taxonID	type	description	language	source
1F1B8784F716FFF9FF73FB6FFD2D41F0.taxon	type_taxon	Type species. Anakitium storozhenkoi Arakelyan & Yan, sp. nov.	en	Arakelyan, Stepan A., Strelnikova, Olesya D., Felker, Anastasia S., Yan, Evgeny V. (2025): Rare full-body imprint of the giant Asiocoleidae (Coleoptera: Archostemata) from the Lower Triassic sheds light on the morphology of the family. Zootaxa 5715 (1): 41-52, DOI: 10.11646/zootaxa.5715.1.6, URL: https://doi.org/10.11646/zootaxa.5715.1.6
1F1B8784F716FFF9FF73FB6FFD2D41F0.taxon	diagnosis	Diagnosis. Very large beetle, body length (from mandible tips to abdominal apex) 25.6 mm. Head weakly transverse. Occiput with three pairs of closely spaced protuberances: rounded P 2 and oval P 3 on same level, with P 3 separated from P 2 by well-defined sulcus. P 4 present as smallest pair, located near junction of P 2 and P 3, close to pronotal anterior margin. Antennae moniliform; scape enlarged, twice as long as the pedicel. Mandibles bear two incisors. Pronotum transverse, nearly rectangular, with small protruding anterior angles. Apical tarsomeres longest. Elytral veins weakly defined. There are large intercalary rows between the 3 rd and 4 th, and between the 5 th and 6 th primary rows (counting from the sutural margin). Fifth abdominal ventrite (= sternite VII) sharply narrowed, subtriangular in shape.	en	Arakelyan, Stepan A., Strelnikova, Olesya D., Felker, Anastasia S., Yan, Evgeny V. (2025): Rare full-body imprint of the giant Asiocoleidae (Coleoptera: Archostemata) from the Lower Triassic sheds light on the morphology of the family. Zootaxa 5715 (1): 41-52, DOI: 10.11646/zootaxa.5715.1.6, URL: https://doi.org/10.11646/zootaxa.5715.1.6
1F1B8784F716FFF9FF73FB6FFD2D41F0.taxon	etymology	Etymology. The name of the new genus, Anakitium, is derived from the locality Anakit- 3, where the fossil was discovered. The genus name is neuter in gender.	en	Arakelyan, Stepan A., Strelnikova, Olesya D., Felker, Anastasia S., Yan, Evgeny V. (2025): Rare full-body imprint of the giant Asiocoleidae (Coleoptera: Archostemata) from the Lower Triassic sheds light on the morphology of the family. Zootaxa 5715 (1): 41-52, DOI: 10.11646/zootaxa.5715.1.6, URL: https://doi.org/10.11646/zootaxa.5715.1.6
1F1B8784F716FFFDFF73F8BBFBA9434C.taxon	description	(Figs 1 – 3)	en	Arakelyan, Stepan A., Strelnikova, Olesya D., Felker, Anastasia S., Yan, Evgeny V. (2025): Rare full-body imprint of the giant Asiocoleidae (Coleoptera: Archostemata) from the Lower Triassic sheds light on the morphology of the family. Zootaxa 5715 (1): 41-52, DOI: 10.11646/zootaxa.5715.1.6, URL: https://doi.org/10.11646/zootaxa.5715.1.6
1F1B8784F716FFFDFF73F8BBFBA9434C.taxon	materials_examined	Material. Holotype No. 5381 / 478, A well-preserved almost complete counterpart and the upper half of the part. Deposited in the A. A. Borissiak Paleontological Institute of the Russian Academy of Sciences (PIN RAS), Moscow, Russia.	en	Arakelyan, Stepan A., Strelnikova, Olesya D., Felker, Anastasia S., Yan, Evgeny V. (2025): Rare full-body imprint of the giant Asiocoleidae (Coleoptera: Archostemata) from the Lower Triassic sheds light on the morphology of the family. Zootaxa 5715 (1): 41-52, DOI: 10.11646/zootaxa.5715.1.6, URL: https://doi.org/10.11646/zootaxa.5715.1.6
1F1B8784F716FFFDFF73F8BBFBA9434C.taxon	diagnosis	Diagnosis. As for the genus (vide supra). Locality and horizon. Anakit- 3 (Bugarikhta Formation, Dvurogian Horizon).	en	Arakelyan, Stepan A., Strelnikova, Olesya D., Felker, Anastasia S., Yan, Evgeny V. (2025): Rare full-body imprint of the giant Asiocoleidae (Coleoptera: Archostemata) from the Lower Triassic sheds light on the morphology of the family. Zootaxa 5715 (1): 41-52, DOI: 10.11646/zootaxa.5715.1.6, URL: https://doi.org/10.11646/zootaxa.5715.1.6
1F1B8784F716FFFDFF73F8BBFBA9434C.taxon	description	Description. Head. (Fig. 2 A, B) weakly transverse (L / W = 1.25). Narrowed neck region absent. Vertexal region with three adjacent protuberances (Fig. 2 C). Antennal sockets covered by P 1, which has the same size as P 2. Eyes large (1.2 mm long), not protruding beyond outline of head, with partially carinated inner margins. Mandibles bidentate, curved; best-preserved right mandible (base to outer tooth) = 1.4 mm, not protruding forward. Penultimate and apical maxillary palpomere ratio 5: 3. Antennomeres 1 – 9 = 4.7 mm (Fig. 2 D); estimated full length about 6 mm. Scape rounded, 1.4 x wider than pedicel; scape, pedicel, and first flagellomere ratio 2: 1: 2. Second and following flagellomeres moniliform, subequal. Thorax. Pronotum subrectangular, transverse (W / L = 1.6); sides broadly angulate; posterior margin shallowly emarginate in front of scutellum. Dorsal side with faint longitudinal depression. Propleuron acutely triangular (Fig. 2 E). Scutellum oval (L / W = 1.4). Middle coxae transverse, completely separated by spear-shaped anteromedian process of metaventrite. Metaventrite max width about 5.3 mm; Metatrochantin exposed, narrow, blade-like, length about 0.9 mm. Legs. Forelegs: femora about 2.6 mm long; tibiae about 2.85 mm long; tarsomeres 1 – 5 ratio 0.4: 0.35: 0.4: 0.35: 1; last segment narrowest. Midlegs: left femur = 3.0 mm; left tibiae = 3.25; right metatarsomeres (Fig. 2 F) 1 – 5 ratio 0.8: 0.6: 0.4: 0.5: 0.9; first two segments widened anteriorly. Claws = 0.6 mm. Hindlegs: metacoxae lenght = 3.3; metatrochanter = 1.35 mm, tapered basally. Both femora incomplete; estimated length about 3.3 – 3.7 mm. First metatarsomere elongate, = 1.65 mm, second = 0.65 mm Elytra. (Fig. 3 A, B). Total length = 19.40 mm, maximum width at midlength = 5.25 mm. Extends 2.0 mm beyond abdominal apex. Base narrower than the remaining part; epipleuron widened proximally near humerus, elsewhere approximately 0.4 mm wide with traces of small tubercles. Apex rounded. Venation weakly visible: 2 A curved dorsally; M + Cu and Rs bent toward epipleuron (Rs short). Veins divide elytral surface into four fields (Fig. 3 A). Field I (between sutural margin and 2 A) proximally with six window punctures rows, one terminating abruptly at first quarter of elytral length (hereafter intercalary row Ir), another intercalary row represented by three small punctures. Medially four window punctures rows present in this field, apically only two rows remain. Field II (between 2 A and M + Cu) with smoothly curved rows toward humeral margin, two of which terminate abruptly (intercalary rows). Medially field contains four window punctures rows (in transverse section). Field III without intercalary rows, maintaining four rows up to apical part. In distal quarter, window punctures arrangement becomes uniform lacking well-defined rows. Window punctures density 2.5 per mm; longest rows contain approximately 45 – 50 punctures. Punctures rounded, diameter 0.15 – 0.20 mm, each with 3 – 5 maculae (Fig. 3 C). Abdomen. (Fig. 1). Cuticle tuberculate. Ventrite I mostly concealed by coxae; Ventrites I – V combined = 9.5 mm. Dimensions I – V (W / median L (mm )): 7.4 / 1.25; 6.95 / 2.2; 6.50 / 1.75; 5.40 / 1.45; 2.50 / 2.80. 5 ventrite apex rounded.	en	Arakelyan, Stepan A., Strelnikova, Olesya D., Felker, Anastasia S., Yan, Evgeny V. (2025): Rare full-body imprint of the giant Asiocoleidae (Coleoptera: Archostemata) from the Lower Triassic sheds light on the morphology of the family. Zootaxa 5715 (1): 41-52, DOI: 10.11646/zootaxa.5715.1.6, URL: https://doi.org/10.11646/zootaxa.5715.1.6
1F1B8784F716FFFDFF73F8BBFBA9434C.taxon	etymology	Etymology. The new species is named in honor of the entomologist Sergey Y. Storozhenko.	en	Arakelyan, Stepan A., Strelnikova, Olesya D., Felker, Anastasia S., Yan, Evgeny V. (2025): Rare full-body imprint of the giant Asiocoleidae (Coleoptera: Archostemata) from the Lower Triassic sheds light on the morphology of the family. Zootaxa 5715 (1): 41-52, DOI: 10.11646/zootaxa.5715.1.6, URL: https://doi.org/10.11646/zootaxa.5715.1.6
1F1B8784F716FFFDFF73F8BBFBA9434C.taxon	discussion	Remarks. Anakitium storozhenkoi sp. nov. is assigned to the family Asiocoleidae based on the elytral structure: the elytra bear rows of punctures not united in pairs; weakly expressed veins on the elytral surface separate rows with similar orientation (fields I – IV); the rows gradually curve towards the humeral bulge; window punctures rows widen toward the elytral base due to the appearance of intercalary ones. The fossil imprint shows better preservation of the counterimprint. Comparison of symmetrical body parts indicates the beetle underwent significant deformation caused by compression, resulting in multidirectional stretching. For instance, the right mesotibia is 0.4 mm longer than the left one and displays deformation marks. The area between the meso- and prothorax exhibits strong deformation with overlapping ventral and dorsal structures. Some elements likely represent the mesanepisternum and mesoventrite, but poor preservation prevents definitive identification. The middle coxae are poorly discernible but appear weakly transverse. The left metatibia is more completely preserved on direct imprint. The epipleural region of the elytra was likely compressed during fossilization. The short Rs vein probably extended laterally, reaching the basal part of the elytra.	en	Arakelyan, Stepan A., Strelnikova, Olesya D., Felker, Anastasia S., Yan, Evgeny V. (2025): Rare full-body imprint of the giant Asiocoleidae (Coleoptera: Archostemata) from the Lower Triassic sheds light on the morphology of the family. Zootaxa 5715 (1): 41-52, DOI: 10.11646/zootaxa.5715.1.6, URL: https://doi.org/10.11646/zootaxa.5715.1.6
