M 3.6 mm (MNHN-IU- 2014 - 13731), 9 M 1.7 – 4.0 mm, 8 F 2.0 - 3.1 mm (MNHN-IU- 2019 - 2677), 1 M 3.4 mm, 1 ov. F 2.8 mm (MNHN-IU- 2016 - 9634). — Stn DW 1880, 27.9133 ° S, 143.4908 ° W, 90 – 94 m, 6 November 2002: 2 M 2.0 - 2.3 mm, 4 ov. F 1.8 – 2.8 mm (MNHN-IU- 2019 - 2682), 3 M 2.1 – 2.8 mm, 1 ov. F 2.0 mm (MNHN-IU- 2014 - 13857). — Stn DW 1881, 27.9166 ° S, 143.4833 ° W, 112 - 121 m, 6 November 2002: 1 M 2.2 mm (MNHN-IU- 2019 - 2684), 1 ov. F 2.6 mm (MNHN-IU- 2019 - 2680). — Stn DW 1894, 27.6688 ° S, 144.3585 ° W, 100 m, 8 November 2002: 2 M 1.8 - 2.2 mm (MNHN-IU- 2019 - 2664), 1 ov. F 2.4 mm (MNHN-IU- 2019 - 2661). — Stn CP 1908, 27.4166 ° S, 144.0166 ° W, 100 – 118 m, 9 November 2002: 3 M 2.3 – 3.4 mm, 1 ov. F 2.5 mm, 2 F 2.0 – 2.2 mm (MNHN-IU- 2019 - 2683). — Stn CP 1918, 27.0500 ° S, 146.0666 ° W, 130 – 140 m, 12 November 2002: 1 M 2.3 mm (MNHN-IU- 2019 - 2685). — Stn DW 1936, 24.6616 ° S, 145.9515 ° W, 80 - 100 m, 14 November 2002: 1 ov. F 2.0 mm (MNHN-IU- 2019 - 2663). — Stn DW 1939, 23.8278 ° S, 147.6936 ° W, 100 m, 15 November 2002: 1 M 2.0 mm (MNHN-IU- 2019 - 2679). — Stn DW 2013, 22.6428 ° S, 152.8288 ° W, 80 – 93 m, 25 November 2002: 1 M 2.3 mm (MNHN-IU- 2019 - 2678).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF9AE6604F9CFF2970CCBF4E.taxon	etymology	Etymology. From the name Poeas, an Argonaut, son of Thaumacus and father of Philoctetes. The name is considered a substantive in apposition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF9AE6604F9CFF2970CCBF4E.taxon	description	Description. Carapace: as long as or slightly longer than broad; transverse ridges with dense short setae and few scattered long and thick setae. Gastric region slightly convex with 4 transverse ridges: epigastric ridge distinct with 1, 2 or 3 (in largest specimens) pairs of spines in transverse row, innermost pair always the largest, outer pairs (when present) smaller, sometimes indicated by granules; anterior protogastric ridge non interrupted medially, nearly extending laterally to carapace margin; anterior mesogastric ridge not medially interrupted, laterally interrupted by cervical groove, laterally continuing to first branchial spine; anterior metagastric ridge scale-like. Mid-transverse ridge uninterrupted, preceded by distinct cervical groove, followed by 2 uninterrupted or minutely interrupted ridges, interspersed with 1 short lateral ridge. Lateral margins convex, with 6 – 7 spines: first anterolateral spine well-developed, reaching anteriorly to level of lateral orbital spine, second spine (hepatic) small to obsolescent, slightly dorsomesially from lateral margin, and followed by 4 – 5 branchial spines behind distinct anterior cervical groove (3 anterior and 1 – 2 posterior). Rostrum leaf-like, horizontal, dorsally flattish or slightly concave, [1.6] – 1.8 × as long as broad, length [0.4] – 0.5 and breadth [0.2] – 0.3 that of carapace; lateral margins serrated and convex, with well-developed supraocular basal spines and small subapical spines. Pterygostomian flap with anterior spine; upper margin smooth. Thoracic sternum: As wide as long. Sternite 3 moderately broad, 1.5 – [2.2] × as wide as long, anterior margin convex, often serrated, moderately produced anterolaterally. Sternite 4 widely contiguous to sternite 3; surface depressed in midline, smooth; greatest width 2.7 – [2.8] × that of sternite 3, 2.4 – [2.7] × as wide as long. Pleon: Elevated ridges with short setae and a few scattered long setae. Tergite 2 with anterior and posterior transverse elevated ridges; tergites 3 – 4 with anterior transverse ridge only; tergites 5 – 6 smooth. Eye: Eyestalk length about 0.8 × broader than long, peduncle distally setose, not distinctly expanded proximally; maximum corneal diameter 0.9 × rostrum width, as wide as eyestalk. Antennule: Article 1 slightly longer than wide, with 5 distal spines: distomesial spine well-developed; proximal lateral spine small, always present. Antenna: Article 1 with prominent mesial process, distally falling well short of lateralmost antennular spine. Article 2 with well-developed distomesial and distolateral spines. Article 3 with distomesial spine. Article 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus [0.6] – 0.7 × length of ischium, with well-developed distal spine on extensor and flexor margins. P 1 (lost in holotype): 2.8 – 3.1 (males), 2.4 – 2.5 (females) × carapace length; subcylindrical, spiny and with dense long stiff setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus 1.0 – 1.1 length of carapace, 1.7 – 1.9 × as long as carpus. Carpus 1.4 – 1.8 × as long as wide. Palm 1.2 – 1.3 × carpus length, 1.8 – 2.4 × as long as broad. Fingers 0.7 – 0.8 × palm length; fixed finger with several proximal spines; movable finger with 1 well-developed basal spine and several small spines or granules along dorsal margin. P 2 – 4: Densely setose and spinose. Meri successively shorter posteriorly: P 3 merus 0.8 × length of P 2 merus, P 4 merus 0.9 – 1.0 × length of P 3 merus. P 2 merus, 0.6 × carapace length, [4.0] – 4.4 × as long as broad, [1.2] – 1.6 × as long as P 2 propodus; P 3 merus 3.6 – 4.0 × as long as broad, 1.3 – 1.4 × as long as P 3 propodus; P 4 merus 2.7 – [3.8] × as long as broad, [1.1] – 1.2 × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, unarmed; flexor margin irregular, with distal spine on P 2 – 3, distal spine absent in P 4. Carpi with 0 – 2 prominent spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, smaller on P 4; row of small spines below extensor margin on lateral surface of P 2 – 3, unarmed on P 4; flexor margin unarmed. Propodi moderately stout, [5.0 – 5.5] 4.5 – 6.0 × as long as broad; extensor margin irregular, armed with 1 – 4 distinct well-developed spines; flexor margin with 3 - 4 slender movable spines in addition to distal pair. Dactyli [0.6] – 0.8 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 4 – 6 movable spines. Eggs: Ov. F carried approximately 10 – 30 eggs of 0.3 mm diameter. Live colour. Body light orange to dark orange, covered by orange spots. P 1 dark orange, covered by light orange setae. P 2 – 4 light orange, darker on extensor and flexor margins. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF9AE6604F9CFF2970CCBF4E.taxon	distribution	Distribution. French Polynesia, almost all specimens were found between 50 and 160 m. A single specimen was found in a station of 580 – 820 m depth (MNHN-IU- 2014 - 13737) that we consider as an anomaly.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF9AE6604F9CFF2970CCBF4E.taxon	discussion	Remarks. Phylladiorhynchus poeas is morphologically undistinguishable from P. pusillus from Australia, Tasmania and New Zealand and they can be considered as cryptic species. However, genetically they are very different for mitochondrial genes. Schnabel & Ahyong (2019) revised the type material and numerous topotypic specimens of P. pusillus and found that one of the diagnostic characters of the species (4 spines on the epigastric ridge) showed a certain variability. After examination of a large amount of material from Australia, Tasmania and New Zealand, Schnabel & Ahyong (2019) reported a variation in the number and size of the epigastric spines, from 1 to 3 pairs of epigastric spines, with the lateralmost being granules in small specimens. The new species from French Polynesia has a similar variation in this character, with specimens having 2 to 6 spines and / or granules, recommending caution when using this character for species delimitation. Unfortunately, we don’t know the colour pattern of P. pusillus to compare with the colouration of the new species. Therefore, future studies will confirm the existence of additional differences to separate morphologically both species. A subtle character useful to distinguish both species might be the thoracic sternite 3 moderately produced anterolaterally in P. poeas; whereas with broad granule or square in P. pusillus (Schnabel & Ahyong 2019). The genetic divergences between P. poeas and P. pusillus were 8 % (COI) and 2 % (16 S). No intraspecific divergence was observed in both genes. The specimens of P. poeas range from 1.8 to 3.5 mm postorbital carapace length.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF99E6674F9CFF297563B8CE.taxon	description	(Figs. 44, 50 C)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF99E6674F9CFF297563B8CE.taxon	materials_examined	Type material. Holotype. New Caledonia. LIFOU Stn 1451, 20.7883 ° S, 167.1133 ° E, 10 – 21 m, 19 November 2000: ov. F 3.5 mm (MNHN- IU- 2014 - 13797) Paratype. New Caledonia, Recif Mbere, 22.3316 ° S, 166.2200 ° E, 10 m, 5 May 1993: 1 M 3.8 mm (MNHN-IU- 2019 - 2698).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF99E6674F9CFF297563B8CE.taxon	etymology	Etymology. From the name Pollux, an Argonaut, son of Zeus and Leda. The name is considered a substantive in apposition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF99E6674F9CFF297563B8CE.taxon	description	Description. Carapace: As long as or slightly longer than broad; transverse ridges with dense short setae, and few scattered short and thick setae. Gastric region slightly convex, with some transverse ridges: epigastric ridge armed with 4 epigastric spines; anterior protogastric ridge not medially interrupted, nearly extending laterally to carapace margin, followed by a posterior protogastric ridge not medially interrupted; anterior mesogastric ridge not medially interrupted, laterally continuing to first branchial spine, posterior mesogastric ridge absent or scale like; anterior metagastric ridge not medially interrupted, laterally continuing to second branchial spine, followed by posterior median metagastric scale; secondary scales can be present between ridges. Mid-transverse ridge not interrupted, medially depressed, preceeded by distinct cervical groove, followed by 2 uninterrupted or minutely interrupted ridge, interspersed with 2 scale-like ridges. Lateral margins slightly convex, with 5 spines: first anterolateral spine well-developed, overreaching level of lateral orbital spine, second spine (hepatic) well-developed, slightly dorsomesial from lateral margin, and followed by 3 branchial spines (2 anterior and 1 posterior). Rostrum leaf-like, horizontal, dorsally flattish, narrow [1.7] – 1.9 × as long as broad, length 0.4 and breadth 0.2 that of carapace; lateral margins smooth and convex, with well-developed supraocular basal supraocular spines, subapical spines minutes. Pterygostomian flap ending in blunt tooth, upper margin slightly irregular. Thoracic sternum: As wide as long. Sternite 3 moderately broad, 2.0 – [3.0] × as wide as long, anterior margin nearly straight. Sternite 4 widely contiguous to sternite 3; surface depressed in midline, smooth; greatest width [3.5] – 4.6 × that of sternite 3, 2.6 – [2.7] × as wide as long. Pleon: Elevated ridges with with short setae and a few scattered thick setae. Tergite 2 – 3 with anterior and posterior transverse elevated ridges; tergite 4 with anterior transverse ridge; tergites 5 – 6 smooth. Eye: Eyestalk length about 0.9 – [1.0] × broader than long, peduncle distally setose, not distinctly expanded proximally; maximum corneal diameter [1.1] – 1.2 × rostrum width, as wide as eyestalk. Antennule: Article 1 slightly longer than wide, with 4 – 5 well-developed distal spines: distomesial spine welldeveloped; proximal lateral spine absent or minute. Antenna: Article 1 with prominent mesial process, distally falling well short of lateral antennular spine. Article 2 with small distomesial and distolateral spines. Articles 3 and 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus and ischium subequal in size, with well-developed distal spine on extensor and flexor margins. P 1: 3.4 (male), [2.1] (female) × carapace length; subcylindrical, spiny and densely covered with plumose setae and scattered long stiff setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus 1.2 – 1.3 length of carapace, twice as long as carpus. Carpus twice as long as wide. Palm 1.3 × carpus length, 2.8 × as long as broad. Fingers 0.7 × palm length; fixed finger unarmed; movable finger with small basal spine. P 2 – 3 (P 4 lost in both specimens): Moderately stout, subcylindrical, highly setose and spinose, densely covered with plumose setae: P 3 merus 0.6 – [0.9] × length of P 2 merus. P 2 merus, 0.8 × carapace length, 4.4 – [4.7] × as long as broad, 1.2 – [1.4] × as long as P 2 propodus; P 3 merus [4.3] – 4.6 × as long as broad, [1.1] – 1.2 × as long as P 3 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; flexor margin irregular, with distal spine on P 2 – 3. Carpi with 3 – 4 spines on extensor margin on P 2 – 3; distal spine prominent; row of small spines below extensor margin on lateral surface of P 2 – 3; flexor margin unarmed or with small distal spine. Propodi moderately stout, 5.0 – 6.5 × as long as broad; extensor margin irregular, armed proximally with 2 – 4 small spines on P 2 – 3; flexor margin with 3 – 4 slender movable spines in addition to distal pair. Dactyli 0.5 – 0.6 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 5 – 6 movable spines. Eggs. No data. The holotype carried one egg of 0.5 mm diameter. Live colour. Unknown. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF99E6674F9CFF297563B8CE.taxon	distribution	Distribution. New Caledonia, between 10 and 21 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF99E6674F9CFF297563B8CE.taxon	discussion	Remarks. Phylladiorhynchus pollux belongs to the group of species having 4 epigastric spines and Mxp 3 merus with one prominent spine along the flexor margin. The group of species includes P. australis, from New Zealand and Southern Australia, P. integrus from Japan to Chesterfield Islands, P. lenzi from Chile, P. nui from southeastern Australia, Tasmania and New Zealand, P. poeas, from French Polynesia, P. porteri, from Chile, and P. pusillus from the southwestern Pacific. However, the new species is easily distinguished from these species by the number of spines on the anterior branchial margin: 2 spines in P. pollux and 3 spines in the other species. Furthermore, the posterior protogastric ridge is present and entire in P. pollux, being absent or scale-like in the other species. The sequences of P. pollux were more than 22 % (COI) and 11 % (16 S) divergent from all other species.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF9FE6654F9CFDA97537BA0A.taxon	description	(Figs. 45, 50 D)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF9FE6654F9CFDA97537BA0A.taxon	materials_examined	Type material. Holotype. Chile, Valparaiso, coll. Porter, 1899: M 2.7 mm (MNHN-IU- 2019 - 2598 (Ga- 763 )). Paratypes. Chile, Valparaiso, coll. Porter, 1899: 3 M 2.0 – 2.7 mm, 4 ov. F 2.4 – 2.8 mm (MNHN-IU- 2014 - 23831 (Ga- 763 )).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF9FE6654F9CFDA97537BA0A.taxon	etymology	Etymology. Named after C. E. Porter, Chilean naturalist, who collected the specimens.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF9FE6654F9CFDA97537BA0A.taxon	description	Description. Carapace: As long as broad; transverse ridges with dense short setae, and few scattered long thick setae. Gastric region slightly convex with 4 transverse ridges: epigastric ridge distinct with 2 or 3 (if specimen is massive) pairs of spines in transverse row, innermost pair always the largest, outer pair (s) prominent, smaller or absent, sometimes indicated by granules; anterior protogastric ridge medially uninterrupted, nearly extending laterally to carapace margin; anterior mesogastric ridge medially interrupted, with several median scales, laterally interrupted by cervical groove, laterally continuing uninterrupted to first branchial spine; anterior metagastric ridge scale-like. Mid-transverse ridge uninterrupted, preceded by distinct cervical groove, followed by 2 uninterrupted or minutely interrupted ridges, interspersed with 1 short lateral ridge. Lateral margins convex, with 7 spines: first anterolateral spine well-developed, reaching anteriorly to level of lateral orbital spine, second spine (hepatic) small to obsolescent, slightly dorsomesially from lateral margin, and followed by 5 branchial spines behind distinct anterior cervical groove (3 anterior and 2 posterior). Rostrum leaf-like, lanceolate, horizontal, dorsally flattish or slightly concave, 1.4 – [1.8] × as long as broad, length 0.4 and breadth [0.2] – 0.3 that of carapace; lateral margins smooth and convex, with well-developed supraocular basal spines and small subapical spines. Pterygostomian flap ending in blunt tooth; upper margin unarmed. Thoracic sternum: As wide as long. Sternite 3 moderately broad, 1.8 – [2.0] × as wide as long, anterior margin convex. Sternite 4 narrowly contiguous to sternite 3; surface depressed in midline, smooth; greatest width [2.9] – 3.0 × that of sternite 3, 2.0 – [3.0] × as wide as long. Pleon: Elevated ridges with short setae and a few scattered long setae. Tergite 2 with anterior and posterior transverse elevated ridges; tergites 3 – 4 with anterior transverse ridge only; tergites 5 – 6 smooth. Eye: Eyestalk length about [0.9] – 1.0 × broader than long, peduncle distally setose, not distinctly expanded proximally, with few short transverse striae on lateral surfaces; maximum corneal diameter 0.7 – [0.8] × rostrum width, as wide as eyestalk (0.9 × maximum peduncle width). Antennule: Article 1 slightly longer than wide, with 5 distal spines: distomesial spine well-developed; proximal lateral spine small, always present. Antenna: Article 1 with prominent mesial process, distally falling well short of lateralmost antennular spine. Article 2 with well-developed distal spines laterally and mesially. Article 3 with distomesial and distolateral spines. Article 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus 0.6 – [0.7] × length of ischium, with well-developed distal spine on extensor margin and one much larger spine on flexor margin. P 1 (only in males, all missing in females): [3.0] × carapace length, subcylindrical, spiny and with dense long stiff setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus 1.4 length of carapace, [1.6] – 2.3 × as long as carpus. Carpus 1.5 – [1.7] × as long as wide. Palm [1.2] – 1.4 × carpus length, 1.8 – [1.9] × as long as broad. Fingers 0.8 × palm length; movable and fixed fingers with several proximal marginal spines. P 2 – 4: (attached legs only) Stout, densely setose and spinose. Meri successively shorter posteriorly: P 3 merus 0.6 × length of P 2 merus, P 4 merus 0.8 × length of P 3 merus. P 2 merus, 0.8 × carapace length, 2.7 – [3.6] × as long as broad, 1.3 – [1.6] × as long as P 2 propodus; P 3 merus 4 twice as long as broad, 1.3 × as long as P 3 propodus; P 4 merus 3.6 × as long as broad, 1.2 × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, unarmed; flexor margins irregular, with distal spine on P 2 – 3, distal spine absent in P 4. Carpi with prominent spines on extensor margin on P 2 – 3, unarmed on 4; distal spine prominent on P 2 – 3, smaller on P 4; row of small spines below extensor margin on lateral surface of P 2 – 3, unarmed on P 4; flexor margin unarmed. Propodi stout, [3.5] 3.3 – 4.3 × as long as broad; extensor margin irregular usually armed with 1 – 4 well-developed spines; flexor margin with 4 slender movable spines in addition to distal pair. Dactyli 0.6 – [0.8] × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 4 – 6 movable spines. Eggs. No data. Live colour. Unknown. Genetic data. No data.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF9FE6654F9CFDA97537BA0A.taxon	distribution	Distribution. Chile, Valparaíso, unknown depth.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF9FE6654F9CFDA97537BA0A.taxon	discussion	Remarks. Phylladiorhynchus porteri belongs to the group of species having usually 4 spines on the epigastric ridge, the anterior metagastric ridge scale-like, a very small hepatic spine, the anterior margin of the thoracic sternite 3 convex and one spine on the flexor margin of the Mxp 3 merus. Phylladiorhynchus porteri is closely related to P. lenzi (Rathbun, 1907), from Chile. However, they can be easily distinguished by the following aspects: - The rostrum is more lanceolate in P. porteri than in P. lenzi. - The anterior mesogastric ridge is medially interrupted, with few medial scales in P. porteri, whereas this ridge is uninterrupted in P. lenzi. - The distomesial and the proximal lateral spines of the antennular article 1 are small or minute in P. lenzi, whereas these spines are well developed in P. porteri. - The spines on the P 2 – 4 meri are stronger in P. porteri than in P. lenzi. Furthermore, the extensor margin of the propodus has well developed spines in P. porteri, being unarmed in P. lenzi. The specimens of Phylladiorhynchus porteri range from 2.0 to 2.8 mm postorbital carapace length.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF9DE6684F9CFCE574D3BFA2.taxon	description	(Figs. 37 C – D, 46, 50 E)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF9DE6684F9CFCE574D3BFA2.taxon	materials_examined	Type material. Holotype. Mariana Islands, Maug Islands, Stn MAU- 385, 20.02973294 ° N, 145.2084746 ° E, 17 m, 04 May 2014: M 2.2 mm (UF 54423). Paratypes: Northern Mariana Islands. Maug Islands. Stn MAU- 302, 20.01416942 ° N, 145.2342278 ° E, 15 m, 03 May 2014: 4 M 1.4 – 1.8 mm, 3 ov. F 1.6 – 1.9 mm, 1 F 1.7 mm (UF 54435). — Stn MAU- 303, 20.01416942 ° N, 145.2342278 ° E, 15, 03 May 2014: 4 M 1.4 – 2.3 mm (UF 54431). — Stn MAU- 304, 20.01416942 ° N, 145.2342278 ° E, 15 m, 3 May 2014: 2 M 1.8 – 2.1 mm, 5 ov. F 1.8 – 2.2 mm (UF 54434). — Stn MAU- 305, 20.01416942 ° N, 145.2342278 ° E, 15 m, 3 May 2014: 1 M 1.6 mm, 2 ov. F 1.8 – 2.1 mm (UF 54433). Other material. Northern Mariana Islands, Maug Islands. Stn MAU- 354, 20.01416942 ° N, 145.2342278 ° E, 15 m, 3 May 2014: 1 M 1.4 mm, 4 ov. F 2.0 – 2.3 mm (UF 54432). — Stn MAU- 355, 20.01416942 ° N, 145.2342278 ° E, 15 m, 3 May 2014: 1 M 2.4 mm, 1 ov. F 2.0 mm (UF 54424), 1 M 2.1 mm (UF 54425). — Stn MAU- 357, 20.01416942 ° N, 145.2342278 ° E, 15 m, 3 May 2014: 8 M 1.3 – 2.2 mm, 11 ov. F 1.5 – 2.3 mm, 6 F 1.4 – 1.9 mm (UF 54430). — Stn MAU- 476, 20.01416942 ° N, 145.2342278 ° E, 15 m, 3 May 2014: 1 F 1.9 mm (UF 4470). — Stn MAU- 478, 20.01416942 ° N, 145.2342278 ° E, 15 m, 3 May 2014: 2 M 1.4 – 1.9 mm, 1 F 1.5 mm (UF 54429). — Stn MAU- 479 20.01416942 ° N, 145.2342278 ° E, 15 m, 3 May 2014: 1 M 1.8 mm 3 ov. F 1.8 – 2.4 mm (UF 54427). — Stn MAU- 383, 20.02973294 ° N, 145.2084746 ° E, 17 m, 4 May 2014: 4 M 1.4 – 1.9 mm, 4 ov. F 1.6 – 2.1 mm (UF 54428). — Stn MAU- 384, 20.02973294 ° N, 145.2084746 ° E, 17 m, 04 May 2014: 1 M 2.1 mm, 4 ov. F 2.0 – 2.4 mm (UF 54426). — Stn MAU- 385, 20.02973294 ° N, 145.2084746 ° E, 17 m, 04 May 2014: 4 M 2.1 – 2.8 mm, 5 ov. F 2.0 – 2.1 mm, 1 F 1.9 mm (UF 54422) Northern Mariana Islands, Pagan Island. Stn PAG- 407, 18.10734 ° N, 145.78587 ° E, 11 m, 20 April 2014: 1 M 1.3 mm, 2 ov. F 2.0 – 2.1 mm (UF 54453). — Stn PAG- 480, 18.10734 ° N, 145.78587 ° E, 11 m, 20 April 2014: 1 ov. F 2.0 mm (UF 54450). — Stn PAG- 483 18.10734 ° N, 145.78587 ° E, 11 m, 20 April 2014: 7 M 1.4 – 2.4 mm, 3 ov. F 2.2 – 2.5 mm (UF 54471). — Stn PAG- 565, 18.10734 ° N, 145.78587 ° E, 11 m, 20 April 2014: 1 ov. F 2.2 mm (UF 54476). — Stn PAG- 566, 18.10734 ° N, 145.78587 ° E, 11 m, 20 April 2014: 2 M 1.8 – 2.3 mm, 1 ov. F 2.3 mm, 1 F 1.6 mm (UF 54472). — Stn PAG- 567, 18.10734 ° N, 145.78587 ° E, 11 m, 20 April 2014: 2 M 2.3 – 2.8 mm (UF 54473). — Stn PAG- 568, 18.10734 ° N, 145.78587 ° E, 11 m, 20 April 2014: 3 M 1.3 – 1.5 mm (UF 54452). — Stn PAG- 571, 18.10734 ° N, 145.78587 ° E, 11 m, 20 April 2014: 1 F 1.8 mm (UF 54443). — Stn PAG- 573 18.10734 ° N, 145.78587 ° E, 11 m, 20 April 2014: 2 M 1.8 – 1.9 mm (UF 54449). — Stn PAG- 603, 18.10734 ° N, 145.78587 ° E, 11 m, 20 April 2014: 1 ov. F 2.1 mm (UF 54438). — Stn PAG- 694, 18.0706862 ° N, 145.7137933 ° E, 11 m, 20 April 2014: 1 M 1.4 mm, 1 ov. F 2.1 mm (UF 54440), 1 M 1.6 mm (UF 54441). — Stn PAG- 697, 18.0706862 ° N, 145.7137933 ° E, 11 m, 20 April 2014: 1 M 1.8 mm, 1 ov. F 1.9 mm (UF 54448). — Stn PAG- 762, 18.0706862 ° N, 145.7137933 ° E, 11 m, 20 April 2014: 1 M 1.7 mm, 1 ov. F 2.0 mm (UF 54439). — Stn PAG- 809, 18.0706862 ° N, 145.7137933 ° E, 11 m, 20 April 2014: 2 M 1.4 – 1.6 mm, 1 F 1.5 mm (UF 54446). — Stn PAG- 810, 18.0706862 ° N, 145.7137933 ° E, 11 m, 20 April 2014: 1 ov. F 1.8 mm (UF 54445). — Stn PAG- 840, 18.11947952 ° N, 145.7555622 ° E, 12 m, 23 April 2014: 2 M 2.0 – 2.3 mm (UF 54447). — Stn PAG- 841 18.11947952 ° N, 145.7555622 ° E, 12 m, 23 April 2014: 1 M 2.0 mm, 1 ov. F 2.1 mm (UF 54437). — Stn PAG- 865, 18.11947952 ° N, 145.7555622 ° E, 12 m, 23 April 2014: 1 ov. F 1.8 mm (UF 54451). — Stn PAG- 896, 18.11947952 ° N, 145.7555622 ° E, 12 m, 23 April 2014: 2 M 1.8 – 1.9 mm, 1 ov. F 2.0 mm, 1 F 1.8 mm (UF 54442). — Stn PAG- 900, 18.11947952 ° N, 145.7555622 ° E, 12 m, 23 April 2014: 1 M 1.8 mm (UF 54436). — Stn PAG- 957, 18.11947952 ° N, 145.7555622 ° E, 12 m, 23 April 2014: 1 M 2.0 mm, 1 ov. F 1.7 mm, 2 F 1.6 – 2.0 mm (UF 54444). Mariana Islands, Guam Island. 13.5167 ° N, 144.8 ° E, 10 – 25 m, 27 June 2003: 1 ov. F 1.6 mm (UF 4173). — Stn GUA- 601, 13.57847 ° N, 144.82831 ° E, 11 m, 25 March 2014: 1 F 2.1 mm (UF 54455). — Stn GUA- 645, 13.57847 ° N, 144.82831 ° E, 11 m, 25 March 2014: 2 M 1.6 – 2.0 mm (UF 54456). — Stn GUA- 695, 13.57847 ° N, 144.82831 ° E, 11 m, 25 March 2014: 1 ov. F 1.9 mm (UF 54454). Marshall Islands, Wake Island. 19 ° 17.259 ’ N, 166 ° 36.782 ’ E, 11.5 – 13.5 m, 18 November 2005: 1 ov. F 2.0 mm (UF 8548). — no position, 15 – 21 m, 24 March 2009: 1 ov. F 1.8 mm (UF 36081). — Stn WAK- 083, 19.2917761 ° N, 166.6072896 ° E, 14 m, 17 March 2014: 1 ov. F 2.2 mm (UF 54468). — Stn WAK- 084, 19.2917761 ° N, 166.6072896 ° E, 14 m, 17 March 2014: 5 M 1.6 – 2.2 mm, 4 ov. F 1.8 – 2.2 mm (UF 54467). — Stn WAK- 084, 19.2917761 ° N, 166.6072896 ° E, 14 m, 17 March 2014: 1 M 2.3 mm (UF 54541). — Stn WAK- 114, 19.2917761 ° N, 166.6072896 ° E, 14 m, 17 March 2014: 1 M 2.2 mm (UF 54458). — Stn WAK- 117, 19.2917761 ° N, 166.6072896 ° E, 14 m, 17 March 2014: 1 ov. F 1.7 mm (UF 54474). — Stn WAK- 118, 19.2917761 ° N, 166.6072896 ° E, 14 m, 17 March 2014: 15 M 1.5 – 2.0 mm, 6 ov. F 1.8 – 2.1 mm (UF 54469). — Stn WAK- 152, 19.31627096 ° N, 166.5983406 ° E, 12 m, 16 March 2014: 5 M 1.9 – 2.3 mm, 4 ov. F 1.8 – 2.1 mm, 5 F 1.9 – 2.3 mm (UF 54466). — Stn WAK- 189, 19.27067533 ° N, 166.6516393 ° E, 14 m, 19 March 2014: 3 ov. F 2.0 – 2.5 mm, 1 F 2.0 mm (UF 54460). — Stn WAK- 218, 19.27067533 ° N, 166.6516393 ° E, 14 m, 19 March 2014: 2 M 1.9 – 2.2 mm (UF 54459). — Stn WAK- 219, 19.27067533 ° N, 166.6516393 ° E, 14 m, 19 March 2014: 3 M 2.5 – 2.6 mm (UF 54462). — Stn WAK- 220, 19.27067533 ° N, 166.6516393 ° E, 14 m, 19 March 2014: 1 ov. F 2.0 mm (UF 54457). — Stn WAK- 221, 19.27067533 ° N, 166.6516393 ° E, 14 m, 19 March 2014: 1 M 1.9 mm, 2 ov. F 1.9 – 2.0 mm (UF 54463). — Stn WAK- 222, 19.27067533 ° N, 166.6516393 ° E, 14 m, 19 March 2014: 5 M 1.4 – 2.2 mm, 8 ov. F 1.7 – 2.3 mm (UF 54461). — Stn WAK- 252, 19.27067533 ° N, 166.6516393 ° E, 14 m, 19 March 2014: 16 M 1.3 – 2.3 mm, 21 ov. F 1.6 – 2.5 mm (UF 54475). — Stn WAK- 256, 19.27067533 ° N, 166.6516393 ° E, 14 m, 19 March 2014: 2 M 2.0 – 2.1 mm, 2 ov. F 1.8 – 2.1 mm, 1 F 1.7 mm (UF 54465). — Stn WAK- 280, 19.27067533 ° N, 166.6516393 ° E, 14 m, 19 March 2014: 6 M 1.6 – 2.3 mm, 9 ov. F 1.5 – 2.2 mm, 1 F 1.7 mm (UF 4464).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF9DE6684F9CFCE574D3BFA2.taxon	etymology	Etymology. From the name Priasus, an Argonaut, son of Caeneus and Phocus. The name is considered a substantive in apposition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF9DE6684F9CFCE574D3BFA2.taxon	description	Description. Carapace: Slightly longer than broad; transverse ridges with dense short setae and few scattered long and thick iridescent setae. Gastric region with 4 transverse ridges: epigastric ridge distinct with 2 median spines, usually 1 median produced scale usually with thick plumose setae before epigastric ridge; anterior protogastric ridge not medially interrupted, nearly extending laterally to carapace margin; anterior mesogastric ridge not medially interrupted, laterally continuing to first branchial spine, sometimes followed by some short small scales; anterior metagastric ridge medially interrupted. Mid-transverse ridge not interrupted, cervical groove indistinct, followed by 2 uninterrupted or minutely interrupted ridges, interspersed with 1 short lateral ridge and sometimes few, short scattered scales. Lateral margins slightly convex, with 4 distinct spines: first anterolateral spine well-developed, reaching anteriorly to level of lateral orbital spine, hepatic margin unarmed; anterolateral spine followed by 3 branchial spines (2 anterior and 1 posterior). Rostrum bottle-shaped, horizontal, dorsally concave, [1.1] – 1.4 × as long as broad, length [0.3] – 0.4 and breadth 0.25 – [0.3] that of carapace; lateral margins serrated, straight proximally and distally slightly convex, with well-developed supraocular basal spines and small subapical spines. Pterygostomian flap ending in blunt tooth, upper margin smooth. Thoracic sternum: As wide as long. Sternite 3 quadrangular, slightly wider than long (1.5 × as wide as long), anterior margin with a median blunted projection, lateral margins rounded. Sternite 4 widely contiguous to sternite 3; surface depressed in midline, smooth; greatest width [2.9] – 3.3 × that of sternite 3, 2.1 – [2.9] × as wide as long. Pleon: Elevated ridges with short setae and a few scattered long setae. Tergite 2 with anterior and posterior transverse elevated ridges; tergites 3 – 4 with anterior transverse ridge; tergites 5 – 6 smooth. Eye: Eyestalk length about [0.9] × broader than long, peduncle distally setose, slightly expanded proximally; maximum corneal diameter [0.6] – 0.7 × rostrum width, as narrower than eyestalk (0.8 – 0.9 × maximum peduncle width). Antennule: Article 1 slightly longer than wide, with 5 distal spines: distomesial spine well-developed; proximal lateral spine small, always present. Antenna: Article 1 with prominent mesial process distally falling well short of lateralmost antennular spine. Article 2 with well-developed distomesial and distolateral spines. Article 3 with distomesial spine. Article 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus [0.7] 0.6 – 0.8 × length of ischium, with well-developed distal spine on extensor and flexor margins. P 1: [2.2] – 2.9 (males), 1.8 – 2.1 (females) × carapace length; subcylindrical, spiny and with dense long stiff setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, mesial spines usually stronger than others. Merus [0.8] – 0.9 length of carapace, [2.2] 1.8 – 2.3 × as long as carpus. Carpus [1.3] 1.1 – 1.4 × as long as wide. Palm [1.2] – 1.4 × carpus length, [1.3] – 1.7 × as long as broad. Fingers 0.6 – [1] × palm length; fixed finger with 1 – 2 basal spines; movable finger with 1 basal spine. P 2 – 4: Stout, setose and spinose. Meri successively shorter posteriorly: P 3 merus [0.7] 0.7 × length of P 2 merus, P 4 merus 0.8 – [0.9] × length of P 3 merus. P 2 merus, 1.6 – [1.7] × carapace length, 3.8 – [5.0] × as long as broad, 1.2 – [1.4] × as long as P 2 propodus; P 3 merus 3.8 – [4.2] × as long as broad, [1.1] – 1.4 × as long as P 3 propodus; P 4 merus 3.3 – [3.4] × as long as broad, 1.0 – [1.1] × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, unarmed; flexor margin irregular, with distal spine on P 2 – 4, P 4 lateral surface with row of 2 – 4 spines. Carpi with 1 – 3 prominent spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 4; row of small spines or granules below extensor margin on lateral surface of P 2 – 3, unarmed on P 4; flexor margin unarmed. Propodi stout, [4.0 – 5.0] 3.7 – 5.6 × as long as broad; extensor margins irregular, armed with 1 – 3 spines on proximal half or unarmed; flexor margins with 3 – 4 slender movable spines in addition to distal pair. Dactyli [0.5 – 0.7] 0.5 – 0.8 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 5 – 6 movable spines. Eggs: Ov. F carried approximately 10 – 35 eggs of 0.3 – 0.5 mm diameter. Live colour. Unknown. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF9DE6684F9CFCE574D3BFA2.taxon	distribution	Distribution. Mariana Islands, Guam Island, Marshall Islands, Wake Island, between 10 and 25 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF9DE6684F9CFCE574D3BFA2.taxon	discussion	Remarks. Phylladiorhynchus priasus belongs to the group of species with 2 epigastric spines, hepatic margin unarmed and 2 spines on the anterior branchial margin. The closest species are P. integrirostris, from Hawaii, and P. lynceus, from Chagos, Western Australia, Kiribati and Samoa. The 3 species are barely distinguishable morphologicaly although they are clearly different genetically. However, these species can be distigushed by the following subtle traits: - Phylladiorhynchus priasus and P. integrirostris usually have a median produced scale, behind epigastric ridge, usually with thick setae [not in the neotype neither in the material examined by Schnabel & Ahyong (2019)]. However, this scale is always absent in all specimens examined of P. lynceus. - The rostrum is proximally straight (bottle-shaped) in P. priasus and P. integrirostris, whereas it is convex (leaf-like) in P. lynceus. - The carapace has numerous thick iridescent setae in P. lynceus and P. priasus, whereas these setae are usually absent in P. integrirostris. The genetic divergences between P. priasus and P. lynceus were 7 % (COI) and 2 % (16 S). These divergences were larger between P. priasus and P. integrirostris, 23 % (COI) and 12 % (16 S). Phylladiorhynchus priasus is also close to P. orpheus (see the differences under the Remarks of P. orpheus). The eight and six sequences of P. priasus from Northern Mariana Islands, Guam and Wake Island diverged 0.4 % (COI) and 0.1 % (16 S). The specimens of P. priasus) range from 1.3 to 2.8 mm postorbital carapace length.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF90E66F4F9CF8BD7382BB07.taxon	description	(Figs. 47, 50 F)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF90E66F4F9CF8BD7382BB07.taxon	materials_examined	Type material. Holotype. French Polynesia, MUSORSTOM 9 Stn CP 1156, 07.9833 ° S, 140.7283 ° W, 80 m, 23 August 1997: M 1.6 mm (MNHN-IU- 2014 - 13849). Paratypes. Philippines. MUSORSTOM 3 Stn DR 117, 12.5166 ° N, 120.6500 ° E, 92 – 97 m, 3 June 1985: 1 M 1.4 – 1.8 mm, 2 ov. F 1.8 – 2.0 mm (MNHN-IU- 2014 - 13848), 1 M 1.4 mm (MNHN-IU- 2017 - 1342). Vanuatu. SANTO Stn DS 103, 15.5683 ° S, 167.2666 ° E, 10 – 80 m, 14 October 2006: 1 M 1.5 mm (MNHN-IU- 2014 - 13868). French Polynesia. MUSORSTOM 9 Stn CP 1239, 09.7033 ° S, 139.0600 ° W, 89 – 95 m, 31 August 1997: 1 ov. F 2.0 mm (MNHN-IU- 2014 - 13733). — no station data: 1 M 1.8 mm (MNHN-IU- 2019 - 2675).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF90E66F4F9CF8BD7382BB07.taxon	etymology	Etymology. From the latin pulcher, beautiful, in reference to the beauty of this species.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF90E66F4F9CF8BD7382BB07.taxon	description	Description. Carapace: Robust or massive, [0.9] – 1.1 × as long as broad; transverse ridges elevated, serrated, with few short setae, and few scattered long thick iridescent setae. Gastric region convex (upraised dorsally) with some transverse ridges: epigastric ridge indistinct, with 5 spines (1 median and 2 pairs of spines laterally); anterior protogastric ridge not medially interrupted, laterally interrupted with few scales to carapace margin, posterior protogastric ridge scale-like; anterior mesogastric ridge scale-like, laterally interrupted by anterior branch of cervical groove, laterally continuing with some scales; anterior metagastric ridge scale-like, followed by some scales on posterior metagastric area. Mid-transverse ridge not interrupted, medially depressed, preceded by cervical groove, followed by 1 – 2 not interrupted or minutely interrupted ridges, interspersed with 2 short lateral ridges. Lateral margins clearly convex, with 7 spines: first anterolateral spine well-developed, reaching end of lateral orbital spine, second spine (hepatic) well-developed, slightly dorsomesially from lateral margin, and followed by 5 branchial spines (3 anterior and, 2 posterior). Rostrum leaf-like, horizontal, dorsally flattish or slightly concave, [1.1] 1.5 – 1.6 × as long as broad, length 0.4 and breadth 0.3 that of carapace; lateral margins serrated and convex, with well-developed supraocular spines, subapical spines well-developed. Pterygostomian flap ending in anterior spine, upper margin smooth or serrated. Thoracic sternum: As wide as long or slightly wider than long, lateral margins of posterior half slightly divergent. Sternite 3 broad, [2.2] – 2.5 × as wide as long, anterior margin convex. Sternite 4 widely contiguous to sternite 3; surface flattened, smooth; greatest width 2.7 – [2.9] × that of sternite 3, 2.6 – [2.9] × as wide as long. Pleon: Elevated ridges upraised dorsally, with a few scattered short setae. Tergites 2 – 3 with anterior and posterior transverse elevated ridges; tergite 4 with anterior transverse ridge; tergites 5 – 6 smooth. Eye: Eyestalk length about [1.3] × broader than long, peduncle distally setose, not distinctly expanded proximally; maximum corneal diameter [0.7] × rostrum width, as wide as eyestalk. Antennule: Article 1 slightly longer than wide, with 4 distal spines: distomesial spine well-developed; proximal lateral spine absent. Antenna: Article 1 with prominent mesial process distally not reaching lateral antennular spine. Article 2 with small distal spines laterally and mesially. Articles 3 and 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus [0.5] – 0.6 × length of ischium at midlength, with 0 – 1 median small spine and and 1 well-developed distal spines on extensor margin and 1 strong spine on flexor margin. P 1 (lost in holotype): 2.5 – 3.0 (males) × carapace length; subcylindrical, spiny and with scattered long stiff setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus 1.1 × length of carapace, 2.5 – 2.7 × as long as carpus. Carpus 1.8 – 2.4 × as long as wide. Palm 1.3 – 1.6 × carpus length, 2.1 – 2.5 × as long as broad. Fingers unarmed, 0.8 × palm length. P 2 – 4 (lost in most specimens): Subcylindrical, moderately setose and spinose, with few scattered plumose setae. Meri successively shorter posteriorly: P 3 merus 0.8 × length of P 2 merus, P 4 merus [0.8] – 0.9 × length of P 3 merus; P 2 merus, [0.6] 0.8 × carapace length, [3.5] – 4.7 × as long as broad, [1.0] – 1.4 × as long as P 2 propodus; P 3 merus [4.0] – 5.5 × as long as broad, [1.1] – 1.3 × as long as P 3 propodus; P 4 merus [4.1] – 5.2 × as long as broad, [0.9] – 1.1 × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, unarmed other than small distal spine otherwise serrated on proximal half; flexor margins of all legs irregular, with distal spine. Carpi with 2 or 3 spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, smaller on P 4; row of small spines below extensor margin on lateral surface of P 2 – 3, unarmed on P 4; flexor margins of P 2 – 4 irregular, each with distal spine. Propodi stout, [4.2 – 5.2] 4 – 5 × as long as broad; extensor margin irregular, unarmed or with minute proximal spines; flexor margin with 3 – 4 slender movable spines in addition to distal pair. Dactyli [0.7] – 0.8 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 5 – 6 movable spines. Eggs. Ov. F carried approximately 5 – 10 eggs of 0.3 mm diameter. Live colour. Unknown. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF90E66F4F9CF8BD7382BB07.taxon	distribution	Distribution. French Polynesia, Philippines and Vanuatu, from 10 to 97 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF90E66F4F9CF8BD7382BB07.taxon	discussion	Remarks. Phylladiorhynchus pulchrus belongs to the group of species having 5 epigastric spines, 3 spines on the anterior branchial margin, gastric region strongly convex, the rostrum leaf-like (margins clearly convex and subapical spines present), and carapace ridges upraised dorsally. The new species is closely related to P. koumac from New Caledonia and they can be distinguished by subtle differences: - The proximal half of the extensor margin of the P 2 – 4 propodi has well-developed spines in P. koumac, whereas these spines are obsolescent or absent in P. pulchrus. - The P 2 – 4 propodi are slender (6 – 7 × as long as wide) in P. koumac, whereas they are stout (4 – 5 × as long as wide) in P. pulchrus. The sequences of P. pulchrus were 11 % (COI) and 10 % (16 S) divergent from P. koumac. The specimens of P. pulchrus range in size from 1.4 to 2.0 mm postorbital carapace length.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF97E66D4F9CFDD473D9BC2E.taxon	description	(Figs. 48, 50 G, 56 B)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF97E66D4F9CFDD473D9BC2E.taxon	materials_examined	Type material. Holotype. New Caledonia, LIFOU Stn 1648, 20.9017 ° S, 167.0550 ° E, 150 – 200 m, 7 November 2000: ov. F 2.8 mm (MNHN-IU- 2014 - 13844). Paratypes. New Caledonia. LIFOU Stn 1647, 20.70750 ° S, 167.1333 ° E, 150 – 200 m, 6 November 2000: 1 M 3.2 mm (MNHN-IU- 2014 - 13845). — Stn 1648, 20.9017 ° S, 167.0550 ° E, 150 – 200 m, 7 November 2000: 1 F 1.8 mm (MNHN-IU- 2014 - 13843). New Caledonia. Koumac 2.3 Stn KL 17, 20.55685 ° S, 164.0732667 ° E, 92 m, 14 November 2019: 1 M 1.7 mm (MNHN-IU- 2014 - 20101).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF97E66D4F9CFDD473D9BC2E.taxon	etymology	Etymology. From the Latin punctum, spot, in reference of the red spots covering the pleonal tergites of this species.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF97E66D4F9CFDD473D9BC2E.taxon	description	Description. Carapace: As long as broad, transverse ridges with few short setae. Gastric region flattened, with 4 transverse ridges: epigastric ridge indistinct, with 5 spines (1 median and 2 pairs of spines laterally); anterior protogastric ridge not medially interrupted, laterally interrupted with few scales; anterior mesogastric ridge scalelike (with 1 to 3 scales), laterally interrupted by anterior branch of cervical groove, laterally continuing uninterrupted to first branchial spine; anterior metagastric ridge scale-like and followed by few scales on posterior metagastric area. Mid-transverse ridge not interrupted, medially depressed, preceded by a distinct cervical groove, followed by 2 uninterrupted or minutely interrupted ridges, interspersed with 1 short lateral ridge and few short, scattered scales. Lateral margins straight or slightly convex, with 6 spines: first anterolateral spine well-developed, reaching end of lateral orbital spine, second spine (hepatic) well-developed, slightly dorsomesially from lateral margin, and followed by 4 branchial spines (2 anterior and 2 posterior). Rostrum triangular, horizontal, dorsally flattish or slightly concave, 1.4 – [1.6] × as long as broad, length [0.4] 0.4 and breadth [0.3] 0.3 that of carapace; lateral margins smooth and straight, with well-developed supraocular spines, subapical spines absent. Pterygostomian flap ending in small spine, upper margin smooth. Thoracic sternum: As wide as long. Sternite 3 sharply broad, [3.4] 3.3 – 4.0 × as wide as long, anterior margin straight or moderately convex, with a median deep notch, moderately produced anterolaterally. Sternite 4 widely contiguous to sternite 3; surface depressed in midline, smooth; greatest width 1.5 – [2.4] × that of sternite 3, [3.3] – 4.2 × as wide as long. Pleon: Elevated ridges with short setae and with few scattered long setae. Tergite 2 with anterior and posterior transverse elevated ridges; tergites 3 – 4 with anterior transverse ridge; tergites 5 – 6 smooth. Eye: Eyestalk length about 1.3 × broader than long, peduncle distally setose, not distinctly expanded proximally; maximum corneal diameter 1.0 × rostrum width, as wide as eyestalk. Antennule: Article 1 1.3 × longer than wide, with 5 spines: distomesial spine small; proximal lateral spine well developed, always present. Antenna: Article 1 with prominent mesial process, distally not reaching lateralmost antennular spine. Article 2 with small distomesial and distolateral spines. Articles 3 and 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus 0.5 × length of ischium, with well-developed distal spine on extensor margin and 2 strong spines on flexor margin. P 1 (lost in holotype and in males): 3.5 (females) × carapace length; subcylindrical, spiny and with scattered long stiff setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus 1.4 length of carapace, 2.0 – 2.5 × as long as carpus. Carpus 2.5 – 2.9 × as long as wide. Palm 1.1 × carpus length, 2.2 – 3.5 × as long as broad. Fingers subequal in length to palm; fixed finger unarmed; movable finger with well-developed basal spine. P 2 – 4: Slender, subcylindrical, moderately setose and spinose. Meri successively shorter posteriorly: P 3 merus 0.9 – [1.0] × length of P 2 merus, P 4 merus 0.9 × length of P 3 merus. P 2 merus, [0.7] 0.8 × carapace length, [6.0] – 8.0 × as long as broad, [1.0] – 1.3 × as long as P 2 propodus; P 3 merus [6.0] – 7.0 × as long as broad, 1.0 × as long as P 3 propodus; P 4 merus 4.5 – [5.5] × as long as broad, as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, with small distal spine; flexor margins of P 2 – 4 irregular, each with distal spine; P 4 lateral surface with median row of 3 small spines, absent in P 2 – 3. Carpi with 3 or 4 spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, smaller on P 4; row of small spines below extensor margin on lateral surface of P 2 – 3, unarmed on P 4; flexor margins on P 2 - 4 with distal spine. Propodi slender, 7.0 – [9.0] × as long as broad; extensor margin usually armed with 1 – 2 small spines on proximal half of P 2 – 4; flexor margin with 4 slender movable spines in addition to distal pair. Dactyli [0.6] – 0.7 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 6 – 7 movable spines. Live colour. Body light orange; carapacecovered by orange and golden-yellow spots. Pleonal tergites 1 – 4 light orange; tergites 3 – 4 each with two symmetrical vertical white stripes and 2 red spots. P 2 – 4 clear with distinct red bars. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF97E66D4F9CFDD473D9BC2E.taxon	distribution	Distribution. New Caledonia and Chesterfield Islands, from 92 to 200 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF97E66D4F9CFDD473D9BC2E.taxon	discussion	Remarks. Phylladiorhynchus punctatus belongs to the group of species having 5 epigastric spines, 2 spines on the anterior branchial margin and the Mxp 3 merus with 2 prominent spines along the flexor margin. Phylladiorhynchus punctatus resembles P. kermadecensis Schnabel & Ahyong, 2019 from New Zealand, but they can be distinguished by the following characters: - The rostral margins are slightly concave in P. kermadecensis, whereas these margins are straight in P. punctatus. - The anterior mesogastric ridge is scale-like in P. punctatus, whereas this ridge is not medially interrupted in P. kermadecensis. - The anterior metagastric ridge is not medially interrupted in P. punctatus, whereas it is medially interrupted in P. kermadecensis. The sequences of P. punctatus were 17 % (COI) and 7 % (16 S) divergent from P. kermadecensis. The specimens of P. punctatus range in size from 1.7 to 3.2 mm postorbital carapace length.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF95E6924F9CFA3C7280BD2D.taxon	diagnosis	Diagnosis (modified from Schnabel & Ahyong 2019). Rostrum lateral margins convex; subapical spines present. Carapace with 4 epigastric spines of which outermost may be present as a granule and much smaller than inner pair (rarely with 6 spines); anterior mesogastric ridge not medially interrupted; anterior metagastric ridge scalelike, medially interrupted. Pleonal tergite 3 usually without posterior transverse ridge. Thoracic sternite 3 anterior margin convex, median projection absent. Antennular article 1 with 5 spines: distomesial spine over-reaching lower distolateral spine, slightly to distinctly overreached by upper distolateral and first lateral spines. Antennal article 1 mesial process not reaching level of second lateral antennular spine; article 2 distolateral and distomesial spines subequal; article 3 with distinct distomesial spine. Flexor margin of Mxp 3 merus with one prominent spine only. P 2 – 4 dactylus extensor margin without upright spines at bases of movable spines. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF95E6924F9CFA3C7280BD2D.taxon	distribution	Distribution. After the complete redescription by Schnabel & Ahyong (2019), the species is found along the southeastern, southern and western Australia and northern New Zealand, between 10 and 274 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF95E6924F9CFA3C7280BD2D.taxon	discussion	Remarks. Schnabel & Ahyong (2019) provided a complete description of the species, with a comparison with other species. As these authors pointed out, P. pusillus shows a certain variability in the number of epigastric spines, recommending caution in the use of this character. The species is very close to P. poeas, from French Polynesia (see the differences under the Remarks of P. poeas).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF6AE6964F9CFB3D74F7B9CA.taxon	description	(Figs. 49, 50 H)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF6AE6964F9CFB3D74F7B9CA.taxon	description	Galathea serrirostris. — Miyake & Baba, 1966 a: 67, fig. 8 (Amami-oshima, Ryukyu Islands, Japan, intertidal).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF6AE6964F9CFB3D74F7B9CA.taxon	materials_examined	Type material. Lectotype. Japan, Bonin Islands (Ogasawara), Port Loyd, Dr. Sixten Bocks Japan exp. 1914, 23 / 7, Djup (depth): dykare (diver), Botten (bottom): korall (coral), det. G. Melin: ov. F 2.9 mm (UPSZTY 183822). Paralectotypes. Dr. Sixten Bocks Japan exp. 1914. Bonin Islands (Ogasawara), Port Loyd. 23 / 7. Djup: dykare. Botten: korall. Det. G. Melin: 12 M 1.9 – 3.0 mm, 11 ov. F 2.1 – 3.0 mm, 1 F parasitized 1.9 mm, 1 broken (UPSZTY 2531).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF6AE6964F9CFB3D74F7B9CA.taxon	description	Description. Carapace: As long as or slightly longer than broad; transverse ridges with dense short setae and few scattered long and thick setae. Gastric region slightly convex with some transverse ridges: epigastric ridge distinct with 2 median spines, often followed by few short scales; anterior protogastric ridge not medially interrupted, nearly extending laterally to carapace margin, followed by a posterior protogastric scale-like ridge; anterior mesogastric ridge not medially interrupted, often followed by some scales on posterior mesogastric area; anterior metagastric ridge entire or minutely medially interrupted, followed by a short median scale. Mid-transverse ridge not interrupted, cervical groove indistinct, followed by 2 – 3 not interrupted or minutely interrupted ridges, interspersed with 1 – 2 interrupted ridges. Lateral margins distinctly convex, with 4 distinct spines: first anterolateral spine well-developed, reaching anteriorly to level of lateral orbital spine, hepatic margin unarmed; anterolateral spine followed by 3 branchial spines (2 anterior and 1 posterior). Rostrum leaf-like, horizontal, dorsally concave, [1.4] 1.2 – 1.7 × as long as broad, length [0.3] – 0.4 and breadth [0.2] – 0.3 that of carapace; lateral margins serrated and slightly convex, with well-developed supraocular basal spines and small subapical spines. Pterygostomian flap ending in acute tooth, upper margin often slightly serrated. Thoracic sternum: As wide as long. Sternite 3 moderately broad, [2.0] – 2.1 × as wide as long, anterior margin serrated and convex, slightly produced anterolaterally. Sternite 4 widely contiguous to sternite 3; anterolaterally serrated, surface depressed in midline, smooth; greatest width 2 – [3] × that of sternite 3, [2.7] × as wide as long. Pleon: Elevated ridges with short setae and a few scattered long setae. Tergite 2 – 3 with anterior and posterior transverse elevated ridges; tergite 4 with anterior transverse ridge only; tergites 5 – 6 smooth. Eye: Eyestalk length about [0.8] – 0.9 × broader than long, peduncle distally setose, not distinctly expanded proximally; maximum corneal diameter [0.6] × rostrum width, as wide as eyestalk. Antennule: Article 1 slightly longer than wide, with 5 distal spines: distomesial spine well-developed; proximal lateral spine small, always present. Antenna: Article 1 with prominent mesial process, distally falling well short lateralmost antennular spine. Article 2 with well-developed distomesial and distolateral spines. Article 3 with distomesial spine. Article 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus [0.6] – 0.7 × length of ischium, with well-developed distal spine on extensor and flexor margins. P 1: 2.4 – 3.0 (males), [2.4] – 3.0 (females) × carapace length; subcylindrical, densely spiny and densely with long stiff setae; merus, carpus and palm with several rows of spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus 0.6 – [0.9] length of carapace, 1.6 – [2.0] × as long as carpus. Carpus 1.4 – [1.6] × as long as wide. Palm [1.1] – 1.4 × carpus length, 1.4 – [1.6] × as long as broad. Fingers [0.8] – 0.9 × palm length fixed finger with 2 basal spines; movable finger with 1 basal spines and spine at mid-length. P 2 – 4: Stout, setose and spinose. Meri successively shorter posteriorly: P 3 merus 0.7 × length of P 2 merus, P 4 merus 0.8 × length of P 3 merus. P 2 merus, 0.4 – [0.6] × carapace length, [4.2] 3.5 – 4.5 × as long as broad, [1.2] 0.9 – 1.3 × as long as P 2 propodus; P 3 merus [3.1] – 4 × as long as broad, [1.2] 0.9 – 1.3 × as long as P 3 propodus; P 4 merus [3.8] × as long as broad, [0.9] × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, with 1 – 2 spines,; flexor margin irregular, with distal spine on P 2 – 4; P 4 lateral surface with 2 – 3 spines. Carpi with 2 – 4 well-developed spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, smaller on P 4; flexor margin unarmed. Propodi stout, [4.0 – 5.0] 3.6 – 5.0 × as long as broad; extensor margin irregular, armed with 1 – 4 spines on P 2 – P 4; flexor margin with 3 – 4 slender movable spines in addition to distal pair. Dactyli [0.5 – 0.6] 0.5 – 0.8 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 4 – 6 movable spines. Eggs: Ov. F carried approximately 10 – 50 eggs of 0.2 – 0.4 mm diameter. Live colour. Unknown. Genetic data. No data.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF6AE6964F9CFB3D74F7B9CA.taxon	distribution	Distribution. Japan: Bonin Islands, Ryukyu Islands, intertidal to 128 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF6AE6964F9CFB3D74F7B9CA.taxon	discussion	Remarks. Phylladiorhynchus serrirostris was described as Galathea serrirostris by Melin (1939), from specimens collected in Bonin Islands, Japan. Tirmizi (1966) considered G. serrirostris identical to G. pusilla, but Miyake & Baba (1965, 1967) considered G. pusilla and G. serrirostris as close, but not identical, species. After, Baba (1991) included P. serrirostris as a junior synonym of P. integrirostris. The study of the type material of P. serrirostris and its comparison with the numerous specimens of P. integrirostris suggests that they can be considered as different species. Therefore, we have considered P. serrirostris as a valid species. The 2 species belong to the group of species having 2 epigastric spines, hepatic margin unarmed and 2 spines on the anterior branchial margin. Phylladiorhynchus serrirostris is easily differentiable from P. integrirostris by the following characters: - The anterior metagastric ridge is not medially interrupted in P. serrirostris, whereas it is medially interrupted in P. integrirostris. - The posterior gastric ridges (posterior protogastric, mesogastric and metagastric) are usually present in P. serrirostris, whereas they are absent in P. integrirostris. - The pleonal tergite 3 has 2 ridges (anterior and posterior) in P. serrirostris, whereas there is only the anterior ridge in P. integrirostris. - The P 1 is densely spinose in P. serrirostris, whereas they are scarcely spinose in P. integrirostris. The specimens of P. serrirostris range in size from 1.9 to 3.0 mm postorbital carapace length.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF6EE6974F9CFE9870CBB8CE.taxon	description	(Figs. 30 D – F, O – P, 50 I)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF6EE6974F9CFE9870CBB8CE.taxon	materials_examined	Material examined. New Caledonia CHALCAL 1 Stn DC 26, 19.1783 ° S, 158.5833 ° E, 48 m, 18 July 1984: 1 M 2.6 mm (MNHN-IU- 2013 - 9439 (Ga- 2055 )). — Stn DC 55, 21.3983 ° S, 158.9933 ° E, 55 m, 25 July 1984: 1 M 2.9 mm (MNHN-IU- 2016 - 473 (Ga- 2056 )). — Ouen Is. Prony Bay. Stn 121, 22.4666 ° S, 166.7183 ° E, 12 m, August 1984: 1 ov. F 3.7 mm (MNHN-IU- 2019 - 2613). — Stn 242, 22.3666 ° S, 167.0366 ° E, 25 m, October 1984: 1 M 3.4 mm (MNHN-IU- 2019 - 2619). — Lagon Est. Stn 713, 21.3766 ° S, 166.0116 ° E, 34 – 35 m, 11 August 1986: 1 M 3.3 mm (MNHN-IU- 2016 - 492), 1 M 2.7 mm (MNHN-IU- 2016 - 491). — Stn 782, 21.1016 ° S, 165.6116 ° E, 30 m January 1987: 1 M 2.7 mm (MNHN-IU- 2019 - 2621). Chesterfield Islands. CORAIL 2 Stn DW 88, 19.1000 ° S, 158.9300 ° E, 32 m 26 July 1988: 1 F 2.9 mm (MNHN- IU- 2016 - 469 (Ga- 2052 )). — Stn DW 160, 19.7666 ° S, 158.3833 ° E, 35 – 41 m, 1 September 1988: 1 M 2.5 mm (MNHN-IU- 2016 - 470 (Ga- 2054 )).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF6EE6974F9CFE9870CBB8CE.taxon	diagnosis	Diagnosis (modified from Schnabel & Ahyong 2019). Rostrum dagger-like, lateral margins nearly straight; subapical spines present. Carapace with 2 epigastric spines; metagastric ridge between third branchial marginal spines medially continuous; anterior branchial margin with 3 spines; hepatic spine present. Pleonal tergites 2 – 4 with anterior and posterior ridges. Thoracic sternite 3 anterior margin biconcave, with obtuse median projection. Antennular article 1 with 5 spines: distomesial spine large; second (proximal) lateral spine long, slender. Antennal article 1 mesial process distally falling well short of second lateral antennular spine; article 2 distolateral spine distinctly larger than distomesial spine; article 3 with small to minute distomesial spine, distolaterally unarmed. Flexor margin of Mxp 3 merus with one prominent spine only. P 1 very slender (P 1 male more than 3.5 × as long as carapace). P 2 – 4 dactylus extensor margin with sharp upright spines at bases of movable spines.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF6EE6974F9CFE9870CBB8CE.taxon	distribution	Distribution. Eastern Australia from Queensland, the Norfolk Ridge and Norfolk Island, at 4 – 81 m (one record at 570 – 578 m). New Caledonia and Chesterfield Islands, from 12 to 55 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF6EE6974F9CFE9870CBB8CE.taxon	discussion	Remarks. Phylladiorhynchus spinosus belongs to the group of species characterized by the presence of 2 epigastric spines, 1 hepatic spine, 3 spines on the anterior branchial margin, the rostrum dagger-like (margin straights or nearly straight) and the presence of dactylar spines along the flexor margin of the P 2 – 4 dactyli. There are 4 closely related species: P. asclepius from Western Australia, P. euryalus from Eastern Australia (Queensland), New Caledonia and Chesterfield Islands, P. lini from Taiwan and P. spinosus Schnabel & Ahyong, 2019 (also found in New Caledonia). The species from this complex are morphologically very similar, although genetically very distinct. However, they can be distinguished by the following subtle characters (sometimes overlapping): - The dorsal carapace, pleonal ridges and ocular peduncles have many thick iridescent setae in P. asclepius, P. euryalus and P. spinosus, whereas these iridescent setae are absent in P. lini. - The carapace is always wider than long in P. asclepius, whereas it is usually as long as or slightly longer than wide in the rest of the species. - The rostrum is twice longer than wide in P. spinosus and P. asclepius, whereas it is less than twice longer than wide in P. euryalus and P. lini. - The antennule proximal lateral spine is prominent in P. euryalus, reaching the mid-length of the lateral spine, whereas this spine is smaller in the other species. - The anterior margin of thoracic sternite 3 is anterolaterally projected (biconcave with a median blunted triangular projection) in P. spinosus and P. euryalus, whereas this margin has a median projection and it is anterolaterally rounded in the remaining species. - The spines of the antennal article 2 are subequal in P. asclepius, P. euryalus and P. lini, whereas the distomesial spine is minute in P. spinosus. Furthermore, the antennal article 3 is usually armed with a small to distinct distomesial spine in P. asclepius, whereas this spine is absent in the remaining species. - The P 1 is very slender in P. spinosus (P 1 male> 3.5 as long as carapace), moderately slender in and P. euryalus and P. lini (P 1 male about 3.0 as long as carapace), whereas it is moderately stout in P. asclepius (P 1 male <3.0 as long as carapace). The genetic divergences between P. spinosus and other closely related species were around 8 % (COI) and 6 – 7 % (16 S). The subtle morphological differences among specimens of P. spinosus from different localities were mentioned by Schnabel & Ahyong (2019). Unfortunately, we have not examined the type specimens of P. spinosus, but we suggest that this species is probably restricted to Queensland, Norfolk Ridge and Norfolk Islands area. Their material from Western Australia is probably referable to P. asclepius, although it should be confirmed in the future.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF6FE6954F9CFDA9756FBD46.taxon	description	(Figs. 50 J, 51)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF6FE6954F9CFDA9756FBD46.taxon	materials_examined	Type material. Holotype. Western Australia, Montilivet Island, Stn 117 / K 12, 14.2883 ° S, 125.2255 ° E, 0 – 11 m, 25 October 2012: 1 ov. F 1.9 mm (WAM C 51399). Paratypes. Western Australia. Adele Island Stn 01 / K 09 - adhoc, 15.5234 ° S, 123.2053 ° E, 0 – 20 m, 13 October 2009: 1 ov. F 2.4 mm (WAM C 43997). — Stn 02 / K 09 - adhoc, 15.5192 ° S, 123.1957 ° E, 0 – 14 m, 14 October 2009: 1 ov. F 1.8 mm (WAM C 43826). — Ningaloo Reef. 22.7581 ° S, 113.6491 ° E, 13 m, 1 May 2009: 1 F 1.8 mm (UF 217029). — 22.6083 ° S 113.6249 ° E, 10 m, 1 May 2009: 1 M 2.1 mm (UF 22293). Other material. Indonesia. Rumphius Exp. II. 1975, NE coast Marsegu Is., on coral. 18 January 1975: 1 M 2.7 mm (MNHN-IU- 2014 - 23832 (Ga- 1154 )). Papua New Guinea. PAPUA NIUGINI Stn PB 39, 05.2650 ° S, 145.7850 ° E, 18 – 27 m, 6 December 2012: 1 M 2.1 mm (MNHN-IU- 2019 - 2637). — KAVIENG Stn KB 36, 02.6466 ° S, 150.6400 ° E, 3 – 8 m, 13 June 2014: 1 M 2.3 mm, 1 F 2.2 mm (MNHN-IU- 2014 - 13664). — Stn KB 58, 02.5683 ° S, 150.6250 ° E, 12 m, 22 June 2014: 1 F 2.5 mm (MNHN-IU- 2016 - 5858). — Stn KB 60, 02.5416 ° S, 150.5883 ° E, 20 m, 23 June 2014: 1 F 1.8 mm (MNHN-IU- 2016 - 476).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF6FE6954F9CFDA9756FBD46.taxon	etymology	Etymology. From the name Talaus, an Argonaut, son of Bias and Pero. The name is considered a substantive in apposition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF6FE6954F9CFDA9756FBD46.taxon	description	Description. Carapace: As long as or slightly longer than broad; transverse ridges with dense short setae and few scattered long and thick iridescent setae. Gastric region slightly convex with 4 transverse ridges: epigastric ridge distinct with 2 median spines and some lateral short scales; anterior protogastric ridge not medially interrupted, nearly extending laterally to carapace margin; anterior mesogastric ridge not medially interrupted, laterally continuing to first branchial spine; anterior metagastric scale-like or medially and laterally minutely interrupted, sometimes followed by short scales. Mid-transverse ridge not interrupted, medially slightly depressed, cervical groove indistinct, followed by 2 not interrupted or minutely interrupted ridges, interspersed with 2 short lateral ridges and few short scales. Lateral margins convex slightly convex, with 7 distinct spines: first anterolateral spine welldeveloped, reaching anteriorly to level of lateral orbital spine, second spine (hepatic) small, slightly dorsomesially from lateral margin, and followed by 5 branchial spines (3 anterior and 2 posterior). Rostrum leaf-like, horizontal, dorsally convex, [1.2] – 1.3 × as long as broad, length [0.3] – 0.4 and breadth 0.3 that of carapace; lateral margins smooth or minutely serrated and convex, with well-developed supraocular basal spines and small subapical spines (tridentiform). Pterygostomian flap with anterior spine, upper margin smooth. Thoracic sternum: As wide as long. Sternite 3 quadrangular, 1.5 – [1.7] × as wide as long, anterior margin convex, anterolaterally rounded. Sternite 4 widely contiguous to sternite 3; anterolaterally serrated, surface depressed in midline, smooth; greatest width [2.5] – 3.3 × that of sternite 3, 3.3 – [3.5] × as wide as long. Pleon: Elevated ridges with short setae and a few scattered long setae. Tergite 2 with anterior and posterior transverse elevated ridges; tergite 3 – 4 with anterior transverse ridge, posterior transverse ridge absent; tergites 5 – 6 smooth. Eye: Eyestalk length about 0.9 – [1.1] × broader than long, peduncle distally setose, not distinctly expanded proximally; cornea expanded distally, maximum corneal diameter [1.1] – 1.2 × rostrum width, as wide as eyestalk. Antennule: Article 1 slightly longer than wide, with 5 distal spines: distomesial spine well-developed; proximal lateral spine small, always present. Antenna: Article 1 with prominent mesial process, distally falling short or overreaching lateral antennular spine. Article 2 with well-developed distomesial and distolateral spines. Articles 3 with small distomesial spine. Article 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus 0.7 – [0.8] × length of ischium, with well-developed distal spine on extensor and flexor margins. P 1: 2.3 (males), [2.0] – 2.5 (females) × carapace length; subcylindrical, spiny and with long stiff setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus [0.8] 0.6 – 0.9 length of carapace, [1.9] 1.6 – 2.1 × as long as carpus. Carpus [1.6] 1.5 – 1.9 × as long as wide. Palm [1.1] – 1.3 × carpus length, [1.5] – 2.3 × as long as broad. Fingers unarmed, 0.7 – [1.1] × palm length. P 2 – 4 (P 3 lost in holotype): Setose and spinose. Meri successively shorter posteriorly: P 3 merus 0.8 – 0.9 × length of P 2 merus, P 4 merus 0.7 – 0.9 × length of P 3 merus. P 2 merus, [0.5] – 0.6 × carapace length, [3.5] – 4.5 × as long as broad, [1.1] – 1.3 × as long as P 2 propodus; P 3 merus 3.5 – 4.0 × as long as broad, 0.9 × as long as P 3 propodus; P 4 merus 3.0 – [3.3] × as long as broad, 0.9 – [1.1] × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, unarmed; flexor margin irregular, with distal spine on P 2 – 4. Carpi with 4 spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, absent on P 4; granules below extensor margin on lateral surface of P 2 – 4; flexor margin unarmed. Propodi stout, [4.3 – 4.7] 5.0 – 5.8 × as long as broad; extensor margin irregular, armed with 3 spines; flexor margin with 3 – 4 slender movable spines in addition to distal pair. Dactyli [0.7 – 0.8] 0.6 – 0.8 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 5 – 6 movable spines. Eggs: Ov. F carried approximately 15 – 40 eggs of 0.4 – 0.5 mm diameter. Live colour. Unknown. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF6FE6954F9CFDA9756FBD46.taxon	distribution	Distribution. Indonesia, Marsegu Island (Seram Island), Papua New Guinea, and Western Australia, from 0 to 27 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF6FE6954F9CFDA9756FBD46.taxon	discussion	Remarks. Phylladiorhynchus talaus belongs to the group of species having 2 epigastric spines, 1 hepatic spine and 3 spines on anterior branchial margin. The closest species is P. triginta Schnabel & Ahyong, 2019 from Tasman Sea and Norfolk Island, but they are easily differentiated by several characters (see the differences under the Remarks of this species). The two sequences of P. talaus from Western Australia and Papua New Guinea diverged by 2.3 % (COI) and 0.2 % (16 S), respectively. The specimens of P. talaus range from 1.8 to 2.7 mm postorbital carapace length.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF6DE69B4F9CFB147200BC76.taxon	description	(Figs. 50 K, 52)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF6DE69B4F9CFB147200BC76.taxon	materials_examined	Type material. Holotype. New Caledonia, SURPRISES Stn DW 1395, 18.2933 ° S, 163.0316 ° E, 34 – 36 m, 13 May 1999: M 2.4 mm (MNHN-IU- 2014 - 13794). Paratypes. New Caledonia. CHALCAL 2 Stn DW 80, 23.4500 ° S, 168.0333 ° E, 80 – 160 m, 31 October 1986: 2 M 2.4 – 2.9 mm, 1 ov. F 2.1 mm (MNHN-IU- 2016 - 477).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF6DE69B4F9CFB147200BC76.taxon	etymology	Etymology. From the name Tiphys, an Argonaut, son of Hagnias. The name is considered a substantive in apposition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF6DE69B4F9CFB147200BC76.taxon	description	Description. Carapace: As long as or slightly longer than broad; transverse ridges with dense short setae and few scattered long and thick iridescent setae. Gastric region slightly convex with 4 transverse ridges: epigastric ridge distinct with 2 median spines, scale between spines with thick setae, short scales laterally; anterior protogastric ridge not medially interrupted, nearly extending laterally to carapace margin; anterior mesogastric ridge not medially interrupted, laterally continuing to first branchial spine, interrupted by interception with cervical groove; anterior metagastric ridge scale-like. Mid-transverse ridge not interrupted, medially slightly depressed, preceded by a shallow or indistinct cervical groove, followed by 2 uninterrupted or minutely interrupted ridges, interspersed with 1 short lateral ridge and few short scales. Lateral margins straight or slightly convex, with 6 distinct spines: first anterolateral spine well-developed, reaching anteriorly to level of lateral orbital spine, second spine (hepatic) small, slightly dorsomesially from lateral margin, and followed by 4 branchial spines (3 anterior and 1 posterior). Rostrum leaf-like, horizontal, dorsally convex, [1.2] – 1.7 × as long as broad, length 0.4 and breadth 0.3 that of carapace; lateral margins minutely serrated and convex, with well-developed supraocular basal spines and small subapical spines. Pterygostomian flap with anterior spine, upper margin smooth. Thoracic sternum: As wide as long. Sternite 3 moderately broad, 1.2 – [1.5] × as wide as long, anterior margin convex, anterolaterally projected. Sternite 4 widely contiguous to sternite 3; anterolaterally smooth, surface depressed in midline, smooth; greatest width [3.0] – 3.5 × that of sternite 3, 2.5 – 3.0 × as wide as long. Pleon: Elevated ridges with short setae and a few scattered long setae. Tergite 2 with anterior and posterior transverse elevated ridges; tergite 3 with anterior transverse ridge, posterior transverse ridge interrupted; tergite 4 with anterior transverse ridge; tergites 5 – 6 smooth. Eye: Eyestalk length about [1.1] × broader than long, peduncle distally setose, not distinctly expanded proximally, with few short transverse striae on lateral surfaces; cornea expanded distally, maximum corneal diameter [0.6] × rostrum width, as wide as eyestalk. Antennule: Article 1 slightly longer than wide, with 5 distal spines: distomesial spine well-developed; proximal lateral spine small, always present. Antenna: Article 1 with prominent mesial process distally not reaching lateralmost antennular spine. Article 2 with well-developed distomesial and distolateral spines. Articles 3 and 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus [0.8] × length of ischium, with well-developed distal spine on extensor and flexor margins. P 1: [2.4] – 3.0 (males), 2.3 (females) × carapace length; subcylindrical, spiny and with long stiff setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines usually stronger than others. Merus [0.8] – 0.9 length of carapace, [1.3] – 2.0 × as long as carpus. Carpus 2.0 – [2.5] × as long as wide. Palm [1.1] – 1.2 × carpus length, 2.0 – [2.2] × as long as broad. Fingers unarmed, [0.7] – 0.8 × palm length, spine at hingepoint of palm-finger joint. P 2 – 4: Stout, setose and spinose. Meri successively shorter posteriorly: P 3 merus [0.9] × length of P 2 merus, P 4 merus [0.7] × length of P 3 merus. P 2 merus, [0.7] × carapace length, [3.5] – 4.7 × as long as broad, 1.2 × as long as P 2 propodus; P 3 merus [3.9] × as long as broad, [1.1] × as long as P 3 propodus; P 4 merus [3.2] – 3.5 × as long as broad, 0.9 × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, with some spines; flexor margin irregular, with distal spine on P 2 – 4. Carpi with 4 spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, absent on P 4; granules below extensor margin on lateral surface of P 2 – 4; flexor margin unarmed. Propodi stout, [4.5 – 5.5] 5.5 – 6.0 × as long as broad; extensor margin irregular, armed with 3 spines; flexor margin with 3 – 4 slender movable spines in addition to distal pair. Dactyli [0.6 – 0.7] 0.5 – 0.7 × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 5 – 6 movable spines. Eggs: Ov. F (MNHN-IU- 2016 - 477) carried 17 eggs of 0.3 mm diameter. Live colour. Unknown. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF6DE69B4F9CFB147200BC76.taxon	distribution	Distribution. New Caledonia, between 34 and 160 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF6DE69B4F9CFB147200BC76.taxon	discussion	Remarks. Phylladiorhynchus tiphys belongs to the group of species having 2 epigastric spines, 1 hepatic spine, and 3 spines on the anterior branchial margin. Phylladiorhynchus tiphys closely resembles to P. zetes from the French Polynesia (see the differences under the Remarks of this species). The specimens range in size from 2.1 to 2.9 mm postorbital carapace length.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF63E6984F9CFA137472B8B2.taxon	diagnosis	Diagnosis (modified from Schnabel & Ahyong 2019). Rostrum lateral margins distinctly convex; subapical spines minute. Carapace with 2 epigastric spines; anterior branchial margin with 3 spines; metagastric ridge between third branchial marginal spines interrupted medially; anterior branchial margin with 3 spines; hepatic spine present. Anterior upper margin of pterygostomian flap usually smooth. Pleonal tergite 3 without posterior transverse ridge. Thoracic sternite 3 moderately broad, anterior margin biconcave, with obtuse median projection. Antennular article 1 with 5 spines: distomesial spine large; second lateral spine long, slender. Antennal article 1 mesial process falling well short of second lateral antennular spine; article 2 distolateral and distomesial spines well developed, subequal in length; article 3 with small to minute distomesial spine, distolaterally unarmed. Maxilliped 3 merus with 1 prominent spine on flexor margin. P 2 – 4 dactylus extensor margin without upright spines at bases of movable spines. Genetic data. COI, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF63E6984F9CFA137472B8B2.taxon	distribution	Distribution. Tasman Sea, Lord Howe Island, Middleton Reef and Norfolk Island at 10 – 84 m (Schnabel & Ahyong 2019).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF63E6984F9CFA137472B8B2.taxon	discussion	Remarks. The species belongs to the group of species with 2 epigastric spines, hepatic margin armed with 1 small spine and 3 spines along the anterior branchial margin. The closest relative is P. talaus, from Indonesia, Papua New Guinea, and Western Australia. The two species can be differentiated by the following characters: - The rostrum has well-developed subapical spines in P. talaus, whereas these spines are minute in P. triginta. - Thoracic sternite 3 is quadrangular (less than twice wider than long) in P. talaus, compared to moderately broad (twice as wide as long) in P. triginta. The genetic divergence between P. triginta and P. talaus is very high, 27 % (COI).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF60E69D4F9CFE4072DBBE6E.taxon	description	(Figs. 50 L, 53, 56 C)	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF60E69D4F9CFE4072DBBE6E.taxon	materials_examined	Type material. Holotype. French Polynesia, Marquesas Islands, Nuku Hiva, Ohotea Point, W side of Taiohae Bay, 08.9309 ° S, 140.0978 ° W, 36 m, 28 November 2012: M parasitized 2.1 mm (UF 30059). Paratypes: French Polynesia. Marquesas Islands. PAKAIHI I TE MOANA Stn MQ 2 - GR-B, 08.9371 ° S, 140.1206 ° W, 20 – 23 m, 7 January 2012: 4 M 1.2 – 1.6 mm, 1 ov. F 2.3 mm, 3 F 1.2 – 1.7 mm (MNHN-IU- 2019 - 2639). Other material. French Polynesia. Marquesas Islands. PAKAIHI I TE MOANA Stn MQ 11 - GRS, 10.0140 ° S, 139. 10 ° 00.845 ’ S, 139 ° 07.345 ’ W 1224 ° W, 6 – 12 m, 15 January 2012: 1 F 1.7 mm (MNHN- IU- 2019 - 2602). — Stn MQ 15 - GR-B, 10.4718 ° S, 138.6780 ° W, 0 – 28 m, 17 June 2012: 2 M 2.0 – 2.2 mm (MNHN-IU- 2019 - 2593). — Stn MQ 19 - B, 09.7612 ° S, 138.8449 ° W, 10 – 25 m, 21 January 2012: 2 M 1.8 – 2.3 mm (MNHN-IU- 2019 - 2600). — Stn MQ 27 - GR-B, 08.6781 ° S, 140.6205 ° W, 5 – 22 m, 25 January 2012: 1 M 1.5 mm (MNHN-IU- 2019 - 2589). French Polynesia. BENTHAUS Stn DW 1926, 24.6360 ° S, 146.0136 ° W, 50 – 90 m 13 November 2002: 1 F parasitized 2.4 mm (MNHN-IU- 2014 - 13873). — Society Islands. Moorea Islands. 17.5044 ° S, 149.7584 ° W, 65 – 66 m, 26 January 2012: 1 M 1.5 mm (UF 33537). — 17.5044 ° S, 149.7584 ° W, 65 – 66 m, 26 January 2012: ov. F 1.4 mm (UF 33536). — 17.5526 ° S, 149.7735 ° W, 57 m, 27 January 2012: 1 M 1.5 mm (UF 33636). French Polynesia. Marquesas Islands. MUSORSTOM 9 Stn DW 1203,09.8783 ° S, 139.0366 ° W, 60 m, 28 August 1997: 1 M 1.8 mm (MNHN-IU- 2014 - 13874). — Stn CP 1264, 09.3550 ° S, 140.1283 ° W, 53 – 57 m, 03 September 1997: 1 M 2.6 mm (MNHN-IU- 2019 - 2640).	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF60E69D4F9CFE4072DBBE6E.taxon	etymology	Etymology. From the name Zetes, an Argonaut, son of Boreas and Oreithyia. The name is considered a substantive in apposition.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF60E69D4F9CFE4072DBBE6E.taxon	description	Description. Carapace: As long as or slightly wider than long; transverse ridges with dense short setae and few long and thick iridescent setae. Gastric region slightly convex with 4 transverse ridges: epigastric ridge indistinct with 2 median spines and some, and several lateral short scales; anterior protogastric ridge not medially interrupted, nearly extending laterally to carapace margin; anterior mesogastric ridge not medially interrupted, laterally continuing to first branchial spine; anterior metagastric scale-like, often followed by small scale. Midtransverse ridge not interrupted, medially slightly depressed, preceeded by shallow or indistinct cervical groove, followed by 2 not interrupted or laterally interrupted ridges, interspersed with 1 short lateral ridge and few short scales. Lateral margins straight or slightly convex, with 6 distinct spines: first anterolateral spine well-developed, reaching anteriorly to level of lateral orbital spine, second spine (hepatic) small, slightly dorsomesial from lateral margin, and followed by 3 – 4 branchial spines (3 anterior and 0 – 1 posterior). Rostrum leaf-like, horizontal, dorsally convex, 1.3 – [1.5] × as long as broad, length 0.4 and breadth 0.3 that of carapace; lateral margins minutely serrated and convex, with well-developed supraocular basal spines and subapical spines small. Pterygostomian flap with anterior spine, upper margin smooth. Thoracic sternum: As wide as long. Sternite 3 moderately broad, 1.9 – [2.0] × as wide as long, anterior margin convex. Sternite 4 widely contiguous to sternite 3; anterolaterally smooth, surface depressed in midline, smooth; greatest width 2.5 – [2.7] × that of sternite 3, [2.7] – 2.8 × as wide as long. Pleon: Elevated ridges with short setae and a few scattered long setae. Tergite 2 with anterior and posterior transverse elevated ridges; tergite 3 with anterior transverse ridge, posterior transverse ridge interrupted; tergite 4 with anterior transverse ridge; tergites 5 – 6 smooth. Eye: Eyestalk length about [1.1] × broader than long, peduncle distally setose, not distinctly expanded proximally, with few short transverse striae on lateral surfaces; cornea expanded distally, maximum corneal diameter 0.7 – [1.0] × rostrum width, as wide as eyestalk. Antennule: Article 1 slightly longer than wide, with 5 distal spines: distomesial spine well-developed; proximal lateral spine small, always present. Antenna: Article 1 with prominent mesial process. Article 2 with well-developed distomesial and distolateral spines. Articles 3 and 4 unarmed. Mxp 3: Ischium with distinct distal spines on flexor and extensor margins. Merus 0.6 – [0.8] × length of ischium, with well-developed distal spine on extensor and flexor margins. P 1: [2.6] – 3.0 (males), 2.3 – 2.4 (females) × carapace length; subcylindrical, spiny and with long stiff setae; merus, carpus and palm with spines along mesial, dorsal and lateral surfaces, distal and mesial spines stronger than others. Merus 0.8 – [1.0] length of carapace, 1.6 – [2.0] × as long as carpus. Carpus [1.7] – 2.5 × as long as wide. Palm 1.0 – [1.2] × carpus length, [1.8] 1.8 × as long as broad. Fingers 0.7 – [0.8] × palm length fixed finger with 2 basal spines; movable finger with 1 – 2 basal spines. P 2 – 4: Stout, setose and spinose. Meri successively shorter posteriorly: P 3 merus 0.9 – [0.9] × length of P 2 merus, P 4 merus 0.8 – [0.9] × length of P 3 merus. P 2 merus, 0.6 – [0.7] × carapace length, 3.0 – [3.5] × as long as broad, 1.1 – [1.4] × as long as P 2 propodus; P 3 merus 3.6 – [3.7] × as long as broad, 1.0 – [1.3] × as long as P 3 propodus; P 4 merus 3.5 – [3.8] × as long as broad, 0.9 – [1.2] × as long as P 4 propodus; extensor margin of P 2 and P 3 with row of spines, proximally diminishing, with prominent distal spine; P 4 extensor margin irregular, unarmed; flexor margin irregular, with distal spine on P 2 – 4. Carpi with 1 – 2 spines on extensor margin on P 2 – 3, unarmed on P 4; distal spine prominent on P 2 – 3, absent on P 4; granules below extensor margin on lateral surface of P 2 – 4; flexor margin unarmed. Propodi stout, [3.5 – 3.7] 3.1 – 3.9 × as long as broad; extensor margin irregular, armed with 0 – 3 spines; flexor margin with 3 – 4 slender movable spines in addition to distal pair. Dactyli 0.6 – [0.7] × length of propodi, ending in incurved, strong, sharp spine; flexor margin with 5 – 6 movable spines. Eggs: Ov. F carried approximately 12 – 20 eggs of 0.2 – 0.3 mm diameter. Live colour. Body light orange, with small orange spots and wittish patches and bands. P 1 whitish, palm and fingers orange, distal tip finger darker. P 2 – 4 light whitish-translucent. Genetic data. COI and 16 S, Table 1.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF60E69D4F9CFE4072DBBE6E.taxon	distribution	Distribution. French Polynesia, 20 to 23 m.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
2B7E87C3FF60E69D4F9CFE4072DBBE6E.taxon	discussion	Remarks. Phylladiorhynchus zetes belongs to the group of species having 2 epigastric spines, 1 hepatic spine, 3 spines on the anterior branchial margin and the pleonal tergite 3 with the posterior ridge interrupted. Phylladiorhynchus zetes closely resembles P. tiphys from New Caledonia but they can be distinguished by the following characters: - One scale with thick setae between the epigastric spines in P. tiphys, whereas this scale is absent in P. zetes. - Thoracic sternite 3 is quadrangular (less than twice as long as wide) in P. tiphys, whereas this sternite is moderately broad (more than twice as long as broad) in P. zetes. - The P 2 – 4 propodi are usually stout in P. zetes (3.1 – 3.9 × as long as broad), whereas they are more slender in P. tiphys (3.5 – 4.5 as long as broad). Phylladiorhynchus zetes is also highly similar to P. medea, also found in French Polynesia and New Caledonia. Both species can be distinguished by the following characters: - The rostrum has small or minute subapical spines in P. zetes, whereas these spines are obsolescent in P. medea. - Pleonal tergite 3 has a posterior ridge in P. zetes, whereas this ridge is absent in P. medea. - The P 2 – 4 propodi are usually stout in P. zetes (3.1 – 3.9 × as long as broad), whereas they are more slender in P. medea (3.5 – 4.5 as long as broad). The genetic divergences among P. zetes, P. tiphys and P. medea ranged from 7 – 11 % (COI) and 3 – 6 % (16 S). The two sequences of P. zetes from French Polynesia diverged by 0.2 % (COI) and 0 % (16 S). The specimens of P. zetes range from 1.2 to 2.6 mm postorbital carapace length.	en	Rodríguez-Flores, Paula C., Macpherson, Enrique, Machordom, Annie (2021): Revision of the squat lobsters of the genus Phylladiorhynchus Baba, 1969 (Crustacea, Decapoda, Galatheidae) with the description of 41 new species. Zootaxa 5008 (1): 1-159, DOI: 10.11646/zootaxa.5008.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5008.1.1
