identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
2B77423D341BFFBFD99AFB3863EF0467.text	2B77423D341BFFBFD99AFB3863EF0467.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heterococcus	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Key to adult females of the  Heterococcus group </p>
            <p> 1. Multilocular pores absent.....................................................................  Stachycoccus</p>
            <p>- Multilocular pores present at least around vulva............................................................. 2</p>
            <p> 2. Circuli numbering 4; tubular ducts small, short and stout............................................  Annulococcus</p>
            <p>- Circuli numbering 0-3; tubular ducts elongate............................................................... 3</p>
            <p>3. Trilocular pores totally absent from body................................................................... 4</p>
            <p>- Trilocular pores present at least around spiracle opening, in each cerarius and on ostioles............................. 6</p>
            <p> 4. Tarsal claw without a denticle; anterior ostioles absent............................................  Pseudorhodania</p>
            <p>- Tarsal claw with a denticle; anterior ostiole present........................................................... 5</p>
            <p> 5. Anal lobe cerarii each with more than 2 enlarged setae; quinquelocular pores only in cerarii and on ostioles.....  Boreococcus</p>
            <p> - Anal lobe cerarii each with only 2 slender cerarian setae; quinquelocular pores scattered all over body........  Heterococcus</p>
            <p>6. Ostioles completely absent.............................................................................. 7</p>
            <p>- At least posterior ostiole present.......................................................................... 8</p>
            <p>7. Circuli oval, cerari absent..................................................................... Laingicoccus</p>
            <p> - Circuli absent; cerarii present at least last two abdominal segments...................................  Paramococcus</p>
            <p> 8. Circuli present, numbering 3; legs stout........................................................  Heterococcopsis</p>
            <p>- Circuli numbering 0 or 1; legs slender..................................................................... 9</p>
            <p> 9. Tarsal claw without a denticle; anterior ostiole absent; antennae 6 or 7 segmented...........................  Brevennia</p>
            <p> - Tarsal claw with a denticle; anterior ostiole present; antennae 8 or 9 segmented...............  Heterobrevennia gen. nov.</p>
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	https://treatment.plazi.org/id/2B77423D341BFFBFD99AFB3863EF0467	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kaydan, Bora	Kaydan, Bora (2011): Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey. Zootaxa 2970: 49-62, DOI: 10.5281/zenodo.203960
2B77423D341BFFBCD99AF8996590006F.text	2B77423D341BFFBCD99AF8996590006F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heterococcopsis Borchsenius	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Heterococcopsis Borchsenius</p>
            <p> Type species:  Heterococcopsis lonicerae Borchsenius, 1948</p>
            <p> Heterococcopsis lonicerae Borchsenius 1948: 955 , by original designation. </p>
            <p>Generic diagnosis. Adult female. Body elongate to oval, 1.8–2.0 mm long, 0.82–1.28 mm wide. Antennae 9 segmented. Eyes present on margin. Labium 3 segmented. Posterior pair of spiracles always larger than anterior spiracles. Legs well developed, with translucent pores on hind tibia; tibia broad, tarsus shorter than tibia, claw with a denticle; tarsal digitules setose, claw digitules capitate, thicker than tarsal digitules. Posterior ostioles well developed, anterior ostioles very weakly developed or not detected. Heptalocular pores smaller than other multilocular pores, scattered on venter; absent from dorsum. Oral-collar tubular ducts elongate, generally situated on body margin. Anal ring with 6 setae.</p>
            <p>Dorsum. Cerarii numbering 2 pairs, situated on posterior 2 abdominal segments. Dorsal body setae spine-like. Multilocular pores present on abdominal segments and near body margin. Quinquelocular pores scattered over body. Trilocular pores present on last abdominal segment (VIII) in cerarii and around ostioles. Oral-collar tubular ducts few, generally present on body margin.</p>
            <p>Venter. Body setae slender, generally short, longest setae present medially on head; apical setae long. Three circuli present. Multilocular disc pores present on posterior abdominal segments (IV–VIII+IX) plus a few on thorax and head. Heptalocular pores and quinquelocular pores about same size, scattered over venter. Minute discoidal pores scattered on body surface. Oral-collar tubular ducts few.</p>
            <p> Comment. Hitherto,  Heterococcopsis included three species:  H. desertus Bazarow &amp; Nurmamatov ,  H. lonicera Borchsenius and  H. opertus . Herein,  H. desertus is transferred to  Phenacoccus , and  H. opertus is transferred to  Heterobrevennia gen. nov. , leaving  Heterococcopsis as a monotypic genus. </p>
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	https://treatment.plazi.org/id/2B77423D341BFFBCD99AF8996590006F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kaydan, Bora	Kaydan, Bora (2011): Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey. Zootaxa 2970: 49-62, DOI: 10.5281/zenodo.203960
2B77423D3418FFBAD99AFC9065AE0245.text	2B77423D3418FFBAD99AFC9065AE0245.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heterococcopsis lonicerae Borchsenius	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Heterococcopsis lonicerae Borchsenius</p>
            <p> Material examined: Lectotype adult female (here designated) + 2 paralectotype adult females (on 1 slide): Kazakhstan, Alma-Ata,  Lonicera tatarica (  Caprifoliaceae ), Saygurov, 02.vi.36 (ZIAS: 80/37). Lectotype specimen in circle on left side, other 2 specimens indicated as paralectotypes. </p>
            <p>Adult female (Fig. 1)</p>
            <p>Live appearance: Body oval.</p>
            <p>Mounted material. Body elongate oval, 1.80–2.00 mm long, 0.82–1.28 mm wide. Eyes located on margin, each 30 µm wide. Antenna 9 segmented, 270–235 µm long; apical segment 35–45 µm long, 17.5–20.0 µm wide, with apical setae 27.5–30.0 µm long and with 3 fleshy setae, each 27.5–30.0 µm long. Clypeolabral shield 150–165 µm long, 110 µm wide. Labium 80–90 µm long, 62.5–70.0 µm wide. Anterior spiracles 37.5–45.0 µm long, 20–25 µm wide across atrium; posterior spiracles 50 µm long, 22.5–25.0 µm wide across atrium. Circuli numbering 3, each oval in shape, each 35 µm, 32.5 µm and 20 µm wide respectively. Legs well developed; coxa 70–80 µm long, hind trochanter + femur 130–145 µm long, hind tibia + tarsus 155–160 µm long, hind claw 20.0–22.5 µm long. Ratio of lengths of hind tibia + tarsus to hind trochanter + femur 1.10–1.19:1, ratio of lengths of hind tibia to tarsus 1.71–1.83:1, ratio of length of hind trochanter + femur to greatest width of femur 3.71–4.14:1. Tarsal digitules setose, each 15–20 µm long. Claw digitules simalar and capitate, each 17.5–20.0 μm long, knobbed and thicker than tarsal digitules. Translucent pores present on tibia of hind legs, totalling 32–43. Posterior pairs of ostioles present, each with 4 trilocular pores and 3 setae; anterior ostiole, not easy to detect or poorly developed. Anal ring 55 µm wide, with 6 anal ring setae, each seta 50–60 µm long.</p>
            <p>Dorsum. Cerarii numbering 2 pairs, situated on posterior 2 abdominal segments: anal lobe cerarii each with 2 conical setae (each 14–16 µm long) plus 5 trilocular pores and C17 each with 2 conical setae (each 15 µm long) plus 2 trilocular pores. Dorsal body setae spine-like, each 5.0–7.5 µm long. Multilocular pores each 6.25–7.5 µm wide with more than 10 loculi, present across posterior 2 abdominal segments and along body margin. Quinquelocular pores each 5 µm in diameter, scattered over dorsum but absent from last abdominal segment. Trilocular pores each 4–5 µm in diameter, restricted to last abdominal segment, cerarii and posterior ostioles. Oral-collar tubular ducts, each 9–11 µm long, 2–3 µm wide, present in a transverse row across last abdominal IV-VIII segments and along body margin.</p>
            <p>Venter. Setae slender, each 20–55 µm long, longest setae present medially on head; apical setae of anal lobe 75–105 µm long. Multilocular disc pores each 7.5–8.0 µm in diameter, present across abdominal segments IV −VIII + IX as follows: 0–6 pores on posterior margins of segment IV, 1–12 pores on posterior margins of V, 18–22 pores on posterior margins of VI, 30–32 across full width of VII, 19–34 across full width of VIII + IX. Quinquelocular and heptalocular pores each 4–5 µm wide, scattered over venter. Oral-collar tubular ducts each 8–11 µm long, 2.5– 3.0 µm wide, present in a row on last abdominal segment and on margins and submargins of each abdominal segment; ducts distributed as follows: 2–4 on segment I; 2–4 on II; 1–4 on III; 3–4 on IV; 4 on V; 4 on VI; 4–6 on VII; and 4–6 on VIII and a few on the metathorax and the mesothorax.</p>
            <p> Comment: For discussion, see under  Heterobrevinia gen. nov. below. </p>
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	https://treatment.plazi.org/id/2B77423D3418FFBAD99AFC9065AE0245	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kaydan, Bora	Kaydan, Bora (2011): Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey. Zootaxa 2970: 49-62, DOI: 10.5281/zenodo.203960
2B77423D341EFFBBD99AFE636730016D.text	2B77423D341EFFBBD99AFE636730016D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heterobrevennia Kaydan	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Heterobrevennia Kaydan ,  gen. nov.</p>
            <p> Type species:  Heterobrevennia opertus (Borchsenius, 1949 ) </p>
            <p>Generic diagnosis. Adult female: Body elongate, 2.82–3.20 mm long, 1.56–2.20 mm wide. Antennae generally 8 or 9 segmented (but occasionally 7 when 2nd and 3rd segments fused). Eyes present on margin. Labium 3 segmented. Posterior spiracles slightly larger than anterior spiracles. Legs well developed, with translucent pores on hind femur and tibia; claw with a denticle; tarsal digitules setose, claw digitules capitate, broader than tarsal digitules. Both anterior and posterior ostioles well developed. Multilocular disc pores limited to margins of both body surfaces and to posterior abdominal segments. Quinquelocular pores smaller than multilocular pores, scattered over both body surfaces. Trilocular pores present on dorsum. Oral-collar tubular ducts elongate, generally situated on body margin. Anal ring with 6 setae.</p>
            <p>Dorsum. Cerarii numbering always 5 or more pairs, situated on posterior abdominal segments and on head. Dorsal body setae spine-like. Multilocular pores situated on body margin. Quinquelocular pores scattered throughout. Trilocular pores scattered throughout, sometimes few, but always present at least on cerarii, ostioles and near spiracle opening. Minute discoidal pores scattered throughout. Oral-collar tubular ducts elongate scattered throughout.</p>
            <p>Venter. Body setae slender, longest setae present medially on head; apical setae long; spine-like setae present submarginally on body. Multilocular disc pores present on posterior abdominal segments and on margin of thorax and head. Quinquelocular pores scattered over body. Trilocular pores present around atrium of spiracle, absent elsewhere. Minute discoidal pores scattered over body. Oral-collar tubular ducts in rows on posterior abdominal segment and scattered on thorax and head.</p>
            <p> Comment.  Heterobrevennia gen. nov. is most similar to  Heterococcopsis in having trilocular pores on the dorsum and 8- or 9-segmented antennae. The two genera differ as follows (features of  Heterococcopsis in parentheses): i) no circuli (with 3 circuli), ii) trilocular pores scattered on dorsum (trilocular pores on dorsum restricted to cerarii, ostioles and posterior abdominal segments), iii) at least five cerarii, of which 3 are present on head (only 2 cerarii, restricted to posterior abdominal segments), and iv) legs slender (legs stout). </p>
            <p> Heterobrevennia is also similar to all the other genera in the  Heterococcus -group. The affinities of  Heterobrevennia with other related genera are discussed below. </p>
            <p> Asphodelococcus can be readily separated from  Heterobrevennia by the following combination of features (features of  Heterobrevennia in parenthesis): (i) antennae 6 segmented (antennae 9 segmented), (ii) multilocular pores of unusual shape and often quadrate, and with heavily sclerotised hexagonal pores (pores normal), and (iii) cerarii numbering 4 pairs and restricted to abdomen (cerarii numbering 5 or more pairs, present on both abdomen and head). </p>
            <p> Annulococcus differs from  Heterobrevennia in the following combination of features (features of  Heterobrevennia in parenthesis): (i) cerarii poorly developed, present along the body margin (cerarii numbering 5 or more pairs, present on both abdomen and head), (ii) quinquelocular pores larger than other multilocular pores (quinquelocular pores smaller than other multilocular pores), (iii) oral-collar tubular ducts very short and stout (oral-collar tubular ducts elongate), and (iv) trilocular pores absent (trilocular pores present) (Miller, 1975; Foldi and Cox, 1989). </p>
            <p> Brevennia Goux shares many morphological features with  Heterobrevennia , namely, the presence of trilocular pores on the dorsum and venter and the abundance of quinquelocular pores on both dorsum and venter, but the two genera differ in the absence of claw denticles and  Brevennia also lacks anterior ostioles. </p>
            <p> Boreococcus Danzig ,  Heterococcus Borchsenius and  Pseudorhodania can be readily distinguished from  Heterobrevennia due to the complete absence of trilocular pores in these three genera. </p>
            <p> Laingococcus Williams differs from  Heterobrevennia in having the following combination of features (features of  Heterobrevennia in parenthesis): (i) ostioles absent (ostioles present), (ii) cerarii absent (5 or more cerarii present on both abdomen and head); (iii) a circulus present (absent); (iv) dorsal body setae similar in size to those on venter (dorsal body setae spinose, ventral setose); and (v) in having a rotund body (body elongate oval). </p>
            <p> In having trilocular pores on the dorsum and venter,  Heterobrevennia is somewhat similar to Paramacoccus Foldi &amp; Cox but Paramacoccus is unique in lacking ostioles. </p>
            <p> On the other hand, in having trilocular pores on the dorsum and venter, and spine-like setae on dorsum,  Heterobrevennia is somewhat similar to  Phenacoccus Cockerell but new genus differs in having many quinquelocular pores on the dorsum. </p>
            <p> Etymology. The name  Heterobrevennia refers to its similarity to the genera  Heterococcus and  Brevennia . Gender female. </p>
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	https://treatment.plazi.org/id/2B77423D341EFFBBD99AFE636730016D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kaydan, Bora	Kaydan, Bora (2011): Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey. Zootaxa 2970: 49-62, DOI: 10.5281/zenodo.203960
2B77423D341FFFBBD99AFD9C63EF000D.text	2B77423D341FFFBBD99AFD9C63EF000D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heterobrevennia	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Key to  Heterobrevennia species </p>
            <p>1. Multilocular pores scattered throughout dorsum; coxa without transculent pores.................................... 2</p>
            <p> - Multilocular pores present dorsally only on abdominal segments VI and VII: coxa with transculent pores...............................................................................................  H. kozari Kaydan spec. nov.</p>
            <p> 2. Cerarii totalling 6 pairs; with two groups of multilocular pores on dorsum of head; many trilocular pores present on dorsum.............................................................................  H. gullanae Kaydan spec. nov.</p>
            <p> - Cerarii totalling 7 pairs; without multilocular pores on dorsum of head; trilocular pores few on dorsum, restricted to thorax, on posterior abdominal segments, in cerarii and on ostioles.....................................  H. opertus Borchsenius</p>
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	https://treatment.plazi.org/id/2B77423D341FFFBBD99AFD9C63EF000D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kaydan, Bora	Kaydan, Bora (2011): Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey. Zootaxa 2970: 49-62, DOI: 10.5281/zenodo.203960
2B77423D341FFFB9D99AFC30644307BE.text	2B77423D341FFFB9D99AFC30644307BE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heterobrevennia gullanae Kaydan	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Heterobrevennia gullanae Kaydan sp. nov.</p>
            <p> Material examined: Holotype adult female, Turkey, VAN-Gevaş, N: 38°16’663’’, E: 043°03’898’’, 1834 m altitude, on the leaf sheaths of  Poaceae, M.B. Kaydan , 09.vi.2009 (CCVT: 4530). Deposited in CCVT. </p>
            <p>Paratype adult females, same data as holotype, 5 specimens on 5 slides (4 Slides in CCVT: 4530 plus 1 in ZIAS).</p>
            <p>Adult female (Fig. 2)</p>
            <p>Live appearance: Body oval. In the leaf sheaths of grasses.</p>
            <p>Mounted material. Body elongate oval, 2.80–3.18 mm long, 1.12–1.25 mm wide. Eye marginal, 30–35 µm wide. Antenna 8 segmented (rarely 7 segmented, when 2nd - and 3rd -antennal segments fused), 255–270 µm long; apical segment 40–45 µm long, 20.0–22.5 µm wide, with apical seta 30 µm long, plus 3 fleshy setae, each 15.0– 27.5 µm long. Clypeolabral shield 130–140 µm long, 102.5–110.0 µm wide. Labium 67.5–70.0 µm long, 60.0– 67.5 µm wide. Anterior spiracles 40.0–47.5 µm long, 20–25 µm wide across atrium; posterior spiracles 45–55 µm long, 30–35 µm wide across atrium. Legs well developed; coxa 80–95 µm long, hind trochanter + femur 175–195 µm long, hind tibia + tarsus 210–230 µm long, hind claw 20.0–22.5 µm long. Ratio of lengths of hind tibia + tarsus to hind trochanter + femur 1.15–1.24:1, ratio of lengths of hind tibia to tarsus 1.91–2.21:1, ratio of length of hind trochanter + femur to greatest width of femur 5.0–5.3:1. Tarsal digitules setose, each 20–25 µm long. Claw digitules subequal, each 22.5–25.0 µm long, knobbed and thicker than tarsal digitules. Translucent pores present on femur and tibia of hind legs, totalling 12–26. Both pairs of ostioles present, posterior ostioles each with 7–10 trilocular pores plus 0–3 setae, anterior ostioles with 4–6 trilocular pores and 1 seta. Anal ring 55–60 µm wide, with 6 anal-ring setae, each 55–65 µm long.</p>
            <p>Dorsum. Cerarii numbering 6 pairs, with pairs on posterior 3 abdominal segments plus 3 anteriorly on head; anal lobe cerarii, each with 2 conical setae, each 15.0–20 µm long, plus 4 or 5 trilocular pores; C17 each with 2 conical setae, each 15.0–17.5 µm long, plus 2 trilocular pores; C16 each with 2 conical setae, each 10.0–12.5 µm long, plus 1 trilocular pore; C1 each with 2 conical setae, each 7.5–10 µm long, plus 2 trilocular pores; C2 each with 2 conical setae, each 7.5–10 µm long, plus 2 trilocular pores; C3 each with 3 conical setae, each 7.5–10 µm long, plus 2 trilocular pores. Dorsal body setae spine-like, each 7.5–15.0 µm long, becoming shorter posteriorly. Multilocular pores each 6.0–7.5 µm wide with more than 10 loculi, in rows on posterior 3 abdominal segments, and on margin of remaining abdominal segments and thorax; also with 2 pairs groups on head: 1 pair each with 12–19 pores close to antennae, and other pairs each with 11–15 pores nearer anterior ostioles. Quinquelocular pores, each 5–6 µm wide, scattered throughout. Trilocular pores, each 5–6 µm in diameter, scattered throughout. Minute discodial pores few, each 2 µm in diameter, scattered on body surface. Oral-collar tubular ducts, each 9–11 µm long, 2–3 µm wide, scattered on all abdominal segments and along body margin on thorax.</p>
            <p>Venter. Body setae slender, each 10–45 µm long, longest setae medially on head; apical anal lobe setae 120.0– 137.5 µm long. Multilocular disc pores each 7.5–8.0 µm in diameter, present in rows or bands on posterior abdominal segments (IV – VIII + IX): 6–8 pores on segment IV, 26–37 pores on segment V, 48–69 pores on segment VI, 58–62 on segment VII, 34–42 on segments VIII + IX. Quinquelocular pores each 4–5 µm wide, scattered throughout. Trilocular pores each 5–6 µm in diameter, restricted to around spiracle atrium. Minute discodial pores, each 2 µm in diameter, scattered throughout. Oral-collar tubular ducts each 9–11 µm long, 2.5–3.0 µm wide, present in rows or bands on posterior abdominal segments plus a few near margins on head and thorax; distributed as follows: 2–4 on each side of abdominal segments I–III; 8–12 on IV; 31–38 on V; 40–53 on VI; 34–42 on VII, and 4–10 on VIII.</p>
            <p> Comment. The adult female of  Heterobrevennia gullanae Kaydan is unique for a species in the Heterococcusgroup in having the following combination of features: (i) many quinquelocular pores on both the dorsum and venter, (ii) short hair-like tarsal digitules, (iii) claw digitules capitate, thicker than tarsal digitules, (iv) many multilocular pores on venter, (v) translucent pores present on both femur and tibia, (vi) two groups of multilocular pores on dorsum of head, (viii) six pairs of cerarii, 3 on head and 3 posteriorly on abdomen, (ix) many oral-collar tubular ducts on the venter and dorsum, and (x) the presence of trilocular pores throughout the dorsum. </p>
            <p> Heterobrevennia gullanae resembles  H. opertus in having most of the above features but  H. gullanae differs in having the following combination of characters (characters of  H. opertus in brackets): (i) two groups of multilocular pores on the head (absent), (ii) cerarii numbering 6 pairs (7), (iii) many trilocular pores present throughout dorsum (trilocular pores on dorsum few, generally restricted to posterior abdominal segment, and in cerarii and ostioles), (iv) antennae 8, rarely 7 segmented (9 segmented).  Heterobrevennia gullanae is also similar to  H. kozari Kaydan spec. nov. , described below, but differs in having (character states for  H. kozari in brackets): (i) 6 pairs of cerarii (5), (ii) translucent pores on coxa absent (present), and (iii) one size of oral-collar tubular ducts on dorsum (two sizes medially on posterior abdominal segment). </p>
            <p>Etymology. The new species is named in honor of Prof. Dr. Penny Gullan (Division of Evolution, Ecology &amp; Genetics Research School of Biology, The Australian National University, Canberra, Australia), for her valuable studies on all aspects of coccidology.</p>
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	https://treatment.plazi.org/id/2B77423D341FFFB9D99AFC30644307BE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kaydan, Bora	Kaydan, Bora (2011): Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey. Zootaxa 2970: 49-62, DOI: 10.5281/zenodo.203960
2B77423D341DFFB7D99AFAFA6449056A.text	2B77423D341DFFB7D99AFAFA6449056A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heterobrevennia kozari Kaydan	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Heterobrevennia kozari Kaydan spec. nov.</p>
            <p> Material examined. Holotype adult female, Turkey, Bitlis-Siirt road, N: 38°11’772’’, E: 041°49’067’’, 748 m altitude, on the leaf sheaths of  Cynodon dactylon (Poaceae) , M.B. Kaydan, 26.v.2008 (CCVT: 4325). Deposited in CCVT. </p>
            <p> Paratype adult females: Turkey, Diyarbakır-Silvan road, N: 38°07’722’’, E: 041°10’530’’, 748 m altitude, on the leaf sheaths of  Cynodon dactylon, M.B. Kaydan and Ferenc Kozár , 26.v.2008. Six parasitized specimens deposited as follows: 5 in CCVT: 4327 and 1 in ZIAS. </p>
            <p>Adult female (Fig. 3)</p>
            <p>Description made solely from the holotype. Measurements from other specimens not used because they were parasitized.</p>
            <p> Live appearance. Body oval. In the leaf sheaths of  C. dactylon . </p>
            <p>Mounted material. Body elongate oval, 3.28 mm long, 1.28 mm wide. Eye marginal, 35–40 µm wide. Antenna 9 segmented, 380–390 µm long; apical segment 50–55 µm long, 25–30 µm wide with apical setae 30 µm long, plus 3 fleshy setae, each 25–30 µm long. Clypeolabral shield 135 µm long, 152.5 µm wide. Labium 100 µm long, 75 µm wide. Anterior spiracles 55–60 µm long, 25.0–27.5 µm wide across atrium; posterior spiracles 60–65 µm long, 35 µm wide across atrium. Legs well developed; coxa 140–145 µm long; hind trochanter + femur 230– 235 µm long; hind tibia + tarsus 285–290 µm long, and hind claw 27.5–30.0 µm long. Ratio of lengths of hind tibia + tarsus to hind trochanter + femur 1.21–1.26:1, ratio of lengths of hind tibia to tarsus 2.05–2.21:1, ratio of length of hind trochanter + femur to greatest width of femur 5.17–5.27:1. Tarsal digitules setose, each 27.5–30.0 µm long. Claw digitules subequal, each 27.5–30.0 µm long, knobbed and thicker then tarsal digitules. Translucent pores present on coxa, femur and tibia of hind legs, totalling 58–79. Both pairs of ostioles present, posterior ostioles with 8–12 trilocular pores plus 1 seta, anterior ostioles 6 trilocular pores and 1 setae. Anal ring 70 µm wide, with 6 analring setae, each 120–125 µm long.</p>
            <p>Dorsum. Cerarii numbering 5 pairs,with pairs on posterior 2 abdominal segments, plus 3 pairs on head: anal lobe cerarii each with 2 conical setae, each 17.5–20 µm long, plus 6 or 7 trilocular pores; C17 each with 2 conical setae, each 15 µm long, plus 3–4 trilocular pores; C1 each with 2 conical setae, each 7.5–10 µm long, plus 1 trilocular pore; C2 each with 2 conical setae, each 7.5–10 µm long, plus 1 or 2 trilocular pores; C3 each with 3 conical setae, each 7.5 µm long, with 2 trilocular pores. Dorsal setae spine-like, each 7–11 µm long. Multilocular pores each 6.0–7.5 µm with more than 10 loculi, restricted to a few on abdominal segments VI and VII. Quinquelocular pores each 4–5 µm wide, scattered throughout. Trilocular pores each 4–5 µm in diameter, scattered throughout. Minute discoidal pores few, each 2 µm in diameter, scattered on body surface. Oral-collar tubular ducts of 2 sizes, larger each 7–11 µm long, 4–5 µm wide, present in rows on each abdominal segment as follows: 14 on I; 8 on II; 19 on III; 16 on IV; 25 on V; 28 on VI; 20 on VII; 14 on VIII; also scattered on thorax plus a few on head; smaller ducts few, each 7–11 µm long, 2.5–3 µm wide, present medially on posterior 3 abdominal segments,.</p>
            <p>Venter. Body setae slender, each 10–85 µm long, longest setae medially on head; apical anal lobe setae 210– 220 µm long. Multilocular disc pores each 7.5–9.0 µm in diameter, present in rows or bands on posterior abdominal segments (IV–VIII+IX): 4–6 pores on segment IV, 15 on segment V, 37 on segment VI, 38 on segment VII, 36 on segments VIII + IX. Quinquelocular pores each 4–5 µm wide, scattered throughout. Trilocular pores restricted to around spiracle atrium, each 2.5–3.0 µm in diameter. Minute discoidal pores each 2 µm in diameter, scattered throughout. Oral-collar tubular ducts each 7.5–10 µm long, 2.5–3.0 µm wide, in rows across most abdominal segments plus a few on thorax; distributed as follows: 2 on abdominal segment I; 4 on II; 11 on III; 12 on IV; 28 on V; 26 on VI; 28 on VII, and 15 on VIII.</p>
            <p> Comment. The adult female of  Heterobrevennia kozari Kaydan is unique for a species in the  Heterococcus group in having the following combination of character states: (i) many quinquelocular pores on both dorsum and venter, (ii) hair-like tarsal digitules, (iii) ventral multilocular pores present on posterior abdominal segments, (iv) translucent pores present on coxa, femur and tibia, (v) five pairs of cerarii, and (vi) many oral-collar tubular ducts throughout the dorsum. The most unique characters are the presence of 5 cerarii and pores on hind coxa. </p>
            <p> In having most of the above mentioned features,  H. kozari resembles  H. opertus but differs in having (characters of  H. opertus in brackets): (i) only 5 pairs of cerarii (7), (ii) many trilocular pores scattered throughout dorsum (trilocular pores on dorsum few, generally restricted to the posterior abdominal segment, cerarii and ostioles), (iii) transcluent pores present on hind coxa (absent), and (iv) dorsal multilocular pores present only on abdominal segments VI and VII (scattered all over body).  H. kozari is also similar to  H. gullanae Kaydan , described above, but differs in having (characters of  H. gullanae in brackets): (i) 5 pairs of cerarii (6), (ii) transcluent pores on coxa (absent), (iii) one size of oral-collar tubular ducts on dorsum (two sizes on posterior abdominal segment), (iv) dorsal multilocular pores restricted to abdominal segments VI and VII (also present in two group on head), and (v) antennae 9 segmented (8, rarely 7). </p>
            <p>Etymology. The new species is named in honor of Dr. Ferenc Kozár (Department of Zoology, Plant Protection Institute, Hungarian Academy of Sciences, Budapest, Hungary), who has made valuable studies on many aspects of coccidology throughout the World.</p>
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	https://treatment.plazi.org/id/2B77423D341DFFB7D99AFAFA6449056A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kaydan, Bora	Kaydan, Bora (2011): Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey. Zootaxa 2970: 49-62, DOI: 10.5281/zenodo.203960
2B77423D3413FFB5D99AF99366950738.text	2B77423D3413FFB5D99AF99366950738.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heterobrevennia opertus (Borchsenius) Borchsenius	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Heterobrevennia opertus (Borchsenius) comb.nov.</p>
            <p> Material examined: Syntypic adult females: Tadzikistan, Gissarskii Ridge,  Dactylis glomerata (Poaceae) , H. Borchsenius, 17.vii.1944 (2 specimens, ZIAS: 34-45). </p>
            <p>Adult female (Fig. 4)</p>
            <p>Live appearance: Body elongate oval.</p>
            <p>Mounted material. Body elongate oval, 2.82–3.60 mm long, 1.56–2.20 mm wide. Eye marginal, 32.5–35.0 µm wide. Antenna 9 segmented, 270–290 µm long; apical segment 45–55 µm long, 20–25 µm wide, with apical setae 30–35 µm long, plus 3 fleshy setae, each 30–35 µm long. Clypeolabral shield 145–150 µm long, 110–115 µm wide. Labium 80–85 µm long, 65–70 µm wide. Anterior spiracles 42.5–52.5 µm long, 25.0–27.5 µm wide across atrium; posterior spiracles 50–60 µm long, 25–35 µm wide across atrium. Legs well developed; coxa 90–105 µm long, hind trochanter + femur 215–220 µm long, hind tibia + tarsus 255–260 µm long, hind claw 25.0–27.5 µm long. Ratio of lengths of hind tibia + tarsus to hind trochanter + femur 1.16–1.18:1, ratio of lengths of hind tibia to tarsus 1.8–2.03:1, ratio of length of hind trochanter + femur to greatest width of femur 4.4–4.8:1. Tarsal digitules setose, each 32.5–35.0 µm long. Claw digitules subequal, each 27.5–35.0 µm long, knobbed and broader than tarsal digitules. Translucent pores present on femur and tibia of hind legs, totalling 16–24. Both pairs of ostioles present, posterior ostioles each with 5–7 trilocular pores and 2 or 3 setae, anterior ostioles each with 5 trilocular pores and 2 or 3 setae. Anal ring 60–62.5 µm wide, with 6 anal-ring setae, each 85–95 µm long.</p>
            <p>Dorsum. Cerarii numbering 7 pairs, with pairs on posterior 4 abdominal segments, and on 3 pairs on head; anal lobe cerarii each with 2 conical setae, each 15.0–17.5 µm long, plus 5–7 trilocular pores; C17 each with 2 conical setae, each 15.0–17.5 µm long, plus 4 trilocular pores; C16 each with 2 conical setae, each 15 µm long, plus 2 trilocular pores; C15 each with 2 conical setae, each 11.0–12.5 µm long, plus 2 trilocular pores; C1 each with 2 conical setae, 10 µm long, plus 2 trilocular pores; C2 each with 2 conical setae, each 10 µm long, plus 3 trilocular pores; and C3 each with 3 conical setae, each 7.5–10 µm long, plus 2 trilocular pores. Dorsal body setae spine-like, each 10.0–12.5 µm long. Multilocular pores each 7.5–8.0 µm wide with 11 loculi, scattered throughout. Quinquelocular pores each 5–6 µm, scattered throughout. Trilocular pores each 5–6 µm in diameter, restricted to posterior abdominal segments VII, VIII and IX, and in cerarii, on ostioles and the middle area of thorax. Oral-collar tubular ducts present along body margin, each 9–11 µm long, 2–3 µm wide.</p>
            <p>Venter. Body setae slender, each 15–65 µm long, longest setae medially on head; apical setae of anal lobe 145–165 µm long. Multilocular disc pores each 7.5–8.0 µm in diameter, present in rows or bands on posterior abdominal segments (IV–VIII + IX): 12 pores on segment IV, 12–24 on V, 32–36 on VI, 53–62 on VII, and 27–33 on VIII + IX. Quinquelocular pores each 5–6 µm wide, scattered throughout. Oral-collar tubular ducts each 7.5– 10.0 µm long, 2.5–3.0 µm wide, present in rows on all abdominal segments plus a few on head and thorax; distributed as follows: 2–4 on each side of abdominal segments I–III and a few medially; 4 on IV; 15–25 on V; 14–24 on VI; 19 on VII; 18–23 on VIII in row or band.</p>
            <p> Comment. For a comparison of  H. opertus with other species in the genus, see under  H. gullae and  H. kozari above. </p>
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	https://treatment.plazi.org/id/2B77423D3413FFB5D99AF99366950738	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kaydan, Bora	Kaydan, Bora (2011): Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey. Zootaxa 2970: 49-62, DOI: 10.5281/zenodo.203960
2B77423D3411FFB3D99AFB7C671300C6.text	2B77423D3411FFB3D99AFB7C671300C6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phenacoccus desertus (Bazarov & Nurmamatov) Bazarov & Nurmamatov	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Phenacoccus desertus (Bazarov &amp; Nurmamatov) ,  comb. nov.</p>
            <p> Heterococcopsis desertus Bazarov &amp; Nurmamatov, 1975: 63 . </p>
            <p> Material examined. Holotype adult female: Tadzhikistan, Gbao, Vosim, Gasur Lurgap, Org.  Artemisia demissa (Asteraceae) , L. Kuarzivsheva, 18.vii.1924 (ZIAS). </p>
            <p>Adult female (Fig. 5)</p>
            <p>Live appearance. Body oval.</p>
            <p>Mounted material. Body oval, 2.20 mm long, 1.34 mm wide. Eye marginal, 35.0 µm wide. Antenna 9 segmented, 130–140 µm long; apical segment 47.5 µm long, 20 µm wide, with apical setae 22.5 µm long, plus 3 fleshy setae, each 20–30 µm long. Clypeolabral shield 145 µm long, 120 µm wide. Labium 110 µm long, 75 µm wide. Anterior spiracles 45 µm long, 15 µm wide across atrium; posterior spiracles 45.0–47.5 µm long, 25.0–27.5 µm wide across atrium. Legs well developed; coxa 40 µm long, hind trochanter + femur 150 µm long, hind tibia + tarsus 165 µm long, hind claw 25.0 µm long. Ratio of lengths of hind tibia + tarsus to hind trochanter + femur 1.10, ratio of lengths of hind tibia to tarsus 1.3:1, ratio of length of hind trochanter + femur to greatest width of femur 3.3:1. Tarsal digitules setose, each 10 µm long. Claw digitules subequal, each 17.5 µm long, knobbed and broader than tarsal digitules. Translucent pores present on femur and tibia of hind legs, totalling 16–24. Both pairs of ostioles present, posterior ostioles each with 9 trilocular pores and 1 setae, anterior ostioles each with 4 trilocular pores and 2 or 3 setae. Anal ring 55 µm wide, with 6 anal-ring setae, each 60–65 µm long.</p>
            <p>Dorsum. Cerarii numbering 2 pairs, with pairs on posterior 2 abdominal segments; anal lobe cerarii each with 2 conical setae (broken, as shown on figure), plus 5–7 trilocular pores; C17 each with 2 conical setae, each 10 µm long, plus 2 trilocular pores. Dorsal body setae spine-like, each 5–10 µm long. Multilocular pores each 7.5–9.0 µm wide with 11 loculi, present in laterally in abdominal segment I and in rows or bands on posterior abdominal segments (II–VII): 12 pores on segment on II, 19 on III, 37 on IV, 37 on V, 24 on VI, 30 on VII and no pores on segment VIII. Quinquelocular pores very few (only three pores seen on head and thorax), each 5–6 µm. Trilocular pores each 4–5 µm in diameter, scattered throughout. Oral-collar tubular ducts each 7.5–10.0 µm long, 2–3 µm wide, present in rows on all abdominal segments plus 8 on head and thorax laterally; distributed on abdomen as follows: 3 on I; 5 on II; 6 on III; 6 on IV; 7 on V; 2 on VI; 3 on VII; 4 on VIII in row.</p>
            <p>Venter. Body setae slender, each 12.5–40.0 µm long, longest setae medially on head; apical setae of anal lobe broken. Multilocular disc pores present, each 7.5–8.0 µm in diameter, present laterally on abdominal segments I–II and in rows or bands on posterior abdominal segments (III–VIII + IX): 22 pores on segment III, 34 on IV, 36 on V, 48 on VI, 37 on VII, and 27 on VIII + IX. Quinquelocular pores each 5–6 µm wide, scattered throughout. Oral-collar tubular ducts each 7.5–9.0 µm long, 2.5–3.0 µm wide, present in rows on all abdominal segments plus a few on head and thorax; distributed as follows: 1–2 on each side of abdominal segments I–III; 13 on IV; 14 on V; 8 on VI in row or band and 3 on VII; 3 on VIII laterally.</p>
            <p> Comment. Species in the  Heterococcus -group differ from those in  Phenacoccus Cockerell in having: (i) quinquelocular pores on the dorsum and either no trilocular pores or trilocular pores restricted to around each spiracle. Because  Heterococcopsis desertus lacks both of these character states, it is not congeneric with  Heterococcopsis lonicerae Borchsenius , the type species of the genus  Heterococcopsis but does appear to be congeneric with  Phenacoccus aceris Signoret , the type species of the genus  Phenacoccus .  H. desertus shares the following important features with  P. aceris : (i) presence of many trilocular pores on dorsum and venter, (ii) very few quinquelocular pores (only 3 were found) on dorsum, and (iii) a long labium. Based on the morphological affinities of  H. desertus with  P. aceris , I here transfer  H. desertus to  Phenacoccus , as  Phenacoccus desertus (Bazarov &amp; Nurmamatov) comb. nov. and illustrate the species based on the type material. </p>
            <p> This species is very close to  Phenacoccus tergrigorianae Borchsenius &amp; Ter-Grigorian , and P. p u m i l u s Kiritshenko in having multilocular pores on dorsum, but differs from both in having many quinquelocular pores on the venter. The genus  Phenacoccus has many species and needs further study to clarify the status of the new combination of  P. desertus . </p>
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	https://treatment.plazi.org/id/2B77423D3411FFB3D99AFB7C671300C6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kaydan, Bora	Kaydan, Bora (2011): Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey. Zootaxa 2970: 49-62, DOI: 10.5281/zenodo.203960
