taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
266887BDFF9CA13C0135FADCFC14B2FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4747410/files/figure.png	https://doi.org/10.5281/zenodo.4747410	Figure 1. Onthophagus osculatii. (a, b) Dorsal habitus of male (neotype) and female (scale bar: 1 mm). (g, h) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (k, l) Lamella copulatrix and accessory lamellae; arrow indicating the basolateral carina of the inferior left lobe of O. osculatii. (m) Female genitalia (scale bar: 0.02 mm); arrow indicating the slight medial excavation of the ventral sclerotization of O. osculatii. Onthophagus chacoensis sp. nov. (c, d) Dorsal habitus of male (holotype) and female. (e) Metasternal punctuation (scale bar: 1 mm). (f) Detail of the pronotal punctuation. (i, j) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (n, o) Lamella copulatrix and accessory lamellae; arrow indicating the deep excavation of the external margin of the left lobe of O. chacoensis. (p) Female genitalia (scale bar: 0.02 mm).	Figure 1. Onthophagus osculatii. (a, b) Dorsal habitus of male (neotype) and female (scale bar: 1 mm). (g, h) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (k, l) Lamella copulatrix and accessory lamellae; arrow indicating the basolateral carina of the inferior left lobe of O. osculatii. (m) Female genitalia (scale bar: 0.02 mm); arrow indicating the slight medial excavation of the ventral sclerotization of O. osculatii. Onthophagus chacoensis sp. nov. (c, d) Dorsal habitus of male (holotype) and female. (e) Metasternal punctuation (scale bar: 1 mm). (f) Detail of the pronotal punctuation. (i, j) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (n, o) Lamella copulatrix and accessory lamellae; arrow indicating the deep excavation of the external margin of the left lobe of O. chacoensis. (p) Female genitalia (scale bar: 0.02 mm).	2018-03-07	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario		Zenodo	biologists	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario			
266887BDFF9CA13C0135FADCFC14B2FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4747418/files/figure.png	https://doi.org/10.5281/zenodo.4747418	Figure 5. Geographic distribution of the species of the osculatii complex.	Figure 5. Geographic distribution of the species of the osculatii complex.	2018-03-07	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario		Zenodo	biologists	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario			
266887BDFF85A13F0189FA61FEB9B46A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4747410/files/figure.png	https://doi.org/10.5281/zenodo.4747410	Figure 1. Onthophagus osculatii. (a, b) Dorsal habitus of male (neotype) and female (scale bar: 1 mm). (g, h) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (k, l) Lamella copulatrix and accessory lamellae; arrow indicating the basolateral carina of the inferior left lobe of O. osculatii. (m) Female genitalia (scale bar: 0.02 mm); arrow indicating the slight medial excavation of the ventral sclerotization of O. osculatii. Onthophagus chacoensis sp. nov. (c, d) Dorsal habitus of male (holotype) and female. (e) Metasternal punctuation (scale bar: 1 mm). (f) Detail of the pronotal punctuation. (i, j) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (n, o) Lamella copulatrix and accessory lamellae; arrow indicating the deep excavation of the external margin of the left lobe of O. chacoensis. (p) Female genitalia (scale bar: 0.02 mm).	Figure 1. Onthophagus osculatii. (a, b) Dorsal habitus of male (neotype) and female (scale bar: 1 mm). (g, h) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (k, l) Lamella copulatrix and accessory lamellae; arrow indicating the basolateral carina of the inferior left lobe of O. osculatii. (m) Female genitalia (scale bar: 0.02 mm); arrow indicating the slight medial excavation of the ventral sclerotization of O. osculatii. Onthophagus chacoensis sp. nov. (c, d) Dorsal habitus of male (holotype) and female. (e) Metasternal punctuation (scale bar: 1 mm). (f) Detail of the pronotal punctuation. (i, j) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (n, o) Lamella copulatrix and accessory lamellae; arrow indicating the deep excavation of the external margin of the left lobe of O. chacoensis. (p) Female genitalia (scale bar: 0.02 mm).	2018-03-07	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario		Zenodo	biologists	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario			
266887BDFF85A13F0189FA61FEB9B46A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4747418/files/figure.png	https://doi.org/10.5281/zenodo.4747418	Figure 5. Geographic distribution of the species of the osculatii complex.	Figure 5. Geographic distribution of the species of the osculatii complex.	2018-03-07	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario		Zenodo	biologists	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario			
266887BDFF86A132010AFCF1FDECB29F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4747412/files/figure.png	https://doi.org/10.5281/zenodo.4747412	Figure 2. Onthophagus transisthmius. (a, b) Dorsal habitus of male and female. (c, d) Dorsal habitus of the holotype and original labels (scale bar: 1 mm). (h, i) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (j, k) Lamella copulatrix and accessory lamellae. (l) Female genitalia (scale bar: 0.02 mm). Onthophagus basicarinatus sp. nov. (e, f) Dorsal habitus of male (holotype) and female. (g) Detail of the male horns (scale bar: 1 mm). (m, n) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (o, p) Lamella copulatrix and accessory lamellae (scale bar: 0.02 mm). (q) Female genitalia (scale bar: 0.02 mm).	Figure 2. Onthophagus transisthmius. (a, b) Dorsal habitus of male and female. (c, d) Dorsal habitus of the holotype and original labels (scale bar: 1 mm). (h, i) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (j, k) Lamella copulatrix and accessory lamellae. (l) Female genitalia (scale bar: 0.02 mm). Onthophagus basicarinatus sp. nov. (e, f) Dorsal habitus of male (holotype) and female. (g) Detail of the male horns (scale bar: 1 mm). (m, n) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (o, p) Lamella copulatrix and accessory lamellae (scale bar: 0.02 mm). (q) Female genitalia (scale bar: 0.02 mm).	2018-03-07	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario		Zenodo	biologists	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario			
266887BDFF86A132010AFCF1FDECB29F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4747410/files/figure.png	https://doi.org/10.5281/zenodo.4747410	Figure 1. Onthophagus osculatii. (a, b) Dorsal habitus of male (neotype) and female (scale bar: 1 mm). (g, h) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (k, l) Lamella copulatrix and accessory lamellae; arrow indicating the basolateral carina of the inferior left lobe of O. osculatii. (m) Female genitalia (scale bar: 0.02 mm); arrow indicating the slight medial excavation of the ventral sclerotization of O. osculatii. Onthophagus chacoensis sp. nov. (c, d) Dorsal habitus of male (holotype) and female. (e) Metasternal punctuation (scale bar: 1 mm). (f) Detail of the pronotal punctuation. (i, j) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (n, o) Lamella copulatrix and accessory lamellae; arrow indicating the deep excavation of the external margin of the left lobe of O. chacoensis. (p) Female genitalia (scale bar: 0.02 mm).	Figure 1. Onthophagus osculatii. (a, b) Dorsal habitus of male (neotype) and female (scale bar: 1 mm). (g, h) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (k, l) Lamella copulatrix and accessory lamellae; arrow indicating the basolateral carina of the inferior left lobe of O. osculatii. (m) Female genitalia (scale bar: 0.02 mm); arrow indicating the slight medial excavation of the ventral sclerotization of O. osculatii. Onthophagus chacoensis sp. nov. (c, d) Dorsal habitus of male (holotype) and female. (e) Metasternal punctuation (scale bar: 1 mm). (f) Detail of the pronotal punctuation. (i, j) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (n, o) Lamella copulatrix and accessory lamellae; arrow indicating the deep excavation of the external margin of the left lobe of O. chacoensis. (p) Female genitalia (scale bar: 0.02 mm).	2018-03-07	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario		Zenodo	biologists	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario			
266887BDFF86A132010AFCF1FDECB29F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4747418/files/figure.png	https://doi.org/10.5281/zenodo.4747418	Figure 5. Geographic distribution of the species of the osculatii complex.	Figure 5. Geographic distribution of the species of the osculatii complex.	2018-03-07	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario		Zenodo	biologists	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario			
266887BDFF8BA13501F9F98FFB23B65D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4747412/files/figure.png	https://doi.org/10.5281/zenodo.4747412	Figure 2. Onthophagus transisthmius. (a, b) Dorsal habitus of male and female. (c, d) Dorsal habitus of the holotype and original labels (scale bar: 1 mm). (h, i) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (j, k) Lamella copulatrix and accessory lamellae. (l) Female genitalia (scale bar: 0.02 mm). Onthophagus basicarinatus sp. nov. (e, f) Dorsal habitus of male (holotype) and female. (g) Detail of the male horns (scale bar: 1 mm). (m, n) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (o, p) Lamella copulatrix and accessory lamellae (scale bar: 0.02 mm). (q) Female genitalia (scale bar: 0.02 mm).	Figure 2. Onthophagus transisthmius. (a, b) Dorsal habitus of male and female. (c, d) Dorsal habitus of the holotype and original labels (scale bar: 1 mm). (h, i) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (j, k) Lamella copulatrix and accessory lamellae. (l) Female genitalia (scale bar: 0.02 mm). Onthophagus basicarinatus sp. nov. (e, f) Dorsal habitus of male (holotype) and female. (g) Detail of the male horns (scale bar: 1 mm). (m, n) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (o, p) Lamella copulatrix and accessory lamellae (scale bar: 0.02 mm). (q) Female genitalia (scale bar: 0.02 mm).	2018-03-07	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario		Zenodo	biologists	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario			
266887BDFF8BA13501F9F98FFB23B65D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4747418/files/figure.png	https://doi.org/10.5281/zenodo.4747418	Figure 5. Geographic distribution of the species of the osculatii complex.	Figure 5. Geographic distribution of the species of the osculatii complex.	2018-03-07	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario		Zenodo	biologists	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario			
266887BDFF8CA13701F6FECCFF4AB2D4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4747414/files/figure.png	https://doi.org/10.5281/zenodo.4747414	Figure 3. Onthophagus nyctopus. (a, b) Dorsal habitus of male and female. (c, d) Dorsal habitus of the lectotype and original labels (scale bar: 1 mm). (i, j) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (k, l) Lamella copulatrix and accessory lamellae; arrow indicating the basolateral carina of the inferior left lobe of O. nyctopus. (m) Female genitalia (scale bar: 0.02 mm). Onthophagus steinheili. (e, f) Dorsal habitus of male and female. (g, h) Dorsal habitus of the lectotype and original labels (scale bar: 1 mm). (n, o) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (p, q) Lamella copulatrix and accessory lamellae. (r) Female genitalia (scale bar: 0.02 mm).	Figure 3. Onthophagus nyctopus. (a, b) Dorsal habitus of male and female. (c, d) Dorsal habitus of the lectotype and original labels (scale bar: 1 mm). (i, j) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (k, l) Lamella copulatrix and accessory lamellae; arrow indicating the basolateral carina of the inferior left lobe of O. nyctopus. (m) Female genitalia (scale bar: 0.02 mm). Onthophagus steinheili. (e, f) Dorsal habitus of male and female. (g, h) Dorsal habitus of the lectotype and original labels (scale bar: 1 mm). (n, o) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (p, q) Lamella copulatrix and accessory lamellae. (r) Female genitalia (scale bar: 0.02 mm).	2018-03-07	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario		Zenodo	biologists	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario			
266887BDFF8CA13701F6FECCFF4AB2D4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4747418/files/figure.png	https://doi.org/10.5281/zenodo.4747418	Figure 5. Geographic distribution of the species of the osculatii complex.	Figure 5. Geographic distribution of the species of the osculatii complex.	2018-03-07	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario		Zenodo	biologists	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario			
266887BDFF8EA10901FEFA43FE13B2F9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4747414/files/figure.png	https://doi.org/10.5281/zenodo.4747414	Figure 3. Onthophagus nyctopus. (a, b) Dorsal habitus of male and female. (c, d) Dorsal habitus of the lectotype and original labels (scale bar: 1 mm). (i, j) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (k, l) Lamella copulatrix and accessory lamellae; arrow indicating the basolateral carina of the inferior left lobe of O. nyctopus. (m) Female genitalia (scale bar: 0.02 mm). Onthophagus steinheili. (e, f) Dorsal habitus of male and female. (g, h) Dorsal habitus of the lectotype and original labels (scale bar: 1 mm). (n, o) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (p, q) Lamella copulatrix and accessory lamellae. (r) Female genitalia (scale bar: 0.02 mm).	Figure 3. Onthophagus nyctopus. (a, b) Dorsal habitus of male and female. (c, d) Dorsal habitus of the lectotype and original labels (scale bar: 1 mm). (i, j) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (k, l) Lamella copulatrix and accessory lamellae; arrow indicating the basolateral carina of the inferior left lobe of O. nyctopus. (m) Female genitalia (scale bar: 0.02 mm). Onthophagus steinheili. (e, f) Dorsal habitus of male and female. (g, h) Dorsal habitus of the lectotype and original labels (scale bar: 1 mm). (n, o) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (p, q) Lamella copulatrix and accessory lamellae. (r) Female genitalia (scale bar: 0.02 mm).	2018-03-07	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario		Zenodo	biologists	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario			
266887BDFF8EA10901FEFA43FE13B2F9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4747418/files/figure.png	https://doi.org/10.5281/zenodo.4747418	Figure 5. Geographic distribution of the species of the osculatii complex.	Figure 5. Geographic distribution of the species of the osculatii complex.	2018-03-07	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario		Zenodo	biologists	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario			
266887BDFFB0A10D0110FA60FBC7B1E2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4747416/files/figure.png	https://doi.org/10.5281/zenodo.4747416	Figure 4. Onthophagus confusus stat. nov. (a, b) Dorsal habitus of male and female. (c, d) Dorsal habitus of the lectotype and original labels (scale bar: 1 mm). (j, k) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (l, m) Lamella copulatrix and accessory lamellae. (n) Female genitalia (scale bar: 0.02 mm). Onthophagus nabeleki. (e, f) Dorsal habitus of the lectotype and original labels (scale bar: 1 mm). Onthophagus insularis. (g–i) Dorsal habitus of the lectotype and paralectotype, original labels of the name-bearing type (scale bar: 1 mm). (o, p) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (q, r) Lamella copulatrix and accessory lamellae (scale bar: 0.02 mm).	Figure 4. Onthophagus confusus stat. nov. (a, b) Dorsal habitus of male and female. (c, d) Dorsal habitus of the lectotype and original labels (scale bar: 1 mm). (j, k) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (l, m) Lamella copulatrix and accessory lamellae. (n) Female genitalia (scale bar: 0.02 mm). Onthophagus nabeleki. (e, f) Dorsal habitus of the lectotype and original labels (scale bar: 1 mm). Onthophagus insularis. (g–i) Dorsal habitus of the lectotype and paralectotype, original labels of the name-bearing type (scale bar: 1 mm). (o, p) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (q, r) Lamella copulatrix and accessory lamellae (scale bar: 0.02 mm).	2018-03-07	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario		Zenodo	biologists	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario			
266887BDFFB0A10D0110FA60FBC7B1E2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4747418/files/figure.png	https://doi.org/10.5281/zenodo.4747418	Figure 5. Geographic distribution of the species of the osculatii complex.	Figure 5. Geographic distribution of the species of the osculatii complex.	2018-03-07	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario		Zenodo	biologists	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario			
266887BDFFB7A10001E9FF0AFD21B3AF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4747416/files/figure.png	https://doi.org/10.5281/zenodo.4747416	Figure 4. Onthophagus confusus stat. nov. (a, b) Dorsal habitus of male and female. (c, d) Dorsal habitus of the lectotype and original labels (scale bar: 1 mm). (j, k) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (l, m) Lamella copulatrix and accessory lamellae. (n) Female genitalia (scale bar: 0.02 mm). Onthophagus nabeleki. (e, f) Dorsal habitus of the lectotype and original labels (scale bar: 1 mm). Onthophagus insularis. (g–i) Dorsal habitus of the lectotype and paralectotype, original labels of the name-bearing type (scale bar: 1 mm). (o, p) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (q, r) Lamella copulatrix and accessory lamellae (scale bar: 0.02 mm).	Figure 4. Onthophagus confusus stat. nov. (a, b) Dorsal habitus of male and female. (c, d) Dorsal habitus of the lectotype and original labels (scale bar: 1 mm). (j, k) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (l, m) Lamella copulatrix and accessory lamellae. (n) Female genitalia (scale bar: 0.02 mm). Onthophagus nabeleki. (e, f) Dorsal habitus of the lectotype and original labels (scale bar: 1 mm). Onthophagus insularis. (g–i) Dorsal habitus of the lectotype and paralectotype, original labels of the name-bearing type (scale bar: 1 mm). (o, p) Lateral and dorsal view of the parameres (scale bar: 0.05 mm). (q, r) Lamella copulatrix and accessory lamellae (scale bar: 0.02 mm).	2018-03-07	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario		Zenodo	biologists	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario			
266887BDFFB7A10001E9FF0AFD21B3AF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4747418/files/figure.png	https://doi.org/10.5281/zenodo.4747418	Figure 5. Geographic distribution of the species of the osculatii complex.	Figure 5. Geographic distribution of the species of the osculatii complex.	2018-03-07	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario		Zenodo	biologists	Rossini, Michele;Vaz-de-Mello, Fernando Z.;Zunino, Mario			
