identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
252A879DFF96A17AFF2FFF13FEC7C21A.text	252A879DFF96A17AFF2FFF13FEC7C21A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leusaba Walker	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Genus  Leusaba Walker</p>
            <p> Leusaba Walker, 1857: 144 ; Melichar 1914: 105. </p>
            <p> Type species:  Leusaba marginalis Walker, 1857: 144 , by original designation. </p>
            <p>Description. Relatively large and stout tropiduchids (Figs. 1–4), length (from apex of vertex to tip of anal tube) 8.0– 10.6 mm (N=7).</p>
            <p>Colour. General color green to ochraceous; frons with anterior and lateral margins and medial line suffused with orange to reddish stripes (black medial line may be present), apex of rostrum, eyes and antennae with pedicel suffused reddish to brownish; vertex with posterior marginal carinae, pronotum with median disc and lateral portion all suffused yellowish to reddish; fore wings hyaline, veins and stigma brown; fore tibia and mid tibia with apices, hind tibia with bases, lateral spines with apices, long stripes on hind femora all suffused with black.</p>
            <p>Head and thorax. Head (Figs. 1 A, 1C, 1E, 1G) including eyes slightly narrower than pronotum, vertex approximately twice as broad as long, projecting before eyes for less than half length of eye; apex broadly convex in dorsal view, evenly and broadly rounded to base of frons; vertex and frons not separated, lateral carinae continuous. Eyes oval. Face with frons laterally carinate, carinae attaining frontoclypeal suture, with or without median ridge; slightly longer medially than greatest width and distinctly longer than width at anterior margin; ventrally depressed each side of midline (Figs. 1 B, 2C) or depressed lateroventrally, forming a ‘v’-shaped central elevated area (Figs. 1 D, 3C); disc smooth or covered with microsetae; clypeus (Figs. 1 B, 1D, 1F, 1H) with distinct median carina, without lateral carinae; rostrum short, attaining mesotrochanters, apical segment short, longer than broad; ocelli distinct; antennae (Figs. 2 B, 2C, 4B, 4C) with scape short, ring-like; pedicel cylindrical, covered with many microsetae extending to base of pedicel. Vertex (Figs. 2 A, 4A) subquadrate, distinctly shorter than greatest width, slightly longer than pronotum at midline; lateral margins ridged and subparallel; posterior margin carinate and arched anteriorly; anterior part of vertex smoothly swollen, uniting with base of frons to form evenly rounded surface; posterior part of vertex distinctly depressed, lateral margins of depression incurving anteriorly and meeting medially to form nearly semicircular arc; with a very faint median carina.</p>
            <p>Pronotum (Figs. 2 A, 4A) distinctly shorter than mesonotum in midline; with two contiguous median carinae and oblique and separate lateral carinae; anterior margin slightly carinate, nearly straight; disc projected anteriorly, deeply depressed between median and lateral carinae, with impression on each side; posterior margin obtusely angled excavate. Mesonotum (Figs. 2 A, 4A) tricarinate, median carina straight, reaching to transverse suture; pronotum and mesonotum together medially 4.8–5.9 times as long as median length of vertex. Fore wings (Figs. 1 A, 1C, 1E, 1G) hyaline, surpassing extremity of abdomen by about 0.5– 0.7 times length of body, widening towards tips, 2.7–2.8 times as long as maximum breadth; corium without granulation; veins prominent, thickly covered with short setae dorsally and ventrally (veins in basal area of corium not covered with setae dorsally); costal cell without cross veins; Sc+R and M forking at nodal line; Cu forking after level of junction of claval veins, the latter forked at basal 2/5 of clavus; nodal line forming a zigzag transverse line, subapical line slightly oblique, with 6 subapical and 17–20 apical cells, subapical cells a little longer than latter. Hind wings (Figs. 2 E, 4E) hyaline, veins prominent and thickly covered with short setae dorsally and ventrally, posterior margin strongly sinuate, with three disconnected, oblique transverse veins before apical area, venation as in Figs. 2 E, 4E. Legs elongate; hind tibiae each with 3–4 distinct lateral spines, abnormally with 5 (often on one leg only, basal one very fine); spinal formula of hind leg 6–6–2.</p>
            <p>Male genitalia. Pygofer (Figs. 2 F, 2H, 2I) symmetrical, short, in lateral view forming a perpendicular irregular rectangle, wider dorsally than ventrally, with dorsal posterior margin angularly produced posteriorly; dorsal margin deeply excavated to accommodate anal tube (Fig. 2 H); ventral margin strongly excavated anteriorly to accommodate gonostyli (Fig. 2 I). Xth segment (Figs. 2 F, 2H) very long and slender, apical margin distinctly concave in dorsal view, lateroventral margin with two pairs of distinct long spines (Fig. 2 F); anal style short and small. Gonostyli (Figs. 2 F, 2G, 2I) bilaterally asymmetrical, membranously fused with pygofer at base, left gonostylus forming irregular ellipse, more than 3 times as long as maximum breadth in lateral view, tapering to apex in distal 1/6, dorsal edge with two processes (see Fig. 2 F), right gonostylus forming irregular rectangle, relatively large and broad, more than 3 times as long as maximum breadth in lateral view, apex relatively broad, basal dorsal edge with a process (see Fig. 2 G). Periandrium (Figs. 3 A, 3B) dorsally connected with ventrobasal margin of anal-tube, membranously fused with pygofer at ventral side, irregularly ring-like, distinctly sclerotized, surrounding aedeagus at basal part and visible in lateral view, apical margin with a branched process at ventral side.</p>
            <p>Aedeagus (Figs. 3 A, 3B) asymmetrical, elongate, robust and mostly sclerotized, with a long, sinuous process at basal part, basal 1/4 to 1/2 part of aedeagus expanded into ellipsoid, connected with membranous endosoma apically, with three projections at left side and a prominence at right side; shaft of aedeagus sclerotized, embraced by endosoma, apical part strongly curved and directed posteroventrally, phallotrema hole-like, exposed apically; endosoma with three projections.</p>
            <p>Female genitalia. Anal tube (Figs. 3 C, 3D, 4F, 4G) relatively short, apical margin convex in dorsal view, anal styles reaching beyond apical ventral margin of anal tube. Gonopophyses VIII (first valvulae) (Figs. 3 C, 3F, 4F, 4H) strongly sclerotized with 5 blunt teeth on dorsal margin, 3 blunt apical teeth on ventral margin, and several minute teeth in two angle-like rows laterobasad of middle. Gonopophyses VIII nearly symmetrical or asymmetrical at base in ventral view (Figs. 3 F, 4H). Gonopophyses IX (second valvulae) (Figs. 3 E, 4I) triangular, fused together on inner-lateral margin and strongly reduced but well sclerotized, apical ends confluent or not; gonospiculum (Figs. 3 E, 4I) shorter than median length of triangular part (0.75–0.95:1). Gonoplacs (third valvulae) (Figs. 3 C, 3D, 3F, 4F, 4G) with 3–4 teeth at dorsal margin, dentition simple, forming one line, continue without interruption along ventral margin and involving a further 13 teeth (see Figs. 4 F, 4G); or dentition complex, not forming one line, with 3 outer teeth at apical part, then continue with 12–13 teeth from apical to ventral margin (see Figs. 3 C, 3D, 3F). Gonocoxae VIII (Figs. 3 C, 4F) almost rectangular sclerotized plates with a distinct black spot on caudadorsal edge. Genital opening (gonoporus) singular (monotrysian) occurring between gonopophyses VIII and IX (Figs. 3 F, 4H). Sternite VII with median part sinuous or broad concave at ventral side. Posterior vagina strikingly sclerotized in its ventral part (Figs. 3 F, 4H). Dorsally, a large bursa copulatrix (Figs. 3 C, 4F) opens directly into vagina and an enlarged common oviduct and spermatheca open apically into anterior vagina (Fig. 4 H).</p>
            <p> Distribution. Species of  Leusaba are currently known from Malaysia (Borneo), Philippines and Sri Lanka. </p>
            <p> Discussion.  Leusaba belongs in the tribe Paricanini Melichar, which was redefined by Fennah (1982). In addition to  Leusaba , the tribe includes  Paricana Walker, 1857 ,  Paricanoides Liang, 2003 , and  Stacota Stål, 1859 . Species of  Leusaba are externally similar to those of  Paricana but can be distinguished from the latter by the laterally carinate vertex with a median carina and fore wings with six subapical and 18–20 apical cells, with subapical cells a little longer than the latter.  Leusaba can be distinguished from  Paricanoides by the frons and postclypeus with a median carina and more numerous apical and subapical cells in the forewing and more numerous apical cells in the hindwing (see Liang, 2003). It can be separated from  Stacota by the vertex being distinctly shorter than greatest width (slightly longer medially than greatest width in  Stacota ), and fore wings lacking transverse veinlets in the costal cell. </p>
            <p> In addition to  L. philippina the following other species belong in  Leusaba . Two undescribed females from Malaysia are considered to represent two additional species of  Leusaba but are not formally named because of the lack of male specimens. They indicate that the Malaysian fauna is more widespread and diverse than currently known. </p>
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	https://treatment.plazi.org/id/252A879DFF96A17AFF2FFF13FEC7C21A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Wang, Rong-Rong;Webb, Michael D.;Liang, Ai-Ping	Wang, Rong-Rong, Webb, Michael D., Liang, Ai-Ping (2010): First record of the male of Leusaba philippina Stål (Hemiptera: Fulgoromorpha: Tropiduchidae) with a redescription of the genus and species. Zootaxa 2344: 52-60, DOI: 10.5281/zenodo.275553
252A879DFF93A178FF2FFB49FBA5C224.text	252A879DFF93A178FF2FFB49FBA5C224.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leusaba philippina Stal	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Leusaba philippina Stål</p>
            <p>(Figs. 1 A–B, 2A–I, 3A–F)</p>
            <p> Leusaba philippina Stål, 1870: 749 ; Melichar, 1914: 108. </p>
            <p>Description. Length (from apex of vertex to tip of fore wings): 3 (N=1), 15.0 mm; Ƥ (N=1), 15.5 mm. Body length (from apex of vertex to tip of anal tube): 3, 9.1 mm; Ƥ, 9.4 mm.</p>
            <p>Colour. General color ochraceous; frons with anterior and lateral margins and medial line suffused with orange stripes, apex of rostrum, eyes and antennal pedicel, suffused with brown; vertex and pronotum dark green, vertex with posterior marginal carinae, pronotum with median disc and lateral portion, and mesonotum with median carina, all suffused with orange; fore wings hyaline, veins and stigma brown; fore tibia and mid tibia with apices and bases, hind tibia with bases, lateral spines with apices, long stripes on hind femora and tarsi with apical segments all suffused with black.</p>
            <p>Head and thorax. Head (Figs. 1 A, 2A) projecting beyond eyes for 0.39 times longer than length of eye. Frons (Figs. 1 B, 2C) without median carina, lateral carinae slightly carinate, disc smooth, median length 1.02 times longer than greatest width and 1.45 times longer than breadth at anterior margin of frons. Vertex (Figs. 1 A, 2A) distinctly shorter than greatest width (0.37:1), slightly longer than pronotum at midline (1.11:1).</p>
            <p>Pronotum (Figs. 1 A, 2A) distinctly shorter than mesonotum in midline (0.18:1). Mesonotum (Figs. 1 A, 2A) with median carina uniting with lateral carinae; lateral carinae curvulate for their posterior half and converging sharply anteriorly in anterior half; pronotum and mesonotum together medially 5.9 times as long as median length of vertex. Fore wings (Figs. 1 A, 2D) hyaline, surpassing extremity of abdomen about 0.65 times as long as length of body and 2.8 times as long as maximum breadth; with 18–20 apical cells. Hind tibiae usually with 4 distinct lateral spines, if with 5 spines then often on one leg only and with basal spine very fine.</p>
            <p>Male genitalia. Xth segment with lateroventral margin with two pair of distinctly long spines, one pair situating at apical 1/3 and other situated at level of extremity of apical dorsal margin (Figs. 2 F, 2H); anal style short and small, only slightly surpassing extremity of apical dorsal margin of anal tube. Left gonostylus (Figs. 2 F, 2G, 2I) 3.18 times as long as maximum breadth in lateral view (see Fig. 2 F), basal dorsal edge with a dorsoposteriorly directed prominence, followed by a finger-like, dorsoposteriorly directed process (see Fig.</p>
            <p>2F). Right gonostylus 3.23 times as long as maximum breadth in lateral view, basal dorsal edge with a clublike, dorsolaterally directed process (see Fig. 2 G). Periandrium (Figs. 3 A, 3B) with apical margin with a thin and sinuous branched process at ventral side, with apical part curved dorsally, process forked at middle part, protruding anteriorly, apex curved posterodorsally. Aedeagus (Figs. 3 A, 3B) with a long, sinuous process at basal part, lateroventrally directed, with ventral margin with many denticles, apical part curved posterodorsally; basal 1/4 to 1/2 of aedeagus with three projections at left side: one lamelliform projection I small and short, directed dorsally at dorsal side, one lamelliform projection II relatively long, stout and twisted, directed ventrally at ventral side and one global projection III arising from middle part between two lamelliform projections; with a tooth-like dorsoposteriorly directed prominence arising from dorsal edge at right side; endosoma with three projections: right projection abruptly extended into a falciform plate, directed dorsally, ventral projection spine-like, directed anteroventrally, and apical projection reaching to level of dorsal margin of anal tube, stout and cylindrical, apical part twisted with apex oblique and lanceolate in form, with margin denticulated.</p>
            <p>Female genitalia. Gonopophyses VIII (Figs. 3 C, 3F) asymmetrical at base in ventral view (Fig. 3 F), left produced into lobe with large globular apex, right produced into lobe with smaller globular apex. Gonopophyses IX (Fig. 3 E) with apical ends not confluent; gonospiculum (Fig. 3 E) slightly shorter than median length of triangular part (0.95:1), indistinctly flattened laterally. Gonoplacs (Figs. 3 C, 3D, 3F) with 4 teeth at dorsal margin, dentition complex, not forming one line, with 3 outer teeth at apical part, followed by 12–13 teeth from apical to ventral margin. Sternite VII with median part sinuous at ventral side (Fig. 3 F).</p>
            <p>Material examined. Philippines: 13, P. I. MISAMIS OR., Mt. Empagatao, 1050–1200 m, Rain forest, 19–30.IV.1961, H. Torrevillas (BPBM); 1Ƥ, P. I., MINDANAO, Agusan, Esperanza, 4–11.XI.1959, C. M. Yoshimoto (BPBM).</p>
            <p> Remarks.  L. philippina was described from a single specimen (holotype). The above material has been identified based on a comparison of images sent of this specimen (NHRS). </p>
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	https://treatment.plazi.org/id/252A879DFF93A178FF2FFB49FBA5C224	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Wang, Rong-Rong;Webb, Michael D.;Liang, Ai-Ping	Wang, Rong-Rong, Webb, Michael D., Liang, Ai-Ping (2010): First record of the male of Leusaba philippina Stål (Hemiptera: Fulgoromorpha: Tropiduchidae) with a redescription of the genus and species. Zootaxa 2344: 52-60, DOI: 10.5281/zenodo.275553
252A879DFF91A178FF2FFBACFE0BC3B7.text	252A879DFF91A178FF2FFBACFE0BC3B7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leusaba marginalis Walker	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Leusaba marginalis Walker</p>
            <p> Leusaba marginalis Walker, 1857: 144 , fig. 1; Melichar, 1914: 107. </p>
            <p>Material examined. Malaysia: syntype Ƥ, Sarawak, Wallace, 68.4 (BMNH).</p>
            <p> Remarks. The female type of this species (originally stated as male) is without head. However, the original description shows a complete specimen similar to  L. philippina and indet. sp. 1 below with respect to the distinctly visible frons in dorsal view. However, the terminalia in dorsal view are more similar to indet. sp. 2 (see Remarks below). </p>
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	https://treatment.plazi.org/id/252A879DFF91A178FF2FFBACFE0BC3B7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Wang, Rong-Rong;Webb, Michael D.;Liang, Ai-Ping	Wang, Rong-Rong, Webb, Michael D., Liang, Ai-Ping (2010): First record of the male of Leusaba philippina Stål (Hemiptera: Fulgoromorpha: Tropiduchidae) with a redescription of the genus and species. Zootaxa 2344: 52-60, DOI: 10.5281/zenodo.275553
252A879DFF91A178FF2FFA3BFE9CC1B4.text	252A879DFF91A178FF2FFA3BFE9CC1B4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leusaba rufitarsis (Kirby) Kirby	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Leusaba rufitarsis (Kirby)</p>
            <p>(Figs. 1 G–H)</p>
            <p> Stacota rufitarsis Kirby, 1891: 134 . </p>
            <p> Leusaba rufitarsis (Kirby) , Melichar, 1898: 326. </p>
            <p>Material examined. Sri Lanka: type Ƥ, Pundaloya, Green coll. (BMNH); 13, 1Ƥ, Kandy, IX.1902; 1?, Kandy VI.1902 (all BMNH).</p>
            <p> Remarks. This species can be distinguished by its narrower head and more elongate frons (see Figs. 1 G– H). In addition to the examined material Wilson and Malenovsky (2007) record 13 [no data], and 13, 1Ƥ, Sri Lanka, Peradeniya 29.III.1902 Dr. Uzel (all MMBC). This material was misidentified by Melichar (1903) as  L. marginalis . </p>
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	https://treatment.plazi.org/id/252A879DFF91A178FF2FFA3BFE9CC1B4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Wang, Rong-Rong;Webb, Michael D.;Liang, Ai-Ping	Wang, Rong-Rong, Webb, Michael D., Liang, Ai-Ping (2010): First record of the male of Leusaba philippina Stål (Hemiptera: Fulgoromorpha: Tropiduchidae) with a redescription of the genus and species. Zootaxa 2344: 52-60, DOI: 10.5281/zenodo.275553
252A879DFF90A179FF2FFF59FA80C7A7.text	252A879DFF90A179FF2FFF59FA80C7A7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leusaba	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Leusaba indet. sp. 1 </p>
            <p>(Figs. 1 C–D, 4A–I)</p>
            <p>Material examined. Malaysia (Sabah): 1Ƥ, Tawau Residency, Kalabakan R., Tawau 48 km W, Primary Forest, 18.IX.1958, T. C. Maa (BPBM).</p>
            <p> Remarks. This specimen is externally similar to  L. philippina and  L. marginalis , but can be distinguished from these species by its shorter anal tube and different proportions to the external female genitalia. </p>
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	https://treatment.plazi.org/id/252A879DFF90A179FF2FFF59FA80C7A7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Wang, Rong-Rong;Webb, Michael D.;Liang, Ai-Ping	Wang, Rong-Rong, Webb, Michael D., Liang, Ai-Ping (2010): First record of the male of Leusaba philippina Stål (Hemiptera: Fulgoromorpha: Tropiduchidae) with a redescription of the genus and species. Zootaxa 2344: 52-60, DOI: 10.5281/zenodo.275553
252A879DFF9FA176FF2FFF59FCCDC7D7.text	252A879DFF9FA176FF2FFF59FCCDC7D7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leusaba	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Leusaba indet. sp. 2 </p>
            <p>(Figs. 1 E–F)</p>
            <p>Material examined. W. Malaysia: 1Ƥ, Pahang, Benon Mts, 15 km. E. Kampong Dong, 24.III–15.IV.1998, 300– 1000 m, Dembický and Pacolátko (MMBC).</p>
            <p> Remarks. This specimen is similar to  L. marginalis with respect to the length of the anal tube and proportions of the external female genitalia. However, it does not match the original figure of  L. marginalis with respect to the frons which is barely visible in dorsal view but is clearly visible in  marginalis . Furthermore, the frons in facial view differs from the previous  Leusaba species in that the lateral margins are more parallel adjacent to the eyes (compare Figs. 1 B and 1F). </p>
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	https://treatment.plazi.org/id/252A879DFF9FA176FF2FFF59FCCDC7D7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Wang, Rong-Rong;Webb, Michael D.;Liang, Ai-Ping	Wang, Rong-Rong, Webb, Michael D., Liang, Ai-Ping (2010): First record of the male of Leusaba philippina Stål (Hemiptera: Fulgoromorpha: Tropiduchidae) with a redescription of the genus and species. Zootaxa 2344: 52-60, DOI: 10.5281/zenodo.275553
