occurrenceID	taxonID	catalogNumber	collectionCode	institutionCode	typeStatus	verbatimLabel	sex	individualCount	eventDate	recordedBy	recordNumber	decimalLatitude	decimalLongitude	minimumElevationInMeters	maximumElevationInMeters	minimumDepthInMeters	maximumDepthInMeters	country	stateProvince	municipality	locality	references	associatedOccurrences	associatedReferences	associatedSequences	basisOfRecord	eventRemarks	occurrenceRemarks
2620F071FFA6AC19FF0AF9DFFCCA45A7.mc.1EE14B3AFFA6AC19FF0AF8A9FCCA45A7	2620F071FFA6AC19FF0AF9DFFCCA45A7.taxon			ERMR		Known only from Florida, this is one of the species that lacks transverse striae medially on the pronotum. Johansen (1987) placed cassiae in his “ Group Obesus ” and thus distinguished it from mali on the basis of the claimed differences in the shape of the tube discussed and illustrated above. Almost no structural difference has been detected between specimens identified as cassiae from Southeastern USA, and specimens identified as oribates from Southwestern USA. These two species have the mesonotal transverse striae (Fig. 1) more widely spaced than in mali. However, a single female (in ERMR) from Venezuela has been studied that has the mesonotum closely striate but otherwise shares the character states of cassiae.		1										United States of America	Florida	These	Known	https://treatment.plazi.org/id/2620F071FFA6AC19FF0AF9DFFCCA45A7#1EE14B3AFFA6AC19FF0AF8A9FCCA45A7				MaterialCitation		
2620F071FFA5AC1AFF0AFF3EFD6A4089.mc.1EE14B3AFFA5AC1AFF0AFEF7FD6A4089	2620F071FFA5AC1AFF0AFF3EFD6A4089.taxon			USNM	paratype	This is another member of the genus known only from the western States. It is one of the few species in the genus with a small tooth on the inner margin of the fore tarsus, but is distinguished from two other species with this structure by the presence of several duplicated cilia on the fore wing. The pronotum bears transverse striae, the pronotal mid-lateral setae are present as in fasciculatus, but the mesonotum is closely striate in contrast to that species (Fig. 28). Cott (1956: 103) indicated that distalis had been found breeding in large numbers on a species of Atriplex at Firebaugh (near Fresno) in California. He stated that the plants were severely damaged by this thrips, and made no reference to the possibility that it might be predatory. Johansen described pristinus from Baja California, and distinguished it because of the presence on antennal segment IV of a small accessory sense cone on the external margin. However, as discussed above, the presence of these minute sensilla coeloconica appears to be unstable in some species, and examination of a paratype of pristinus (in USNM) suggests that this species cannot be distinguished satisfactorily from distalis.		1											California	western States. It	The	https://treatment.plazi.org/id/2620F071FFA5AC1AFF0AFF3EFD6A4089#1EE14B3AFFA5AC1AFF0AFEF7FD6A4089				MaterialCitation		
2620F071FFA5AC1AFF0AFC9BFEAD4770.mc.1EE14B3AFFA5AC1AFF0AFB9FFEAD4770	2620F071FFA5AC1AFF0AFC9BFEAD4770.taxon					Described from both sexes on Eriogonum fasciculatum [Polygonaceae] at Claremont, California, this species is known to breed in the flowers of this plant with no evidence that it is predatory (Wiesenborn 2012). Specimens have also been studied from Nevada and Arizona. Despite sharing with purpuratus the presence of a fore tarsal tooth and the absence of duplicated wing cilia, the pronotum of fasciculatus has no prominent sculpture lines but has well-developed mid-lateral setae (Fig. 22). Moreover, the sculpture lines on the mesonotum are not closely spaced, but form an almost reticulate pattern (Fig. 28). The fore tarsal tooth of males is considerably larger than that of females.		1											California		California	https://treatment.plazi.org/id/2620F071FFA5AC1AFF0AFC9BFEAD4770#1EE14B3AFFA5AC1AFF0AFB9FFEAD4770				MaterialCitation		
2620F071FFA5AC1AFF0AFAD3FADF459B.mc.1EE14B3AFFA5AC1AFF0AF9ADFADF459B	2620F071FFA5AC1AFF0AFAD3FADF459B.taxon			ERMR	lectotype	Described from an unspecified number of both sexes taken from an unknown plant near Palm Springs, California, a Lectotype was designated for this species by Pitkin (1978: 282). The number of major sense cones on antennal segment IV was stated by Johansen (1987: 78) to be three plus a minor one. However, re-examination of the lectotype and seven specimens bearing the same data has confirmed that all of them bear four sensilla basiconica (= major sense cones) on this segment. In May, 2015, heliomanes was found in large numbers at Big Morongo Canyon, 50 miles east of Riverside. It was breeding on grasses in association with unidentified mites. Adults collected at the same site but on the leaves of Prosopis trees have shorter antennae with only three sense cones on segment IV and possibly represent papago. Series of heliomanes typically have few or no duplicated cilia on the fore wing, but individuals sometimes have up to 6 cilia on one but not on both fore wings. Moreover, the duplicated cilia on some wings are unusually slender. Samples sometimes include individuals with three major sense cones on segment IV, and if such individuals were collected alone they would be identified as either papago or primigenus, depending on the presence or absence of duplicated cilia. Despite this, judging from the many specimens in ERMR, it appears that heliomanes is a valid species and probably the Western version of the Eastern species mali.		1											California	Big Morongo Canyon	Riverside. It	https://treatment.plazi.org/id/2620F071FFA5AC1AFF0AFAD3FADF459B#1EE14B3AFFA5AC1AFF0AF9ADFADF459B				MaterialCitation		
2620F071FFA4AC14FF0AF9B4FB184332.mc.1EE14B3AFFA4AC14FF0AF90EFB184332	2620F071FFA4AC14FF0AF9B4FB184332.taxon			ERMR	paratype	Described from Arizona, with paratypes from California, all associated with Larrea tridentata [Zygophyllaceae], this species probably feeds on mites within cecidomyiid galls on this plant (Wiesenborn 2015). More than 100 slide mounted specimens from Larrea have been studied (from ERMR), mainly collected in California. When the head is truly horizontal, the compound eyes are clearly longer ventrally than dorsally (but see caveat above). The postocular setae are short, 20 – 25 microns in length, scarcely reaching the posterior margin of the eye. The number of duplicated cilia on the fore wing is variable within any given population, with bilateral asymmetry in the number being common. At least 10 % of adults have no duplicated cilia on one or both fore wings. Antennal segment IV usually has three sense cones, but a very few specimens have been seen with only two on one antenna. Moreover, the small ancillary sense cone (= sensillum coeloconicum) on the external margin of segment IV is present in most examined specimens, but cannot be seen on one or both antennae of a substantial minority of specimens. Despite this variation, the specimens from Larrea consistently have antennal segments III and IV rather short, and antennal segment III with no sense cone. However, not all Leptothrips individuals collected from Larrea are identifiable as larreae. Three specimens from Larrea at Banning, California, have a sense cone on antennal segment III, and antennal segments III + IV more than 160 microns long; the two females have four sense cones on segment IV but the male only three, and the number of duplicated cilia varies from 1 to 4 on these three specimens.		1											Arizona	More	The	https://treatment.plazi.org/id/2620F071FFA4AC14FF0AF9B4FB184332#1EE14B3AFFA4AC14FF0AF90EFB184332				MaterialCitation		
2620F071FFAAAC15FF0AFF3EFF0341EA.mc.1EE14B3AFFAAAC15FF0AFE96FF0341EA	2620F071FFAAAC15FF0AFF3EFF0341EA.taxon			USNM, FSAC, ERMR	lectotype	The Code of Zoological Nomenclature (http: // www. iczn. org / iczn / index. jsp) specifies that a hyphen should be deleted from any compound name, except where the first of the two names is a single letter. Watson described this species from an unspecified number of specimens taken from orange leaves at Gainesville, Florida, in January, 1913. Johansen (1987: 93) stated that he had designated a Lectotype of macroocellatus, but no such specimen has been found in either USNM or FSAC. The only slides found that bear original data are in FSAC, and these include two larvae and six adults, of which only one has an antenna. Based on these specimens, this species is interpreted as having sculpture lines on the pronotum posteriorly and laterally but not medially (Fig. 21), and in having antennal segment IV with three sense cones. Moreover, the external margin of segment IV bears a small ancillary sense cone (= sensillum coeloconicum), although this is not always visible. The compound eyes are large, but the extent to which they are longer ventrally than dorsally depends on the orientation of the head on slide mounted specimens. Samples of this species have been studied (in ERMR) from Florida (Orlando) and South Carolina (Colleton Cty), also (in USNM) from Ft Garland (Colorado). However, a few unidentified specimens mentioned by Mound et al. (2016) that were collected from Jatropha curcas at Chiapas, Mexico can now be identified as this species, together with 50 specimens representing both sexes collected by Francisco Infante in March and April, 2014, at various sites in the Southern parts of Chiapas State. The species confusus was based on a single female from Homestead, Florida (in USNM), and placed in a key between mali and macroocellatus. This specimen was distinguished from macroocellatus because it has the eyes scarcely prolonged ventrally and the major setae rather shorter.		50		April & Southern & Homestead & Florida & This								Mexico	Chiapas	Mound	The Code of Zoological Nomenclature	https://treatment.plazi.org/id/2620F071FFAAAC15FF0AFF3EFF0341EA#1EE14B3AFFAAAC15FF0AFE96FF0341EA				MaterialCitation		
2620F071FFAAAC16FF0AFBA6FD6C434B.mc.1EE14B3AFFAAAC16FF0AFAF8FD6C434B	2620F071FFAAAC16FF0AFBA6FD6C434B.taxon				holotype	Johansen distinguished maliaffinis and septemtrionalis from mali on the grounds that specimens of mali lack a major sense cone on the external apical margin of antennal segment IV - that is, he claimed that mali has only three major sense cones on this segment. However, all available specimens of mali, including specimens identified as this species by Johansen himself, have on segment IV four well-developed sensilla basiconica, even though one of these is sometimes smaller than the others. Johansen further distinguished septemtrionalis from maliaffinis on the basis that the former lacks a small accessory sense cone (sensillum coeloconicum) on the external margin of antennal segment IV, whereas the latter has such a sense cone. However, although this structure is visible on the right antenna of the holotype female of maliaffinis it is not visible on the left antenna, nor on either of the antennae of two available paratypes of maliaffinis. Similarly, this small sense cone is not visible on the holotype of septemtrionalis nor one male paratype from Virginia, but is clearly visible on the left (but not the right) antenna of a female paratype of septemtrionalis from near Blacksburg. There is no evidence from any of the available slides that the small sense cone has been broken off during slide preparation, and the assumption here is therefore that its presence is variable (or that its visibility is variable on slide-mounted specimens). One female from Colorado, identified by Johansen as californicus, has this minute sense cone visible on the left antenna but not the right. As discussed above, gurdus was described by Johansen as having the tube short and conical, but that wording does not seem appropriate to the tube of either the holotype female (Fig. 14) or the paratype male, and these specimens cannot be distinguished from the common species mali. Johansen also placed mcconelli in the same group with mali as having only three major sense cones on antennal segment IV, but placed californicus in the group with maliaffinis as having four such sense cones. The first of these is here considered a valid species, but the second is returned to synonymy with mali. As interpreted here, mali is variable in the colour and length of antennal segments III and IV, with some individuals having these segments largely brown and slender, but others having them shorter and yellow in the basal half. Similar variation in the colour and shape of these antennal segments occurs in the closely related western species, heliomanes.		1											Virginia		Virginia	https://treatment.plazi.org/id/2620F071FFAAAC16FF0AFBA6FD6C434B#1EE14B3AFFAAAC16FF0AFAF8FD6C434B				MaterialCitation		
2620F071FFA9AC16FF0AFAA5FCDB444D.mc.1EE14B3AFFA9AC16FF0AFA3CFCDB444D	2620F071FFA9AC16FF0AFAA5FCDB444D.taxon					This species remains known only from the original specimens: two females and one male collected at Bridgeland, Utah. It is remarkable for the long and deeply retracted maxillary stylets that are close together medially in the head (Fig. 5). Despite this, the species is similar to heliomanes in having the pronotum with particularly strong transverse striae, and the fore wing lacking duplicated cilia.		1											Utah		Utah	https://treatment.plazi.org/id/2620F071FFA9AC16FF0AFAA5FCDB444D#1EE14B3AFFA9AC16FF0AFA3CFCDB444D				MaterialCitation		
2620F071FFA9AC16FF0AF9C4FCC14595.mc.1EE14B3AFFA9AC16FF0AF95FFCC14595	2620F071FFA9AC16FF0AF9C4FCC14595.taxon					This species is based on a single female that is labelled “ Colorado, Allenspark, Pine, 1950 ”. The specimen is slidemounted in Clarite, and as a result it is crushed and distorted, with crumpled wings, and the sense cones eroded and faint. The pronotum has strong transverse striae at least on the anterior third, the mesonotum is transversely striate, the basal pores of four major sense cones are visible on antennal segment IV, but the head is too distorted to decide if the eyes are prolonged ventrally. Antennal segment II is asymmetric and presumably aberrant (Fig. 6), and the specimen is possibly a deformed individual of heliomanes.		1											Colorado		Colorado	https://treatment.plazi.org/id/2620F071FFA9AC16FF0AF9C4FCC14595#1EE14B3AFFA9AC16FF0AF95FFCC14595				MaterialCitation		
2620F071FFA9AC16FF0AFEC6FDA346EF.mc.1EE14B3AFFA9AC16FCF5FDC9FCE44097	2620F071FFA9AC16FF0AFEC6FDA346EF.taxon				lectotype	From these specimens Johansen (1987) designated a female lectotype (in California Academy of Sciences), and listed many specimens of the species from Mexico and also a few in USA from Utah, Idaho and California		1										Mexico	Mexico	Utah	Idaho	https://treatment.plazi.org/id/2620F071FFA9AC16FF0AFEC6FDA346EF#1EE14B3AFFA9AC16FCF5FDC9FCE44097				MaterialCitation		
2620F071FFA9AC16FF0AFEC6FDA346EF.mc.1EE14B3AFFA9AC16FC8AFD01FB214107	2620F071FFA9AC16FF0AFEC6FDA346EF.taxon			ERMR	holotype	He then described primigenus from a single female taken at Yosemite, California, distinguishing it because antennal segment IV of the holotype lacks a small ancillary sense cone (= sensillum coeloconicum) on the external surface. This five micron long ancillary sense cone is present on most of the 150 specimens (in ERMR) that are here identified as mcconelli, although it is certainly absent on both antennae of a few of them, and mcconelli is here interpreted as a common species in California		150										Mexico	California	This	California	https://treatment.plazi.org/id/2620F071FFA9AC16FF0AFEC6FDA346EF#1EE14B3AFFA9AC16FC8AFD01FB214107				MaterialCitation		
2620F071FFA9AC16FF0AFEC6FDA346EF.mc.1EE14B3AFFA9AC16FB49FC91FDA346EF	2620F071FFA9AC16FF0AFEC6FDA346EF.taxon				holotype	It is distinguished from larreae by the presence of a sense cone on antennal segment III, and the greater length, 130 microns, of antennal segments III + IV, and distinguished from mali by the presence on segment IV of three not four major sense cones (sensilla basiconica). However, the number of fore wing duplicated cilia is variable amongst the specimens examined. For example, in a series of 14 individuals taken from Sycamore at Riverside (30. vi. 1958) the number varies from 0 to 7, two of these specimens have no duplicated cilia on either wing, and one male has antennal segment IV with four sense cones. As indicated above, character state variation amongst the five species, heliomanes, larreae, mali, mcconelli and papago is very confusing, with the occasional individual intermediate in structure between two or more of these. Specimens identified as mcconelli have been studied from Alaska, Arizona, California, Colorado, Texas, and Utah from various plants, particularly tree species in the genera Acer, Alnus, Fraxinus, Prunus, and Quercus.		1	1958-06-30									Mexico	Alaska, Arizona, California, Colorado, Texas	Specimens	Utah	https://treatment.plazi.org/id/2620F071FFA9AC16FF0AFEC6FDA346EF#1EE14B3AFFA9AC16FB49FC91FDA346EF				MaterialCitation		
2620F071FFA8AC10FF0AF9C7FD3A43A2.mc.1EE14B3AFFA8AC17FF0AF89DFD544525	2620F071FFA8AC10FF0AF9C7FD3A43A2.taxon					Described from Colorado, Arizona and New Mexico, with one synonym from Oregon, specimens of this species have also been seen from Nevada and California		1										Mexico	Colorado		Colorado	https://treatment.plazi.org/id/2620F071FFA8AC10FF0AF9C7FD3A43A2#1EE14B3AFFA8AC17FF0AF89DFD544525				MaterialCitation		
2620F071FFA8AC10FF0AF9C7FD3A43A2.mc.1EE14B3AFFA8AC10FD5BF8F3FACF42F8	2620F071FFA8AC10FF0AF9C7FD3A43A2.taxon					It is closely associated with species of Pinus, and has been taken several times from immature pine cones. It is distinguished from pini, with which it shares this host association, by the presence on antennal segment IV of four sense cones. Both species lack pronotal sculpture and have the fore wings pale at the base, but the available distribution records suggest that they are adapted to rather different ecological conditions, with pini mainly from the Florida area, but oribates from the Western states		1										Mexico	Western		Florida area	https://treatment.plazi.org/id/2620F071FFA8AC10FF0AF9C7FD3A43A2#1EE14B3AFFA8AC10FD5BF8F3FACF42F8				MaterialCitation		
2620F071FFA8AC10FF0AF9C7FD3A43A2.mc.1EE14B3AFFAFAC10FAA8FF3EFB074282	2620F071FFA8AC10FF0AF9C7FD3A43A2.taxon					Johansen described brevicapitis from a few specimens associated with Pinus from California and Arizona		1										Mexico	California		California	https://treatment.plazi.org/id/2620F071FFA8AC10FF0AF9C7FD3A43A2#1EE14B3AFFAFAC10FAA8FF3EFB074282				MaterialCitation		
2620F071FFA8AC10FF0AF9C7FD3A43A2.mc.1EE14B3AFFAFAC10FB61FF14FD3A43A2	2620F071FFA8AC10FF0AF9C7FD3A43A2.taxon			ERMR	holotype	He stated that these specimens have the eyes prolonged ventrally, in contrast to those of oribates. However, the difference between the dorsal and ventral surfaces in the holotype (see Fig. 73 in Johansen, 1987) is trivial, and involves a difference of no more than the diameter of a single ommatidium. The extent to which the eyes are longer ventrally than dorsally in the available specimens of oribates is here interpreted as being associated with the orientation of the head in slide-mounted specimens (see discussion above). More than 40 specimens of oribates have been studied from various sites in California (in ERMR); two of these were noted to have only three sense cones on segment IV of one antenna (but not on both), and in all of them segment IV lacked a small ancillary sense cone (= sensillum coeloconicum) on the external surface.		40										Mexico	California	Johansen	The	https://treatment.plazi.org/id/2620F071FFA8AC10FF0AF9C7FD3A43A2#1EE14B3AFFAFAC10FB61FF14FD3A43A2				MaterialCitation		
2620F071FFAFAC10FF0AFDEEFD1A479A.mc.1EE14B3AFFAFAC10FF0AFC84FD1A479A	2620F071FFAFAC10FF0AFDEEFD1A479A.taxon			ERMR	holotype	This species was based on a female holotype, with three female and one male paratypes, all collected from cottonwood, at Tucson, Arizona. Measurements of the antennae in the four species listed in the synonymy above are remarkably different. The length of antennal segments III + IV is 145 in the holotype of papago, whereas the measurements provided by Hood indicate only 107 for the holotype of acaciae. The similar measurement for the holotype of robustus is 130, and 110 for the holotype of arizonensis. No other character state differences have been found that correlate with this variation in antennal segment lengths. A further species in which these segments are unusually short is larreae, but in that species antennal segment III lacks a sense cone. Hood described acaciae from Wickenburg, Arizona, based on 17 females and one male collected from Acacia or Prosopis. Johansen described robustus from two specimens from Wyoming and one from Colorado, all three being mounted in Clarite. These specimens are thus seriously distorted, with the antennal sense cones almost completely eroded, and the fore wings of the holotype not available. However, one fore wing of the paratype from Wyoming clearly has no duplicated cilia. Johansen distinguished arizonensis from papago in a key, indicating that the eyes were not prolonged ventrally in the first but prolonged in papago. However, the difference in dorsal and ventral lengths of the eyes in the available specimens is largely dependent on how horizontal the head is on a slide-mounted specimen. Johansen placed robustus in his Group Obesus, presumably because the tube of the holotype is wide at the base (Fig. 16). However, this specimen is mounted in clarite, and is thus crushed with the tube distorted. Individuals identified here as papago have been seen from a wide range of unrelated plants at sites between Utah and California, based on 100 slides in ERMR; the recorded plants include members of the following genera: Adenostema, Ceanothus, Chrysothamnus, Hymenochloa, Purshia and Quercus.	female	17											Arizona	Tucson	The	https://treatment.plazi.org/id/2620F071FFAFAC10FF0AFDEEFD1A479A#1EE14B3AFFAFAC10FF0AFC84FD1A479A				MaterialCitation		
2620F071FFAFAC10FF0AF988FBEA45C1.mc.1EE14B3AFFAFAC10FF0AF960FBEA45C1	2620F071FFAFAC10FF0AF988FBEA45C1.taxon			ERMR, USNM		Described from Florida, and recorded from Virginia and Maryland, in association with species of Pinus, there are specimens in ERMR from Texas, taken on Pinus taeda at Smithville, and on Pinus sp. at Blossom. The following have also been studied (in USNM): one specimen from Arizona, one from Georgia, five from New York and three from South Carolina. This species shares with singularis the presence of only two sense cones on the fourth antennal segment. The pronotum lacks sculptured striae medially but is strongly striate anterolaterally, the fore wing is pale at the base, but the apex of antennal segment III is shaded brown.		1										Georgia	Florida	The	South Carolina. This	https://treatment.plazi.org/id/2620F071FFAFAC10FF0AF988FBEA45C1#1EE14B3AFFAFAC10FF0AF960FBEA45C1				MaterialCitation		
2620F071FFAEAC11FF0AFF3EFF114045.mc.1EE14B3AFFAEAC11FF0AFE88FF114045	2620F071FFAEAC11FF0AFF3EFF114045.taxon					Described from one female and one male, taken on a species of Atriplex at Tempe, Arizona, this species was described as having antennal segments III – VI “ bright yellow ”. The fore tarsus bears a small tooth on the inner margin, as in distalis and fasciculatus, and it shares with distalis the presence of strong transverse striae on the pronotum but lacks mid-lateral setae. In contrast, fasciculatus has a smooth pronotum, and purpuratus is unique among the three species of the distalis group in lacking a sense cone on antennal segment III. Specimens have been studied from Arizona (Sacaton), California (Tecopa, Thermal, Victorville), New Mexico (Rodeo) and Texas (Ysleta).		1											Arizona		Arizona	https://treatment.plazi.org/id/2620F071FFAEAC11FF0AFF3EFF114045#1EE14B3AFFAEAC11FF0AFE88FF114045				MaterialCitation		
2620F071FFAEAC11FF0AFDCCFC004125.mc.1EE14B3AFFAEAC11FF0AFD47FC004125	2620F071FFAEAC11FF0AFDCCFC004125.taxon					Described from three females and five males collected at Pine Key, Florida, this species is similar to pini but has the pronotum and hind tibiae paler. These two both have only two sense cones on antennal segment IV, but both are known from too few specimens to be sure that this character state is constant. A further species from Florida that is similar in structure and lacks sculpture lines across the pronotum is cassiae.		1											Florida		Florida	https://treatment.plazi.org/id/2620F071FFAEAC11FF0AFDCCFC004125#1EE14B3AFFAEAC11FF0AFD47FC004125				MaterialCitation		
