taxonID	type	description	language	source
234487CB8124405E614AF980FAE4F9AD.taxon	type_taxon	Type species Mimofulvius pentatomus Schmitz, 1978 (original designation).	en	Wolski, Andrzej, Yasunaga, Tomohide (2025): First report of little-known cylapine genera Mimofulvius Schmitz and Rhinophrus Hsiao (Insecta: Hemiptera: Heteroptera: Miridae) from Thailand, with description of a new species. European Journal of Taxonomy 987: 61-80, DOI: 10.5852/ejt.2025.987.2859, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2859/12995
234487CB8124405E614AF980FAE4F9AD.taxon	diagnosis	Diagnosis In addition to the characters presented by Schmitz (1978) and Yeshwanth et al. (2016) for defining Mimofulvius, this genus can be distinguished from other cylapine genera by having the rudimentary stridulatory device (exocorium edge-hind femur, cf. Figs 8, 12 – 13) and the endosoma with lobal sclerites (Figs 4, 33).	en	Wolski, Andrzej, Yasunaga, Tomohide (2025): First report of little-known cylapine genera Mimofulvius Schmitz and Rhinophrus Hsiao (Insecta: Hemiptera: Heteroptera: Miridae) from Thailand, with description of a new species. European Journal of Taxonomy 987: 61-80, DOI: 10.5852/ejt.2025.987.2859, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2859/12995
234487CB8124405E614AF980FAE4F9AD.taxon	description	Redescription Macropterous; body elongate to elongate-oval (Figs 1 – 2, 6, 14 – 16, 24 – 27). Dorsum shiny, covered with sparse or dense, simple setae (Figs 6 – 9). HEAD. Subhorizontal (Figs 1 – 2, 6 – 7, 14 – 16, 18, 24 – 27), rugose, covered with long, dense, semirecumbent setae (Figs 1 – 2, 6 – 7, 18); clypeal base situated above ventral margin of eye (Fig. 18); mandibular plate without sulcus posteriorly (Fig. 18); antennal insertion contiguous with sulcus between maxillary and mandibular plates (Figs 7, 18); eyes contiguous with pronotal collar, relatively large, reniform in lateral view (Figs 7, 18); antenna short, about ⅔ times as long as total body length; antennal segment I short, weakly surpassing apex of clypeus, cylindrical, weakly narrowed basally, covered with sparse, thick, erect and semirecumbent setae; segment II weakly broadened toward apex, covered with dense, erect and semirecumbent setae; segments III and IV as thick as segment II, covered with setae similar to those present on segment II; segment IV subdivided medially, forming pseudo- 5 - segmented (Figs 1, 6, 14 – 17, 25, 27); labial segment I subdivided medially; segment II subdivided subapically (Fig. 7). THORAX. Pronotum. Pronotal collar relatively broad, approximately as wide as antennal segment II diameter; pronotal calli from narrow, restricted to medial part of anterior part of pronotum to broad, reaching pronotal lateral margins (Figs 1 – 2, 6, 14 – 16); dorsal surface of pronotum shiny, covered with sparse, semirecumbent setae (Figs 6, 18). Mesoscutum and scutellum. Covered with sparse, erect or semirecumbent setae; scutellum flat (Figs 1 – 2). Thoracic pleura. Covered with sparse, short, semirecumbent setae (Figs 7, 9, 18 – 19); metathoracic scent efferent system with narrow evaporative area and ovoid auricula, occupying ventral and posterior margins; peritreme flat, relatively broad (Figs 9, 19). Hemelytron. Shiny, covered with sparse or dense, simple setae (Figs 1 – 2, 7 – 8); exocorium with notched edge that may represent stridulitrum, or stridulatory device (Figs 8, 20). Legs. Short; covered with sort, semirecumbent and recumbent setae (Figs 1 – 2, 24 – 27); metafemur with short carina, probably plectrum of stridulatory mechanism (Figs 12 – 13, 21); pretarsal claw with tiny, subapical tooth (Fig. 10). ABDOMEN. Covered with dense, reclining and semirecumbent setae (Fig. 7). MALE GENITALIA. Genital capsule with dorsal wall long, weakly shorter than ventral wall, genital opening is terminal in orientation (Figs 7, 11, 28). Left and right paramere curved; paramere body covered with relatively dense setae dorsally (Figs 3, 5, 29 – 32). Left paramere. Apical process elongated, inner surface with small swelling basally (Figs 3, 29 – 30). Aedeagus. Seminal duct long and thin, sclerotized part of seminal duct expanded; endosoma membranous, with two long lobal-sclerites and with ovoid, large, membranous lobe basally embraced by expanded sclerotized part of seminal duct (Figs 4, 33). FEMALE GENITALIA. As described and depicted by Yeshwanth et al. (2016: fig. 30).	en	Wolski, Andrzej, Yasunaga, Tomohide (2025): First report of little-known cylapine genera Mimofulvius Schmitz and Rhinophrus Hsiao (Insecta: Hemiptera: Heteroptera: Miridae) from Thailand, with description of a new species. European Journal of Taxonomy 987: 61-80, DOI: 10.5852/ejt.2025.987.2859, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2859/12995
234487CB8124405E614AF980FAE4F9AD.taxon	discussion	Remarks The present study adds the possession of a (presumed) stridulatory device (a set of a notched margin of the forewing and a metafemoral plectrum or keel) to the diagnostic characters for Mimofulvius (Figs 8, 13, 20 – 21). As stated by Yasunaga et al. (2019) and Tamada et al. (2020), many epigeic plant bug species of the tribe Hallodapini Van Duzee, 1916 (Phylinae) have a similar stridulatory mechanism which is used for intraspecific communication. However, the actual function of the Mimofulvius stridulatory device requires further verification.	en	Wolski, Andrzej, Yasunaga, Tomohide (2025): First report of little-known cylapine genera Mimofulvius Schmitz and Rhinophrus Hsiao (Insecta: Hemiptera: Heteroptera: Miridae) from Thailand, with description of a new species. European Journal of Taxonomy 987: 61-80, DOI: 10.5852/ejt.2025.987.2859, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2859/12995
234487CB8126405A615DF9BBFD30F9B3.taxon	description	urn: lsid: zoobank. org: act: 1 C 3604 E 4 - 0 AC 4 - 42 D 9 - B 521 - 50 EA 19 F 2 A 5 A 5 Figs 1, 3 – 13, 16 – 17, 26 – 27, 50; Table 1	en	Wolski, Andrzej, Yasunaga, Tomohide (2025): First report of little-known cylapine genera Mimofulvius Schmitz and Rhinophrus Hsiao (Insecta: Hemiptera: Heteroptera: Miridae) from Thailand, with description of a new species. European Journal of Taxonomy 987: 61-80, DOI: 10.5852/ejt.2025.987.2859, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2859/12995
234487CB8126405A615DF9BBFD30F9B3.taxon	diagnosis	Diagnosis Currently known only by male specimens. Recognized by its shiny castaneous body; pronotum with narrow, distinct, yellow stipe along posterior margin (Figs 1, 16, 26 – 27); male genitalia as described below (Figs 3 – 5). These characters enables this species to be readily distinguished from the other congener, M. pentatomus (see below).	en	Wolski, Andrzej, Yasunaga, Tomohide (2025): First report of little-known cylapine genera Mimofulvius Schmitz and Rhinophrus Hsiao (Insecta: Hemiptera: Heteroptera: Miridae) from Thailand, with description of a new species. European Journal of Taxonomy 987: 61-80, DOI: 10.5852/ejt.2025.987.2859, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2859/12995
234487CB8126405A615DF9BBFD30F9B3.taxon	etymology	Etymology From Latin, ‘ castaneus ’ (‘ castaneous brown’), referring to the basic body coloration of this new species.	en	Wolski, Andrzej, Yasunaga, Tomohide (2025): First report of little-known cylapine genera Mimofulvius Schmitz and Rhinophrus Hsiao (Insecta: Hemiptera: Heteroptera: Miridae) from Thailand, with description of a new species. European Journal of Taxonomy 987: 61-80, DOI: 10.5852/ejt.2025.987.2859, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2859/12995
234487CB8126405A615DF9BBFD30F9B3.taxon	materials_examined	Type material Holotype THAILAND • ♂; Nakhon Ratchasima, Wang Nam Khiao, Sakaerat Environmental Research Station (SERS); 14 ° 30 ′ 27 ″ N, 101 ° 55 ′ 39 ″ E; 410 m alt.; 17 Mar. 2010; T. Yasunaga leg.; UV (mercury) light trap; DOAT, AMNH _ PBI 00380662. Paratypes THAILAND • 1 ♂; same data as for holotype; 18 Mar. 2010; T. Yasunaga leg.; UV light trap; TYCN • 1 ♂; same data as for holotype; 15 Sep. 2008; T. Yasunaga leg.; UV light trap; TYCN • 1 ♂; same data as for holotype; 12 – 14 Jun. 2009; Yasunaga and Yamada leg.; UV light trap; TYCN.	en	Wolski, Andrzej, Yasunaga, Tomohide (2025): First report of little-known cylapine genera Mimofulvius Schmitz and Rhinophrus Hsiao (Insecta: Hemiptera: Heteroptera: Miridae) from Thailand, with description of a new species. European Journal of Taxonomy 987: 61-80, DOI: 10.5852/ejt.2025.987.2859, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2859/12995
234487CB8126405A615DF9BBFD30F9B3.taxon	description	Description Male COLORATION. Dorsum castaneous with yellow areas (Figs 1, 16, 26 – 27). HEAD. Castaneous broadly tinged with dark brown on vertex, frons, and clypeus (Figs 1, 16 – 17, 26 – 27); antenna dark yellowish brown with dark brown, narrow annulations on apices of all segments and apex of basal half of subdivided segment IV (Figs 1, 16 – 17, 26 – 27); labium yellowish brown (Fig. 17). THORAX. Pronotum. Castaneous with distinct, narrow, yellow stipe along posterior margin (Figs 1, 16, 26 – 27). Mesoscutum and scutellum. Dark castaneous (Figs 1, 16, 26 – 27). Thoracic pleura. Proepimeron and proepisternum castaneous; remaining pleura dark yellowish brown; metathoracic scent gland evaporative areas and peritreme contrastingly yellow (Fig. 17). Hemelytron. Castaneous, with large pale-yellow patch above cuneus (Figs 1, 16, 26 – 27). Legs. Procoxa castaneous; meso- and metacoxae yellow; remaining segments dark yellowish brown (Figs 1, 16 – 17, 26 – 27). ABDOMEN. Dark castaneous (Fig. 17). STRUCTURE, TEXTURE, AND VESTITURE. Body 2.8 – 3.1 times as long as wide, 3.1 times as long as posterior width of pronotum. Dorsum shiny, covered with sparse, erect and semirecumbent setae (Figs 1, 6 – 8). Thorax. As in generic description. Hemelytron. Exocorial edge as in Fig. 8. Legs. Relatively long; metafemur with three narrow keels that are considered as plectra for stridulation (Fig. 13). MALE GENITALIA. Left paramere. Apical process narrowed toward apex, sharply pointed (Fig. 3). Endosoma. Membranous medial lobe occupying half of endosoma; left sclerite (ls) short, narrowed toward apex; right sclerite (rs) long, originating near base of sclerotized portion of seminal duct and terminating at basal two thirds of endosoma (Fig. 4). Measurements See Table 1. Female Unknown.	en	Wolski, Andrzej, Yasunaga, Tomohide (2025): First report of little-known cylapine genera Mimofulvius Schmitz and Rhinophrus Hsiao (Insecta: Hemiptera: Heteroptera: Miridae) from Thailand, with description of a new species. European Journal of Taxonomy 987: 61-80, DOI: 10.5852/ejt.2025.987.2859, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2859/12995
234487CB8126405A615DF9BBFD30F9B3.taxon	biology_ecology	Biology Unknown; all available specimens (e. g., Figs 26 – 27) were collected by UV lighting method (using mercury lamps).	en	Wolski, Andrzej, Yasunaga, Tomohide (2025): First report of little-known cylapine genera Mimofulvius Schmitz and Rhinophrus Hsiao (Insecta: Hemiptera: Heteroptera: Miridae) from Thailand, with description of a new species. European Journal of Taxonomy 987: 61-80, DOI: 10.5852/ejt.2025.987.2859, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2859/12995
234487CB8126405A615DF9BBFD30F9B3.taxon	distribution	Distribution Thailand (Nakhon Ratchasima) (Fig. 50).	en	Wolski, Andrzej, Yasunaga, Tomohide (2025): First report of little-known cylapine genera Mimofulvius Schmitz and Rhinophrus Hsiao (Insecta: Hemiptera: Heteroptera: Miridae) from Thailand, with description of a new species. European Journal of Taxonomy 987: 61-80, DOI: 10.5852/ejt.2025.987.2859, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2859/12995
234487CB81224057612BF999FE0FFB85.taxon	description	Figs 2, 14 – 15, 18 – 25, 28 – 34, 50	en	Wolski, Andrzej, Yasunaga, Tomohide (2025): First report of little-known cylapine genera Mimofulvius Schmitz and Rhinophrus Hsiao (Insecta: Hemiptera: Heteroptera: Miridae) from Thailand, with description of a new species. European Journal of Taxonomy 987: 61-80, DOI: 10.5852/ejt.2025.987.2859, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2859/12995
234487CB81224057612BF999FE0FFB85.taxon	diagnosis	Diagnosis Distinguished by the pronotum black with narrow, indistinct, yellowish stripe along the posterior margin (Figs 2, 14 – 15, 24 – 25); clavus with large, whitish stripe medially, occupying most of the claval surface (Figs 2, 14 – 15, 24 – 25); the female genitalia as presented and depicted by Yeshwanth et al. (2016: fig. 30). The male is herein documented for the first time (Figs 28 – 34).	en	Wolski, Andrzej, Yasunaga, Tomohide (2025): First report of little-known cylapine genera Mimofulvius Schmitz and Rhinophrus Hsiao (Insecta: Hemiptera: Heteroptera: Miridae) from Thailand, with description of a new species. European Journal of Taxonomy 987: 61-80, DOI: 10.5852/ejt.2025.987.2859, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2859/12995
234487CB81224057612BF999FE0FFB85.taxon	materials_examined	Type material Holotype INDIA • ♀; Mysore State, Chikkaballapura; 3000 ft.; Apr. 1917; Brit. Mus.; 1928 - 503, Pr? 679 - 120; BMNH. Additional examined material NEPAL • 1 ♂; Makawanpur District, Chitwan National Park, Machan Wildlife Resort; 27 ° 32 ′ 06 ″ N, 84 ° 44 ′ 17 ″ E; 7 – 9 Nov. 2005; T. Yasunaga et al. leg.; UV light trap; NMTU, AMNH _ PBI 00380661.	en	Wolski, Andrzej, Yasunaga, Tomohide (2025): First report of little-known cylapine genera Mimofulvius Schmitz and Rhinophrus Hsiao (Insecta: Hemiptera: Heteroptera: Miridae) from Thailand, with description of a new species. European Journal of Taxonomy 987: 61-80, DOI: 10.5852/ejt.2025.987.2859, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2859/12995
234487CB81224057612BF999FE0FFB85.taxon	description	Redescription Male Overall coloration and basic structure as in female (Figs 14, 24 – 25), but body more elongate and slenderer, parallel-sided (Figs 14, 24 – 25); basic coloration coffee brown; dorsum comparatively shining (Figs 14, 24 – 25); head weakly porrect, with eyes enlarged, reminiscent of those in Psallopinae Schuh, 1976 (Figs 14, 24 – 25; apical corium and anterior margin of cuneus tinged with red around white macula; posterior margin of metafemur with a keel (Fig. 21). GENITALIA. Apex of pygophore as in Fig. 28; left paramere elongate, C-shaped, with a median tooth inward (Figs 29 – 30); right paramere broadly margined apically (Figs 31 – 32); endosoma with two rather developed lobal-sclerites (Fig. 33). Measurements See Table 1.	en	Wolski, Andrzej, Yasunaga, Tomohide (2025): First report of little-known cylapine genera Mimofulvius Schmitz and Rhinophrus Hsiao (Insecta: Hemiptera: Heteroptera: Miridae) from Thailand, with description of a new species. European Journal of Taxonomy 987: 61-80, DOI: 10.5852/ejt.2025.987.2859, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2859/12995
234487CB81224057612BF999FE0FFB85.taxon	biology_ecology	Biology Unknown; a male individual (e. g., Figs 24 – 25) was collected by UV lighting method (using fluorescent lamps).	en	Wolski, Andrzej, Yasunaga, Tomohide (2025): First report of little-known cylapine genera Mimofulvius Schmitz and Rhinophrus Hsiao (Insecta: Hemiptera: Heteroptera: Miridae) from Thailand, with description of a new species. European Journal of Taxonomy 987: 61-80, DOI: 10.5852/ejt.2025.987.2859, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2859/12995
234487CB81224057612BF999FE0FFB85.taxon	distribution	Distribution India (Yeshwanth et al. 2016) and Nepal (Makawanpur) (this paper) (Fig. 50).	en	Wolski, Andrzej, Yasunaga, Tomohide (2025): First report of little-known cylapine genera Mimofulvius Schmitz and Rhinophrus Hsiao (Insecta: Hemiptera: Heteroptera: Miridae) from Thailand, with description of a new species. European Journal of Taxonomy 987: 61-80, DOI: 10.5852/ejt.2025.987.2859, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2859/12995
234487CB81224057612BF999FE0FFB85.taxon	discussion	Remarks Mimofulvius pentatomus is distinguished from M. castaneus by the black pronotum and large, medial, whitish patch on the clavus, and shape of the metafemoral plectrum (a single narrow keel) (Figs 2, 14 – 15, 21, 24 – 25).	en	Wolski, Andrzej, Yasunaga, Tomohide (2025): First report of little-known cylapine genera Mimofulvius Schmitz and Rhinophrus Hsiao (Insecta: Hemiptera: Heteroptera: Miridae) from Thailand, with description of a new species. European Journal of Taxonomy 987: 61-80, DOI: 10.5852/ejt.2025.987.2859, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2859/12995
234487CB812E4056615CFEDFFE0CFC7B.taxon	diagnosis	Diagnosis Easily distinguished from other cylapines by the following combination of characters: body strongly elongate (Figs 35, 43); head with protruding, pointed frons and compressed clypeus (Figs 43 – 45); procoxae much longer than meso- and metacoxae (Fig. 44); scent gland efferent system weakly developed, restricted to posterior angle of metepisternum (Fig. 46); central row of tiles on pretarsal unguitractor absent (Fig. 47).	en	Wolski, Andrzej, Yasunaga, Tomohide (2025): First report of little-known cylapine genera Mimofulvius Schmitz and Rhinophrus Hsiao (Insecta: Hemiptera: Heteroptera: Miridae) from Thailand, with description of a new species. European Journal of Taxonomy 987: 61-80, DOI: 10.5852/ejt.2025.987.2859, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2859/12995
234487CB812E4056615CFEDFFE0CFC7B.taxon	discussion	Remarks As misplaced previously under a unique hallodapine genus Acrorrhinium (Gorczyca 1994), Rhinophrus is characterized primarily by the remarkably slender body shape (Fig. 43) and a projection on the frons (Fig 44). However, the plausible systematic position of Rhinophorus is yet to be determined (see Discussion below).	en	Wolski, Andrzej, Yasunaga, Tomohide (2025): First report of little-known cylapine genera Mimofulvius Schmitz and Rhinophrus Hsiao (Insecta: Hemiptera: Heteroptera: Miridae) from Thailand, with description of a new species. European Journal of Taxonomy 987: 61-80, DOI: 10.5852/ejt.2025.987.2859, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2859/12995
234487CB812E40516143FBDEFAADFEC8.taxon	description	Figs 35 – 50	en	Wolski, Andrzej, Yasunaga, Tomohide (2025): First report of little-known cylapine genera Mimofulvius Schmitz and Rhinophrus Hsiao (Insecta: Hemiptera: Heteroptera: Miridae) from Thailand, with description of a new species. European Journal of Taxonomy 987: 61-80, DOI: 10.5852/ejt.2025.987.2859, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2859/12995
234487CB812E40516143FBDEFAADFEC8.taxon	materials_examined	Type material examined Holotype Malaysia • ♀; Sandakan Borneo (Baker); no further data; USNM. Additional material examined THAILAND • 1 ♀; Nakhon Ratchasima, Wang Nam Khiao, Sakaerat Environmental Research Station (SERS); 14 ° 30 ′ 27 ″ N, 101 ° 55 ′ 39 ″ E; 410 m alt.; 19 – 21 Aug. 2008; Yasunaga and Shishido leg.; UV light trap; DOAT.	en	Wolski, Andrzej, Yasunaga, Tomohide (2025): First report of little-known cylapine genera Mimofulvius Schmitz and Rhinophrus Hsiao (Insecta: Hemiptera: Heteroptera: Miridae) from Thailand, with description of a new species. European Journal of Taxonomy 987: 61-80, DOI: 10.5852/ejt.2025.987.2859, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2859/12995
234487CB812E40516143FBDEFAADFEC8.taxon	description	Description See Hsiao (1944) for further diagnostic characters. FEMALE GENITALIA. Bursa copulatrix nearly ovoid; sclerotized ring rounded; dorsal labiate plate (DLP) narrow, elongate toward caudal direction; seminal depository strongly inflated, globose, with pair of large, ovoid rings (Figs 37, 39 – 40); posterior wall of bursa copulatrix membranous, covered with minute, brush-like microprocesses (Fig. 38); gonapophysis 8 broadened subapically and distinctly pointed apically (Fig. 41); gonapophysis 9 pointed apically, strongly serrate subapically (Fig. 42). Rhinophrus borneensis can be distinguished from R. hsiaoi by black antennal segment I (brown in R. hsiaoi) and the shape of the parameres (Gorczyca 1996).	en	Wolski, Andrzej, Yasunaga, Tomohide (2025): First report of little-known cylapine genera Mimofulvius Schmitz and Rhinophrus Hsiao (Insecta: Hemiptera: Heteroptera: Miridae) from Thailand, with description of a new species. European Journal of Taxonomy 987: 61-80, DOI: 10.5852/ejt.2025.987.2859, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2859/12995
234487CB812E40516143FBDEFAADFEC8.taxon	distribution	Distribution Malaysia (Sandakan, Borneo) (Hsiao 1944); Thailand (Nakhon Ratchasima) (this paper) (Fig. 50).	en	Wolski, Andrzej, Yasunaga, Tomohide (2025): First report of little-known cylapine genera Mimofulvius Schmitz and Rhinophrus Hsiao (Insecta: Hemiptera: Heteroptera: Miridae) from Thailand, with description of a new species. European Journal of Taxonomy 987: 61-80, DOI: 10.5852/ejt.2025.987.2859, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2859/12995
