taxonID	type	description	language	source
2050F30FBA4BFFC3FE3F20FCFC43FC72.taxon	discussion	Remarks Bouchet et al. (2017) treated Hypselostomatinae as a subfamily of Gastrocoptidae Pilsbry, 1918. The systematic relationships of the genera classified in this (sub) family are largely unclear. Here, to be consistent with our previous papers (i. e. Páll-Gergely et al. 2015, 2020, 2022), we use Hypselostomatidae at the family level. According to Pilsbry (1917), Boysidia Ancey, 1881 possesses a concrescent angular and parietal lamella, whereas they are separate in Bensonella and Paraboysidia. This systematics has been widely used since (i. e. Schileyko 1998; Panha and Burch 2005). However, it was already noted by Pilsbry (1917) that there are species showing intermediate character states. For example, Boysidia strophostoma (Möllendorff, 1885) possesses clearly separated angular and parietal lamellae, but is still classified in Boysidia. Solving the homologies of apertural barriers and the systematics of the related hypselostomatid genera is beyond the scope of the present paper, but should be done as soon as possible.	en	Páll-Gergely, Barna, White, Tom S. (2023): Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History 56 (45 - 48): 2011-2029, DOI: 10.1080/00222933.2022.2152750, URL: http://dx.doi.org/10.1080/00222933.2022.2152750
2050F30FBA4BFFC5FE89254EFCDCF92A.taxon	description	Boysidia (Paraboysidia) Pilsbry, 1917 (in Pilsbry 1916 – 1918): 174, 201.	en	Páll-Gergely, Barna, White, Tom S. (2023): Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History 56 (45 - 48): 2011-2029, DOI: 10.1080/00222933.2022.2152750, URL: http://dx.doi.org/10.1080/00222933.2022.2152750
2050F30FBA4BFFC5FE89254EFCDCF92A.taxon	type_taxon	Type species Pupa plicidens Benson, 1849 (Fig. 3 (a – e) herein), by original designation.	en	Páll-Gergely, Barna, White, Tom S. (2023): Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History 56 (45 - 48): 2011-2029, DOI: 10.1080/00222933.2022.2152750, URL: http://dx.doi.org/10.1080/00222933.2022.2152750
2050F30FBA4BFFC5FE89254EFCDCF92A.taxon	discussion	Remarks According to Pilsbry ̾ s (1917) original description of Paraboysidia, this genus (at that time a subgenus of Boysidia) differs from Bensonella by having normal (not hooked) apertural barriers. However, this claim is incorrect because the type species of Bensonella (Pupa plicidens) has normal barriers. The type species of Paraboysidia (Boysidia paviei Bavay and Dautzenberg, 1912, fig. 3 (f – k )) is very similar to that of Bensonella (Pupa plicidens) in general shell shell and aperture shape, and in the arrangement of the apertural barriers. The main difference is the presence of some additional ̍ supplementary ̾ apertural folds (i. e. tiny plicae between main plicae) in P. plicidens, which does not seem to justify the distinction of two genera. Therefore, herein Paraboysidia is treated as a junior synonym of Bensonella. Gittenberger et al. (2021) also treated Paraboysidia as a synonym of Bensonella but did not explain their decision to do so. Bensonella species with hooked and normal barriers in the Himalaya (i. e. B. hooki sp. nov. and B. plicidens) are very similar to each other in all other shell characters. Moreover, hooked and normal apertural barriers are characteristic of otherwise extremely similar and close-occurring species in the genera Acinolaemus F. G. Thompson and Upatham 1997; Anauchen Pilsbry, 1917; and Hypselostoma Benson, 1856 (Vermeulen et al. 2007, 2019). This suggests that the hooked or normal apertural barriers are subjects of rapid, probably adaptive (see Wada and Chiba 2013) evolutionary changes. Since it is probable that the hooked apertural barriers have been evolved multiple times in different groups within the Hypselostomatidae, it is unlikely that species with apertural barriers would form a monophyletic unit. Therefore, it is not recommended to use this character state to distinguish between genera. We thus classify species with both hooked (B. hooki sp. nov., B. lakainguta Hwang, 2014) and normal apertural barriers (i. e. B. plicidens) in Bensonella. In the last three decades, several Boysidia (Bensonella) species from China have been newly described or transferred into the (sub) genus: Boysidia dilamellaris D. - N. Chen, Y. - H. Liu and W. - X. Wu, 1995 (transferred to Bensonella by Chen et al. 1999); Boysidia (Bensonella) jinpingensis M. Tian, B. Fao and Y. - X. Chen, 2015; Boysidia (Bensonella) nanjiangensis Z. - L. Zhang, W. - H. Zhang and T. - C. Luo, 2011; Boysidia (Bensonella) qingliangfengensis F. Fang, J. Wang and Y. Chen, 2015; Boysidia (Bensonella) shilinensis D. - N. Chen, M. Wu and G. - Q. Zhang, 1999; Boysidia (Bensonella) tianxingqiaoensis T. - C. Luo, D. - N. Chen and G. - Q. Zhang, 2000 (Luo et al. 2000); Boysidia (Bensonella) tongguanensis D. - N. Chen and W. - H. Zhang, 2002; Boysidia (Bensonella) xingyinensis Y. - H. Guo, W. - C. Zhou and T. - C. Luo, 2006 (Guo et al. 2006); and Boysidia (Bensonella) xiuwenensis W. - H. Zhang, T. - C. Luo and W. - C. Zhou, 2010. Of these, B. dilamellaris, B. nanjiangensis, B. shilinensis, B. tongguanensis and B. xiuwenensis were classified as within Boysidia Ancey, 1881 (without subgeneric classification) by Zhang et al. (2014). We agree with this classification because these species are similar in shell shape and apertural dentition (i. e. fused angular and palatal lamella) to the type species, Pupa dorsata Ancey, 1881. We further classify B. jinpingensis in Boysidia for the same reason (i. e. that species is a member of Boysidia and not Bensonella). Boysidia (Bensonella) tianxingqiaoensis (treated as a Boysidia species by Zhang et al. 2014) and Boysidia (Bensonella) xingyinensis are here both transferred to Gyliotrachela Tomlin, 1930 due to the free and keeled last whorl and the separate parietal and angular lamella all being similar to those of the type species of that genus: Hypselostoma hungerfordianum Möllendorff, 1891. Boysidia (Boysidia) fengxianensis D. - N. Chen, Y. - H. Liu and W. - X. Wu, 1995 was transferred to Bensonella by Schileyko (1998) and to Boysidia (Bensonella) by Zhang et al. (2014). Judging as best we can from the poor-quality drawing in the original description (mostly the three parietal lamellae), we agree that this species belongs to Bensonella. Lastly, the Sumatran Bensonella karoensis Maassen, 1999 is retained in Bensonella herein, due to the arrangement of the apertural barriers (separate angular and parietal lamella, two palatal lamellae, a basal and a columellar lamella) that largely match with the situation in Pupa plicidens Benson, 1849 and Boysidia paviei Bavay and Dautzenberg, 1912, the type species of Bensonella and Paraboysidia, respectively.	en	Páll-Gergely, Barna, White, Tom S. (2023): Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History 56 (45 - 48): 2011-2029, DOI: 10.1080/00222933.2022.2152750, URL: http://dx.doi.org/10.1080/00222933.2022.2152750
2050F30FBA4FFFCCFE5021C1FCF3FA3F.taxon	description	(Figures 3 (a – e), 4, 5, 9 (a, b )).	en	Páll-Gergely, Barna, White, Tom S. (2023): Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History 56 (45 - 48): 2011-2029, DOI: 10.1080/00222933.2022.2152750, URL: http://dx.doi.org/10.1080/00222933.2022.2152750
2050F30FBA4FFFCCFE5021C1FCF3FA3F.taxon	description	Boysidia plicidens – Sowerby, 1876: pl. 16, fig. 151 Pupa (Odontocyclas) plicidens – Pfeiffer, 1879 (1878 – 1881): 350	en	Páll-Gergely, Barna, White, Tom S. (2023): Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History 56 (45 - 48): 2011-2029, DOI: 10.1080/00222933.2022.2152750, URL: http://dx.doi.org/10.1080/00222933.2022.2152750
2050F30FBA4FFFCCFE5021C1FCF3FA3F.taxon	description	Bensonella landourensis and plicidens – Budha and Backeljau 2017 (partim: figs 2 B, 3, 5 A and F show B. plicidens) Types examined UMZC I. 103325. A, lectotype of P. plicidens (selected herein from a series of 15 syntypes in the UMZC), R. McAndrew coll. (ex- W. H. Benson coll.), Rockville, Landour, Himalaya; NHMUK 1954.6. 2.937, paralectotype of P. plicidens, J. S. Hawkins coll. (ex- W. H. Benson coll.), W Himalaya (labelled as paratype); ANSP 16721, holotype of B. landourensis (photos examined: see Figure 5).	en	Páll-Gergely, Barna, White, Tom S. (2023): Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History 56 (45 - 48): 2011-2029, DOI: 10.1080/00222933.2022.2152750, URL: http://dx.doi.org/10.1080/00222933.2022.2152750
2050F30FBA4FFFCCFE5021C1FCF3FA3F.taxon	materials_examined	Additional material examined NHMUK 1856.9.15.69 / A, 36 shells, Capt. T. Hutton coll., Moussorie; NHMUK 20191111 / A, 10 shells, Cherra, India, mixed sample that includes NHMUK 20191111 / B, identified as B. hooki sp. nov.; NHMUK 1903.7.1.2882 / B, six shells, Godwin-Austen coll., Moussorie, NW Himalaya, mixed sample that includes NHMUK 1903.7.1.2882 / C, identified as B. hooki sp. nov.; NHMUK 1906.2.2.190 / B, 7 shells, W. T. Blanford coll., Cherra Poonje, Khasi Hills, mixed sample that includes NHMUK 1906.2.2.190 / C, identified as B. hooki sp. nov.	en	Páll-Gergely, Barna, White, Tom S. (2023): Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History 56 (45 - 48): 2011-2029, DOI: 10.1080/00222933.2022.2152750, URL: http://dx.doi.org/10.1080/00222933.2022.2152750
2050F30FBA4FFFCCFE5021C1FCF3FA3F.taxon	diagnosis	Extended diagnosis Shell triangular (pyramidal), slightly ovoid, height somewhat variable; shell colour greyishyellowish and corneous; sculpture with inconspicuous growth lines and very fine spiral striation (its strength variable, in some shells nearly absent); aperture with numerous barriers; parietal wall with a parietal and an angular lamella, both being lower at their middles, and a third, short, deeply situated one between parietal lamella and columellarparietal junction; palatal wall with three long main plicae with elevated central parts, usually (but not always) with additional short plicae between them; columellar lamella similar to palatal plicae; between the lowest palatal plicae and the columellar lamella there are two short plicae; an additional, blunt tubercle sits on the edge of the palatal lip, at the position where the palatal lip forms a little sinus; lower part of columellar lip with two very low, blunt denticles (swellings). Differential diagnosis For differences with B. hooki sp. nov., see under that species. Bensonella plicidens differs from B. lakainguta in the following traits: shell shape is similar, but B. plicidens is wider triangular; the apex is less pointed; the sculpture is clearly densely spirally striated, whereas B. lakainguta is smooth; the arrangement of the apertural barriers is similar, but the barriers of B. plicidens are elongated ridges, which are highest at their middle parts, whereas in B. lakainguta the central part of each barrier is a hook pointing anteriorly (outside of the shell), with an anteriorly elongated low ridge before the hook.	en	Páll-Gergely, Barna, White, Tom S. (2023): Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History 56 (45 - 48): 2011-2029, DOI: 10.1080/00222933.2022.2152750, URL: http://dx.doi.org/10.1080/00222933.2022.2152750
2050F30FBA4FFFCCFE5021C1FCF3FA3F.taxon	discussion	Remarks The original spelling of B. landourensis was B. landurensis by Pilsbry (1915), although the type locality was spelled Landour (town in Mussoorie, India). Therefore, the spelling ̍ landourensis ̾ by van Benthem Jutting (1950) is a justified emendation, and ̍ landurensis ̾ is an incorrect original spelling (under ICZN Art. 32.5). Pilsbry (1915) distinguished B. landourensis from B. plicidens by the uninterrupted angular lamella in the former. Additional differences were also mentioned, such as the length of the angular lamella, which does not emerge to the edge of the peristome (i. e. it stops before the peristome edge) in B. landourensis but does in B. plicidens, and the parietal lamella, which is shorter in B. plicidens than in B. landourensis. Although the original description of B. landourensis does not mention hooked apertural barriers in B. plicidens, it is clear that Pilsbry ̾ s (1915) new species was not compared with correctly identified B. plicidens but with the species that is described here as new (B. hooki sp. nov.), because B. hooki sp. nov. has an interrupted angular lamella, whereas that of B. plicidens is not interrupted. Comparison of the holotype of B. landourensis with syntypes of B. plicidens reveal no significant differences, other than the somewhat more slender shell of B. landourensis, which can be explained by intraspecific variability. Pilsbry (1915) mentioned that the single shell of B. landourensis (= B. plicidens in our current understanding) was found amongst shells of B. plicidens (= B. hooki sp. nov. in our current understanding), all of which had been received from Benson. We were unable to examine the shells studied by Pilsbry, other than the holotype of B. landourensis. It is possible that Benson had both B. plicidens (a species with normal barriers) and B. hooki sp. nov. (a species with hooked barriers) when he described B. plicidens. However, neither Benson (1849) nor subsequent authors mentioned hooked barriers as a character of that species, and the syntypes in the original Benson collection in the UMZC all have normal barriers (Preece et al., in litteris). Even the name ̍ plicidens ̾ indicates that the apertural barriers were plica-like. We do not know how Pilsbry received Benson ̾ s original specimens, or whether it is really true that he did. It is remarkable that all of the syntypes of B. plicidens that remain in England (UMZC and the shell from the J. S. Hawkins collection in the NHM) have normal barriers, but the lot Pilsbry received, according to this analysis, was entirely hooked with only one normal B. plicidens that was redescribed as B. landourensis by Pilsbry (1915). It is possible that the ̍ B. plicidens ̾ specimens that Pilsbry (1915) referred to as originating from Benson were not part of the type series, or were not directly from William Benson. Here we select the syntype in Cambridge (UMZC) as a lectotype to avoid further nomenclatural problems. Consequently, B. plicidens is a species with normal apertural barriers and B. hooki sp. nov. is a species with hooked barriers.	en	Páll-Gergely, Barna, White, Tom S. (2023): Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History 56 (45 - 48): 2011-2029, DOI: 10.1080/00222933.2022.2152750, URL: http://dx.doi.org/10.1080/00222933.2022.2152750
2050F30FBA44FFCEFE6E2493FE64FC78.taxon	description	(Figures 6, 9 (c, d )) Boysidia (Bensonella) plicidens – Pilsbry, 1917 (in Pilsbry 1916 – 1918): 198 (partim), pl. 34, figs 1, 2, 4	en	Páll-Gergely, Barna, White, Tom S. (2023): Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History 56 (45 - 48): 2011-2029, DOI: 10.1080/00222933.2022.2152750, URL: http://dx.doi.org/10.1080/00222933.2022.2152750
2050F30FBA44FFCEFE6E2493FE64FC78.taxon	materials_examined	Type material Holotype. NHMUK 20191114.1, 1 shell (SW: 1.72 mm, SH: 2.01 mm, AW: 0.83 mm, AH: 0.86 mm), Cherrapunji, Godwin-Austen coll. Paratypes. NHMUK 20191114.2, 782 shells, same data as for holotype; NHMUK 20191111 / B, 3 shells from a mixed sample identified as B. plicidens (NHMUK 20191111 / A), Cherra, India; NHMUK 1903.7.1.2882 / C, 1 shell from a mixed sample with B. plicidens, Moussorie, NW Himalaya, Godwin-Austen coll.; NHMUK 1906.2.2.190 C, 6 shells from a mixed sample identified as B. plicidens (NHMUK 1906.2.2.190 B), Cherra Poonje, Khasi Hills, W. T. Blanford coll.; NHMUK 20191107, 16 shells, Khasi, R. Beddome coll., no. 5024; NHMUK 1888.12.4.592 – 8, seven shells, Cherra, W. Theobald coll.; NHMUK 1903.7. 1.2062, three shells, Diyung valley; NHMUK 20191113, 13 shells, Cherrapoonji, Assam, GodwinAusten coll.; NHMUK 20191110, six shells, Teria Ghat, Assam, H. F. Blanford coll.; NHMUK 20191112, five shells, Khasi Hills, A. S. Kennard coll.; NHMUK 20191109, 13 shells, Himalaya, originally glued to two pieces of paper, some detached); NHMW-MO- 69532 (2 shells), Khasi Hills, Ind., coll. Oberwimmer, ex coll. Stoliczka; NHMW-MO- 113727 (2 shells), Khasi, coll. Stoliczka, 1880. xv. 473; NHMW-MO- 113725 (1 shell), Cachar, coll. Stoliczka 1870. Additional material. NHMUK 20191114.3, 9 juveniles, same data as for holotype.	en	Páll-Gergely, Barna, White, Tom S. (2023): Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History 56 (45 - 48): 2011-2029, DOI: 10.1080/00222933.2022.2152750, URL: http://dx.doi.org/10.1080/00222933.2022.2152750
2050F30FBA44FFCEFE6E2493FE64FC78.taxon	diagnosis	Diagnosis. A Bensonella species with a triangular-ovoid, reddish brown shell and hooked apertural barriers.	en	Páll-Gergely, Barna, White, Tom S. (2023): Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History 56 (45 - 48): 2011-2029, DOI: 10.1080/00222933.2022.2152750, URL: http://dx.doi.org/10.1080/00222933.2022.2152750
2050F30FBA44FFCEFE6E2493FE64FC78.taxon	description	Description. Shell triangular-ovoid, height somewhat variable; apex rather blunt, shell colour reddish brown; sculpture with occasionally strong, widely spaced growth lines; spiral striae absent; aperture subrectangular with clearly isolated sinulus due to angular lamella and parietal tubercle; peristome white, strongly widened, expanded, not reflected; aperture with numerous barriers; parietal wall with an angular lamella having two parts: its outer part reaches peristome, its inner part forms an outside-pointing hook; parietal lamella deeply situated, hooked; third lamella between parietal lamella and columellarparietal junction is ca. half as high as parietal lamella, and also forms a hook; palatal wall with three long main plicae (suprapalatal plica, upper palatal plica, lower palatal plica, however it is possible that the uppermost plica is the upper palatal plicae) with hooked central parts, their outer parts very slender, low, reaching peristome; columellar lamella similar to palatal plicae, hooked; between the lowest palatal plicae and the columellar plicae there are two or three short, hooked plicae; there are additional short, low, deeply situated plicae above the suprapalatal plica inside the sinulus; parietal tubercle blunt, knob-like, sits on edge of palatal lip, at the position where the palatal lip forms a little sinus; lower part of columellar lip widened, sometimes a low denticle-like swelling discernible; umbilicus very narrow, rounded. Measurements (in mm). SW = 1.58 – 1.77, SH = 1.84 – 2.3, AW = 0.74 – 0.83, AH = 0.79 – 0.92 (NHMUK 20191114).	en	Páll-Gergely, Barna, White, Tom S. (2023): Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History 56 (45 - 48): 2011-2029, DOI: 10.1080/00222933.2022.2152750, URL: http://dx.doi.org/10.1080/00222933.2022.2152750
2050F30FBA44FFCEFE6E2493FE64FC78.taxon	diagnosis	Differential diagnosis. This new species differs from B. plicidens in the following traits: Bensonella hooki has a shell that is larger and darker in colour, with more widely spaced and stronger growth lines; spiral striation absent; aperture with wider peristome, and the palatal tubercle on the peristome edge is less pointed, rather knob-like. The most important difference is in the apertural barriers, which are always hooked in the new species, but are never so in B. plicidens. Bensonella lakainguta has a more triangular shell (with straight side lines) and a more pointed apex, the shell colour is yellowish corneous (reddish brown in B. hooki sp. nov.), its parietal callus is adnate to the penultimate whorl (diverges to a greater distance from the penultimate whorl in B. hooki sp. nov.), the peristome is overall thinner and the parietal tooth on the peristome edge is more pointed. The dentition is strikingly similar.	en	Páll-Gergely, Barna, White, Tom S. (2023): Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History 56 (45 - 48): 2011-2029, DOI: 10.1080/00222933.2022.2152750, URL: http://dx.doi.org/10.1080/00222933.2022.2152750
2050F30FBA44FFCEFE6E2493FE64FC78.taxon	etymology	Etymology. This new species is named after Captain Hook, a fictional character from J. M. Barrie ̾ s Peter Pan, referring to the hook-like apertural barriers. It is a Latin singular, masculine noun in the genitive case.	en	Páll-Gergely, Barna, White, Tom S. (2023): Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History 56 (45 - 48): 2011-2029, DOI: 10.1080/00222933.2022.2152750, URL: http://dx.doi.org/10.1080/00222933.2022.2152750
2050F30FBA44FFCEFE6E2493FE64FC78.taxon	distribution	Distribution. Shells of this species were collected in the southwest (Mussoorie) and the southeast (Assam) Himalaya.	en	Páll-Gergely, Barna, White, Tom S. (2023): Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History 56 (45 - 48): 2011-2029, DOI: 10.1080/00222933.2022.2152750, URL: http://dx.doi.org/10.1080/00222933.2022.2152750
2050F30FBA44FFCEFE6E2493FE64FC78.taxon	discussion	Remarks. Although shells with hooked and normal barriers have sometimes been included in mixed lots from the Himalaya, there are also several lots with only one ̍ type ̾ of apertural barrier. This, and the additional differences between the two types (i. e. B. plicidens and B. hooki sp. nov.) indicates that contrary to the previous view (Budha and Backeljau 2017), these types are not morphological varieties of the same species, but represent two distinct species.	en	Páll-Gergely, Barna, White, Tom S. (2023): Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History 56 (45 - 48): 2011-2029, DOI: 10.1080/00222933.2022.2152750, URL: http://dx.doi.org/10.1080/00222933.2022.2152750
2050F30FBA46FFCFFE5E255FFDC8FC90.taxon	description	(Figures 7 – 8, 9 (e – h )) Boysidia (Bensonella) plicidens – Pilsbry, 1917 (in Pilsbry 1916 – 1918): 198 (partim), pl. 34, figs 3, 9 – 10	en	Páll-Gergely, Barna, White, Tom S. (2023): Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History 56 (45 - 48): 2011-2029, DOI: 10.1080/00222933.2022.2152750, URL: http://dx.doi.org/10.1080/00222933.2022.2152750
2050F30FBA46FFCFFE5E255FFDC8FC90.taxon	materials_examined	Material examined NHMUK 201901108, 12 shells, Omi, Japan, Hirase coll. 457; NHMUK 20191128, three shells, Rijozen, Omi, Japan; NHMW-MO- 38313 (1 shell), Mt. Fujiwara, 300 m, Japan, coll. Blume ex coll. Azuma, 24 September 1951; NHMW-MO- 55810 (5 shells), Japan, Mie Pref., Fujiwara-dake, 300 m, coll. Edlauer ex coll. Kuiper; NHMW-MO- 38645 (4 shells), Yoro, Mino; NHMW-MO- 69480 (2 shells), Riozen, Omi, Japan, coll. Oberwimmer ex coll. Jetschin; NHMW-MO- 113726 (3 shells), Omi, Japan, coll. Rušnov; HNHM 105323 (figured shell), Japan, Kochi Prefecture, Kami-shi, Ryugado Prefectural National Park, above the cave, 500 m along a side-road, 290 m, 33.601917 ° N, 133.746217 ° E, Leg. Hunyadi, A., Murányi, D. and Páll-Gergely, B., 08 August 2016.	en	Páll-Gergely, Barna, White, Tom S. (2023): Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History 56 (45 - 48): 2011-2029, DOI: 10.1080/00222933.2022.2152750, URL: http://dx.doi.org/10.1080/00222933.2022.2152750
2050F30FBA46FFCFFE5E255FFDC8FC90.taxon	discussion	Remarks In the original description, Hwang (2014) mentioned three differences between Indian B. plicidens and the subspecies Bensonella plicidens lakainguta. The most important of them was the submarginal elevations of columellar, basal and palatal barriers, i. e. anterior to the hooks those barriers become more elevated. However, this trait can also be seen in Indian B. hooki sp. nov. (Figure 9 (c, d )); see differences between B. lakainguta and B. hooki sp. nov. outlined under the description of the latter. Boysidia (Bensonella) qingliangfengensis (Figure 7 (e, f )) was described from the border region of the Chinese Anhui and Zhejiang Provinces (very close to Pilsbry ̾ s (1917) record from Hangzhou), and is claimed to be larger than B. plicidens, yet looks identical for all other characters. Therefore, it is treated here as a junior synonym of B. lakainguta Hwang, 2014. Maassen (1999) mentioned that he compared Bensonella karoensis with Japanese shells of Bensonella plicidens (= Bensonella lakainguta). Furthermore, he erroneously stated that Bensonella plicidens inhabited Thailand.	en	Páll-Gergely, Barna, White, Tom S. (2023): Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History 56 (45 - 48): 2011-2029, DOI: 10.1080/00222933.2022.2152750, URL: http://dx.doi.org/10.1080/00222933.2022.2152750
