identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
144D8797FFB17524FF0BFF2AFC4CE78C.text	144D8797FFB17524FF0BFF2AFC4CE78C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhinophis karinthandani Sampaio & Narayanan & Cyriac & Venu & Gower 2020	<div><p>Rhinophis karinthandani sp. nov.</p><p>Figs. 1–3, 6–9, Tables 1–3, Appendices 1–4 urn:lsid:zoobank.org:act: 95B061B8-AC6E-4996-8740-4D8F2BDA8CCD</p></div>	https://treatment.plazi.org/id/144D8797FFB17524FF0BFF2AFC4CE78C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sampaio, Filipa L.;Narayanan, Surya;Cyriac, Vivek Philip;Venu, Govindappa;Gower, David J.	Sampaio, Filipa L., Narayanan, Surya, Cyriac, Vivek Philip, Venu, Govindappa, Gower, David J. (2020): A new Indian species of Rhinophis Hemprich, 1820 closely related to R. sanguineus Beddome, 1863 (Serpentes: Uropeltidae). Zootaxa 4881 (1): 1-24, DOI: 10.11646/zootaxa.4881.1.1
144D8797FFBF752AFF0BFAC5FA04E08F.text	144D8797FFBF752AFF0BFAC5FA04E08F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhinophis	<div><p>Key to the Indian species of Rhinophis</p><p>This is an updated version of the key presented by Cyriac et al. (2020), revised with the addition of R. karinthandani sp. nov. We have removed the relative rostral length character used in Cyriac et al.’s (2020) key because we are less convinced of its diagnostic value and because it can be difficult to measure. All six species are endemic to India and are the only members of this genus known to occur in India.</p><p>1 Number of dorsal scale rows just behind midbody 17......................................................... 2</p><p>- Number of dorsal scale rows just behind midbody 15......................................................... 3</p><p>2 Ventral shields fewer than 160............................................................... R. travancoricus</p><p>- Ventral shields more than 180.................................................................... R. goweri</p><p>3 Ventral surface with dark blotches in irregular, staggered, transverse half-bands.................................... 4</p><p>- Ventral surface with irregular dark speckles or spots.......................................................... 5</p><p>4 Only known specimen a female with 195 ventral and 5,5 subcaudal shields (from Cardamom Hills of Western Ghats, South of Palghat Gap)............................................................................. R. fergusonianus</p><p>- Ventral shields more than 215; subcaudals 6 or more pairs (from Wayanad region of Western Ghats, North of Palghat Gap)......................................................................................... R. melanoleucus</p><p>5 Subcaudals 5 (more typically 6)–11 (5– 7 females; 8– 11 males); venter background colour bright red in life (paler, more orangeyellow in smaller specimens).................................................................. R. sanguineus</p><p>- Subcaudals 4–8 (4– 5 females; 6– 8 males); venter background colour whitish (to pale pink anteriorly) in life.............................................................................................. R. karinthandani sp. nov.</p></div>	https://treatment.plazi.org/id/144D8797FFBF752AFF0BFAC5FA04E08F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sampaio, Filipa L.;Narayanan, Surya;Cyriac, Vivek Philip;Venu, Govindappa;Gower, David J.	Sampaio, Filipa L., Narayanan, Surya, Cyriac, Vivek Philip, Venu, Govindappa, Gower, David J. (2020): A new Indian species of Rhinophis Hemprich, 1820 closely related to R. sanguineus Beddome, 1863 (Serpentes: Uropeltidae). Zootaxa 4881 (1): 1-24, DOI: 10.11646/zootaxa.4881.1.1
144D8797FFA37536FF0BF827FA9AE0A1.text	144D8797FFA37536FF0BF827FA9AE0A1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhinophis fergusonianus Boulenger 1896	<div><p>Rhinophis fergusonianus</p><p>BMNH 1946.1.16.77 (formerly BMNH 95.7.29.1) holotype, Cardamom Hills, “Travancore” [probably Kerala] .</p></div>	https://treatment.plazi.org/id/144D8797FFA37536FF0BF827FA9AE0A1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sampaio, Filipa L.;Narayanan, Surya;Cyriac, Vivek Philip;Venu, Govindappa;Gower, David J.	Sampaio, Filipa L., Narayanan, Surya, Cyriac, Vivek Philip, Venu, Govindappa, Gower, David J. (2020): A new Indian species of Rhinophis Hemprich, 1820 closely related to R. sanguineus Beddome, 1863 (Serpentes: Uropeltidae). Zootaxa 4881 (1): 1-24, DOI: 10.11646/zootaxa.4881.1.1
144D8797FFA07535FF0BFF2BFC1BE7A4.text	144D8797FFA07535FF0BFF2BFC1BE7A4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhinophis goweri Aengals & Ganesh 2013	<div><p>Rhinophis goweri</p><p>BNHS 3465, near Solakkadu, Kolli Hills, Namakkal district, Tamil Nadu.</p></div>	https://treatment.plazi.org/id/144D8797FFA07535FF0BFF2BFC1BE7A4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sampaio, Filipa L.;Narayanan, Surya;Cyriac, Vivek Philip;Venu, Govindappa;Gower, David J.	Sampaio, Filipa L., Narayanan, Surya, Cyriac, Vivek Philip, Venu, Govindappa, Gower, David J. (2020): A new Indian species of Rhinophis Hemprich, 1820 closely related to R. sanguineus Beddome, 1863 (Serpentes: Uropeltidae). Zootaxa 4881 (1): 1-24, DOI: 10.11646/zootaxa.4881.1.1
144D8797FFA07535FF0BFEBFFB1DE5BC.text	144D8797FFA07535FF0BFEBFFB1DE5BC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhinophis karinthandani Sampaio & Narayanan & Cyriac & Venu & Gower 2020	<div><p>Rhinophis karinthandani sp. nov.</p><p>BMNH 79.7.4.2, holotype, Manantoddy, Wynad, 2,000 ft [= Mananthavady, Wayanad district, Kerala] .</p><p>BNHS 3540, BNHS 3542, 3544, 3545 and 3546, paratypes, Lakkidi, Wayanad district, Kerala .</p><p>ZSI/ WGRC /V-3143, paratype, Chandanathodu near Mananthawady, Wayanad district, Kerala .</p><p>BNHS 3541 and 3543, Lakkidi, Wayanad district, Kerala.</p><p>BMNH 79.7.4.3, Manantoddy, Wynad, 2,000 ft [= Mananthavady, Wayanad district, Kerala][originally in same jar as holotype BMNH 79.7.4.2]; lacks head and anterior end of body .</p><p>BMNH 74.4.29.109, Wynad [Wayanad district, Kerala].</p><p>BNHS 3605 and BNHS 3606, near Pookode, Wayanad district, Kerala. One of these is from nr. Pozhutana and one from Sugandhagiri.</p><p>BNHS 3153, Anakkampoil, Wayanad district, Kerala; listed as R. sanguineus in Cyriac et al. (2000) .</p><p>ZMB 10358, Wayanad, India; listed as R. sanguineus in Cyriac et al. (2000) .</p><p>MNHN 1895.75, “ Sri Lanka ” [presumably in error]; listed as R. sanguineus in Cyriac et al. (2000) .</p></div>	https://treatment.plazi.org/id/144D8797FFA07535FF0BFEBFFB1DE5BC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sampaio, Filipa L.;Narayanan, Surya;Cyriac, Vivek Philip;Venu, Govindappa;Gower, David J.	Sampaio, Filipa L., Narayanan, Surya, Cyriac, Vivek Philip, Venu, Govindappa, Gower, David J. (2020): A new Indian species of Rhinophis Hemprich, 1820 closely related to R. sanguineus Beddome, 1863 (Serpentes: Uropeltidae). Zootaxa 4881 (1): 1-24, DOI: 10.11646/zootaxa.4881.1.1
144D8797FFA07535FF0BFD47FCC7E49C.text	144D8797FFA07535FF0BFD47FCC7E49C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhinophis melanoleucus	<div><p>Rhinophis melanoleucus</p><p>BNHS 3534, holotype, Wayanad Wild resort, Lakkidi, Wayanad district, Kerala .</p><p>BNHS 3537 and BNHS 3538, paratypes, Lakkidi, Wayanad district, Kerala .</p><p>BNHS 3535, BNHS 3536 and ZSI /WGRC/IR/ V/3101, paratypes, Kerala Veterinary and Animal Sciences University, Pookode, Wayanad district, Kerala .</p><p>ZSI/ WGRC /IR/ V/3100, paratype, Vythiri road, Pookode, Wayanad district, Kerala (female) , Kerala .</p><p>BNHS 3539, Wayanad Wild resort, Lakkidi, Wayanad district, Kerala.</p></div>	https://treatment.plazi.org/id/144D8797FFA07535FF0BFD47FCC7E49C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sampaio, Filipa L.;Narayanan, Surya;Cyriac, Vivek Philip;Venu, Govindappa;Gower, David J.	Sampaio, Filipa L., Narayanan, Surya, Cyriac, Vivek Philip, Venu, Govindappa, Gower, David J. (2020): A new Indian species of Rhinophis Hemprich, 1820 closely related to R. sanguineus Beddome, 1863 (Serpentes: Uropeltidae). Zootaxa 4881 (1): 1-24, DOI: 10.11646/zootaxa.4881.1.1
144D8797FFA07535FF0BFBA7FADDE104.text	144D8797FFA07535FF0BFBA7FADDE104.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhinophis sanguineus	<div><p>Rhinophis sanguineus</p><p>BMNH 1946.1.16.54 (formerly BMNH 64.3.9.2) lectotype (see Pyron et al. 2016), Cherambady [Cherambadi, Nilgiris district, Tamil Nadu] .</p><p>BMNH 1946.1.16.76 (formerly BMNH 74.4.29.110) holotype of R. microlepis, Wayanad district, Kerala .</p><p>BMNH 74.2.29.695–697, Wynad [Wayanad district, Kerala].</p><p>BMNH 68.4.3.38, Nilgherries [possibly Wayanad district, Kerala or Nilgiris district, Tamil Nadu] .</p><p>BMNH 1922.5.25.6, Nellacolla [Nellacotta?, Nilgiris district, Tamil Nadu].</p><p>BMNH 74.4.29.108, Wynad [Wayanad district, Kerala].</p><p>BMNH 66.12.15.3, Wynad, Madras Presidency [possibly Wayanad district, Kerala or Nilgiris district, Tamil Nadu] .</p><p>ZMB 5536, Cherambady [Cherambadi, Nilgiris district, Tamil Nadu].</p><p>VPRS0918092 and VPRS0918093, Meppadi, Wayanad district, Kerala.</p><p>BUB S-2, BUB S-4, BUB S-15, BUB S-1584, BUB S-1588, Regional Agricultural Research Station (RARS), Ambalavayal, Wayanad district, Kerala.</p><p>MNHN 1897.246, “ Sri Lanka ” [probably in error]. See Discussion for comments on colour pattern and identification of this specimen.</p><p>MCZ 3865 and MCZ 3854b, “Madras” [probably the historical administrative region of Madras Presidency, which included parts of present-day Wayanad]. See Discussion for comments on colour pattern and identification of MCZ 3854b.</p></div>	https://treatment.plazi.org/id/144D8797FFA07535FF0BFBA7FADDE104	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sampaio, Filipa L.;Narayanan, Surya;Cyriac, Vivek Philip;Venu, Govindappa;Gower, David J.	Sampaio, Filipa L., Narayanan, Surya, Cyriac, Vivek Philip, Venu, Govindappa, Gower, David J. (2020): A new Indian species of Rhinophis Hemprich, 1820 closely related to R. sanguineus Beddome, 1863 (Serpentes: Uropeltidae). Zootaxa 4881 (1): 1-24, DOI: 10.11646/zootaxa.4881.1.1
144D8797FFA07532FF0BF91FFB4EE23C.text	144D8797FFA07532FF0BF91FFB4EE23C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhinophis travancoricus	<div><p>Rhinophis travancoricus</p><p>BMNH 92.10.5.2, holotype, 6 miles along Trivandrum-Vambayam road, Thiruvananthapuram district, Kerala .</p><p>BMNH 1903.4.7.1–2 and BMNH 99.11.16.1, Trivandrum, Thiruvananthapuram district, Kerala.</p><p>BMNH 94.3.15.1, Piermed [Peermade, Peermedu], Idukki district, Kerala.</p><p>CAS 244341, Sakrapani Mukku, near Kulithurai river, Kanyakumari district, Tamil Nadu.</p><p>CAS 39620, Trivandrum, Thiruvananthapuram district, Kerala .</p><p>MW219 and MW221, Palod, Thiruvananthapuram district, Kerala.</p><p>MW 2180, MW 2182, MW 2183 and MW 2184, Maramalai, Kanyakumari district, Tamil Nadu.</p><p>VPRT1115073, near Konni, Pathanamthitta district, Kerala.</p><p>APPENDIX 2. GenBank accession numbers for sequences of uropeltids used in phylogenetic analyses.All sequences were previously published except for those accession codes in bold text. Species identifications follow those reported on GenBank and/or in papers in which data were reported, except for: * (listed as R. blythii in LSMNS catalogue but identified by Jins et al. 2018 as R. drummondhayi on basis of DNA data), and ** (following Gower 2020; listed as R. homolepis in previous studies).Abbreviations not listed in the Materials and Methods or by Pyron et al. (2016) or Cyriac &amp; Kodandaramaiah (2017) as follows: KA: Karnataka; KL: Kerala; LSUMNS: Louisiana State University Museum of Natural Science; NMSL: National Museum, Sri Lanka, Colombo field tag; NP: National Park; UK: University of Kerala field tag; WHT: World Heritage Trust specimens housed in NMSL. VP: V.P. Cyriac field tag. Blank cell indicates lack of data. In the Sample column, numbers in parentheses indicate voucher code where this is known to differ from tissue code.</p><p>......continued on the next page</p><p>APPENDIX 2. (Continued)</p><p>APPENDIX 3. Coordinates for Rhinophis records mapped in Fig. 3. * indicates type locality; ** indicates possible approximate location of Mr Minchin’s Estate, type locality of R. microlepis . Coordinates reported to less than three decimal places are estimated from online maps.</p><p>APPENDIX 4. Dorsal scale row reductions in holotype (*) and paratypes (†) and referred specimens of Rhinophis karinthandani sp. nov., and comparative specimens of R. sanguineus . Dorsal scale rows were not counted anterior to approximately the tenth ventral scale, or adjacent to approximately last five ventrals anterior to the vent.</p></div>	https://treatment.plazi.org/id/144D8797FFA07532FF0BF91FFB4EE23C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Sampaio, Filipa L.;Narayanan, Surya;Cyriac, Vivek Philip;Venu, Govindappa;Gower, David J.	Sampaio, Filipa L., Narayanan, Surya, Cyriac, Vivek Philip, Venu, Govindappa, Gower, David J. (2020): A new Indian species of Rhinophis Hemprich, 1820 closely related to R. sanguineus Beddome, 1863 (Serpentes: Uropeltidae). Zootaxa 4881 (1): 1-24, DOI: 10.11646/zootaxa.4881.1.1
