taxonID	type	description	language	source
D592ED85A49D15D5C5D43944EC23EA0C.taxon	discussion	The genus Puntius Hamilton is diagnosed by the following combination of characters: absence or presence of maxillary only or rostral and maxillary barbels; dorsal fin with last simple ray serrate or entire, branched rays usually 8; anal fin with last simple ray entire, branched rays usually 5; lateral line complete or incomplete, lateral-line scales 17 - 36 in row; cephalic cutaneous papillae minute or absent; pharyngeal teeth in 3 rows, usually 2,3,5 / 5,3,2; colour pattern extremely variable (Roberts, 1989). According to Kottelat (1999), Puntius is a catchall ‘ genus’ in which a large number of unrelated small barbs have been placed. Manipur, with its varying forms of water bodies harbours many species of Puntius. Hamilton (1822) described Cyprinus ticto (now Puntius) from the Gangetic basin. While describing Barbus stoliczkanus (now Puntius) from the Chindwin basin, Pegu and Moulmein, Day (1878) wrote that the species supersedes the distribution of P. ticto in the east. Hora et al (1937), Talwar and Jhingran (1991) and Jayaram (1991) considered P. stoliczkanus to be a junior synonym of P. ticto. Hora (1921), without giving any collection data, reported P. ticto to be very common in the Imphal valley. Menon (1953) recorded the species from fields near Kanglatombi and from the banks of Imphal River (Chindwin basin). Menon (1954) recorded the species from Manipur without discussing its distribution by basin. In order to clarify the identity and distribution of Puntius ticto and P. stoliczkanus, several collections have been made from the Barak and Chindwin drainages for a detailed study. The present work reveals that two species are distinct. Puntius ticto is restricted to the Barak basin while P. stoliczkanus is restricted to the Chindwin basin. Puntius ticto, as reported by Vishwanath & Juliana (2004) from the Imphal River is P. ater, described here as a new species. Another species from the Khuga River is also described here as P. khugae.	en	I. Linthoingambi, W. Vishwanath (2007): Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa 1450: 45-56, URL: http://www.zoobank.org/urn:lsid:zoobank.org:pub:EAD51382-7DB4-421C-8412-F8F0C093A8AF
223FCC2790E30AE471C441FB4B0E4C42.taxon	description	(Fig. 1)	en	I. Linthoingambi, W. Vishwanath (2007): Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa 1450: 45-56, URL: http://www.zoobank.org/urn:lsid:zoobank.org:pub:EAD51382-7DB4-421C-8412-F8F0C093A8AF
223FCC2790E30AE471C441FB4B0E4C42.taxon	diagnosis	Diagnosis. A species of Puntius with the following combination of characters: dorsal fin edge black, its spine weak, serrated posteriorly with 13 - 17 serrae, spine length 16.8 - 23.3 % SL; predorsal scales 10 or 11; lateral line incomplete with 5 - 11 pored scales, 25 - 29 scales in lateral-line row; transverse scales ½ 4 / 1 / 4 ½; a black longitudinal stripe covering the upper half of the 1 st scale row and lower half of the 2 nd scale row above lateral-line row of scales; a black spot extending over 19 th and 20 th scales of lateral-line row at the level above the posterior end of the anal fin base; intestine long and coiled, its relative gut length 2.0 - 2.5 TL. Other differentiating characters from nearest congeners are in Table 1.	en	I. Linthoingambi, W. Vishwanath (2007): Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa 1450: 45-56, URL: http://www.zoobank.org/urn:lsid:zoobank.org:pub:EAD51382-7DB4-421C-8412-F8F0C093A8AF
223FCC2790E30AE471C441FB4B0E4C42.taxon	description	Description. Table 2 presents morphometric data, and Table 3 presents frequency distributions of meristic characters. Body elongated. Head large. Eyes large. Barbels absent. Lips thin, no lateral fold on snout. Dorsal fin origin opposite that of pelvic fin, inserted midway between tip of snout and base of caudal fin; 3 simple and 8 branched rays; third simple ray spinous, weak, and serrated posteriorly with 13 - 17 serrae. Pectoral fin with 1 simple posteriorly serrated ray and 12 branched rays, almost reaching pelvic fin origin. Pelvic fin with 1 simple and 8 branched rays. Anal fin with 3 simple and 5 branched rays. Caudal fin with 10 + 9 principal rays and 9 + 8 branched rays. Scales large, 25 - 29 in lateral-line row; lateral line incomplete with 5 - 11 pored scales; ½ 4 scales between dorsal fin origin and lateral line row, 4 ½ scales between lateral line and pelvic fin origin. Predorsal scales 10 - 11. Preanal scales 20. Predorsal bones 5 and predorsal neural spines 5. First pterygiophore inserted between 9 th and 10 th vertebrae. Number of total vertebrae 4 + 28 - 29. The infraorbital 3 + 4 is much broader and has greatly elevated margins compared to other bones of the infraorbital series. Intestine long and coiled (Fig. 2 a), its relative length 2.0 - 2.5 TL.	en	I. Linthoingambi, W. Vishwanath (2007): Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa 1450: 45-56, URL: http://www.zoobank.org/urn:lsid:zoobank.org:pub:EAD51382-7DB4-421C-8412-F8F0C093A8AF
223FCC2790E30AE471C441FB4B0E4C42.taxon	description	Colouration. In formalin, specimens have a black longitudinal stripe covering the upper half of the 1 st scale row and lower half of the 2 nd scale row above lateral-line row of scales, and a black spot on the 19 th and 20 th scales of lateral-line row, at the level above the posterior end of the anal fin base. Dorsal fin reddish, edged black. Pectoral, pelvic and anal fins reddish.	en	I. Linthoingambi, W. Vishwanath (2007): Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa 1450: 45-56, URL: http://www.zoobank.org/urn:lsid:zoobank.org:pub:EAD51382-7DB4-421C-8412-F8F0C093A8AF
223FCC2790E30AE471C441FB4B0E4C42.taxon	etymology	Etymology. Named after the black longitudinal stripe on the body, derived from ‘ ater’ (Latin), black.	en	I. Linthoingambi, W. Vishwanath (2007): Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa 1450: 45-56, URL: http://www.zoobank.org/urn:lsid:zoobank.org:pub:EAD51382-7DB4-421C-8412-F8F0C093A8AF
223FCC2790E30AE471C441FB4B0E4C42.taxon	distribution	Distribution. India: Manipur: Iril and Imphal rivers, ponds and lakes in Imphal valley (Fig. 3).	en	I. Linthoingambi, W. Vishwanath (2007): Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa 1450: 45-56, URL: http://www.zoobank.org/urn:lsid:zoobank.org:pub:EAD51382-7DB4-421C-8412-F8F0C093A8AF
436EF7C87956DC80CC4F3DF69DF13DF5.taxon	description	(Fig. 4)	en	I. Linthoingambi, W. Vishwanath (2007): Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa 1450: 45-56, URL: http://www.zoobank.org/urn:lsid:zoobank.org:pub:EAD51382-7DB4-421C-8412-F8F0C093A8AF
436EF7C87956DC80CC4F3DF69DF13DF5.taxon	diagnosis	Diagnosis. A species of Puntius with the following combination of characters: dorsal fin edge plain, its spine serrated posteriorly with 10 - 12 serrae, spine length 16.0 - 20.9 % SL; predorsal scales 11 - 12; lateral line incomplete with 8 - 11 pored scales; 28 - 30 scales in lateral-line row; transverse scales ½ 5 / 1 / 4 ½; black blotch on caudal peduncle at level of one scale behind posterior end of anal origin; intestine coiled, its relative length 0.9 - 1.2 TL. Other differentiating characters from nearest congeners are in Table 1.	en	I. Linthoingambi, W. Vishwanath (2007): Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa 1450: 45-56, URL: http://www.zoobank.org/urn:lsid:zoobank.org:pub:EAD51382-7DB4-421C-8412-F8F0C093A8AF
436EF7C87956DC80CC4F3DF69DF13DF5.taxon	description	Description. Table 2 presents morphometric data, and Table 3 presents frequency distributions of meristic characters. Body small and slender. Head large. Eyes large. Barbels absent. Lips thin. Dorsal fin origin opposite that of pelvic fin, inserted midway between tip of snout and base of caudal fin; 3 simple and 8 branched rays; third simple ray spiny and serrated posteriorly with 10 - 12 serrae. Pectoral fin with 1 simple, posteriorly serrated ray and 12 branched rays. Pelvic fin with 1 simple and 8 branched rays. Caudal fin with 10 + 9 principal rays, 9 + 8 branched. Scales large. Lateral line incomplete, 28 - 30 scales in the row with 8 - 11 pored. ½ 5 scales between dorsal fin origin and lateral-line row, 4 ½ between lateral line and pelvic fin base. Predorsal scales 10 - 12. Preanal scales 19. Predorsal bones 5; predorsal neural spines 5; first pterygiophore inserted between 9 th and 10 th vertebrae; total vertebrae 4 + 28 - 29. The infraorbital 3 + 4 is broad and has greatly elevated margins compared to other bones of infraorbital series. Intestine coiled (Fig. 2 b), its relative length 0.9 - 1.2 TL. Colouration. In formalin, specimens have a blue-black longitudinal stripe occupying the upper half of the 1 st scale row and lower half of the 2 nd scale row above the lateral-line scale row. A black spot is present on the 21 st scale of the lateral-line row; dorsal fin with a red band, edge plain. Pectoral, pelvic and anal fins reddish.	en	I. Linthoingambi, W. Vishwanath (2007): Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa 1450: 45-56, URL: http://www.zoobank.org/urn:lsid:zoobank.org:pub:EAD51382-7DB4-421C-8412-F8F0C093A8AF
436EF7C87956DC80CC4F3DF69DF13DF5.taxon	etymology	Etymology. Named after the Khuga River, the type locality for the species.	en	I. Linthoingambi, W. Vishwanath (2007): Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa 1450: 45-56, URL: http://www.zoobank.org/urn:lsid:zoobank.org:pub:EAD51382-7DB4-421C-8412-F8F0C093A8AF
436EF7C87956DC80CC4F3DF69DF13DF5.taxon	distribution	Distribution. India: Manipur: Khuga River (Fig. 3).	en	I. Linthoingambi, W. Vishwanath (2007): Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa 1450: 45-56, URL: http://www.zoobank.org/urn:lsid:zoobank.org:pub:EAD51382-7DB4-421C-8412-F8F0C093A8AF
36C949C93526B0F0490025DFF24F959F.taxon	description	(Fig. 5)	en	I. Linthoingambi, W. Vishwanath (2007): Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa 1450: 45-56, URL: http://www.zoobank.org/urn:lsid:zoobank.org:pub:EAD51382-7DB4-421C-8412-F8F0C093A8AF
36C949C93526B0F0490025DFF24F959F.taxon	materials_examined	Material examined. (All from Brahmaputra basins) MUMF 6115, 1, 38.0 mm SL; India: Manipur: Jiri R. at Jiribam Subdivision; W. Vishwanath, 8 March 2000. MUMF 6116 - 20, 5, 40.0 - 46.0 mm SL; India: Assam: Wetlands of Hajo, North Kamrup District, A. Boishya, 20 April 2004. Uncat, 9, 35.0 - 44.0 mm SL; India: Assam: Brahmaputra R. at Guwahati; W. Vishwanath, 3 August, 2000. Uncat., 3, 40.0 - 41.3 mm SL; India: Nagaland: Dikhu R. at Moalenden; Bendangkokpa Jamir, 10 September 2005. Uncat., 21, 34.5 - 45.8 mm SL; India: Assam: Brahmaputra R. at Guwahati, W. Vishwanath, K. Nebeshwar and I. Linthoingambi, 21 July 2005.	en	I. Linthoingambi, W. Vishwanath (2007): Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa 1450: 45-56, URL: http://www.zoobank.org/urn:lsid:zoobank.org:pub:EAD51382-7DB4-421C-8412-F8F0C093A8AF
36C949C93526B0F0490025DFF24F959F.taxon	diagnosis	Diagnosis. Body deep, its depth 36.8 - 45.0 % SL; last simple ray of dorsal fin long, spiny and serrated posteriorly with 15 - 17 serrae, spine length 20.6 - 28.0 % SL, 78.3 - 88.6 % HL; lateral line incomplete with 22 - 26 scales in the row, 6 - 11 pored; preanal scales 16; two spots along lateral line scale row, one on the 4 th scale and second on 17 th - 20 th scales; two complete black bands on dorsal fin.	en	I. Linthoingambi, W. Vishwanath (2007): Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa 1450: 45-56, URL: http://www.zoobank.org/urn:lsid:zoobank.org:pub:EAD51382-7DB4-421C-8412-F8F0C093A8AF
36C949C93526B0F0490025DFF24F959F.taxon	description	Description. Table 3 presents frequency distributions of meristic characters, and Table 4 presents morphometric data. Body short, compressed and deep. Barbels absent. Lips thick, lateral fold on snout well developed. Dorsal fin origin a little behind that of pelvic fin, inserted midway between tip of snout and base of caudal fin; 3 simple and 8 branched rays; third simple ray long, spiny and finely serrated posteriorly with 15 - 17 serrae. Pectoral fin with 1 posteriorly serrated simple ray and 12 branched rays, almost reaching the pelvic fin origin. Pelvic fin with 8 - 9 rays, the 1 st and last two undivided. Anal fin with 3 simple and 5 branched rays. Caudal fin with 10 + 9 principal rays, 9 + 8 branched. Scales large. Lateral line incomplete, 22 - 26 in the row with 6 - 11 pored. ½ 5 scales between dorsal fin origin and lateral-line row, 5 ½ between lateral line and pelvic fin origin. Predorsal scales 9 - 10. Preanal scales 16. Predorsal bones 4, predorsal neural spines 4. First pterygiophore inserted between 8 th and 9 th vertebrae. Total vertebrae 4 + 26. Colouration. A black spot on the 4 th scale and one on 17 th - 20 th scales of lateral line row. Two black bands on dorsal fin, the first at middle length on the membranes between the branched rays and the second on the branched rays at about 3 / 4 length of the fin from its base.	en	I. Linthoingambi, W. Vishwanath (2007): Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa 1450: 45-56, URL: http://www.zoobank.org/urn:lsid:zoobank.org:pub:EAD51382-7DB4-421C-8412-F8F0C093A8AF
36C949C93526B0F0490025DFF24F959F.taxon	distribution	Distribution. India: Ganges (Hamilton, 1822). India: Assam: Brahmaputra River (Sen, 1985). It is here reported from Jiri R., Manipur state (Fig. 3) and Dikhu R., Nagaland state, India.	en	I. Linthoingambi, W. Vishwanath (2007): Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa 1450: 45-56, URL: http://www.zoobank.org/urn:lsid:zoobank.org:pub:EAD51382-7DB4-421C-8412-F8F0C093A8AF
B5139B2BDC91ED0B729BF8AC36521332.taxon	description	(Fig. 6)	en	I. Linthoingambi, W. Vishwanath (2007): Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa 1450: 45-56, URL: http://www.zoobank.org/urn:lsid:zoobank.org:pub:EAD51382-7DB4-421C-8412-F8F0C093A8AF
B5139B2BDC91ED0B729BF8AC36521332.taxon	materials_examined	Material examined. MUMF 6120, 10, 46.6 - 50.5 mm SL; India: Manipur: Lokchao R at Moreh, (Indo-Myanmar border 94 ° 18 ’ 11 ” E, 24 ° 14 ’ 56 ” N, 212 m asl); W. Vishwanath, 2 April 2004. MUMF 7250, 15, 47.5 - 53.9 mm SL; same data as MUMF 6120; L. Sakuntala, 5 July 1999. MUMF 9500 - 8, 9, 41.2 - 53.0 mm SL; same data as MUMF 6120; A. Darshan and S. Sanjabihari, 14 July 2005. MUMF 10001, 3, 41.0 - 45.5 mm SL; India: Manipur: Maklang R. at Ukhrul district; Kingson, 15 July 2005.	en	I. Linthoingambi, W. Vishwanath (2007): Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa 1450: 45-56, URL: http://www.zoobank.org/urn:lsid:zoobank.org:pub:EAD51382-7DB4-421C-8412-F8F0C093A8AF
B5139B2BDC91ED0B729BF8AC36521332.taxon	diagnosis	Diagnosis. Maximum head width 55.5 - 66.0 % of head length; last simple dorsal ray short, spiny and finely serrated posteriorly with 12 - 16 serrae, spine length 16.0 - 20.0 % SL and 58.0 - 65.4 % HL; lateral line complete with 21 - 24 scales in row and 19 - 23 pored scales, if incomplete, reaches at least to the caudal peduncle; preanal scalesl 8; presence of an oblong black mark above pectoral fin and one black blotch on 17 th- 19 th scales of the lateral-line row; two black bands on dorsal fin, incomplete one at middle length on the membrane between 3 rd - 5 th branched rays and another one on the branched rays at margin.	en	I. Linthoingambi, W. Vishwanath (2007): Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa 1450: 45-56, URL: http://www.zoobank.org/urn:lsid:zoobank.org:pub:EAD51382-7DB4-421C-8412-F8F0C093A8AF
B5139B2BDC91ED0B729BF8AC36521332.taxon	description	Description. Table 3 presents frequency distributions of meristic characters, and Table 4 presents morphometric data. Body short, compressed and deep. Barbels absent. Lips thick, lateral fold on snout well developed. Dorsal fin origin opposite that of pelvic fin, inserted midway between tip of snout and base of caudal fin; 3 simple and 8 branched rays; third simple ray short, spiny and serrated posteriorly with 12 - 16 serrae. Pectoral fin with 1 simple posteriorly serrated ray and 12 branched rays; almost reaching the pelvic fin origin. Pelvic fin with 1 simple and 8 branched rays. Anal fin with 3 simple and 5 branched rays. Caudal fin with 10 + 9 principal rays, 9 + 8 branched. Scales large. Lateral line complete, 21 - 24 in the row with 19 - 23 pored scales; ½ 5 scales between dorsal fin origin and lateral-line row, 5 ½ between lateral line and pelvic fin base. If incomplete, the lateral line at least reaches the caudal peduncle. Predorsal scales 8 - 10. Preanal scales 18. Predorsal bone 4 and predorsal neural spine 4. First pterygiophore inserted between 8 th and 9 th vertebrae. Total vertebrae count 4 + 25 - 26. Colouration. An oblong black mark above pectoral fin on the 3 rd scale of the lateral line and one black blotch on 17 th - 19 th scales of the lateral-line row. Two black bands on the dorsal fin, the first at middle length and on the membrane between 3 rd - 5 th branched rays only, the second on the branched rays at the margin of the fin.	en	I. Linthoingambi, W. Vishwanath (2007): Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa 1450: 45-56, URL: http://www.zoobank.org/urn:lsid:zoobank.org:pub:EAD51382-7DB4-421C-8412-F8F0C093A8AF
B5139B2BDC91ED0B729BF8AC36521332.taxon	distribution	Distribution. Myanmar: Chindwin Basin (Day, 1871). India: Chindwin basin in Manipur (Vishwanath and Juliana, 2004) (Fig. 3). Myanmar, Thailand, Laos (Kottelat, 2001).	en	I. Linthoingambi, W. Vishwanath (2007): Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa 1450: 45-56, URL: http://www.zoobank.org/urn:lsid:zoobank.org:pub:EAD51382-7DB4-421C-8412-F8F0C093A8AF
533FB8EC3A7B60783329DDC42A83D774.taxon	discussion	Discussion Puntius ater and P. khugae are unique among species of Puntius in Manipur in having one black longitudinal stripe on the side of the body. The former species is distinguished from the latter in having a black edge on the dorsal fin, 13 - 17 serrae (modally 13) on the dorsal fin spine, 10 - 11 (modally 10) predorsal scales, modally 26 or 27 (range = 25 - 29) lateral-line scales, and a longer gut (2.0 - 2.5 times total length). Puntius khugae has no black edge on the dorsal fin, 10 - 12 serrae (modally 10) on the dorsal fin spine, 9 - 10 (modally 9) predorsal scales, modally 29 (range = 28 - 30) lateral-line scales, and a shorter gut (0.9 - 1.2 times total length) (Tables 1 & 3). Taki et al (1978) classified species of Puntius having broad and elevated infraorbital 3 + 4 in the Puntius conchonius-group. The two new species have similar structures and thus may be included in the group. Puntius ater inhabits sluggish streams of the Imphal River and its tributaries, while P. khugae inhabits the Khuga, a comparatively faster clear-water stream. The latter species has a shorter gut (Fig. 2 a & b) and probably is a sympatric species adapted to a different environment with a different food habit. Hamilton (1822) described P. ticto from Gangetic basin. He observed the pelvic fin of the species to have one undivided ray in front and two behind. Similar observations were also made in the present study. Hora et al. (1937) observed variations in P. ticto and treated P. punctatus of Peninsular India and P. stoliczkanus of the Irrawady as its junior synonyms. Silas (1952), however, regarded P. punctatus and P. stoliczkanus as subspecies of P. ticto. Jayaram (1991) did not give subspecies status to P. punctatus and P. stolizckanus as the fishes showed great variation in morphology. He also considered P. ticto to be widely distributed and not restricted to any definite geographical region. Day (1871) described P. stoliczkanus from the Chindwin basin, Pegu and Moulmein. He reported the species to have a less compressed body and weaker serrations in the dorsal spine compared to P. ticto. Hora (1936) distinguished P. stoliczkanus from P. ticto in having weaker serrations on the dorsal spine, and also in differences in the positions of lateral spots and the numbers of predorsal scales. Jayaram (1981) considered P. stoliczkanus to be a subspecies of P. ticto. Kottelat (2001) reported on the occurrence of P. stoliczkanus in the Irrawady, Salween, Mekong and Chao Phraya basins. Early workers had access to very few, poorly preserved specimens. Thus, species were poorly described. Variations in ontogenic development were not considered, and several species from different geographically isolated habitats were put together as highly variable species. Various workers (Kottelat, 1996; Kottelat & Lim, 1993, 1995; Ng & Dodson, 1999; Ng & Kottelat, 2000; Chakrabarty & Ng, 2005; Ng, 2005) reexamined such ‘ highly variable’ widely distributed species and concluded that they were in fact aggregates of distinct, often not even closely related species. Puntius ticto and P. stoliczkanus were also examined and found to be distinct (Table 1). None of the fish collections from the Chindwin basins of the state contained P. ticto. The species reported as P. ticto by previous workers from the valley were probably misidentifications of P. stoliczkanus or P. ater. It may be concluded that P. ticto is a Brahmaputra form, and P. stoliczkanus is a Chindwin form.	en	I. Linthoingambi, W. Vishwanath (2007): Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa 1450: 45-56, URL: http://www.zoobank.org/urn:lsid:zoobank.org:pub:EAD51382-7DB4-421C-8412-F8F0C093A8AF
