taxonID	type	description	language	source
1A53878787561F0DFF704AC6863BF89A.taxon	materials_examined	Type species. Neoapterocis mexicanus sp. nov. Included species. Neoapterocis chilensis sp. nov., Neoapterocis mexicanus sp. nov.	en	Lopes-Andrade, Cristiano (2007): Neoapterocis, a new genus of apterous Ciidae (Coleoptera: Tenebrionoidea) from Chile and Mexico. Zootaxa 1481: 35-47, DOI: 10.5281/zenodo.176885
1A53878787561F0DFF704AC6863BF89A.taxon	etymology	Etymology. " Neo " means “ new ”, and “ apterocis ” refers to an apterous Hawaiian ciid genus with a very similar body form. However, this does not suggest any phylogenetic relationship, but just morphological similarities. The noun is masculine.	en	Lopes-Andrade, Cristiano (2007): Neoapterocis, a new genus of apterous Ciidae (Coleoptera: Tenebrionoidea) from Chile and Mexico. Zootaxa 1481: 35-47, DOI: 10.5281/zenodo.176885
1A53878787561F0DFF704AC6863BF89A.taxon	diagnosis	Diagnosis. The genus may be distinguished by the combination of distinct oval, convex body, small eyes and scutellum, absence of hind wings, short prosternum with short and broad intercoxal process, short metaventrite, and first abdominal ventrite longer than the next two together.	en	Lopes-Andrade, Cristiano (2007): Neoapterocis, a new genus of apterous Ciidae (Coleoptera: Tenebrionoidea) from Chile and Mexico. Zootaxa 1481: 35-47, DOI: 10.5281/zenodo.176885
1A53878787561F0DFF704AC6863BF89A.taxon	description	Description. Length 1.11 – 1.84 mm. Body oval, between 1.75 and 2 X as long as wide, convex (GD / EW = 0.72 – 0.86), pubescent, vestiture consisting of small shiny yellowish setae best seen in lateral view. Head, pronotum and elytra punctation single, irregularly distributed; each puncture bearing a short, procumbent to semi-erect seta. Labial palpi (Fig. 11) small, inserted at the apex of prementum; apical segment cylindrical. Maxilla (Fig. 11) with galea small and semioval; lacinia semicircular. Eyes small and prominent, coarsely facetted, each with no more than 20 ommatidia, the individual facets strongly convex. Antennae (Figs 4, 10) inserted beyond eyes, 10 - segmented, third antennomere long and slender; funicle subglabrous; club three-segmented, loose and more setose than funicle; each segment of the antennal club bearing four sensillifers, each formed by a group of small and cylindrical, not well-organized sensilla. Lateral pronotal margins conspicuous, finely crenulate, visible for most of their lengths from above; anterior pronotal angles distinctly produced forward and broadly rounded. Pronotum with anterior margin simple in both sexes; surface densely punctate and setose; punctures small, their diameters smaller than the length of a seta; interstices between punctures microreticulate. Scutellum triangular, indistinct, barely visible at magnifications of less than 60 X. Elytra more sparsely punctate than pronotum, hence less setose; punctures similar to those on pronotum; elytra fused to each other, strongly convex, humeri rounded; lateral margins strong, with conspicuous epipleura below wider anteriorly and tapering to apex. Hind wings absent. Procoxae transverse, not projecting below the plane of the intercoxal process of prosternum. Tibiae not expanded to apices, with a row of spines at apex, simple outer edge and outer apical angle rounded (Figs 5, 12). Tarsi 4 - segmented, appearing as 3 - segmented at magnifications lower than 100 X (Fig. 12, arrows). Prosternum short, biconcave, with a broad longitudinal carina at midline; intercoxal process short and broad, as long as the prosternum length at its midline (Figs 3, 9). Mesocoxae narrowly separated, metacoxae not contiguous. Metaventrite shorter than the first abdominal ventrite at midline; discrimen absent, or around one seventh to one fifth the metaventrite length at midline. First abdominal ventrite as long as at least the next two abdominal ventrites together; male with a distinct setose patch at the midline near the posterior margin. Male genitalia (Figs 6, 13) with ninth segment V-shaped, its apices lanceolate; anterior margin of eighth sternite broadly emarginate; aedeagus subquadrate, barely sclerotized; tegmen longer than median lobe, with a longitudinal emargination from apex to halfway along its length, forming two lateral lobes; median lobe (penis) cylindrical. Female genitalia (Fig. 14) with a distinct pair of gonostyli at apex; gonocoxites transversely divided into four parts; proximal gonocoxites with a pair of transversal bacula; paraprocts and proctiger as long as gonostyli and gonocoxites together; both paraprocts and proctiger with a pair of distinct longitudinal bacula; spiculum ventrale just slightly longer than gonostyli, gonocoxites and paraprocts together.	en	Lopes-Andrade, Cristiano (2007): Neoapterocis, a new genus of apterous Ciidae (Coleoptera: Tenebrionoidea) from Chile and Mexico. Zootaxa 1481: 35-47, DOI: 10.5281/zenodo.176885
1A53878787561F0DFF704AC6863BF89A.taxon	distribution	Distribution. Known from the state of Oaxaca (Mexico) and Concepción Province (Chile).	en	Lopes-Andrade, Cristiano (2007): Neoapterocis, a new genus of apterous Ciidae (Coleoptera: Tenebrionoidea) from Chile and Mexico. Zootaxa 1481: 35-47, DOI: 10.5281/zenodo.176885
1A53878787561F0DFF704AC6863BF89A.taxon	discussion	Comments. The suprageneric classification of Ciidae deserves a revision, but as it stands Neoapterocis gen. nov. would be placed in the tribe Ciini of the subfamily Ciinae. It lacks strongly projecting procoxae and tibiae spinose on outer edge that characterize both Orophiini (Lawrence 1974) and Xylographellini (Kawanabe & Miyatake 1996) and the distinctive type of antennal club, prementum and male genitalia found in the latter tribe.	en	Lopes-Andrade, Cristiano (2007): Neoapterocis, a new genus of apterous Ciidae (Coleoptera: Tenebrionoidea) from Chile and Mexico. Zootaxa 1481: 35-47, DOI: 10.5281/zenodo.176885
1A53878787511F0FFF704EE683C7FBE1.taxon	etymology	Etymology. The specific epithet refers to Chile, terra typica of this species.	en	Lopes-Andrade, Cristiano (2007): Neoapterocis, a new genus of apterous Ciidae (Coleoptera: Tenebrionoidea) from Chile and Mexico. Zootaxa 1481: 35-47, DOI: 10.5281/zenodo.176885
1A53878787511F0FFF704EE683C7FBE1.taxon	diagnosis	Diagnosis. This species is easily distinguished from Neo. mexicanus sp. nov. by the presence of two small tubercles on the male frontoclypeal ridge (Figs 1 – 3, arrows) and the lack of emarginations on the apical lobes of male tegmen (Fig. 6). It is generally smaller than Neo. mexicanus sp. nov., but with the abdominal setose patch of the male proportionally larger and marginated.	en	Lopes-Andrade, Cristiano (2007): Neoapterocis, a new genus of apterous Ciidae (Coleoptera: Tenebrionoidea) from Chile and Mexico. Zootaxa 1481: 35-47, DOI: 10.5281/zenodo.176885
1A53878787511F0FFF704EE683C7FBE1.taxon	description	Description. Holotype. ɗ, measurements in mm: TL 1.26; PL 0.42; PW 0.58; EL 0.79; EW 0.68; GD 0.53. Ratios: PL / PW 0.73; EL / EW 1.15; EL / PL 1.88; GD / EW 0.77; TL / EW 1.85. Upper surfaces brown; undersurfaces brown, but antennae, mouthparts, legs and ventrites yellowish brown. Abdominal ventrites and legs densely covered with setae; each seta more than four times longer than those on the elytra. Head with frontoclypeal ridge bearing two small tubercles (Figs 1 – 3, arrows), separated by a distance of an eye width; lateral margins broadly rounded. Length of the antennomeres (in mm): 0.055; 0.033; 0.041; 0.024; 0.012; 0.012; 0.012; 0.035; 0.034; 0.058. Pronotum with lateral margins visible from above, except for posterior onethird. Epipleura rather smooth, with small sparse punctures anteriorly and larger punctures posteriorly. Prosternum distinctly elevated at midline; intercoxal process subparallel sided, margined; punctures large and coalescent, giving a rugose impression to the surface. Disc of mesoventrite barely visible, punctation similar to that of the intercoxal process of prosternum. Metaventrite without distinct discrimen; disc with sparse punctures, subglabrous. First abdominal ventrite as long as the next three abdominal ventrites together; setose patch circular, marginate, its diameter one-fourth the ventrite length at the midline. Aedeagus (Fig. 6) with apical lobes of tegmen simple. Median lobe of aedeagus (penis) wider near halfway its length, apex sclerotized and subtriangular. Females. Unknown.	en	Lopes-Andrade, Cristiano (2007): Neoapterocis, a new genus of apterous Ciidae (Coleoptera: Tenebrionoidea) from Chile and Mexico. Zootaxa 1481: 35-47, DOI: 10.5281/zenodo.176885
1A53878787511F0FFF704EE683C7FBE1.taxon	materials_examined	Type series. Holotype. ɗ, Chile: labeled “ Chile / Concepcion Prov / Periquillo / 30. ii. 97 / leg. T. Cekalovic ” (MZSP). Paratypes (three specimens; LAPC). Chile: 2 ɗɗ, same data as holotype; 1 ɗ labeled “ Chile / Concepcion Prov / Toma / 12. xii. 1982 / leg. T. Cekalovic ”.	en	Lopes-Andrade, Cristiano (2007): Neoapterocis, a new genus of apterous Ciidae (Coleoptera: Tenebrionoidea) from Chile and Mexico. Zootaxa 1481: 35-47, DOI: 10.5281/zenodo.176885
1A53878787511F0FFF704EE683C7FBE1.taxon	description	Variation. Measurements in mm (n = 4, including the holotype): TL 1.11 – 1.32 (1.24 ± 0.09); PL 0.42 – 0.47 (0.45 ± 0.03); PW 0.53 – 0.63 (0.57 ± 0.05); EL 0.68 – 0.79 (0.76 ± 0.05); EW 0.63 – 0.74 (0.68 ± 0.04); GD 0.47 – 0.63 (0.54 ± 0.07). Ratios: PL / PW 0.73 – 0.90 (0.79 ± 0.08); EL / EW 1.08 – 1.15 (1.12 ± 0.04); EL / PL 1.63 – 1.88 (1.71 ± 0.11); GD / EW 0.75 – 0.86 (0.79 ± 0.05); TL / EW 1.75 – 1.85 (1.81 ± 0.05).	en	Lopes-Andrade, Cristiano (2007): Neoapterocis, a new genus of apterous Ciidae (Coleoptera: Tenebrionoidea) from Chile and Mexico. Zootaxa 1481: 35-47, DOI: 10.5281/zenodo.176885
1A53878787511F0FFF704EE683C7FBE1.taxon	biology_ecology	Biology. No information was available.	en	Lopes-Andrade, Cristiano (2007): Neoapterocis, a new genus of apterous Ciidae (Coleoptera: Tenebrionoidea) from Chile and Mexico. Zootaxa 1481: 35-47, DOI: 10.5281/zenodo.176885
1A53878787511F0FFF704EE683C7FBE1.taxon	distribution	Distribution. Known from Chile, province of Concepción (VIII región del Biobío).	en	Lopes-Andrade, Cristiano (2007): Neoapterocis, a new genus of apterous Ciidae (Coleoptera: Tenebrionoidea) from Chile and Mexico. Zootaxa 1481: 35-47, DOI: 10.5281/zenodo.176885
1A53878787521F00FF704CE6843DFD01.taxon	etymology	Etymology. The specific epithet refers to Mexico, terra typica of this species.	en	Lopes-Andrade, Cristiano (2007): Neoapterocis, a new genus of apterous Ciidae (Coleoptera: Tenebrionoidea) from Chile and Mexico. Zootaxa 1481: 35-47, DOI: 10.5281/zenodo.176885
1A53878787521F00FF704CE6843DFD01.taxon	diagnosis	Diagnosis. This species is easily distinguished from Neo. chilensis sp. nov. by the absence of tubercles on the male frontoclypeal ridge (Figs 7 – 9) and the presence of emarginations on the outer edge of the apical lobes of tegmen (Fig. 13, arrows). It is generally larger than Neo. chilensis sp. nov., but with the abdominal setose patch of the male proportionally smaller and not marginate.	en	Lopes-Andrade, Cristiano (2007): Neoapterocis, a new genus of apterous Ciidae (Coleoptera: Tenebrionoidea) from Chile and Mexico. Zootaxa 1481: 35-47, DOI: 10.5281/zenodo.176885
1A53878787521F00FF704CE6843DFD01.taxon	description	Description. Holotype. ɗ, measurements in mm: TL 1.63; PL 0.58; PW 0.79; EL 1.00; EW 0.89; GD 0.68. Ratios: PL / PW 0.73; EL / EW 1.12; EL / PL 1.73; GD / EW 0.76; TL / EW 1.82. Upper surfaces dark brown, almost black; undersurfaces dark brown, but antennae, mouthparts, legs, and ventrites reddish brown. Ventrites and legs bearing sparse setae; each seta more than four times longer than those on the elytra. Head with frontoclypeal ridge simple (Figs 7 – 9); lateral margins slightly angulate, but angles rounded. Length of the antennomeres (in mm): 0.053; 0.042; 0.044; 0.029; 0.021; 0.018; 0.015; 0.032; 0.033; 0.067. Pronotum with lateral margins visible from above, except for anterior angles. Epipleura with small sparse punctures; subglabrous, setae small, indistinct. Prosternum elevated at midline, lateral concavities not very distinct; intercoxal process marginate, slightly wider near apex; punctures distinct, deep, very close to each other but not coalescent. Disc of mesoventrite easily visible, bearing punctures similar to those on the intercoxal process of prosternum. Metaventrite with a small discrimen, from the posterior margin to one-seventh the metaventrite length at midline. First abdominal ventrite longer than the next two abdominal ventrites together; setose patch semicircular, not marginate, its median length approximately one seventh of the median length of the ventrite. Aedeagus with tegmen bearing a small emargination on the outer edge of both apical lobes (Fig. 13, arrows). Median lobe (penis) of aedeagus cylindrical, with apex not sclerotized and broadly rounded. Females. First abdominal ventrite devoid of setose patch.	en	Lopes-Andrade, Cristiano (2007): Neoapterocis, a new genus of apterous Ciidae (Coleoptera: Tenebrionoidea) from Chile and Mexico. Zootaxa 1481: 35-47, DOI: 10.5281/zenodo.176885
1A53878787521F00FF704CE6843DFD01.taxon	materials_examined	Type series. Holotype. ɗ, Mexico: labeled " Mexico: Oaxaca / 70 Km S Valle Nacional / 2600 m, 3. x. 1986 / R. Baranowski ", " sifting litter, ± at hollow dec. trees, pine-oak forest " (MZLU). Paratypes (14 specimens; 7 in LAPC, 7 in MZLU). Mexico: 11 specimens labeled " Mexico: Oaxaca 26 Km / E Teotitlan del Campo / 2250 m, 26. ix. 1990 / leg. R. Baranowski ", " Sifting litter, mixed oak forest "; one specimen " Mexico: Oaxaca / 70 Km S Valle Nacional / 2500 m, 10. ix. 1986 / R. Baranowski ", " sifting litter, pine-oak forest "; one specimen " Mexico: Oaxaca / 70 Km S Valle Nacional / 2600 m, 3. x. 1986 / R. Baranowski ", " sifting litter, ± at hollow dec. trees, pine-oak forest "; one specimen " Mexico: Oaxaca, 87 Km N / Oaxaca City, 2700 m / 13. xi. 1989 / leg. R. Baranowski ", " Sifting litter, mixed pine forest ".	en	Lopes-Andrade, Cristiano (2007): Neoapterocis, a new genus of apterous Ciidae (Coleoptera: Tenebrionoidea) from Chile and Mexico. Zootaxa 1481: 35-47, DOI: 10.5281/zenodo.176885
1A53878787521F00FF704CE6843DFD01.taxon	description	Variation. Measurements in mm (n = 6, including the holotype): TL 1.58 – 1.84 (1.68 ± 0.10); PL 0.53 – 0.58 (0.54 ± 0.03); PW 0.74 – 0.84 (0.77 ± 0.04); EL 1.00 – 1.26 (1.11 ± 0.11); EW 0.79 – 0.95 (0.87 ± 0.06); GD 0.63 – 0.68 (0.67 ± 0.03). Ratios: PL / PW 0.69 – 0.73 (0.71 ± 0.02); EL / EW 1.12 – 1.35 (1.27 ± 0.10); EL / PL 1.73 – 2.30 (2.03 ± 0.21); GD / EW 0.72 – 0.81 (0.77 ± 0.03); TL / EW 1.82 – 2.00 (1.93 ± 0.07). Three specimens of the type series are slightly yellowish or pale greenish, probably not fully pigmented adults. In some specimens, the prosternum may be more distinctly carinate at midline, the lateral concavities more easily seen, and the punctures both on the prosternum disc and on its intercoxal process not so deep. The metaventrite discrimen varies from one-seventh to one-fifth the metaventrite length at midline. Such variation is found in other Ciidae species, and is probably due to the age of adult specimens when alive (for instance, older adults generally have less conspicuous carinae and punctures).	en	Lopes-Andrade, Cristiano (2007): Neoapterocis, a new genus of apterous Ciidae (Coleoptera: Tenebrionoidea) from Chile and Mexico. Zootaxa 1481: 35-47, DOI: 10.5281/zenodo.176885
1A53878787521F00FF704CE6843DFD01.taxon	biology_ecology	Biology. The specimens were collected sifting litter under shrubs, at logs and at hollows of deciduous trees, in mixed pine and oak forests.	en	Lopes-Andrade, Cristiano (2007): Neoapterocis, a new genus of apterous Ciidae (Coleoptera: Tenebrionoidea) from Chile and Mexico. Zootaxa 1481: 35-47, DOI: 10.5281/zenodo.176885
1A53878787521F00FF704CE6843DFD01.taxon	distribution	Distribution. Known from the state of Oaxaca (Mexico).	en	Lopes-Andrade, Cristiano (2007): Neoapterocis, a new genus of apterous Ciidae (Coleoptera: Tenebrionoidea) from Chile and Mexico. Zootaxa 1481: 35-47, DOI: 10.5281/zenodo.176885
