identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
2324A34FFD6E32290EF7FED71AF3C921.text	2324A34FFD6E32290EF7FED71AF3C921.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemiosus molanoi González-Rodríguez & García-Hernández & Clarkson 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Hemiosus molanoi González-Rodríguez &amp; García-Hernández sp. nov.</p>
            <p>(Figs. 1‒7)</p>
            <p> Type Material.  Holotype (male) (MAUQ-14459) : “ Colombia: Quindío Department,/ Municipality: Córdoba,/ Neighborhood:  Río Verde / 04°24' 12.0'', 075° 43' 26.6'', 1108 m m.a.s.l. / 31-01-2015 / L.M. González-Rodríguez. and A.L. García-Hernández coll. // Habitat: along riverbanks and in temporary ponds,/ substrates: stony/sandy, muddy/stony.” Condition of holotype: stored in 90% ethanol with the dissected male genitalia stored in microvials with glycerin. (Deposited in MAUQ).  Paratypes (24): (same data as holotype: 3 males, 4 females; MAUQ (MAUQ-14460-14461)), (3 males, 3 females; UPTC), (3 males, 3 females; CZUT) ,  (3 males, 3 females; DZUP) . </p>
            <p>Diagnosis. Total body length 2.8‒3.2 mm in males, 3.0‒4.0 mm in females. Pronotum and elytra testaceous with a pair of small black round spots in anteromedial position (Fig. 1). Elytra testaceous with melanic areas (Figs. 1, 6). Mesoventral process subrhomboid; in lateral view with three slightly raised tooth, anterior and posterior tooth shorter than median teeth (Figs. 2, 3). Metaventral process wide, anterior carina as high as mesoventral process (Fig. 4). Ventrite 1 with linear broad medial carina. Apical notch of ventrite 5 with triangular tooth (Fig. 5). Aedeagus (Fig. 7) with parameres subtriangular; median lobe shorter than parameres, appendices acuminate and shorter than median lobe; phallobase shorter than parameres in length, strongly asymmetrical in basal two-fifths, manubrium distinct.</p>
            <p>Description. Size and form (Measurements of the holotype). Total length 3.00 mm; maximum width 1.40 mm; maximum width of head 0.80 mm; maximum width of pronotum 0.90 mm; humeral width 1.10 mm; elytral length 2.15 mm. Body widely oval, strongly convex in lateral view (Fig. 1).</p>
            <p>Color. Head melanic with strong metallic sheen (Fig. 1). Mouthparts dark brown; labial and maxillary palps testaceous, darkened at apex and at apical third respectively; antennae pale brown (Fig. 1). Pronotum testaceous with a pair of small black round spots in anteromedial position, separated by 4 times the width of one spot, without metallic sheen; elytra testaceous with melanic areas, punctures on elytral striae black (Figs. 1, 6). Ventral face of thorax dark-brown to black. Anterior coxae dark-brown, trochanter and base of femora slightly lighter than coxae, remainder of anterior legs testaceous; middle and hind coxae, trochanter and basal third of femora dark brown, remainder of middle and hind legs testaceous.</p>
            <p>Head. Head flattened in lateral view. Ground punctation on frons and clypeus strongly impressed and densely distributed, distance between punctures lesser than width of one puncture. Mentum shiny, glabrous, with scattered punctures and scattered setae only on lateral portions. Maxillary palps nearly as long as maximum width of head; labial palps nearly as long as mentum width.</p>
            <p>Thorax. Pronotal margins smooth. Ground punctation on pronotal disc evenly distributed, punctures as large as those on head; distance between punctures slightly larger than width of one puncture. Serial punctures moderate in size, round, deeply impressed; about 3 times the size of punctures on pronotal disc; inner interstriae flat, about 3 times as wide as striae; outer interstriae narrow and convex; ground and systematic punctures on inner interstriae about half of the size of serial punctures on striae (Fig. 6). Mesoventral process raised along anterior portion, subrhomboid in ventral view, longitudinally depressed along median-posterior portion, posterior portion slightly broad and long (Fig. 2); in lateral view with three slightly raised tooth, anterior and posterior tooth shorter than median teeth (Fig. 3). Metaventral process wide, anteriorly with carina projected between mesocoxae almost reaching the posterior margin of, and as elevated as, mesoventral process; posteriorly with edge of process produced, narrow and blunt; posterolateral margins not produced posteriorly (Fig. 4).</p>
            <p>Abdomen. Ventrite 1 with strongly raised linear broad medial carina, anteriorly extended between metacoxae, posteriorly reaching posterior margin; lateral carinae narrow, blunt, slightly raised, extending along basal half. Ventrite 2 with basal blunt carina extending along basal two-thirds. Apical notch of ventrite 5 with short median triangular tooth, lateral portions smooth, slightly raised (Fig. 5). Aedeagus with each paramere subtriangular, inner margins slightly concave along apical half, narrowly rounded and slightly turned inwards at apex; median lobe shorter than parameres, gradually narrowing along medial two-fourths, rounded at apex; appendices acuminate and nearly as long as median lobe; phallobase elongated, nearly as long as parameres, maximum width subapically, strongly asymmetrical in basal two-fifths with manubrium distinct, narrow and rounded at base (Fig. 7).</p>
            <p>Variation. Coloration: there are variations on the coloration across specimens with some being lighter with darker melanic areas. Sexual: no obvious sexual dimorphisms were observed other than females being generally larger than males.</p>
            <p>Etymology. The species epithet is attributed to honor Fredy Molano Rendón, for his passion and dedication to teaching entomology.</p>
            <p>Natural history. Specimens were collected along riverbanks and in temporary ponds on muddy leaf litter as well as on muddy/stony and stony/sandy substrates (Fig. 15).</p>
            <p> Remarks.  Hemiosus molanoi sp. nov. is fairly similar to  H. fittkaui Oliva (northern Brazil) in the coloration pattern (head melanic with strong metallic sheen; maxillary palps darkened at apex; pronotum and elytra with melanic areas), in some features of the male genitalia (phallobase asymmetrical and median lobe rounded at apex), and in the aspect of the ventrite 5 (median portion of apical notch extended as a triangular tooth). The new species differs from  H. fittkaui by the overall morphology of the meso- and metaventral processes; ventrite 1 with linear broad medial carina (Y-shaped and narrow in  H. fittkaui ); ventrite 4 without medial carina (rudimentary carina in  H. fittkaui ); phallobase nearly as long as parameres (shorter than parameres in  H. fittkaui ); parameres narrowly rounded at apex (sharply acute in  H. fittkaui ); median lobe shorter than parameres (hardly shorter than parameres in  H. fittkaui ) and appendices of the median lobe acuminate and nearly as long as median lobe (appendices lanceolate and shorter than median lobe in  H. fittkaui ). </p>
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	https://treatment.plazi.org/id/2324A34FFD6E32290EF7FED71AF3C921	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	González-Rodríguez, Liza M.;García-Hernández, Andrea L.;Clarkson, Bruno	González-Rodríguez, Liza M., García-Hernández, Andrea L., Clarkson, Bruno (2019): Two new species of the water scavenger beetle genus Hemiosus Sharp (Coleoptera: Hydrophilidae) from Colombian Andes. Zootaxa 4565 (4): 493-500, DOI: 10.11646/zootaxa.4565.4.3
2324A34FFD6F322F0EF7F9121B41CE76.text	2324A34FFD6F322F0EF7F9121B41CE76.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemiosus quindiensis González-Rodríguez & García-Hernández & Clarkson 2019	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Hemiosus quindiensis González-Rodríguez &amp; Clarkson sp. nov.</p>
            <p>(Figs. 8‒14)</p>
            <p> Type Material.  Holotype (male) (MAUQ-14456) : “ Colombia: Quindío Department, Municipality: La Tebaida,/  Neighborhood :  Pizamal / 04°41' 27.6'', 075° 85' 37.7'', 1044 msnm/ 10-10-2014 / Liza María G.R. col./ Lorena G.H y Liza María G.R. det. // Habitat: temporary ponds,/ substrates: muddy/stony, stony/sandy. (Deposited in MAUQ). Paratypes (24): (same data as Holotype: 3 males, 4 females; MAUQ (MAUQ-14457-14458)), (3 males, 3 females; UPTC), (3 males, 3 females; CZUT), (3 males, 3 females; DZUP). </p>
            <p>Diagnosis: Total body length 2.8‒3.0 mm in males, 2.8‒3.5 mm in females. Pronotum testaceous with a pair of</p>
            <p>black longitudinal markings nearly at medial position. Elytra testaceous with melanic areas. Mesoventral process raised, subrhomboid with raised anterior tooth. Metaventral process narrow. Ventrite 1 with fine acuminate medial carina. Apical notch in ventrite 5 with median portion slightly raised. Aedeagus with each paramere gradually tapering towards apex, rounded at apex; median lobe shorter than parameres, appendices shorter than median lobe, acuminate. Phallobase shorter than parameres in length, strongly asymmetrical in basal half with manubrium distinct.</p>
            <p>Description. Size and form (Measurements of the holotype). Total length 3.00 mm; maximum width 1.40 mm; maximum width of head 0.85 mm; maximum width of pronotum 1.00 mm; humeral width 1.30 mm; elytral length 2.15 mm. Body oval, elongate, strongly convex in lateral view (Fig. 8).</p>
            <p>Color. Head melanic with strong metallic sheen (Fig. 8). Mouthparts dark brown; maxillary and labial palps testaceous, maxillary palps darkened at apex (Fig. 8); antennae testaceous. Pronotum testaceous, with a pair of black longitudinal markings nearly at medial position, separated by half the width of one spot, without metallic sheen; elytra testaceous with melanic areas (Fig. 8). Ventral face of thorax and abdomen dark-brown to black. Anterior, middle and hind legs testaceous.</p>
            <p>Head. Head flattened in lateral view. Ground punctation on frons and clypeus strongly impressed and densely distributed, distance between punctures lesser than width of one puncture; mentum width 0.28 mm, with scattered punctures, shiny and glabrous; maxillary palps nearly as long as maximum width of head; labial palps nearly as long as mentum width.</p>
            <p>Thorax. Pronotal margins smooth. Ground punctation on pronotal disc densely distributed, punctures larger than those on head; distance between punctures half width of one puncture. Elytra with serial punctures, moderately impressed; about twice the size of ground punctures on pronotal disc; inner interstriae flat, about 3 times width of a stria; outer interstriae narrow and convex; ground and systematic punctures on inner interstriae about half the size of serial punctures on striae (Fig. 13). Mesoventral process raised, subrhomboid in ventral view, depressed in median portion, posterior portion narrow and long (Fig. 9), projected between mesocoxae, raised anterior portion as a tooth-like projection in lateral view (Fig. 10). Metaventral process narrow, anteriorly with narrow raised carina not projected between mesocoxae, almost reaching the posterior margin of, and as elevated as, mesoventral process; posteriorly with edge of process produced, wide and slightly acuminate; posterolateral margins not produced posteriorly (Fig. 11).</p>
            <p>Abdomen. Ventrite 1 with fine acuminate medial carina, slightly broadening posteriorly, anteriorly extended between metacoxae, restricted to anterior portion of the ventrite. Lateral carinae narrow, slightly raised, blunt, about two-thirds the length of the ventrite. Ventrite 2 with fine medial basal carina not reaching posterior margin of the ventrite. Apical notch of ventrite 5 with median portion slightly raised, lateral portions smooth (Fig. 12). Aedeagus with outer margins of parameres subparallel to slightly convex along basal half, gradually tapering distally along apical half, rounded at apex, slightly turned inwards; median lobe shorter than parameres, subcylindrical, rounded at apex; appendices rather shorter than median lobe, acuminate; phallobase elongated, nearly as long as parameres, maximum width at apex, strongly asymmetrical at basal half with manubrium distinct, narrow and truncate at base (Fig. 14).</p>
            <p>Variation. Coloration: specimens with lighter-colored and darker melanic areas. Sexual: No obvious sexual dimorphisms other than females being generally larger than males.</p>
            <p>Etymology. The species epithet is a reference to the Quindío Department, from where the type specimens were collected.</p>
            <p>Natural history. Specimens were collected along riverbanks and in temporary ponds on muddy leaf litter as well as on muddy/stony and stony/sandy substrates (Fig. 16).</p>
            <p> Remarks.  Hemiosus quindiensis sp. nov. is fairly similar to  H. multimaculatus, (Jensen-Haarup) (distributed in Argentina, Chile, and Bolivia) in the coloration pattern (head with strong metallic sheen, maxillary palp darkened at apex, pronotum and elytra with melanic areas), and in some features of the male genitalia (median lobe shorter than parameres; phallobase asymmetrical). The new species differs from  H. multimaculatus by the morphology of the metaventral process, with a narrow anterior carina (blunt carina in  H. multimaculatus ); the ventrite 1 exhibits a fine acuminate medial carina, slightly broadening posteriorly, not bifurcate posteriorly (carina narrowly Y-shaped, bifurcate posteriorly, in  H. multimaculatus ); ventrite 2 with fine carina (wide carina in  H. multimaculatus ); ventrites 3 and 4 flat, without carina (vestigial carinae in  H. multimaculatus ); and aedeagus with parameres rounded at apex (apex acuminate in  H. multimaculatus ) and appendices of the median lobe shorter than median lobe (as long as median lobe in  H. multimaculatus ). </p>
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	https://treatment.plazi.org/id/2324A34FFD6F322F0EF7F9121B41CE76	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	González-Rodríguez, Liza M.;García-Hernández, Andrea L.;Clarkson, Bruno	González-Rodríguez, Liza M., García-Hernández, Andrea L., Clarkson, Bruno (2019): Two new species of the water scavenger beetle genus Hemiosus Sharp (Coleoptera: Hydrophilidae) from Colombian Andes. Zootaxa 4565 (4): 493-500, DOI: 10.11646/zootaxa.4565.4.3
