identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
200A8781FFD9F35600F09073977E21EE.text	200A8781FFD9F35600F09073977E21EE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus clermonti Paulian 1931	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Onthophagus clermonti Paulian, 1931</p>
            <p> Onthophagus clermonti Paulian, 1931: 271 (type locality: Hoa-Binh, Tonkin [Vietnam])  Onthophagus parvidens Frey, 1971: 98 syn.n. (type locality: Hoa-Binh, Tonkin [Vietnam]) </p>
            <p> Type material examined. Holotype of  O. clermonti (MNHN), female bearing the following labels: </p>
            <p>1. White, printed: Hoa-Binh (Tonkin) (A.de Cooman) Coll.J.Clermont</p>
            <p>2. Red, printed: TYPE</p>
            <p> 3. White, handwritten:  Clermonti type </p>
            <p> 4. Orange: HOLOTYPE  O. clermonti Paulian , 931 [handwritten] S. Tarasov det. 2009 [printed] </p>
            <p> Holotype of  O. parvidens (NHB), male bearing the following labels: </p>
            <p>1. White, printed: HOA-BINH TONKIN VII. [handwritten] 1934 A. DE COOMAN</p>
            <p>2. Red, handwritten: Type</p>
            <p> 3. White, handwritten: Type  Onthophagus [printed]  parvidens n.sp. ♂ det.G.Frey,196 [printed] </p>
            <p> 4. Orange: HOLOTYPE  O. parvidens Frey, 1971 [handwritten] S. Tarasov det. 2009 [printed] </p>
            <p> 5. White:  Onthophagus clermonti Paulian [handwritten] S. Tarasov det. 2009 [printed] </p>
            <p>Additional material examined: 1♂, Vietnam, Hoa Binh Lac Tho, leg. A. de Cooman (MNHN)</p>
            <p> Remarks. Apparently, Frey did not take into account the insufficient and obscure description of  O. clermonti based on one female and did not examine the holotype of the former. In the original description (Frey 1971) he compares  O. parvidens with  O. deflexicollis Lansberge, 1883 . The type series of  O. clermonti is comprised only of the holotype female, therefore it is not possible to examine the critical structures of the aedeagus for species identification in  Onthophagus . However, I conclude that the holotypes of  O. clermonti and  O. parvidens are conspecific because they share the same principle diagnostic character states of external morphology (e.g. the groundplan of the head, coloration, pronotum punctation, etc.). Moreover, both type series were found at the same locality Hoa-Binh in Vietnam. </p>
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	https://treatment.plazi.org/id/200A8781FFD9F35600F09073977E21EE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tarasov, Sergey I.	Tarasov, Sergey I. (2010): Seven new synonyms within the genus Onthophagus (Coleoptera: Scarabaeidae) from the Oriental Region. Zootaxa 2566: 45-48, DOI: 10.5281/zenodo.197274
200A8781FFD8F35600F094309613258E.text	200A8781FFD8F35600F094309613258E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus cognatus Boucomont 1921	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Onthophagus cognatus Boucomont, 1921</p>
            <p> Onthophagus cognatus Boucomont (in Boucomont &amp; Gillet), 1921: 30 (type locality: Laos, prov. Tran Ninh [Laos, Xieng Khouang]) </p>
            <p> Onthophagus jucundus Arrow, 1931: 188 syn.n. (type locality: India Or Manipur) </p>
            <p> Onthophagus jucundus Arrow ; Bacchus, 1978: 104 (lectotype designation) </p>
            <p> Onthophagus (Serrophorus) chulapornus Masumoto, Ochi &amp; Hanboonong, 2008 syn.n. (type locality: Thailand, Chaiyaphum Prov., Chulaporn Dam) </p>
            <p> Type material examined. Two syntypes of  O. cognatus (MNHN), male and female bearing geographical label “ LAOS, prov. Tran Ninh” and indicated by A. Boucomont as “  Onthophagus cognatus n.sp”. </p>
            <p> Lectotype of  O. jucundus (BMNH), female bearing following labels: </p>
            <p>1. Round label: LECTOTYPE</p>
            <p>2. White, printed: Doherty</p>
            <p>3. Yellow, handwritten: [first character illegible] 4311</p>
            <p>4. White, printed: India Or Manipur</p>
            <p>5. White, printed: Fry Coll 1905-100.</p>
            <p> 6. White, handwritten:  Onthophagus jucundus . type arrow </p>
            <p> 7. White:  Onthophagus Ψ  jucundus arrow [handwritten], M.E. Bacchus det 1975 [printed] 5 [handwritten]. LECTOTYPE [printed] </p>
            <p> 8. White:  Onthophagus cognatus Bouc. [handwritten] S. Tarasov det. 2009 [printed] </p>
            <p> Also, 1Ψ paralectotype of  O. jucundus bearing geographical label “Birmah Momeit”. </p>
            <p> Remarks. I have examined large material of  O. cognatus and similar species from Indochina.  Onthophagus cognatus is widespread in Indochina and can be easily separated from similar species by external characters and structure of the aedeagus. Although I have not examined type specimens of  O. chulapornus Masumoto, Ochi &amp; Hanboonsong, 2008 , the original description of this species is well illustrated with a photograph of the holotype and illustration of the aedeagus. These illustrations match the character states seen in  O. cognatus and lead me to conclude that these names should be synonymized. It seems that  O. chulapornus was described as a new species simply because the authors did not know the identity of  O. cognatus . Likewise, the lectotype of  O. jucundus was examined and matched  O. cognatus in diagnostic external and aedeagal character states and these names are here synonymized. </p>
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	https://treatment.plazi.org/id/200A8781FFD8F35600F094309613258E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tarasov, Sergey I.	Tarasov, Sergey I. (2010): Seven new synonyms within the genus Onthophagus (Coleoptera: Scarabaeidae) from the Oriental Region. Zootaxa 2566: 45-48, DOI: 10.5281/zenodo.197274
200A8781FFD8F35500F091D09137204E.text	200A8781FFD8F35500F091D09137204E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus gracilipes Boucomont 1914	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Onthophagus gracilipes Boucomont, 1914</p>
            <p> Onthophagus gracilipes Boucomont, 1914: 219 (type locality: Carin Cheba [Myanmar])  Onthophagus (Digithonthophagus) laosensis Frey, 1971: 95 syn.n. (type locality: Laos, Sala Nam Chan Chin) </p>
            <p> Type material examined. Lectotype of  O. gracilipes (MNHN) here designated, male bearing the following labels: </p>
            <p>1. White: Carin Cheba [printed] 500–1000m [handwritten] L. Fea [printed] XII-87 [handwritten]</p>
            <p>2. White, printed: Mus. Civ. Genova</p>
            <p>3. White, printed: MUSÉUM PARIS 1936 COLL. BOUCOMONT</p>
            <p>4. Red, printed: Typus</p>
            <p> 5. White: Boucomont det. 191 [printed] 4 onthoph.  gracilipes n.sp. ♂ </p>
            <p> 6. Orange printed: Lectotypus  Onthophagus gracilipes Boucomont, 1914 S. Tarasov design. 2009 </p>
            <p> Also, 1Ψ paralectotype of  O. gracilipes (MNHN), bearing labels identical to lectotype with the exception of the first label “Carin Ghecu 1300–1400m L. Fea II–III.88 ”. The lectotype is designated in order to preserve the stability of zoological nomenclature according to the article 74.7 (ICZN 1999). </p>
            <p> Holotype of  O. laosensis (NHB), female bearing the following labels: </p>
            <p>1. White, handwritten: Laos: Sala Nam Chan Chin</p>
            <p>2. White printed: VITALIS 1919–1920</p>
            <p> 3. White:  Onthophagus (Digitonthophagus) n.sp. Ψ [handwritten] Dr. V. Balthasar det. [handwritten] 60. [printed] </p>
            <p>4. Red, printed: TYPE</p>
            <p> 5. White: TYPE Ψ [handwritten]  Onthophagus [printed]  Digitonthophagus laosensis n.sp. [handwritten] det.G.Frey, 196 [printed] </p>
            <p> 6. Orange: HOLOTYPE  Onthophagus laosensis Frey, 1971 [handwritten] S. Tarasov det. 2009 [printed] </p>
            <p> 7. White:  Onthophagus gracilipes Bouc. [handwritten] S. Tarasov det. 2009 [printed] </p>
            <p> Remarks. It seems that Frey (1971) did not thoroughly examine the description of  O. gracilipes and did not see the type series of the former. In the original description he compares  O. laosensis with  O. troniceki Balthasar, 1933 . The type series of Frey’s  O. laosensis includes only a female holotype, however that was not an obstacle for identification because the type series of  O. gracilipes includes a female. I synonymize these two names because the type series of  O. laosensis and  O. parvidens are conspecific based on their same principle diagnostic character states of external morphology: the unique color pattern of the elytra and body, the unique groundplan of head, and the identical pattern of pronotum punctation. Additionally I have examined large material of  O. gracilipes from other localities in Indochina and these also match the two series of both names in the characters detailed above. </p>
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	https://treatment.plazi.org/id/200A8781FFD8F35500F091D09137204E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tarasov, Sergey I.	Tarasov, Sergey I. (2010): Seven new synonyms within the genus Onthophagus (Coleoptera: Scarabaeidae) from the Oriental Region. Zootaxa 2566: 45-48, DOI: 10.5281/zenodo.197274
200A8781FFDBF35500F0949091A725EE.text	200A8781FFDBF35500F0949091A725EE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus orientalis Harold 1868	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Onthophagus orientalis Harold, 1868</p>
            <p> Onthophagus orientalis Harold, 1868: 83 (type locality: original description “ Hongkong, Bengal”; lecotype designated by Zunino (1976) “ Hong Kong ”) </p>
            <p> Onthophagus orientalis Harold ; Zunino, 1976: 79 (lectotype designation) </p>
            <p> Onthophagus perroti Paulian, 1978: 40 syn.n. (type locality: Saigon [Vietnam]) </p>
            <p> Type material examined. Lectotype of  O. orientalis (MNHN), male bearing the following labels: </p>
            <p>1. White, printed: Ex Musaeo E Harold</p>
            <p>2. White, printed: Hong Kong</p>
            <p> 3. Red: Lecto [handwritten] TYPUS [printed]  Onthophagus orientalis Har. [handwritten] M. ZUNINO 19 [printed] 76 [handwritten] </p>
            <p> Also, 1Ψ paralectotype of  O. orientalis (MNHN), from the same locality. </p>
            <p> Holotype of  O. perroti (MNHN), male bearing the following labels: </p>
            <p>1. White, printed: MUSÉUM PARIS SAIGON 1950 [handwritten] J. BARBIER</p>
            <p> 2. White:  Onthophagus perroti n.sp. [handwritten] R. Paulian det. [printed] </p>
            <p>3. Red, printed: HOLOTYPE</p>
            <p> 4. Orange:  Onthophagus perroti Paulian, 1978 [handwritten] S. Tarasov det. 2009 [printed] </p>
            <p> 5. White:  Onthophagus orientalis Har. [handwritten] S. Tarasov det. 2009 [printed] </p>
            <p> Additional material examined: 1♂ (MNHN) bearing the same geographical label as the holotype and labeled by someone later as a paratype of  O. perroti Paulian, 1978 . The original description of  O. perroti was based on a single specimen (Paulian 1978) and therefore this paratype label is erroneous. </p>
            <p> Remarks. Based on my observations,  O. perroti represents individuals exhibiting the minor form of the earlier described and common Oriental species  O. orientalis Harold, 1868 . The character states of external morphology of  O. perroti (e.g. vertex horn, pronotum punctation, etc.) are compatible with the range of variation of  O. orientalis . Morover, the holotype of  O. perroti has the same aedeagus as the lectotype of  O. orientalis . </p>
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	https://treatment.plazi.org/id/200A8781FFDBF35500F0949091A725EE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tarasov, Sergey I.	Tarasov, Sergey I. (2010): Seven new synonyms within the genus Onthophagus (Coleoptera: Scarabaeidae) from the Oriental Region. Zootaxa 2566: 45-48, DOI: 10.5281/zenodo.197274
200A8781FFDBF35500F09030972C260E.text	200A8781FFDBF35500F09030972C260E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus pseudojavanus Paulian 1931	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Onthophagus pseudojavanus Paulian, 1931</p>
            <p> Onthophagus pseudojavanus Paulian, 1931: 272 (type locality: Hoa Binh, Tonkin [Vietnam])  Onthophagus demaak Masumoto, 1989: 93 syn.n. (type locality: Doi Suthep/Pui, Chiang Mai Prov. [Thailand]) </p>
            <p> Type material examined. Holotype of  O. pseudojavanus (MNHN), male bearing the following labels: </p>
            <p>1. White, printed: Hoa-Binh (Tonkin) (A.de Cooman) Coll.J.Clermont</p>
            <p>2. Red, printed: TYPE</p>
            <p> 3. White, handwritten:  Onthophagus sp.? V. de javanus Lansb. </p>
            <p> 4. White, handwritten:  O. pseudojavanus type </p>
            <p> 4. Orange: HOLOTYPE  Onthophagus pseudojavanus Paulian, 1931 [handwritten] S. Tarasov det. 2009 [printed] </p>
            <p> Paratype male of  O. demaak (MNHN), with locality label “Doi Suthep/Pui Chiang Mai Prov. 30.VII.–5.VIII.1988 K. MASUMOTO leg.” </p>
            <p> Additional material examined: 1♂ and 1Ψ (MNHN) bearing the same geographical label as that of the holotype of  O. pseudojavanus Paulian, 1931 and indicated by R. Paulian as “  O. pseudojavanus ? m.”. </p>
            <p> Remarks. The examined paratype of  O. demaak is conspecific with the type series of  O. pseudojavanus because they share the same aedeagus, structure of the head, vertex horn, pronotum punctation and body coloration. It seems that  O. demaak was described as a new species simply because the author did not know the identity of  O. pseudojavanus . </p>
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	https://treatment.plazi.org/id/200A8781FFDBF35500F09030972C260E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tarasov, Sergey I.	Tarasov, Sergey I. (2010): Seven new synonyms within the genus Onthophagus (Coleoptera: Scarabaeidae) from the Oriental Region. Zootaxa 2566: 45-48, DOI: 10.5281/zenodo.197274
200A8781FFDBF35400F0925093A4230E.text	200A8781FFDBF35400F0925093A4230E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus vladimiri Frey 1957	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Onthophagus vladimiri Frey, 1957</p>
            <p> Onthophagus vladimiri Frey, 1957: 687 (type locality: Süd Indien, Anamalai Hills, Cinchona)  Onthophagus anamalaiensis Balthasar, 1974: 183 syn.n. (type locality: S. Indien, Anamalai Hills, Cinchona) Type material examined. One male and one female paratype of  O. vladimiri (NMP) bearing the following two geographical labels: </p>
            <p>1. Anamalai Hills Cinchona 3500 ft.V.56</p>
            <p>2. Süd Indien leg. Nathan</p>
            <p> Holotype of  O. anamalaiensis (NMP), male bearing the following labels: </p>
            <p>1. White, printed: Anamalai Hills Cinchona S. Ind. 3500 ft.1959</p>
            <p> 2. Rose: Onth  anamalaiensis Holo Balt [handwritten] Allotypus [printed, crossed with hand] </p>
            <p> 3. White:  Onthophagus vladimiri Frey, 1957 [handwritten] S. Tarasov det. 2009 [printed] </p>
            <p> Remarks. The examined holotype of  O. anamalaiensis is conspecific with the type series of  O. pseudojavanus because they share the same aedeagus and principle diagnostic characters of the external morphology (structure of head and vertex horns, pronotum punctation, body coloration, etc.). Moreover, both type series were collected at the same locality in southern India. It seems that  O. anamalaiensis was described as a new species because the author did not know the identity of  O. vladimiri . </p>
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	https://treatment.plazi.org/id/200A8781FFDBF35400F0925093A4230E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Tarasov, Sergey I.	Tarasov, Sergey I. (2010): Seven new synonyms within the genus Onthophagus (Coleoptera: Scarabaeidae) from the Oriental Region. Zootaxa 2566: 45-48, DOI: 10.5281/zenodo.197274
