taxonID	type	description	language	source
A0EB04E2D8BC1E267B83AA12D002D2C8.taxon	diagnosis	Diagnosis. Paronellidae with finely denticulate scales covering dorsum of head and body, and ventral face of furcula; Ant. 4 sometimes annulated, never subdivided in two; labial seta L 2 normal, not reduced; eyes 0 - 8; EOS present; Abd. 2 - 4 with 2, 3, 3 bothriothricha; manubrium without spines, dens with 1 - 2 rows of spines; mucro short, with 3 - 5 more or less evenly spaced teeth.	en	Soto-Adames, Felipe N., Taylor, Steven J. (2013): The dorsal chaetotaxy of Trogolaphysa (Collembola, Paronellidae), with descriptions of two new species from caves in Belize. ZooKeys 323: 35-74, DOI: http://dx.doi.org/10.3897/zookeys.323.4950, URL: http://dx.doi.org/10.3897/zookeys.323.4950
FF2F56857627DAFBB37849B35FB7FD6E.taxon	diagnosis	Diagnosis. Trogolaphysa giordanoae sp. n. is unique among species with 6 - 8 eyes in having 5 dorsal head macrochaetae, 3 metathoracic macrochaetae and 4 inner macrochaetae on Abd. 4. Among species with known dorsal chaetotaxy, the new species is most similar to Trogolaphysa riopedrensis, but the two species are easily distinguished by the combination of characters given above and by the presence of a relatively shorter mucro in the new species (Table 2). Additional diagnostic characters distinguishing the new species from all other New World Trogolaphysa with 6 - 8 eyes and capitate / spatulate tenent hair are presented in Table 2. Among the species described before the introduction of chaetotaxy, the new species is most similar to Trogolaphysa distinguenda (Denis, 1931), but the two species can be separated by the presence of a relatively long mucro with 5 teeth in distinguenda, and a 4 - toothed short mucro in Trogolaphysa giordanoae sp. n. Trogolaphysa belizeana is the only other New World Trogolaphysa with 3 metathoracic macrochaetae. However, Trogolaphysa belizeana is a troglobiont (sensu Sket 2008, Culver and Pipan 2009) - blind, with long antennae and modified ungues.	en	Soto-Adames, Felipe N., Taylor, Steven J. (2013): The dorsal chaetotaxy of Trogolaphysa (Collembola, Paronellidae), with descriptions of two new species from caves in Belize. ZooKeys 323: 35-74, DOI: http://dx.doi.org/10.3897/zookeys.323.4950, URL: http://dx.doi.org/10.3897/zookeys.323.4950
FF2F56857627DAFBB37849B35FB7FD6E.taxon	description	Description. Size. Body length up to 2.1 mm. Color. Pattern, if any, obscured by green dye present in the alcohol in which specimens were preserved (Fig. 1). Scale distribution. Scales dark brown, present on Ant. 1 - 2 and base of Ant. 3, more abundant on dorsal face than on ventral face of segment. Scales absent from ventral tube, legs and dorsal face of manubrium. Head. Antenna / cephalic diagonal ratio 2.0 - 2.5 (Fig. 1). Apical bulb of Ant. 4 absent; subapical sensillum capitate (Fig. 2), fully contained in circular depression; guard sensillum absent. Sense organ of Ant. 3 (Fig. 3) with sensilla 1 and 4 acumi nate, thin-walled and translucent; sensillum 5 acuminate, dark (light dense), shorter than 1 and 4; sensilla 2 - 3 wide, leaf-like, resting in shallow grooves. Eyes 6 + 6 (Fig. 4), chaetotaxy of eyepatch well with 4, sometimes 6 ciliate setae, and 1 seta posterior to eye F. Head dorsally with 5 macrochaetae (A 0, A 2, M 3, Pa 5 and Pm 3 - Figs 4 - 5). Series M with 2 setae (M 3 - 4); series S with 5 setae (S 1 - 5); seta M 0 seen only in one individual; S 0 absent. Prelabral setae serrate (Fig. 6). Labral setae smooth: setae in rows A and C subequal; seta B 2 distinctly shorter than setae B 0 and B 1 (Fig. 7). Distal margin of labrum with 1 + 1 medial hooks, papillae absent (Fig. 8). Apical and subapical setae of maxillary palp smooth; sublobular plate with 2 seta-like appendages. Lateral process of labial papilla E weakly bent dorsally, barely reaching apex of papilla (Fig. 9). Labial triangle setae as M 1 M 2 rEL 1 - 2 A 1 - 5 (Fig. 10); r short, stout and sparsely ciliate; L 1 inserted close to E and distant from L 2 when compared to other entomobryoids (Fig. 11). Postlabium covered by setae and scales, all postlabial setae ciliate, modified setae absent. Columns ICELO with 42221 setae (Fig. 11): col umn I with posterior seta detached from main group and much longer than anterior setae. Ventral cervical setae usually 8 + 8. Body. Mesothoracic hood not developed. Complete dorsal macrochaetae as 32 / 73 / 0245 + 0 + 9. Mesothorax with 1 anterior (a 5) and 6 posterior (p 3 complex) macrochaetae arranged as is typical for genus (Fig. 12); microchaetae m 2, m 4, p 1, p 2, p 4, p 5 and p 6 present. Inner chaetotaxy of metathorax with 3 macro- and 1 microchaetae (Fig. 13). First abdominal segment with 1 anterior (a 6) and 4 posterior setae arranged in a single row (Fig. 14). Second abdominal segment (Fig. 15) inner bothriotrix with 3 fan-shaped setae, one microsensillum and macrochaeta m 3; outer bothriotrix with 3 fan-shaped setae and macrochaeta m 5; setae a 6, m 6 and p 5 present. Third abdominal segment (Fig. 16) inner bothriotrix complex with 2 fan-shaped setae, 1 sensillum, and macrochaeta m 3; external bothriotricha with 7 fan-shaped setae, and macrochaetae am 6, pm 6 and p 6; sensillum d 2 present, inserted near pm 6. Fourth abdominal segment with 5 inner and 9 outer macrochaetae (Fig. 17): large inner macrochaetae A 4, A 5, B 4, and B 5 present; B 6 a small macrochaeta; large outer macrochaetae D 3, E 2, E 3, F 1, F 2, and F 3 present; macrochaetae E 4, F 4 and one other seta probably homologous to Fe 4, small. Anterior and medial bothriotricha with 7 and 3 fan-shaped supplementary setae, respectively. Posterior bothriotrix, corresponding to D 4, without associated supplementary setae. Posterior setae 19 - 21 + 19 - 21. Intersegmental membrane between Abd. 4 - 5 with 4 - 10 lenticular organs (as in Trogolaphysa riopedrensis, Fig. 60). Legs. Trochanteral organ with up to 36 setae. Metathoracic claw complex as in Fig. 18. Tenent hair weakly spatulate. Smooth posterior setae on metathoracic legs 0.76 x as long as unguiculus. Unguis with 4 inner teeth: 1 basal tooth sometimes ap pearing slightly larger than other, both paired teeth ending near middle of inner edge; proximal unpaired tooth as large as basal paired teeth, ending on distal half of inner edge; distal unpaired tooth smallest of all inner teeth and ending on distal fourth of inner edge. Outer tooth ending on basal quarter of outer ungual edge. Unguiculus lanceolate, with outer margin serrate. Ventral tube. Anterior face with 3 + 3 or 4 + 4 distal macrochaetae; lateral and posterior setae not seen clearly. Furcula. Dens with 2 rows of ciliate spines: inner row with 35 - 42 spines; outer row with 25 - 28 spines. Basal outer spines longest (Figs 19 - 20). Mucro with 4 short, stout teeth (Fig. 21), ratio mucro length / width of dens tip 1.2 - 1.8 x; basal outer tooth reaches to at least half length of basal inner tooth.	en	Soto-Adames, Felipe N., Taylor, Steven J. (2013): The dorsal chaetotaxy of Trogolaphysa (Collembola, Paronellidae), with descriptions of two new species from caves in Belize. ZooKeys 323: 35-74, DOI: http://dx.doi.org/10.3897/zookeys.323.4950, URL: http://dx.doi.org/10.3897/zookeys.323.4950
FF2F56857627DAFBB37849B35FB7FD6E.taxon	etymology	Etymology. This species is dedicated to Rosanna Giordano, the senior author's wife, for her years of support and contributions to science.	en	Soto-Adames, Felipe N., Taylor, Steven J. (2013): The dorsal chaetotaxy of Trogolaphysa (Collembola, Paronellidae), with descriptions of two new species from caves in Belize. ZooKeys 323: 35-74, DOI: http://dx.doi.org/10.3897/zookeys.323.4950, URL: http://dx.doi.org/10.3897/zookeys.323.4950
FF2F56857627DAFBB37849B35FB7FD6E.taxon	distribution	Distribution. The species is known only fromBelize	en	Soto-Adames, Felipe N., Taylor, Steven J. (2013): The dorsal chaetotaxy of Trogolaphysa (Collembola, Paronellidae), with descriptions of two new species from caves in Belize. ZooKeys 323: 35-74, DOI: http://dx.doi.org/10.3897/zookeys.323.4950, URL: http://dx.doi.org/10.3897/zookeys.323.4950
AFDB54D96169FD6F35FC33EAF47CC404.taxon	diagnosis	Diagnosis. Trogolaphysa jacobyi sp. n. is the only member of the genus that is blind, has 3 - toothed mucro and unguis, and has a single macrochaeta on the metathorax. Trogolaphysa belizeana is the only other New World Trogolaphysa lacking eyes and having a 3 - toothed mucro, but it differs from Trogolaphysa jacobyi sp. n. in having 3 metathoracic macrochaetae (1 in Trogolaphysa jacobyi sp. n.), in the arrangement and identity of inner macrochaetae on Abd. 4 (cf. Figs 38, 49 see discussion below), in having few postlabial scales (absent in Trogolaphysa jacobyi sp. n.) and setae (many in Trogolaphysa jacobyi sp. n., cf. Figs 30, 46), in the presence of sensillum d 2 on Abd. 3 (absent in Trogolaphysa jacobyi sp. n.), in the absence of unpaired ungual teeth (1 tooth in Trogolaphysa jacobyi sp. n.) and in having a typical lanceolate unguiculus (basally swollen in Trogolaphysa jacobyi sp. n.). Table 3 provides a list of characters that distinguish Trogolaphysa jacobyi sp. n. from all other New World Trogolaphysa lacking eyes and having paired basal ungual teeth inserted near the basal fourth of the inner edge.	en	Soto-Adames, Felipe N., Taylor, Steven J. (2013): The dorsal chaetotaxy of Trogolaphysa (Collembola, Paronellidae), with descriptions of two new species from caves in Belize. ZooKeys 323: 35-74, DOI: http://dx.doi.org/10.3897/zookeys.323.4950, URL: http://dx.doi.org/10.3897/zookeys.323.4950
AFDB54D96169FD6F35FC33EAF47CC404.taxon	description	Description. Size. Body length up to 2.0 mm. Color. Living specimens yellowish, with pigment only on a small eyepatch and mesothorax (Fig. 24). Specimens in alcohol white, without trace of pigment. Scale distribution. Scales transparent, present on Ant. 1 - 2. Scales absent from postlabial region of head, ventral tube, legs and dorsal face of manubrium. Head. Antenna / cephalic diagonal ratio up to 5.8 (Fig. 24). Fourth antennomere with incomplete but clear constriction near middle, with many shallow whorls of setae (Fig. 25); apical bulb absent; subapical sensillum not seen. Sense organ of Ant. 3 with sensilla 1 and 4 short, acuminate, thin-walled and translucent; sensillum 5 acuminate, dark and shorter than 1 and 4; sensilla 2 - 3 broad, leaf-like, resting in shallow grooves. Eyes not seen on slide-mounted specimens, but 1 - 2 pigment patches visible in life (Fig. 24). Head dorsally with 8 macrochaetae (A 0, A 2, A 3, M 3, S 3, S 5, Pa 5 and Pm 3 - Fig. 26). Seta M 4 displaced laterally towards cephalic sulcus. Series S with setae S 1 - 5; S 0 absent, macrochaeta S 3 displaced anteriorly, away from cephalic sulcus (cf. Figs 5, 26). Prelabral and all labral setae smooth: setae within row A and C subequal; seta B 2 shorter than B 0 and B 1 (Fig. 27). Distal margin of labrum smooth, papillae absent. Apical and subapical setae of maxillary palp smooth; sublobular plate without seta-like appendages. Lateral process of labial papilla E weakly bent dorsally and not nearly reaching apex of papilla (Fig. 28). Labial triangle setae as M 1 M 2 rEL 1 - 2 A 1 - 5 (Fig. 29), seta M 1 ciliate, all others smooth; r short; A 2 close to r, L 1 close to E and distant from L 2. Postlabium without scales, polychaetotic, uniformly covered with many large and small, weakly ciliate or smooth setae (Fig. 30); modified setae absent. Columns ICELO ill defined due to polychaetosis. Ventral cervical setae usually 6 + 6. Body. Mesothoracic hood not developed. Complete dorsal macrochaetae as 62 / 41 / 0244 + 0 + 9 - 11. Mesothorax with 1 anterior (a 5) and usually 3 posterior (p 1 - 3) macrochaetae forming an arch (Fig. 31); some individuals with only mesothoraxic macrochaeta p 2 (Fig. 32); microchaetae m 1, m 2, m 4, p 4 and p 5 present. Metathorax with 1 macro- and 5 microchaetae (Fig. 33). First abdominal segment seta a 6 absent; 4 posterior setae arranged in a single row (Fig. 34 - 36). Inner bothriotrix complex of Abd. 2 with 3 fan-shaped setae, one microsensillum and macrochaeta m 3; outer bothriotrix with 3 fan-shaped setae and macrochaeta m 5; setae a 6, m 6 and p 5 present. Inner bothriotrix complex of Abd. 3 with 3 fan-shaped setae, one sensillum and macrochaetae m 3; external bothriotrichal complex (Fig. 37) with 6 - 7 fan-shaped setae, macrochaetae am 6, pm 6 and p 6; sensillum d 2 absent. Fourth abdominal segment with 4 inner (Fig. 38) and 9 - 11 outer (Fig. 39) macrochaetae: inner macrochaetae A 5, B 4, and B 5 large, B 6 small; B 5 displaced towards A 6 instead of B 6; (Fig. 38). Outer macrochaetae D 3, E 2, E 3, F 1, and F 2 large; small outer macrochaetae E 4, F 3, F 4 and 3 others probably belonging to series Fe present. Abd. 4 anterior and medial bothriotricha with 4 and 2 fan-shaped supplementary setae, respectively (Fig. 39). Posterior bothriotrix corresponds to D 4, without associated supplementary setae. Posterior setae 6 - 7 + 6 - 7. Intersegmental membrane between Abd. 4 - 5 with 4 - 7 lenticular organs. Legs. Trochanteral organ with up to 25 setae. Claw complex as in Figs 41 - 42. Tenent hair acuminate, longer on L 1 than L 3. Smooth posterior setae on metathoraxic legs as long as unguiculus. Unguis with 3 inner teeth: basal teeth small, subequal and ending on basal fourth of inner edge; unpaired tooth distinctly larger than basal teeth, ending near middle of inner ungual edge. Outer tooth absent on all claws; lateral teeth present only on pro- and mesothoracic legs, and ending on basal quarter of outer edge of unguis (Fig. 41). Unguiculus basally swollen, with basal fifth of outer margin weakly serrate.	en	Soto-Adames, Felipe N., Taylor, Steven J. (2013): The dorsal chaetotaxy of Trogolaphysa (Collembola, Paronellidae), with descriptions of two new species from caves in Belize. ZooKeys 323: 35-74, DOI: http://dx.doi.org/10.3897/zookeys.323.4950, URL: http://dx.doi.org/10.3897/zookeys.323.4950
AFDB54D96169FD6F35FC33EAF47CC404.taxon	distribution	Distribution. The species is known only from caves in southern Belize	en	Soto-Adames, Felipe N., Taylor, Steven J. (2013): The dorsal chaetotaxy of Trogolaphysa (Collembola, Paronellidae), with descriptions of two new species from caves in Belize. ZooKeys 323: 35-74, DOI: http://dx.doi.org/10.3897/zookeys.323.4950, URL: http://dx.doi.org/10.3897/zookeys.323.4950
AFDB54D96169FD6F35FC33EAF47CC404.taxon	etymology	Etymology. This species is dedicated to JoAnn Jacoby, the junior author's wife, in gratitude for her enthusiasm and assistance in the planning and execution of field-work in the caves of Belize and in many earlier excursions.	en	Soto-Adames, Felipe N., Taylor, Steven J. (2013): The dorsal chaetotaxy of Trogolaphysa (Collembola, Paronellidae), with descriptions of two new species from caves in Belize. ZooKeys 323: 35-74, DOI: http://dx.doi.org/10.3897/zookeys.323.4950, URL: http://dx.doi.org/10.3897/zookeys.323.4950
