taxonID	type	description	language	source
2B77423D341BFFBCD99AF8996590006F.taxon	materials_examined	Type species: Heterococcopsis lonicerae Borchsenius, 1948	en	Kaydan, Bora (2011): Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey. Zootaxa 2970: 49-62, DOI: 10.5281/zenodo.203960
2B77423D341BFFBCD99AF8996590006F.taxon	discussion	Comment. Hitherto, Heterococcopsis included three species: H. desertus Bazarow & Nurmamatov, H. lonicera Borchsenius and H. opertus. Herein, H. desertus is transferred to Phenacoccus, and H. opertus is transferred to Heterobrevennia gen. nov., leaving Heterococcopsis as a monotypic genus.	en	Kaydan, Bora (2011): Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey. Zootaxa 2970: 49-62, DOI: 10.5281/zenodo.203960
2B77423D3418FFBAD99AFC9065AE0245.taxon	materials_examined	Material examined: Lectotype adult female (here designated) + 2 paralectotype adult females (on 1 slide): Kazakhstan, Alma-Ata, Lonicera tatarica (Caprifoliaceae), Saygurov, 02. vi. 36 (ZIAS: 80 / 37). Lectotype specimen in circle on left side, other 2 specimens indicated as paralectotypes. Adult female (Fig. 1) Live appearance: Body oval. Mounted material. Body elongate oval, 1.80 – 2.00 mm long, 0.82 – 1.28 mm wide. Eyes located on margin, each 30 µm wide. Antenna 9 segmented, 270 – 235 µm long; apical segment 35 – 45 µm long, 17.5 – 20.0 µm wide, with apical setae 27.5 – 30.0 µm long and with 3 fleshy setae, each 27.5 – 30.0 µm long. Clypeolabral shield 150 – 165 µm long, 110 µm wide. Labium 80 – 90 µm long, 62.5 – 70.0 µm wide. Anterior spiracles 37.5 – 45.0 µm long, 20 – 25 µm wide across atrium; posterior spiracles 50 µm long, 22.5 – 25.0 µm wide across atrium. Circuli numbering 3, each oval in shape, each 35 µm, 32.5 µm and 20 µm wide respectively. Legs well developed; coxa 70 – 80 µm long, hind trochanter + femur 130 – 145 µm long, hind tibia + tarsus 155 – 160 µm long, hind claw 20.0 – 22.5 µm long. Ratio of lengths of hind tibia + tarsus to hind trochanter + femur 1.10 – 1.19: 1, ratio of lengths of hind tibia to tarsus 1.71 – 1.83: 1, ratio of length of hind trochanter + femur to greatest width of femur 3.71 – 4.14: 1. Tarsal digitules setose, each 15 – 20 µm long. Claw digitules simalar and capitate, each 17.5 – 20.0 μm long, knobbed and thicker than tarsal digitules. Translucent pores present on tibia of hind legs, totalling 32 – 43. Posterior pairs of ostioles present, each with 4 trilocular pores and 3 setae; anterior ostiole, not easy to detect or poorly developed. Anal ring 55 µm wide, with 6 anal ring setae, each seta 50 – 60 µm long. Dorsum. Cerarii numbering 2 pairs, situated on posterior 2 abdominal segments: anal lobe cerarii each with 2 conical setae (each 14 – 16 µm long) plus 5 trilocular pores and C 17 each with 2 conical setae (each 15 µm long) plus 2 trilocular pores. Dorsal body setae spine-like, each 5.0 – 7.5 µm long. Multilocular pores each 6.25 – 7.5 µm wide with more than 10 loculi, present across posterior 2 abdominal segments and along body margin. Quinquelocular pores each 5 µm in diameter, scattered over dorsum but absent from last abdominal segment. Trilocular pores each 4 – 5 µm in diameter, restricted to last abdominal segment, cerarii and posterior ostioles. Oral-collar tubular ducts, each 9 – 11 µm long, 2 – 3 µm wide, present in a transverse row across last abdominal IV-VIII segments and along body margin. Venter. Setae slender, each 20 – 55 µm long, longest setae present medially on head; apical setae of anal lobe 75 – 105 µm long. Multilocular disc pores each 7.5 – 8.0 µm in diameter, present across abdominal segments IV − VIII + IX as follows: 0 – 6 pores on posterior margins of segment IV, 1 – 12 pores on posterior margins of V, 18 – 22 pores on posterior margins of VI, 30 – 32 across full width of VII, 19 – 34 across full width of VIII + IX. Quinquelocular and heptalocular pores each 4 – 5 µm wide, scattered over venter. Oral-collar tubular ducts each 8 – 11 µm long, 2.5 – 3.0 µm wide, present in a row on last abdominal segment and on margins and submargins of each abdominal segment; ducts distributed as follows: 2 – 4 on segment I; 2 – 4 on II; 1 – 4 on III; 3 – 4 on IV; 4 on V; 4 on VI; 4 – 6 on VII; and 4 – 6 on VIII and a few on the metathorax and the mesothorax.	en	Kaydan, Bora (2011): Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey. Zootaxa 2970: 49-62, DOI: 10.5281/zenodo.203960
2B77423D3418FFBAD99AFC9065AE0245.taxon	discussion	Comment: For discussion, see under Heterobrevinia gen. nov. below.	en	Kaydan, Bora (2011): Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey. Zootaxa 2970: 49-62, DOI: 10.5281/zenodo.203960
2B77423D341EFFBBD99AFE636730016D.taxon	materials_examined	Type species: Heterobrevennia opertus (Borchsenius, 1949) Generic diagnosis. Adult female: Body elongate, 2.82 – 3.20 mm long, 1.56 – 2.20 mm wide. Antennae generally 8 or 9 segmented (but occasionally 7 when 2 nd and 3 rd segments fused). Eyes present on margin. Labium 3 segmented. Posterior spiracles slightly larger than anterior spiracles. Legs well developed, with translucent pores on hind femur and tibia; claw with a denticle; tarsal digitules setose, claw digitules capitate, broader than tarsal digitules. Both anterior and posterior ostioles well developed. Multilocular disc pores limited to margins of both body surfaces and to posterior abdominal segments. Quinquelocular pores smaller than multilocular pores, scattered over both body surfaces. Trilocular pores present on dorsum. Oral-collar tubular ducts elongate, generally situated on body margin. Anal ring with 6 setae. Dorsum. Cerarii numbering always 5 or more pairs, situated on posterior abdominal segments and on head. Dorsal body setae spine-like. Multilocular pores situated on body margin. Quinquelocular pores scattered throughout. Trilocular pores scattered throughout, sometimes few, but always present at least on cerarii, ostioles and near spiracle opening. Minute discoidal pores scattered throughout. Oral-collar tubular ducts elongate scattered throughout. Venter. Body setae slender, longest setae present medially on head; apical setae long; spine-like setae present submarginally on body. Multilocular disc pores present on posterior abdominal segments and on margin of thorax and head. Quinquelocular pores scattered over body. Trilocular pores present around atrium of spiracle, absent elsewhere. Minute discoidal pores scattered over body. Oral-collar tubular ducts in rows on posterior abdominal segment and scattered on thorax and head.	en	Kaydan, Bora (2011): Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey. Zootaxa 2970: 49-62, DOI: 10.5281/zenodo.203960
2B77423D341EFFBBD99AFE636730016D.taxon	discussion	Comment. Heterobrevennia gen. nov. is most similar to Heterococcopsis in having trilocular pores on the dorsum and 8 - or 9 - segmented antennae. The two genera differ as follows (features of Heterococcopsis in parentheses): i) no circuli (with 3 circuli), ii) trilocular pores scattered on dorsum (trilocular pores on dorsum restricted to cerarii, ostioles and posterior abdominal segments), iii) at least five cerarii, of which 3 are present on head (only 2 cerarii, restricted to posterior abdominal segments), and iv) legs slender (legs stout). Heterobrevennia is also similar to all the other genera in the Heterococcus - group. The affinities of Heterobrevennia with other related genera are discussed below. Asphodelococcus can be readily separated from Heterobrevennia by the following combination of features (features of Heterobrevennia in parenthesis): (i) antennae 6 segmented (antennae 9 segmented), (ii) multilocular pores of unusual shape and often quadrate, and with heavily sclerotised hexagonal pores (pores normal), and (iii) cerarii numbering 4 pairs and restricted to abdomen (cerarii numbering 5 or more pairs, present on both abdomen and head).	en	Kaydan, Bora (2011): Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey. Zootaxa 2970: 49-62, DOI: 10.5281/zenodo.203960
2B77423D341EFFBBD99AFE636730016D.taxon	etymology	Etymology. The name Heterobrevennia refers to its similarity to the genera Heterococcus and Brevennia. Gender female.	en	Kaydan, Bora (2011): Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey. Zootaxa 2970: 49-62, DOI: 10.5281/zenodo.203960
2B77423D341FFFB9D99AFC30644307BE.taxon	materials_examined	Material examined: Holotype adult female, Turkey, VAN-Gevaş, N: 38 ° 16 ’ 663 ’’, E: 043 ° 03 ’ 898 ’’, 1834 m altitude, on the leaf sheaths of Poaceae, M. B. Kaydan, 09. vi. 2009 (CCVT: 4530). Deposited in CCVT. Paratype adult females, same data as holotype, 5 specimens on 5 slides (4 Slides in CCVT: 4530 plus 1 in ZIAS). Adult female (Fig. 2) Live appearance: Body oval. In the leaf sheaths of grasses. Mounted material. Body elongate oval, 2.80 – 3.18 mm long, 1.12 – 1.25 mm wide. Eye marginal, 30 – 35 µm wide. Antenna 8 segmented (rarely 7 segmented, when 2 nd - and 3 rd - antennal segments fused), 255 – 270 µm long; apical segment 40 – 45 µm long, 20.0 – 22.5 µm wide, with apical seta 30 µm long, plus 3 fleshy setae, each 15.0 – 27.5 µm long. Clypeolabral shield 130 – 140 µm long, 102.5 – 110.0 µm wide. Labium 67.5 – 70.0 µm long, 60.0 – 67.5 µm wide. Anterior spiracles 40.0 – 47.5 µm long, 20 – 25 µm wide across atrium; posterior spiracles 45 – 55 µm long, 30 – 35 µm wide across atrium. Legs well developed; coxa 80 – 95 µm long, hind trochanter + femur 175 – 195 µm long, hind tibia + tarsus 210 – 230 µm long, hind claw 20.0 – 22.5 µm long. Ratio of lengths of hind tibia + tarsus to hind trochanter + femur 1.15 – 1.24: 1, ratio of lengths of hind tibia to tarsus 1.91 – 2.21: 1, ratio of length of hind trochanter + femur to greatest width of femur 5.0 – 5.3: 1. Tarsal digitules setose, each 20 – 25 µm long. Claw digitules subequal, each 22.5 – 25.0 µm long, knobbed and thicker than tarsal digitules. Translucent pores present on femur and tibia of hind legs, totalling 12 – 26. Both pairs of ostioles present, posterior ostioles each with 7 – 10 trilocular pores plus 0 – 3 setae, anterior ostioles with 4 – 6 trilocular pores and 1 seta. Anal ring 55 – 60 µm wide, with 6 anal-ring setae, each 55 – 65 µm long. Dorsum. Cerarii numbering 6 pairs, with pairs on posterior 3 abdominal segments plus 3 anteriorly on head; anal lobe cerarii, each with 2 conical setae, each 15.0 – 20 µm long, plus 4 or 5 trilocular pores; C 17 each with 2 conical setae, each 15.0 – 17.5 µm long, plus 2 trilocular pores; C 16 each with 2 conical setae, each 10.0 – 12.5 µm long, plus 1 trilocular pore; C 1 each with 2 conical setae, each 7.5 – 10 µm long, plus 2 trilocular pores; C 2 each with 2 conical setae, each 7.5 – 10 µm long, plus 2 trilocular pores; C 3 each with 3 conical setae, each 7.5 – 10 µm long, plus 2 trilocular pores. Dorsal body setae spine-like, each 7.5 – 15.0 µm long, becoming shorter posteriorly. Multilocular pores each 6.0 – 7.5 µm wide with more than 10 loculi, in rows on posterior 3 abdominal segments, and on margin of remaining abdominal segments and thorax; also with 2 pairs groups on head: 1 pair each with 12 – 19 pores close to antennae, and other pairs each with 11 – 15 pores nearer anterior ostioles. Quinquelocular pores, each 5 – 6 µm wide, scattered throughout. Trilocular pores, each 5 – 6 µm in diameter, scattered throughout. Minute discodial pores few, each 2 µm in diameter, scattered on body surface. Oral-collar tubular ducts, each 9 – 11 µm long, 2 – 3 µm wide, scattered on all abdominal segments and along body margin on thorax. Venter. Body setae slender, each 10 – 45 µm long, longest setae medially on head; apical anal lobe setae 120.0 – 137.5 µm long. Multilocular disc pores each 7.5 – 8.0 µm in diameter, present in rows or bands on posterior abdominal segments (IV – VIII + IX): 6 – 8 pores on segment IV, 26 – 37 pores on segment V, 48 – 69 pores on segment VI, 58 – 62 on segment VII, 34 – 42 on segments VIII + IX. Quinquelocular pores each 4 – 5 µm wide, scattered throughout. Trilocular pores each 5 – 6 µm in diameter, restricted to around spiracle atrium. Minute discodial pores, each 2 µm in diameter, scattered throughout. Oral-collar tubular ducts each 9 – 11 µm long, 2.5 – 3.0 µm wide, present in rows or bands on posterior abdominal segments plus a few near margins on head and thorax; distributed as follows: 2 – 4 on each side of abdominal segments I – III; 8 – 12 on IV; 31 – 38 on V; 40 – 53 on VI; 34 – 42 on VII, and 4 – 10 on VIII.	en	Kaydan, Bora (2011): Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey. Zootaxa 2970: 49-62, DOI: 10.5281/zenodo.203960
2B77423D341FFFB9D99AFC30644307BE.taxon	discussion	Comment. The adult female of Heterobrevennia gullanae Kaydan is unique for a species in the Heterococcusgroup in having the following combination of features: (i) many quinquelocular pores on both the dorsum and venter, (ii) short hair-like tarsal digitules, (iii) claw digitules capitate, thicker than tarsal digitules, (iv) many multilocular pores on venter, (v) translucent pores present on both femur and tibia, (vi) two groups of multilocular pores on dorsum of head, (viii) six pairs of cerarii, 3 on head and 3 posteriorly on abdomen, (ix) many oral-collar tubular ducts on the venter and dorsum, and (x) the presence of trilocular pores throughout the dorsum. Heterobrevennia gullanae resembles H. opertus in having most of the above features but H. gullanae differs in having the following combination of characters (characters of H. opertus in brackets): (i) two groups of multilocular pores on the head (absent), (ii) cerarii numbering 6 pairs (7), (iii) many trilocular pores present throughout dorsum (trilocular pores on dorsum few, generally restricted to posterior abdominal segment, and in cerarii and ostioles), (iv) antennae 8, rarely 7 segmented (9 segmented). Heterobrevennia gullanae is also similar to H. kozari Kaydan spec. nov., described below, but differs in having (character states for H. kozari in brackets): (i) 6 pairs of cerarii (5), (ii) translucent pores on coxa absent (present), and (iii) one size of oral-collar tubular ducts on dorsum (two sizes medially on posterior abdominal segment).	en	Kaydan, Bora (2011): Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey. Zootaxa 2970: 49-62, DOI: 10.5281/zenodo.203960
2B77423D341FFFB9D99AFC30644307BE.taxon	etymology	Etymology. The new species is named in honor of Prof. Dr. Penny Gullan (Division of Evolution, Ecology & Genetics Research School of Biology, The Australian National University, Canberra, Australia), for her valuable studies on all aspects of coccidology.	en	Kaydan, Bora (2011): Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey. Zootaxa 2970: 49-62, DOI: 10.5281/zenodo.203960
2B77423D341DFFB7D99AFAFA6449056A.taxon	materials_examined	Material examined. Holotype adult female, Turkey, Bitlis-Siirt road, N: 38 ° 11 ’ 772 ’’, E: 041 ° 49 ’ 067 ’’, 748 m altitude, on the leaf sheaths of Cynodon dactylon (Poaceae), M. B. Kaydan, 26. v. 2008 (CCVT: 4325). Deposited in CCVT. Paratype adult females: Turkey, Diyarbakır-Silvan road, N: 38 ° 07 ’ 722 ’’, E: 041 ° 10 ’ 530 ’’, 748 m altitude, on the leaf sheaths of Cynodon dactylon, M. B. Kaydan and Ferenc Kozár, 26. v. 2008. Six parasitized specimens deposited as follows: 5 in CCVT: 4327 and 1 in ZIAS. Adult female (Fig. 3) Description made solely from the holotype. Measurements from other specimens not used because they were parasitized. Live appearance. Body oval. In the leaf sheaths of C. dactylon. Mounted material. Body elongate oval, 3.28 mm long, 1.28 mm wide. Eye marginal, 35 – 40 µm wide. Antenna 9 segmented, 380 – 390 µm long; apical segment 50 – 55 µm long, 25 – 30 µm wide with apical setae 30 µm long, plus 3 fleshy setae, each 25 – 30 µm long. Clypeolabral shield 135 µm long, 152.5 µm wide. Labium 100 µm long, 75 µm wide. Anterior spiracles 55 – 60 µm long, 25.0 – 27.5 µm wide across atrium; posterior spiracles 60 – 65 µm long, 35 µm wide across atrium. Legs well developed; coxa 140 – 145 µm long; hind trochanter + femur 230 – 235 µm long; hind tibia + tarsus 285 – 290 µm long, and hind claw 27.5 – 30.0 µm long. Ratio of lengths of hind tibia + tarsus to hind trochanter + femur 1.21 – 1.26: 1, ratio of lengths of hind tibia to tarsus 2.05 – 2.21: 1, ratio of length of hind trochanter + femur to greatest width of femur 5.17 – 5.27: 1. Tarsal digitules setose, each 27.5 – 30.0 µm long. Claw digitules subequal, each 27.5 – 30.0 µm long, knobbed and thicker then tarsal digitules. Translucent pores present on coxa, femur and tibia of hind legs, totalling 58 – 79. Both pairs of ostioles present, posterior ostioles with 8 – 12 trilocular pores plus 1 seta, anterior ostioles 6 trilocular pores and 1 setae. Anal ring 70 µm wide, with 6 analring setae, each 120 – 125 µm long. Dorsum. Cerarii numbering 5 pairs, with pairs on posterior 2 abdominal segments, plus 3 pairs on head: anal lobe cerarii each with 2 conical setae, each 17.5 – 20 µm long, plus 6 or 7 trilocular pores; C 17 each with 2 conical setae, each 15 µm long, plus 3 – 4 trilocular pores; C 1 each with 2 conical setae, each 7.5 – 10 µm long, plus 1 trilocular pore; C 2 each with 2 conical setae, each 7.5 – 10 µm long, plus 1 or 2 trilocular pores; C 3 each with 3 conical setae, each 7.5 µm long, with 2 trilocular pores. Dorsal setae spine-like, each 7 – 11 µm long. Multilocular pores each 6.0 – 7.5 µm with more than 10 loculi, restricted to a few on abdominal segments VI and VII. Quinquelocular pores each 4 – 5 µm wide, scattered throughout. Trilocular pores each 4 – 5 µm in diameter, scattered throughout. Minute discoidal pores few, each 2 µm in diameter, scattered on body surface. Oral-collar tubular ducts of 2 sizes, larger each 7 – 11 µm long, 4 – 5 µm wide, present in rows on each abdominal segment as follows: 14 on I; 8 on II; 19 on III; 16 on IV; 25 on V; 28 on VI; 20 on VII; 14 on VIII; also scattered on thorax plus a few on head; smaller ducts few, each 7 – 11 µm long, 2.5 – 3 µm wide, present medially on posterior 3 abdominal segments,. Venter. Body setae slender, each 10 – 85 µm long, longest setae medially on head; apical anal lobe setae 210 – 220 µm long. Multilocular disc pores each 7.5 – 9.0 µm in diameter, present in rows or bands on posterior abdominal segments (IV – VIII + IX): 4 – 6 pores on segment IV, 15 on segment V, 37 on segment VI, 38 on segment VII, 36 on segments VIII + IX. Quinquelocular pores each 4 – 5 µm wide, scattered throughout. Trilocular pores restricted to around spiracle atrium, each 2.5 – 3.0 µm in diameter. Minute discoidal pores each 2 µm in diameter, scattered throughout. Oral-collar tubular ducts each 7.5 – 10 µm long, 2.5 – 3.0 µm wide, in rows across most abdominal segments plus a few on thorax; distributed as follows: 2 on abdominal segment I; 4 on II; 11 on III; 12 on IV; 28 on V; 26 on VI; 28 on VII, and 15 on VIII.	en	Kaydan, Bora (2011): Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey. Zootaxa 2970: 49-62, DOI: 10.5281/zenodo.203960
2B77423D341DFFB7D99AFAFA6449056A.taxon	discussion	Comment. The adult female of Heterobrevennia kozari Kaydan is unique for a species in the Heterococcus group in having the following combination of character states: (i) many quinquelocular pores on both dorsum and venter, (ii) hair-like tarsal digitules, (iii) ventral multilocular pores present on posterior abdominal segments, (iv) translucent pores present on coxa, femur and tibia, (v) five pairs of cerarii, and (vi) many oral-collar tubular ducts throughout the dorsum. The most unique characters are the presence of 5 cerarii and pores on hind coxa. In having most of the above mentioned features, H. kozari resembles H. opertus but differs in having (characters of H. opertus in brackets): (i) only 5 pairs of cerarii (7), (ii) many trilocular pores scattered throughout dorsum (trilocular pores on dorsum few, generally restricted to the posterior abdominal segment, cerarii and ostioles), (iii) transcluent pores present on hind coxa (absent), and (iv) dorsal multilocular pores present only on abdominal segments VI and VII (scattered all over body). H. kozari is also similar to H. gullanae Kaydan, described above, but differs in having (characters of H. gullanae in brackets): (i) 5 pairs of cerarii (6), (ii) transcluent pores on coxa (absent), (iii) one size of oral-collar tubular ducts on dorsum (two sizes on posterior abdominal segment), (iv) dorsal multilocular pores restricted to abdominal segments VI and VII (also present in two group on head), and (v) antennae 9 segmented (8, rarely 7).	en	Kaydan, Bora (2011): Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey. Zootaxa 2970: 49-62, DOI: 10.5281/zenodo.203960
2B77423D341DFFB7D99AFAFA6449056A.taxon	etymology	Etymology. The new species is named in honor of Dr. Ferenc Kozár (Department of Zoology, Plant Protection Institute, Hungarian Academy of Sciences, Budapest, Hungary), who has made valuable studies on many aspects of coccidology throughout the World.	en	Kaydan, Bora (2011): Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey. Zootaxa 2970: 49-62, DOI: 10.5281/zenodo.203960
2B77423D3413FFB5D99AF99366950738.taxon	materials_examined	Material examined: Syntypic adult females: Tadzikistan, Gissarskii Ridge, Dactylis glomerata (Poaceae), H. Borchsenius, 17. vii. 1944 (2 specimens, ZIAS: 34 - 45). Adult female (Fig. 4) Live appearance: Body elongate oval. Mounted material. Body elongate oval, 2.82 – 3.60 mm long, 1.56 – 2.20 mm wide. Eye marginal, 32.5 – 35.0 µm wide. Antenna 9 segmented, 270 – 290 µm long; apical segment 45 – 55 µm long, 20 – 25 µm wide, with apical setae 30 – 35 µm long, plus 3 fleshy setae, each 30 – 35 µm long. Clypeolabral shield 145 – 150 µm long, 110 – 115 µm wide. Labium 80 – 85 µm long, 65 – 70 µm wide. Anterior spiracles 42.5 – 52.5 µm long, 25.0 – 27.5 µm wide across atrium; posterior spiracles 50 – 60 µm long, 25 – 35 µm wide across atrium. Legs well developed; coxa 90 – 105 µm long, hind trochanter + femur 215 – 220 µm long, hind tibia + tarsus 255 – 260 µm long, hind claw 25.0 – 27.5 µm long. Ratio of lengths of hind tibia + tarsus to hind trochanter + femur 1.16 – 1.18: 1, ratio of lengths of hind tibia to tarsus 1.8 – 2.03: 1, ratio of length of hind trochanter + femur to greatest width of femur 4.4 – 4.8: 1. Tarsal digitules setose, each 32.5 – 35.0 µm long. Claw digitules subequal, each 27.5 – 35.0 µm long, knobbed and broader than tarsal digitules. Translucent pores present on femur and tibia of hind legs, totalling 16 – 24. Both pairs of ostioles present, posterior ostioles each with 5 – 7 trilocular pores and 2 or 3 setae, anterior ostioles each with 5 trilocular pores and 2 or 3 setae. Anal ring 60 – 62.5 µm wide, with 6 anal-ring setae, each 85 – 95 µm long. Dorsum. Cerarii numbering 7 pairs, with pairs on posterior 4 abdominal segments, and on 3 pairs on head; anal lobe cerarii each with 2 conical setae, each 15.0 – 17.5 µm long, plus 5 – 7 trilocular pores; C 17 each with 2 conical setae, each 15.0 – 17.5 µm long, plus 4 trilocular pores; C 16 each with 2 conical setae, each 15 µm long, plus 2 trilocular pores; C 15 each with 2 conical setae, each 11.0 – 12.5 µm long, plus 2 trilocular pores; C 1 each with 2 conical setae, 10 µm long, plus 2 trilocular pores; C 2 each with 2 conical setae, each 10 µm long, plus 3 trilocular pores; and C 3 each with 3 conical setae, each 7.5 – 10 µm long, plus 2 trilocular pores. Dorsal body setae spine-like, each 10.0 – 12.5 µm long. Multilocular pores each 7.5 – 8.0 µm wide with 11 loculi, scattered throughout. Quinquelocular pores each 5 – 6 µm, scattered throughout. Trilocular pores each 5 – 6 µm in diameter, restricted to posterior abdominal segments VII, VIII and IX, and in cerarii, on ostioles and the middle area of thorax. Oral-collar tubular ducts present along body margin, each 9 – 11 µm long, 2 – 3 µm wide. Venter. Body setae slender, each 15 – 65 µm long, longest setae medially on head; apical setae of anal lobe 145 – 165 µm long. Multilocular disc pores each 7.5 – 8.0 µm in diameter, present in rows or bands on posterior abdominal segments (IV – VIII + IX): 12 pores on segment IV, 12 – 24 on V, 32 – 36 on VI, 53 – 62 on VII, and 27 – 33 on VIII + IX. Quinquelocular pores each 5 – 6 µm wide, scattered throughout. Oral-collar tubular ducts each 7.5 – 10.0 µm long, 2.5 – 3.0 µm wide, present in rows on all abdominal segments plus a few on head and thorax; distributed as follows: 2 – 4 on each side of abdominal segments I – III and a few medially; 4 on IV; 15 – 25 on V; 14 – 24 on VI; 19 on VII; 18 – 23 on VIII in row or band.	en	Kaydan, Bora (2011): Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey. Zootaxa 2970: 49-62, DOI: 10.5281/zenodo.203960
2B77423D3413FFB5D99AF99366950738.taxon	discussion	Comment. For a comparison of H. opertus with other species in the genus, see under H. gullae and H. kozari above.	en	Kaydan, Bora (2011): Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey. Zootaxa 2970: 49-62, DOI: 10.5281/zenodo.203960
2B77423D3411FFB3D99AFB7C671300C6.taxon	materials_examined	Material examined. Holotype adult female: Tadzhikistan, Gbao, Vosim, Gasur Lurgap, Org. Artemisia demissa (Asteraceae), L. Kuarzivsheva, 18. vii. 1924 (ZIAS). Adult female (Fig. 5) Live appearance. Body oval. Mounted material. Body oval, 2.20 mm long, 1.34 mm wide. Eye marginal, 35.0 µm wide. Antenna 9 segmented, 130 – 140 µm long; apical segment 47.5 µm long, 20 µm wide, with apical setae 22.5 µm long, plus 3 fleshy setae, each 20 – 30 µm long. Clypeolabral shield 145 µm long, 120 µm wide. Labium 110 µm long, 75 µm wide. Anterior spiracles 45 µm long, 15 µm wide across atrium; posterior spiracles 45.0 – 47.5 µm long, 25.0 – 27.5 µm wide across atrium. Legs well developed; coxa 40 µm long, hind trochanter + femur 150 µm long, hind tibia + tarsus 165 µm long, hind claw 25.0 µm long. Ratio of lengths of hind tibia + tarsus to hind trochanter + femur 1.10, ratio of lengths of hind tibia to tarsus 1.3: 1, ratio of length of hind trochanter + femur to greatest width of femur 3.3: 1. Tarsal digitules setose, each 10 µm long. Claw digitules subequal, each 17.5 µm long, knobbed and broader than tarsal digitules. Translucent pores present on femur and tibia of hind legs, totalling 16 – 24. Both pairs of ostioles present, posterior ostioles each with 9 trilocular pores and 1 setae, anterior ostioles each with 4 trilocular pores and 2 or 3 setae. Anal ring 55 µm wide, with 6 anal-ring setae, each 60 – 65 µm long. Dorsum. Cerarii numbering 2 pairs, with pairs on posterior 2 abdominal segments; anal lobe cerarii each with 2 conical setae (broken, as shown on figure), plus 5 – 7 trilocular pores; C 17 each with 2 conical setae, each 10 µm long, plus 2 trilocular pores. Dorsal body setae spine-like, each 5 – 10 µm long. Multilocular pores each 7.5 – 9.0 µm wide with 11 loculi, present in laterally in abdominal segment I and in rows or bands on posterior abdominal segments (II – VII): 12 pores on segment on II, 19 on III, 37 on IV, 37 on V, 24 on VI, 30 on VII and no pores on segment VIII. Quinquelocular pores very few (only three pores seen on head and thorax), each 5 – 6 µm. Trilocular pores each 4 – 5 µm in diameter, scattered throughout. Oral-collar tubular ducts each 7.5 – 10.0 µm long, 2 – 3 µm wide, present in rows on all abdominal segments plus 8 on head and thorax laterally; distributed on abdomen as follows: 3 on I; 5 on II; 6 on III; 6 on IV; 7 on V; 2 on VI; 3 on VII; 4 on VIII in row. Venter. Body setae slender, each 12.5 – 40.0 µm long, longest setae medially on head; apical setae of anal lobe broken. Multilocular disc pores present, each 7.5 – 8.0 µm in diameter, present laterally on abdominal segments I – II and in rows or bands on posterior abdominal segments (III – VIII + IX): 22 pores on segment III, 34 on IV, 36 on V, 48 on VI, 37 on VII, and 27 on VIII + IX. Quinquelocular pores each 5 – 6 µm wide, scattered throughout. Oral-collar tubular ducts each 7.5 – 9.0 µm long, 2.5 – 3.0 µm wide, present in rows on all abdominal segments plus a few on head and thorax; distributed as follows: 1 – 2 on each side of abdominal segments I – III; 13 on IV; 14 on V; 8 on VI in row or band and 3 on VII; 3 on VIII laterally.	en	Kaydan, Bora (2011): Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey. Zootaxa 2970: 49-62, DOI: 10.5281/zenodo.203960
2B77423D3411FFB3D99AFB7C671300C6.taxon	discussion	Comment. Species in the Heterococcus - group differ from those in Phenacoccus Cockerell in having: (i) quinquelocular pores on the dorsum and either no trilocular pores or trilocular pores restricted to around each spiracle. Because Heterococcopsis desertus lacks both of these character states, it is not congeneric with Heterococcopsis lonicerae Borchsenius, the type species of the genus Heterococcopsis but does appear to be congeneric with Phenacoccus aceris Signoret, the type species of the genus Phenacoccus. H. desertus shares the following important features with P. aceris: (i) presence of many trilocular pores on dorsum and venter, (ii) very few quinquelocular pores (only 3 were found) on dorsum, and (iii) a long labium. Based on the morphological affinities of H. desertus with P. aceris, I here transfer H. desertus to Phenacoccus, as Phenacoccus desertus (Bazarov & Nurmamatov) comb. nov. and illustrate the species based on the type material. This species is very close to Phenacoccus tergrigorianae Borchsenius & Ter-Grigorian, and P. p u m i l u s Kiritshenko in having multilocular pores on dorsum, but differs from both in having many quinquelocular pores on the venter. The genus Phenacoccus has many species and needs further study to clarify the status of the new combination of P. desertus.	en	Kaydan, Bora (2011): Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey. Zootaxa 2970: 49-62, DOI: 10.5281/zenodo.203960
