taxonID	type	description	language	source
102387C1FFE4426C66F0FB89A866F9BA.taxon	description	(Figs 1 – 3, 27, 35)	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFE4426C66F0FB89A866F9BA.taxon	materials_examined	Type material. Holotype (Figs 1, 27): ♂, “ Mozambique, 9 m, Maputo Special Reserve, Mangrove Camp (Man- grove – Woodland Mosaic), 26 ° 19 ’ 35.9 ’’ S, 32 ° 42 ’ 35.7 ’’ E, 7 – 9. XII. 2016. MV, Aristophanous, M., Cristóvão, J., László, G., Miles, W. leg., ANHRT: 2017.22 ”, ANHRT unique number: ANHRTUK 00001646, slide No.: AV 3365 ♂ (Coll. ANHRT). Paratypes: 2 ♀, Mozambique, 22 m, Maputo Special Reserve, West Gate, (Sand Thicket), 26 ° 30 ’ 14.2 ’’ S, 32 ° 42 ’ 59.6 ’’ E, 21 – 30. XI. 2016. Light Trap. Aristophanous, M., Cristóvão, J., László, G., Miles, W., leg. ANHRT: 2017.22, ANHRT unique numbers: ANHRTUK 00010945, 00048252, slide Nos: AV 4399 ♀, AV 4930 ♀ (Coll. ANHRT).	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFE4426C66F0FB89A866F9BA.taxon	diagnosis	Diagnosis. Tumicla elephantina (Figs 1 – 3) is the closest relative of T. mbeghai (Figs 9 – 11), from which it differs externally by its considerably smaller size (particularly in females), slightly broader forewing, brick-red body and wing coloration (that is mustard yellow in T. mbeghai), more diffuse pattern and the presence of blackish dashes at hindwing apex (those are absent in T. mbeghai). Tumicla elephantina is most similar externally to the sympatric T. tsonga Volynkin & László, 2018 (Figs 4 – 6, 30, 38) but can be distinguished by its smoothly curved antemedial line (that is angled in T. tsonga), longer longitudinal dashes in the postmedial area, and the presence of blackish dashes at hindwing apex (those are absent in T. tsonga). The new species also resembles externally T. rosea (Durante, 2008) (Figs 7, 39) due to its subapical dashes of hindwing, but differs by its shorter and broader forewing with more rounded apex, brick-red forewing ground colour (that is bright orange with reddish suffusion in T. rosea), more diffuse forewing pattern, smoothly curved antemedial line (that is angled in T. rosea) and longer postmedial longitudinal dashes. In addition, T. elephantina remotely resembles externally T. rubea (Figs 16 – 18, 32, 40), from which, however, it differs clearly by its larger size, broader forewing, longer longitudinal dashes in the postmedial area, and the presence of blackish dashes at hindwing apex. The male genital capsule of T. elephantina (Fig. 27) is similar to that of T. mbeghai (Figs 28, 29), but differs from it by its longer, more robust vinculum and shorter, medially slightly more dilated valva. The aedeagus of T. elephantina (Fig. 27) is slightly shorter and less curved than that of T. mbeghai (Figs 28, 29). The vesica of T. elephantina (Fig. 27) is thicker than that of T. mbeghai (Figs 28), bearing two robust, pointed cornuti, whereas that of T. mbeghai is armed with one cornutus only. The ground plan of the female genitalia of the two species is similar (Figs 35, 36), but T. elephantina has longer, thicker papillae anales, shorter ductus bursae and narrower ostium bursae compared to those of T. mbeghai; the lateral protrusion at the right side of the posterior section of corpus bursae is much longer, somewhat narrower and directed anterio-laterally in the new species, while that is shorter, broader and directed anteriorly in T. mbeghai; the lateral protrusion at the left side of the posterior section of corpus bursae is shorter in T. elephantina than in the related species; the globular anterior section of corpus bursae is weakly scobinated posteriorly only in the new species (that is evenly scobinated in T. mbeghai); and the appendix bursae of T. elephantina is broadly conical, apically rounded, evenly sclerotised and lacks a long area of lateral sclerotisation, whereas in T. mbeghai that is trigonal with pointed apex, heavily sclerotised laterally and membranous with granulation distally.	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFE4426C66F0FB89A866F9BA.taxon	description	Description. External morphology of adults (Figs 1 – 3). Forewing length 11.5 mm in holotype male and 14 – 14.5 mm in females. Male antennae ciliate, female antennae filiform. Intraspecific variability expressed by the different shades of forewing ground colour. Head brick-red with blackish dot on frons. Thorax brick-red, with four black spots medio-laterally. Forewing moderately broad, more elongated in female, with rounded apex and oblique tornus; forewing ground colour brick-red in males and varies from pale brick-red to orange in females; wing pattern blackish, consists of two large subbasal spots, zigzagged ante-, postmedial and medial lines connecting each other, semilunar discal spot, and interrupted subterminal line consisting of long dashes of various length connected to postmedial line; cilia blackish. Hindwing dark brick-red in males and pale brick-red to reddish orange in females, with diffuse short blackish dashes on veins apically and medially; cilia orange-red with admixture of blackish scales apically and medially, and pure orange-red at tornus and along anal margin. Abdomen brick-red in male and pale orange in females. Male genitalia (Fig. 27). Uncus moderately long, narrow, slightly broadened and hook-like curved distally, apically pointed; tuba analis broad, scaphium thin, weakly sclerotised, subscaphium represented by a field of fine spinulose scobination; tegumen moderately long and broad; juxta broad, shield-like; vinculum robust, U-like, apically rounded; valva elongated, medially broadened, its distal half slightly curved ventrad, apically strongly tapered, with pointed tip; costal margin convex, with short irregular dentation at apex; sacculus short, narrow, weakly setose, without processes, with convex ventral margin. Aedeagus short, straight, relatively narrow; carina elongated ventrally, heavily sclerotised. Vesica broad, directed ventrally, membranous, with one small subapical diverticulum and two long and robust, spine-like subbasal cornuti of different length situated on base connected with carina by narrow sclerotised band. Female genitalia (Fig. 35). Ovipositor short and broad; papillae anales rectangular, with rounded angles; apophyses moderately long and thin; apophyses posteriores slightly longer than apophyses anteriores. Ostium bursae broad with slightly arched margin; ductus bursae broad, dorso-ventrally flattened, slightly rugose medially, heavily sclerotised, its anterior margin oblique at connection with corpus bursae. Posterior section of corpus bursae elongated, very broad, dorso-ventrally flattened, moderately sclerotised, weakly rugose, with narrowly conical, long and heavily sclerotised right protrusion latero-anteriorly, and broad but short semi-globular left protrusion laterally. Anterior section of corpus bursae globular, membranous anteriorly and weakly setose posteriorly. Appendix bursae short, broadly conical with rounded apex, heavily sclerotised, situated postero-laterally at left side.	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFE4426C66F0FB89A866F9BA.taxon	distribution	Distribution. Tumicla elephantina is to date, only known from the Maputo Special Reserve in southern Mozambique.	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFE4426C66F0FB89A866F9BA.taxon	etymology	Etymology. The specific epithet refers to the type locality, Maputo Special Reserve which was originally established for conservation of elephants, boasting a thriving population of the threatened African large mammal.	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFE6426966F0F976ACC5F882.taxon	description	(Figs 9 – 11, 28, 29, 36)	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFE6426966F0F976ACC5F882.taxon	materials_examined	Type material. Holotype (Figs 11, 36): ♀, “ Amani P 269 4 / 61 ” / “ Tanzania: Amani, Malaria Institute, G. Pringle coll. B. M. 1966 - 281 ” / “ G. Pringle coll. B. M. 1966 - 281 ” / “ NHMUK 010915998 ”, slide No.: NHMUK 010314594 Volynkin (Coll. NHMUK). Paratypes: 1 ♂, 3 ♀, with same data as the holotype, slide No.: NHMUK 010314593 (♂) Volynkin (Coll. NHMUK); 1 ♂, [Tanzania] D. O. Afrika, Amani, II. 06, Chr. Schröder S. V. / Am. 706 / T 279 / Paratypus ♂ Asura doa by: L. Kühne / L. Kühne Dauerpräparat / e 386 / 2004 (Coll. ZMB).	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFE6426966F0F976ACC5F882.taxon	diagnosis	Diagnosis. Tumicla mbeghai is the closest relative of T. elephantina. The differences between the two species are discussed above in the diagnosis of T. elephantina. Female of T. mbeghai (Fig. 11) differs from that of T. doa (Fig. 8) by its larger size, broader forewing with more elongated apex, paler forewing ground colour, finer elements of wing pattern, and hindwing cilia being blackish only at apex (whereas in T. doa the cilia is blackish along the entire outer margin). Externally, T. mbeghai (Figs 9 – 11) is most similar to T. similis (Durante, 2008) (Figs 12 – 15), from which it differs by its larger size, longer postmedial longitudinal dashes and the absence of blackish dashes at hindwing apex. In the male genitalia, T. mbeghai (Figs 28, 29) differs clearly from T. similis (Fig. 31) by its narrower uncus, narrower tegumen, wider, U-shaped vinculum (that is V-shaped in T. similis), more elongated valva apex, much longer aedeagus with a sclerotised carina, and broader vesica with a robust cornutus (whereas the vesica of T. similis is narrow, lacking any sclerotised elements). The female genitalia of T. mbeghai (Fig. 36) differ from those of T. doa (Fig. 37) by their broader ostium bursae, broader and more heavily sclerotised ductus bursae having an anterio-lateral sclerotised protrusion to the appendix bursae, and the corpus bursae structure. In T. mbeghai the corpus bursae is subdivided into two sections, the anterior one is globular and membranous with weak scobination, while the posterior one is dilated, dorso-ventrally flattened having conspicuous lateral protrusions, whereas in T. doa the corpus bursae is more or less pear-like with its anterior broadened section heavily sclerotised. In addition, the appendix bursae of T. mbeghai is small, conical, heavily sclerotised, situated postero-laterally at left side, whereas in T. doa that is broadly conical, more weakly sclerotised, rugose and situated laterally at right side. In comparison with those of T. similis (Fig. 41), the female genitalia of T. mbeghai (Fig. 36) have longer and much broader ductus bursae, much broader and sclerotised posterior section of corpus bursae having sclerotised lateral protrusions, and larger and sclerotised appendix bursae.	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFE6426966F0F976ACC5F882.taxon	description	Description. External morphology of adults (Figs 9 – 11). Forewing length 12 – 12.5 mm in males and 16 – 16.5 mm in females. Male antennae ciliate, female antennae filiform. Head pale mustard yellow with blackish dot on frons. Thorax pale mustard yellow, with four black spots medio-laterally and one dot medio-posteriorly. Forewing moderately broad, broader in female, with rounded apex and oblique tornus; forewing ground colour pale mustard yellow; wing pattern blackish, consists of one large subbasal spot, one short longitudinal subbasal dash, zigzagged ante-, postmedial and medial lines connecting each other, semilunar discal spot, and interrupted subterminal line consisting of long dashes of various length connected to postmedial line; cilia blackish. Hindwing pale mustard yellow, lighter than forewing; cilia as wing ground colour, sometimes with admixture of blackish scales at apex. Abdomen pale ochreous. Male genitalia (Figs 28, 29). Uncus moderately long, narrow, distally hook-like curved and slightly broadened, apically pointed; tuba analis broad, scaphium thin, weakly sclerotised, subscaphium represented by a field of fine spinulose scobination; tegumen moderately long and broad; juxta broad, shield-like; vinculum robust, U-shaped; valva elongated, medially broadened, its distal half slightly curved ventrally, apically strongly tapered, with pointed tip; costal margin convex, with short irregular dentation on its distal third; sacculus short, narrow, weakly setose, without processes, with slightly convex ventral margin. Aedeagus short, straight; carina elongated ventrally, heavily sclerotised. Vesica broad, directed ventrad, membranous, with one small subapical diverticulum and one long and robust, spine-like subbasal cornutus situated on base connected to carina by a narrow sclerotised band. Female genitalia (Fig. 36). Ovipositor short and broad; papillae anales rectangular, with rounded angles; apophyses moderately long and thin; apophyses posteriores slightly longer and narrower than apophyses anteriores. Ostium bursae broad with slightly arched margin; ductus bursae broad, dorso-ventrally flattened, slightly rugose medially, heavily sclerotised, it’s anterior left corner heavily sclerotised and protruding to posterior margin of appendix bursae. Posterior section of corpus bursae elongated, very broad, dorso-ventrally flattened, moderately sclerotised, weakly rugose, with trigonal and heavily sclerotised right protrusion latero-anteriorly, and broad but short semi-globular left protrusion laterally. Anterior section of corpus bursae globular, weakly setose. Appendix bursae short, broadly trigonal, heavily sclerotised, situated postero-laterally at left side.	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFE6426966F0F976ACC5F882.taxon	distribution	Distribution. Tumicla mbeghai is only known to date from its type locality in the Usambara Mountains (northeastern Tanzania).	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFE6426966F0F976ACC5F882.taxon	etymology	Etymology. Mbegha is a mythic hero of the Shambaa people inhabiting the Usambara Mountains, considered founder of the Kilindi dynasty.	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFEF426566F0FF52ADFAFC98.taxon	description	(Figs 8, 37)	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFEF426566F0FF52ADFAFC98.taxon	materials_examined	Type material examined. Holotype (Figs 8, 37): ♀, “ D. O. Afrika, Kwedikundi, Nguru Gebg., Reuss S. G. ” / “ 1363 ” / “ T 281 ” / “ Holotype ♀ Asura doa by: L. Kühne ” / “ L. Kühne Dauerpräparat / e 388 / 2004 ” (Coll. ZMB).	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFEF426566F0FF52ADFAFC98.taxon	discussion	Remarks. 1. The type locality handwritten on the holotype label was erroneously cited by Kühne (2007) as “ Nguou Sebg. ” instead of “ Nguru Gebg. ” (Nguru Mts.). 2. The species is currently known by the female holotype only. The male paratype from Amani (the Usambara Mountains in northeastern Tanzania) is not conspecific with the holotype and belongs to T. mbeghai. The second paratype, a female from South Africa (“ Natal ”) illustrated by Kühne (2007, fig. 136) and deposited in the private collection of Lars Kühne has not been examined by us. However, according to its external appearance and collecting locality, this specimen is certainly not conspecific with the holotype of T. doa and supposedly belongs to the narrowly patterned form of T. sagenaria (Wallengren, 1860) distributed in South Africa.	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFEF426566F0FF52ADFAFC98.taxon	diagnosis	Diagnosis. Forewing length is 12.5 mm in the holotype female. The differences between T. doa and T. mbeghai are explained above in the diagnosis of the latter.	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFEF426566F0FF52ADFAFC98.taxon	distribution	Distribution. The species is reliably known from its type locality, the Nguru Mountains in eastern Tanzania only.	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFEF426366F0FC6FAF37FBFB.taxon	description	(Figs 19, 20, 33, 42)	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFEF426366F0FC6FAF37FBFB.taxon	materials_examined	Type material. Holotype (Figs 19, 33): ♂, [Democratic Republic of the Congo, Haut-Katanga Province] “ 20. 22. Luvua River (east bank) 85 miles N. of L. Mweru, ca. 3000 ft., April ’ 22. End of wet season. T. A. Barns ” / “ 006195 ” / “ Joicey Bequest. Brit. Mus. 1934 - 120. ” / “ NHMUK 010916003 ”, slide NHMUK 010314599 Volynkin (Coll. NHMUK). Paratype: ♀, with the same collecting data as the holotype / 005517 / Joicey Bequest. Brit. Mus. 1934 - 120. / NHMUK 010916004, slide NHMUK 010314600 Volynkin (Coll. NHMUK).	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFEF426366F0FC6FAF37FBFB.taxon	diagnosis	Diagnosis. Tumicla admiranda (Figs 19, 20) is an unmistakable species characterized by the presence of two broad dark brown shades on forewings. It resembles somewhat T. fasciata (Durante, 2008) (Figs 21, 22), but differs clearly by its smaller size, narrower forewing, the presence of a wide, dark brown subbasal band (this area is characterized by subbasal spots and distinct antemedial line in T. fasciata), the absence of a medial line (that is present in T. fasciata), its broader outer dark brown area including the discal spot (whereas the discal spot is well separated from the dark outer area in T. fasciata), and wider outer dark brown area of hindwing. The male genitalia of T. admiranda (Fig. 33) are somewhat similar to those of the externally rather different T. sagenaria (Figs 25, 26, 34), but differ by the broader uncus with more curved distal part, the broader apical part of tegumen, the shorter, U-shaped vinculum (that is longer and V-shaped with a rounded apex in T. sagenaria), the basally narrower valva with more strongly dentate distal costal margin, the much shorter and narrower aedeagus, the shorter lateral diverticulum of vesical evenly covered with spine-like cornuti (T. sagenaria has cornuti only in its distal two thirds), and the absence of a small distal diverticulum with spine-like cornuti which is present in T. sagenaria. The male genitalia of T. admiranda also resemble those of T. rubea (Fig. 32) due to the presence of numerous spine-like cornuti in vesica, but differ by the more robust uncus, the broader tegumen, the smaller vinculum, the valva being broadened subapically (that is narrowed subapically in T. rubea), and the diverticulum of vesica is longer and bearing much smaller spine-like cornuti than that of T. rubea. The female genitalia of T. admiranda (Fig. 42) differ clearly from those of T. sagenaria (Fig. 45) by their considerably shorter apophyses anteriores, much shorter ductus bursae, less strongly sclerotised posterior end of corpus bursae, narrower anterior section of corpus bursae bearing a signum (that is absent in T. sagenaria), and larger and less heavily sclerotised appendix bursae. The female genitalia of T. admiranda (Fig. 42) are also similar to those of T. fasciata (Fig. 43) and T. smithi (Fig. 44), but can be easily distinguished by their much shorter ductus bursae, weakly sclerotised posterior section of corpus bursae, narrower anterior section of corpus bursae being membranous and bearing a signum (whereas that is moderately sclerotised, rugose without a signum in T. fasciata and T. smithi), and smaller and less strongly sclerotised appendix bursae.	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFEF426366F0FC6FAF37FBFB.taxon	description	Description. External morphology of adults (Figs 19, 20). Forewing length 10 mm in holotype male and 13 mm in female. Head orange, male antenna ciliate, female antenna filiform. Thorax orange, with medial dark brown spot in male. Forewing elongated, with rounded apex and gently arcuate tornus, ground colour orange at base, in medial area and at apex; antemedial area blackish brown, with rounded outer margin; postmedial area dark brown with slightly S-curved inner margin and W-like curved outer margin at apex; discal spot narrow, semilunar, blackish, indistinct; cilia orange with admixture of dark brown scales. Basal half of hindwing orange, outer area dark brown, with wavy inner margin; cilia dark brown. Abdomen orange, in male with dark brown medial area. Male genitalia (Fig. 33). Uncus moderately long, narrow, distally hook-like curved, apically pointed; tuba analis broad, scaphium thin, weakly sclerotised, subscaphium represented by a field of fine spinulose scobination; tegumen moderately long, its apical two thirds strongly broadened and dorso-ventrally curved; juxta broad, shield-like; vinculum short, U-shaped, with narrow margins. Valva elongated, distally broadened, its distal end slightly curved ventrally, apically strongly tapered, with narrow, elongated and apically truncate tip; costal margin slightly arcuate, with short irregular dentation on its distal third; sacculus short, narrow, weakly setose, without processes, with slightly arcuate ventral margin. Aedeagus short, straight, its proximal end broadened. Vesica slightly broadened subbasally, with elongated diverticulum curved ventrally, covered densely with spine-like cornuti; vesica ejaculatorius originates from subbasal section of vesica, directed dorsally. Female genitalia (Fig. 42). Ovipositor short and broad; papillae anales rectangular, with rounded angles; apophyses moderately long and thin, equal in length. Ostium bursae broad with slightly arched margin; ductus bursae broad and very short, funnel-shaped, dorso-ventrally flattened and heavily sclerotised. Posterior section of corpus bursae tapered and elongated, sclerotised medially; anterior section of corpus bursae broadened, drop-like with weak scobination posteriorly and trigonal dentate signum latero-anteriorly. Appendix bursae short, conical with rounded apex, weakly sclerotised and granulated, situated postero-laterally on the left side of ductus bursae.	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFEF426366F0FC6FAF37FBFB.taxon	distribution	Distribution. Tumicla admiranda is to date, only known from its type locality in the Haut-Katanga province, southeastern Democratic Republic of the Congo.	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFEF426366F0FC6FAF37FBFB.taxon	etymology	Etymology. ‘ Admirandus’ means ‘ astonishing’ in Latin. The specific epithet refers to its unusual and conspicuous wing pattern and coloration.	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFE9426266F0FB40A946FD66.taxon	description	(Figs 23, 24, 44)	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFE9426266F0FB40A946FD66.taxon	materials_examined	Type material. Holotype (Figs 23, 44): ♀, “ Zambia, 1456 m, Changwena Falls, N. Swaka F. R., Central Prov., 13 ° 22 ’ 48 ’’ S, 29 ° 33 ’ 18 ’’ E, 27 – 30. I. 2019, MV Light Trap, Dérozier, V., Mulvaney, L., Takano, H. Leg. ANHRT: 2019.4 ”, ANHRT unique number: ANHRTUK 00073348, slide No.: AV 5243 ♀ (Coll. ANHRT). Paratypes: 3 ♀, with the same data as the holotype, ANHRT unique numbers: ANHRTUK 00072417, 00072418, 00072419 (Coll. ANHRT).	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFE9426266F0FB40A946FD66.taxon	discussion	Remark. Tumicla smithi is to date, only known by females. However, despite its unusual wing pattern, T. smithi certainly belongs to the genus Tumicla due to its legs tibial spur formula 0 - 2 - 2 characteristic for all Tumicla species (Durante 2008), forewing shape, and female genitalia being similar to those of some other species of Tumicla, in particular to T. fasciata and T. admiranda.	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFE9426266F0FB40A946FD66.taxon	diagnosis	Diagnosis. Tumicla smithi (Figs 23, 24) is also an unmistakable species differing clearly from other congeners by its orange forewing lacking transverse lines and having a broad unicolorous black distal band. The female genitalia of T. smithi (Fig. 44) are most similar to those of T. fasciata (Fig. 43), but differ from them by their shorter and less curved ductus bursae without a lateral protrusion (that of T. fasciata bears a well-developed, rounded lateral bulge), more heavily sclerotised posterior section of corpus bursae lacking a lateral protrusion (that is less strongly sclerotised, rugose having a long and broad lateral protrusion in T. fasciata), and shorter appendix bursae having sclerotised basal section and membranous distal section with granulation (that is larger and evenly sclerotised in T. fasciata).	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFE9426266F0FB40A946FD66.taxon	description	Description. External morphology of adults (Figs 23, 24). Forewing length 14.5 – 15 mm in females. Head black; antenna black, filiform in female. Thorax black; patagium orange; tegula orange with large black basal spot. Forewing moderately broad, with rounded apex, outer margin and tornus evenly arcuate. Forewing ground colour bright orange; pattern black, represented by a short basal dash on costa, short V-like subbasal streak, narrow semilu- nar discal spot which may be absent, and broad black terminal area with S-curved inner margin filling in postmedial, subterminal and terminal areas unicolorously; cilia black. Hindwing paler than forewing, orange, with intense black suffusion on subterminal and terminal areas, varying widely in width; cilia black, pale orange along anal margin. Abdomen orange with admixture of brown scales. Female genitalia (Fig. 44). Ovipositor short and broad; papillae anales rectangular, with rounded angles; apophyses moderately long and thin, equal in length. Ostium bursae relatively narrow; ductus bursae moderately long, dorso-ventrally flattened, heavily sclerotised, slightly broadened posteriorly, gently curved to the left, with convex right margin. Posterior section of corpus bursae conically tapered posteriorly, sclerotised; anterior section of corpus bursae globular, moderately sclerotised, rugose. Appendix bursae short, conical, sclerotised basally and membranous with granulation distally, situated postero-laterally on the left side of ductus bursae. Male unknown.	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFE9426266F0FB40A946FD66.taxon	distribution	Distribution. Tumicla smithi is to date, only known from its type locality collected near Changwena Falls in Central Zambia.	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
102387C1FFE9426266F0FB40A946FD66.taxon	etymology	Etymology. The species is dedicated to Mr Richard Smith, founder of the African Natural History Research Trust, organiser of extensive entomological exploratory program in Sub-Saharan Africa.	en	Volynkin, Anton V., László, Gyula M. (2019): Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini). Zootaxa 4668 (3): 421-434, DOI: 10.11646/zootaxa.4668.3.7
