identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
096BB133FF927E7C0061FDE97B5E9233.text	096BB133FF927E7C0061FDE97B5E9233.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Schellencandona Meisch 1996	<div><p>Genus Schellencandona Meisch, 1996</p><p>Diagnosis of the genus Schellencandona Meisch, 1996</p><p>Meisch (1996) gave the following characteristics to the genus Schellencandona: (1) Carapace small (0.4 to 0.6 mm) with six muscle scars of almost equal size and simple hinge. (2) Surface of the carapace smooth or with shallow pits. (3) Eye absent. (4) A2 with a penultimate segment subdivided in males with male bristles. (5) Mdp with 2+3 setae and γ setae smooth. (6) L5 exopodite with 2 plates. (7) L7 protopodite with 2 setae. (8) EII of L7 fused with EIII bearing a g seta. (9) EIV of L7 with two long (h2-h3) and one short (h1) setae. (10) Zenker’s organ with 4+2 rings of spines. (11) Hemipenis with a very flat M-process.</p><p>List of the species: S. schellenbergi (Klie, 1934), S. triquetra (Klie, 1936), S. belgica (Klie, 1937), S. insueta (Klie, 1938), S. mira (Sywula, 1976), S. simililampadis (Danielopol 1978), S. yakushimaensis Smith &amp; Kamiya, 2006, S. tea Karanovic &amp; Lee, 2012, S. dui Ma &amp; Yu, 2018 and S. rhodanensis Issartel &amp; Marmonier (2025) .</p></div>	https://treatment.plazi.org/id/096BB133FF927E7C0061FDE97B5E9233	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Issartel, Colin;Marmonier, Pierre	Issartel, Colin, Marmonier, Pierre (2025): Description of five new species of Schellencandona Meisch, 1996 (Ostracoda: Candoninae) from the southern French Alps, a highly diversified area for groundwater ostracods. European Journal of Taxonomy 1022: 85-133, DOI: 10.5852/ejt.2025.1022.3083, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3083/13745
096BB133FF927E72006DFB2778B293DC.text	096BB133FF927E72006DFB2778B293DC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Schellencandona danielopoli Issartel & Marmonier 2025	<div><p>Schellencandona danielopoli sp. nov.</p><p>urn:lsid:zoobank.org:act: 260AA59F-20E5-4AD0-AF4F-4D7EA84B2B17</p><p>Figs 2–4, 18, 20; Tables 1, 3</p><p>Diagnosis</p><p>Small candonine (L = 0.6 mm) of the genus Schellencandona with a triangular carapace covered with pits and small fossae. Greatest H of LV located just mid-length (H/L = 0.55). For both valves, anterodorsal margin slightly concave. Anterior margin widely rounded and posterior margin more acute. The two valves are strongly asymmetrical: LV with a dorsal hump that overlaps the RV, RV with a dorsal margin straight and slightly inclined backwards. A1 with a reduced number of setae: absence of seta on EIII and only one anterior seta on EVI. A2 with a penultimate segment subdivided in males with male bristles. Only 3 t setae (in female) and 3 z setae, z2 transformed in a claw in male. Mdp with 2+3 setae and γ setae smooth. L5 protopodite with two a setae; exopodite with 2 plates and endopodite transformed in a pair of clasping hook-like organs highly asymmetrical, right one stocky and poorly arched, left one slender and curved. L6 without d and e setae, but with one f and one g setae. L7 protopodite with 2 setae, EII and EIII fused bearing a g seta, EIV with two long (h2–h3) and one medium-sized (h1) setae. Female genital lobe rounded with a flat central part and without dorsal expansion. Zenker’s organ with 6 rings of spines. Hemipenis with outer lobe a large, rounded, dorso-distally oriented, inner lobe b sub-rectangular with a rounded ventral side. Bursa copulatrix (e) rounded with a curved sclerotized distal strip and an internal conical structure. M-process very flat. Ocular structures not visible.</p><p>Etymology</p><p>The new species is named after Dan L. Danielopol for his very important contribution to the systematics of ostracods.</p><p>Type material</p><p>Holotype</p><p>FRANCE • ♂, dissected appendages mounted in glycerine, valves stored in ethanol; Alpes-de-HauteProvence district, Montmaur municipality; 44.5608° N, 5.8952° E; 879 m a.s.l.; Jun.–Jul. 2010; C. Capderrey leg.; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=5.8952&amp;materialsCitation.latitude=44.5608" title="Search Plazi for locations around (long 5.8952/lat 44.5608)">interstitial habitat of the main channel of the Buech River in a large braided sector</a>; MNHN-IU-2023-701.</p><p>Allotype</p><p>FRANCE • ♀; same data as for holotype; MNHN-IU-2023-702.</p><p>Paratypes</p><p>FRANCE • 1 ♂, dissected appendages and valves stored in ethanol; same data as for holotype; MNHNIU-2023-703 • 1 ♂; Drôme district, Verclause municipality; 44.3810° N, 5.4255° E; 534 m a.s.l.; Jun.– Jul. 2010; C. Capderrey leg.; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=5.4255&amp;materialsCitation.latitude=44.381" title="Search Plazi for locations around (long 5.4255/lat 44.381)">interstitial habitat of the main channel of the Eygues River in a braided sector</a>; UCBLZ 2012-3-153-2 • 1 ♀; same data as for holotype; MNHN-IU-2023-704 • 1 juv.; same data as for the holotype; UCBLZ-2012-3-153-1 • 1 juv.; same locality as for holotype; undissected; UCBLZ-2012-3-207 .</p><p>Other material examined</p><p>FRANCE • 23 specs of diverse stages; Lez River at Montjoux; UCBLZ-2012-3-207 • 1 juv.; Lez River at Grignan; UCBLZ-2012-3-207 • 10 specs of diverse stages; Eygues River at Verclause; UCBLZ-2012-3-207 • 2 juvs; Ouvèze River at Entrechaux; UCBLZ-2012-3-207 • 2 juvs; Grand Buech River at Luce la Croix Haute; UCBLZ-2012-3-207 • 60 specs of diverse stages; Petit Buech River at Montmaur; UCBLZ • 1 juv.; Buech River at Sisteron; UCBLZ-2012-3-207 • 5 specs of diverse stages; Les Duyes River at Mallemoisson; UCBLZ.2012.3.207 .</p><p>Description</p><p>MEASUREMENTS. Holotype, ♂ (MNHN-2023-701): LV: L = 600 µm, H = 330 µm (H/L = 0.55). RV = 588 µm, H = 290 µm (H/L = 0.49). W = 210 µm (W/L = 0.35). Range for males (n = 2): L = 575–600 µm, H = 570–588 µm, W = 210–225 µm. Allotype, ♀ (MNHN-2023-702): LV: L = 560 µm, H = 310 µm (H/L = 0.55). RV: L = 550 µm, H = 275 µm (H/L = 0.50). W = 205 µm (W/L = 0.36). Range for females (n = 2): L = 560–565 µm. H = 310–320 µm, W = 200205 µm.</p><p>CARAPACE. Whitish with ornamentation consisting of pits (or small fossae) in central part that vanish progressively toward periphery. General shape of carapace triangular (Figs 2, 18A–B). Two valves strongly asymmetrical. LV overlaps RV with hump-like dorsal margin very variable (Fig. 2A–B) and without marked cardinal angles. Highest H located at middle of L. H slightly superior to ½ L: H/L= 0.55 for both male and female. Carapace viewed dorsally (Fig. 2C, I) moderately compressed, with greatest W at middle of L. Anterior and posterior ends weakly beak-shaped. Posterior end more pointed in female.</p><p>VALVES. For both valves (Fig. 2), anterior margin widely rounded, while posterior margin more acute. Dorsal margin of LV hump-like and variable in males: H/L varying from 0.55 to 0.59 (Fig. 2A–B). Dorso-posterior margin straight or slightly concave in both sexes. Dorso-anterior margin slightly concave in both sexes. Ventral margin slightly convex in both males and females. RV is smaller than LV, trapezoidal with cardinal angles well marked, dorsal margin straight and slightly inclined backward, and ventral margin straight. Inner calcified lamella represents 11% of body length both anteriorly and posteriorly in males and slightly larger anteriorly than posteriorly in females. Fused marginal valve zone moderately large representing 2.5% of body length, with few and straight radial pore canals.</p><p>ANTENNULE, A1 (Figs 3A, 4A). I+II: A-1l(Pu), P-2l(Pu) / III:0 / IV: A-1s /V: A-1l, P-1s / VI: A-1l/ VII: A-2l-1s (α), P-1l /VIII: D-2l-ya-1l(cs). Posterior setae of I+II plumose. Using IV podomere as reference, ratios of podomeres in male 1-1-1.1-0.9-1.2-1.2 from III to VIII. ya aesthetasc long, 5–6.5 × as long as IV podomere.</p><p>ANTENNA, A2 (Figs 3B–D, 4B–D). Protopodite: coxa with 3 setae, 2 long and smooth, 1 short and plumose; basis with 1 long posterior seta; exopodite with 1 long and 2 short setae; EI with 1 posterior aesthetasc Y (equalling 79% of EI length) and distally 2 setae (1s and 1l).</p><p>MALE A2 (Fig. 3B–D). EII and EIII segmented as two individuated podomeres; EII with 1 short aesthetasc (y1) and 4 t setae, t1 long and plumose, t4 short, t2 and t3 transformed in male bristles with length equal to 63% of EI length. EIII with 1 short aesthetasc and 3 external z setae, z1 and z3 shorter than EIV, z2 transformed in claw (170% of EI length), G1 reduced (65% of EI length), G2 well-developed (170% of EI length), G3 reduced to bristle (47% of EI length). EIV with 2 claws, posteriorly 1 long (Gm, 160% of EI length), anteriorly 1 reduced (GM, 90% of EI length), 1 aesthetasc (y3, 50% of EI length) associated with slightly shorter seta, g seta present.</p><p>FEMALE A2 (Fig. 4B–D). EII and EIII fused with anteriorly 2 short aesthetascs (y1 and y2), 3 untransformed t setae, distally 3 z setae (z1 and z2 of medium size, z3 short), reduced G2 claw representing 60% of EI length, and 2 well-developed claws G1 and G3 (both representing 170% of EI length). EIV with anteriorly 1 long (GM, 150% of EI length) and posteriorly 1 reduced claw (Gm, 75% of EI length), 1 aesthetasc y3 (58% of EI length) associated with a subequal seta, g seta present.</p><p>MANDIBLE. Consisting of coxal plate and 4-segmented palp (Mdp). Coxa typically shaped, heavily chitinized with a masticatory part. 1 st podomere of Mdp (Fig. 3F) with externally exopodite plate and 2 long setae, internally with 2 long setae (1 plumose S1) and 2 short setae (1 smooth, α, 1 plumose, S2). 2 nd podomere with externally 2 setae and internally a group of 3 smooth setae and a second group of 2 setae (1 long and plumose, 1 short, β). 3 rd podomere with externally 3 setae, distally 1 long and smooth seta (γ) and internally 3 setae (2 short setae and 1 longer). 4 th podomere with 2 serrated claws (170% of 3 rd podomere length) and 3 small setae.</p><p>MAXILLULAR PALP (Mx1palp, Fig. 3E). Two-segmented. 1 st segment with 4 apical plumose setae on outer corner. 2 nd segment with 2 claw-like setae (4× as long as 2 nd segment) and 4 thinner setae.</p><p>MAXILLA (L5, Figs 3I–J, 4E). With protopodite bearing 2 anterior a setae and 2 exterior setae (b and d), masticatory process (endite) apically with group of 10 setae. Exopodite plate with 2 plumose filaments. Male endopodites transformed in clasping hook-like organs with relatively high asymmetry, right one stocky and poorly arched, left one slender and curved, with 2 short but thick setae on ventral side and thin apical seta. In female, similar set of setae was observed on protopodite and 2 filaments on exopodite. Endopodite with 3 short apical setae.</p><p>WALKING LEG (L6, Figs 3G, 4H). Five-segmented. Protopodite and EI without seta, EII with 1 f seta, EIII with 1 g seta, EIV with 2 short setae (h1 and h3) and long claw (h2) and equalling 160% of EI length.</p><p>CLEANING LEG (L7, Figs 3L, 4F). Four-segmented (with EII and EIII fused). Protopodite with 1 short (d1) and 1 long (dp, 120% of EI length) setae. EI without seta, EII+EIII with short seta (g), EIV with 1 medium-sized seta (h1, 78% of EI length) and 2 long setae (h2, h3, 130 and 140% of EI length, respectively).</p><p>CAUDAL RAMUS (CR, Figs 3M, 4G). Robust with a medium-sized to long sp seta (30% of anterior margin of CR) exceeding the basis of posterior claw, short sa seta and 2 long and curved claws (Ga and Gp). In males, these claws are rather short, representing respectively 72% and 66% of anterior margin of CR, but very long in females (i.e., 96% and 95% respectively), both claws serrated.</p><p>FEMALE GENITAL LOBE (Fig. 4G). rounded with flat central part and without dorsal expansion. Oocyte large (10.5% of valve length) and rounded.</p><p>MALE GENITAL ORGANS. Zenker’s organ (Fig. 3K) with 6 internal rings of spines representing 23% of total length of carapace. Hemipenis (Fig. 3H) with large rounded distal outer lobe (a) dorsally oriented, rectangular-shaped inner lobe (b) with straight distal end, rounded ventral angle and more pointed dorsal angle, with small plication on ventral side. Lobe h not observed. Labyrinth well-sclerotized and divided in 4 sections, section d4 weakly reticulated. Copulatory tube thin located inside rounded bursa copulatrix (e) with well-sclerotized distal strip and internal conical structure. M-process dorsally rounded, linked to C strip, and ventrally thin, joining d3 section of labyrinth.</p><p>OCULAR STRUCTURES. Not visible.</p><p>Ecology and distribution</p><p>Schellencandona danielopoli sp. nov. was collected from 8 sites of the braided river study (Capderrey et al 2013): two sites along the Lez River, one site along each of the Eygues, Ouvèze, Buech and Les Duyes rivers. The first three rivers are left-side tributaries of the Rhône River, flowing at intermediate altitudes (i.e., between 207 and 534 m a.s.l.). The last two rivers flow at higher altitudes (i.e., between 466 and 1018 m a.s.l.) in the Durance catchment, in areas close to the margin of the Würm ice sheet (Fig. 1).</p><p>In these 8 sites, the animals were mainly sampled in areas fed by groundwater upwellings (i.e., for 82% of the 102 individuals collected) with increasing abundances with depth into the river sediment (16% at - 30 cm, 21% at - 60 cm and 62% at - 90 cm). Schellencandona danielopoli sp. nov. was sampled in a wide range of temperature, from 11.1°C in the Petit Buech River at Montmaur to 19.9°C in the Buech River at Sisteron (Fig. 1, Table 1). It was collected at a depth of 90 cm into the bedsediment in interstitial water with a medium electrical conductivity (329–477 µS. cm-1), a pH ranging from 7.4 and 8.1 and a wide range of dissolved oxygen concentrations (3.2–7.1 mg ·L- 1).</p><p>Specialisation to groundwater: the length of ya of A1 and Y of A2 (i.e., more than 5 EIV podomere length of A1 and 79% of EI length of A2, respectively), the lack of eyes and the large size of the oocyte (10.5% of valve length) suggest that Schellencandona danielopoli sp. nov. is specialized for life in groundwater (Danielopol 1973, 1980; Issartel &amp; Marmonier 2025) as other species of Schellencandona, with a wide geographical distribution in the Southern French Alps, a wide altitudinal range and a marked ecological preference for deep river sediment layers.</p><p>Taxonomic remarks</p><p>The general shape of the carapace of Schellencandona danielopoli sp. nov. (triangular) is rather similar to the triangular S. triquetra and S. rhodanensis, but the carapace of the new species differs by its slightly concave dorso-anterior margin, while it is straight in the two other species, and its dorsal margin slightly inclined backwards, while it is parallel to the ventral margin in the two other species (Figs 2, 18A).</p><p>In addition to the carapace shape, the soft parts of Schellencandona danielopoli sp. nov. differ from those of the other European species of the genus (i.e., S. triquetra, S. belgica, S. insueta, S. mira, S. rhodanensis), but seems closely related to S. simililampadis and S. schellenbergi because of (1) the stocky shape of the hemipenis, with a large a lobe dorso-distally oriented, the h lobe not visible and the curved distal sclerotized strip on the bursa copulatrix, (2) the A2 with a z2 seta transformed in a claw in males (see Fig. 19A for A2 of S. simililampadis) and (3) the reduced number of setae on the A1, especially the lack of seta on the podomere III in both males and females (but a seta is present in S. schellenbergi). In contrast, Schellencandona danielopoli differs from S. simililampadis and S. schellenbergi in the following four characteristics: (1) triangular shape of the carapace (trapezoid in the two other species), (2) single anterior seta on podomere VI of A1 (two setae in the others), (3) two a setae on L5 (one in the others), and (4) lack of d and e setae on L6 (present in S. simililampadis).</p><p>The different populations of Schellencandona danielopoli sp. nov. do not show any significant inter-population variability in both the carapace shape (because of a high intra-population variability of the dorsal margin, see Fig. 2A–B) and in the soft-part morphology (similar chaetotaxy in males from the Buech River (holotype, ♂ (MNHN-IU-2023-701)), and from the Eygues River (paratype, ♂ (UCBLZ 2012-3-153-2)).</p><p>Finally, the similarity of Schellencandona danielopoli sp. nov. with the other species described here is detailed in the Discussion.</p></div>	https://treatment.plazi.org/id/096BB133FF927E72006DFB2778B293DC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Issartel, Colin;Marmonier, Pierre	Issartel, Colin, Marmonier, Pierre (2025): Description of five new species of Schellencandona Meisch, 1996 (Ostracoda: Candoninae) from the southern French Alps, a highly diversified area for groundwater ostracods. European Journal of Taxonomy 1022: 85-133, DOI: 10.5852/ejt.2025.1022.3083, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3083/13745
096BB133FF9C7E680060FA327BF59449.text	096BB133FF9C7E680060FA327BF59449.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Schellencandona capderreyae Issartel & Marmonier 2025	<div><p>Schellencandona capderreyae sp. nov.</p><p>urn:lsid:zoobank.org:act: C5B4806B-DBC6-4C1F-818B-7F7BC597D548</p><p>Figs 5–7, 18, 20; Tables 1, 3</p><p>Diagnosis</p><p>Small trapezoid candonine of the genus Schellencandona with a carapace length of about 600 µm covered by fossae. LV overlaps the RV. Greatest H of LV located at 40 and 63% of the animal length (H/L=0.50). Anterior and posterior calcified inner lamella sub-equal amounting to c. 10% of L. A1 without seta on the 3 rd podomere. Male A2: EII and EIII separated with t2 and t 3 male bristles, 3 z setae (z2 transformed in a claw), the longest claw (G2) represents 180% the length of EI. Female A2: EII+III with 3 t and 2 z setae. 2 nd podomere of the mandibular palp bears 3+2 setae. Endopodites of the maxilla (L5) developed in males into prehensile palps slightly curved and asymmetrical. Walking leg (L6) with two g setae. Cleaning leg (L7) 4-segmented, EII and EIII fused, with 2 setae (d1 and dp) on the protopodite. Zenker’s organ with 6 internal rings of spines. The outer lobe a of the hemipenis large and rounded, dorso-distally oriented, the inner lobe b sub-rectangular shaped with anterior and posterior angles rounded. Bursa copulatrix rounded with a well-sclerotized distal strip and an internal conical structure. Female genital lobe widely rounded without any posterior expansion. Ocular structures not visible.</p><p>Etymology</p><p>The new species is named after Cécile Capderrey who collected this species during her PhD.</p><p>Type material</p><p>Holotype</p><p>FRANCE • ♂, dissected appendages mounted in glycerine, valves stored in ethanol; Alpes de Haute Provence district, Noyer-sur-Jabron municipality; 44.1668° N, 5.9251° E; 468 m a.s.l.; Jun.–Jul. 2010; C. Capderrey leg.; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=5.9251&amp;materialsCitation.latitude=44.1668" title="Search Plazi for locations around (long 5.9251/lat 44.1668)">interstitial habitat of the Jabron River before its confluence with the Buech River</a>; MNHN-IU-2023-705.</p><p>Allotype</p><p>FRANCE • ♀; same data as for holotype; MNHN-IU-2023-706.</p><p>Paratypes</p><p>FRANCE • 1 ♂; same data as for holotype; MNHN-IU-2023-707 • 2 ♂♂; same locality as for holotype; undissected; UCBLZ 2012-3-208 • 2 juvs; same data as for holotype; UCBLZ 2012-3-153-3 to UCBLZ 2012-3-153-4 .</p><p>Other material examined</p><p>FRANCE • 8 juvs, undissected; UCBLZ.2012-3-208.</p><p>Description</p><p>MEASUREMENTS. Holotype, ♂ (MNHN-2023-705): LV: L = 600 µm, H = 300 µm (H/L = 0.50). RV = 585 µm, H = 285 µm (H/L = 0.48). W = 205 µm (W/L = 0.34). Range for males (n = 2): L = 595–600 µm, H = 300 µm, W=195–205 µm. Allotype, ♀ (MNHN-2023-706): LV: L=537 µm, H=260 µm (H/L=0.48). RV: L=530 µm, H=232 µm (H/L=0.43). W=180 µm (W/L=0.33).</p><p>CARAPACE. Whitish with ornamentation consisting of fossae on its entire surface. General shape of carapace trapezoid (Figs 5, 18C–D). LV overlaping RV with marked cardinal angles. Highest H located at 40 and 63% of animal length. H/L = 0.50 for male and H/L=0.48 for female. Carapace slightly compressed centrally in dorsal view (Fig. 5C, G), with greatest W at ⅓ and ⅔ L, representing 33% of L. Anterior and posterior ends weakly beak-shaped.</p><p>VALVES. For both valves (Fig. 5), anterior margin widely rounded, while posterior margin more pointed, dorso-posterior margin straight and dorso-anterior margin slightly concave. RV smaller than LV (3% of L difference). LV with ventral margin slightly concave in male and straight in female (Fig. 5D, H). RV with a dorsal margin slightly concave (Fig. 5B, F), while LV is straight (Fig. 5D) or poorly curved (Fig. 5H). Dorsal margin representing 33% of L in both male and female. Anterior and posterior calcified inner lamellas sub-equal, representing 10% of L. Fused marginal valve zone narrow, representing 2.5% of L for both male and female, with straight and dense radial pore canals, more numerous anteriorly.</p><p>ANTENNULE, A1 (Figs 6A, 7A). I+II: A-1l(pu), P-2l(pu) / III: 0/ IV: A-1s/ V: A-1l, P- 1m /VI: A-1l/VII: A-2l-1s(α), P-1l/ VIII: D-2l-ya-1l(cs). Using IV podomere as reference, the ratios of podomeres are 1.4- 1-1.3-1-1.6-1.6 from III to VIII. The ya aesthetasc very long, 7–11 × as long as IV podomere.</p><p>ANTENNA, A2 (Figs 6B–D, 7B–D). Protopodite: coxa with 3 setae, 2 long and smooth, 1 short and plumose; basis with 1 long posterior seta; exopodite with 1 long and 2 short setae; EI with 1 posterior aesthetasc Y (equalling to 60% of the length of EI) and distally 2 setae (1s and 1m).</p><p>MALE A2 (Fig. 6B–D). EII and EIII segmented as 2 individuated podomeres. EII with 1 short aesthetasc (y1) and 4 t setae, t1 medium, t4 short, t2 and t3 transformed in male bristles with length equal to 80% of EI length. EIII with 1 short aesthetasc (y2), 3 external z setae, z1 and z3 equal or slightly longer than EIV length, z2 transformed in long claw (160% of EI length). G1 reduced (56% of EI length), G2 well-developed (180% of EI length), G3 reduced to long bristle (84% of EI length). EIV with 2 claws, posteriorly 1 long (Gm, 170% of EI length) and anteriorly 1 reduced (GM, 52% of EI length), 1 aesthetasc (y3, 60% of EI length) associated with subequal seta, g seta present.</p><p>FEMALE A2 (Fig. 7B–D). EII and EIII fused, with anteriorly 2 short aesthetacs (y1 and y2), 3 t setae, distally 2 z setae (z1 twice the length of EIV and z2 short). G2 claw reduced (55% of EI length). G1 and G3 claws well-developed and sub-equal (170% EI length). EIV with anteriorly 1 long (GM, 140% of EI length) and posteriorly 1 reduced claw (Gm, 66% of EI length), 1 aesthetasc (y3, 45% of EI length) with subequal seta, g seta present.</p><p>MANDIBLE. Consists of coxal plate and 4-segmented palp (Mdp). Coxa typically shaped, heavily chitinized with masticatory part. 1 st podomere of Mdp (Figs 6F, 7F) with externally exopodite plate and 2 long setae, internally with 2 long setae (1 plumose S1) and 2 short setae (1 smooth, α, 1 plumose, S2). 2 nd podomere with externally 2 setae and internally group of 3 smooth setae and second group of 2 setae (1 long and 1 short, β). 3 rd podomere with externally 3 setae, distally 1 long and smooth seta (γ) and internally 3 small setae. 4 th podomere with 2 serrated and long claws (190% of 3 rd podomere length) and 3 small setae.</p><p>MAXILLULAR PALP (Mx1palp, Figs 6E, 7E). Two-segmented: 1 st segment with 4 apical plumose setae on outer corner. 2 nd segment with 2 claw-like setae (4 time as long as 2 nd segment) and 4 thinner setae. MAXILLA (L5, Figs 6I–J, 7H). With protopodite bearing 1 anterior seta (a) and 2 exterior setae (b and d), masticatory process (endite) apically with group of 11 setae. Exopodite plate with 2 filaments. Male endopodites transformed in asymmetrical clasping organs (right one more developed, left one slender), slightly curved with 2 short but thick setae on ventral side and thin apical seta. In female, similar set of setae observed on protopodite and 2 filaments on exopodite. Endopodite with 3 short apical setae.</p><p>WALKING LEG (L6, Figs 6G, 7G). Five-segmented. Protopodite and EI without seta. EII with f seta and EIII with 2 g setae. EIV with 2 short setae (h1 and h3) and long claw (h2) equalling 160% of EI length.</p><p>CLEANING LEG (L7, Figs 6M, 7J). Four-segmented (with EII and EIII fused). Protopodite with 1 short (d1) and 1 long setae (dp, 130% of EI length). EI without seta, EII+EIII with short seta (g). EIV with 3 long setae: h1 (sub-equal to EI length), h2 and h3 (130% and 200% of EI length, respectively).</p><p>CAUDAL RAMUS (CR, Figs 6L, 7I). Robust with medium-sized sp seta (22%–27% of anterior margin of CR, just reaching basis of Gp), short sa seta and 2 long and curved claws (Ga and Gp representing around 80–84% to 64–67% of anterior margin of CR, respectively), both claws serrated.</p><p>FEMALE GENITAL LOBE (Fig. 7I). Widely rounded without posterior expansion. Oocyte large (12% of valve length).</p><p>MALE GENITAL ORGANS. Zenker’s organ (Fig. 6K) with 6 internal rings of spines representing 20% of total length of carapace. Hemipenis (Fig. 6H) with distal outer lobe (a) large and rounded, dorso-distally oriented, inner lobe (b) sub-rectangular shaped with anterior and posterior angles rounded and small plication on ventral side. Lobe h not observed. Labyrinth well-sclerotized and divided in 4 sections, section d4 weakly reticulated. Copulatory tube thin located inside rounded bursa copulatrix (e) with well-sclerotized curved distal strip and internal conical structure. M-process flat with broad rounded dorso-distal part, linked to C strip, and thin basal part reaching d4 section of labyrinth.</p><p>OCULAR STRUCTURES. Not visible.</p><p>Ecology and distribution</p><p>Schellencandona capderreyae sp. nov. was collected from the interstitial habitat of a single river, the Jabron River, a tributary of the Durance River (Fig. 1), in a single sample at an elevation of 468 m a. s.l. It was sampled at the downstream end of a gravel bar (groundwater upwelling zone) at a depth of 90 cm into the riverbed sediment. Water had a temperature of 18.7°C, an electrical conductivity of 382 µS. cm-1, a pH of 7.9 and a rather high dissolved oxygen concentration (i.e., 6.3 mg ·L- 1).</p><p>Specialisation to groundwater: its very long aethetascs (ya on A1, Y and y3 on A2), the lack of visible eyes and the large size of the oocyte (12% of the body length) suggest a specialisation of the species to groundwater life. Schellencandona capderreyae sp. nov. can be considered, for the moment, as an endemic stygobiotic species.</p><p>Taxonomic remarks</p><p>The general shape of the carapace of the new species (trapezoid) is very similar to those of S. schellenbergi and S. simililampadis, but differs by the dorsal margin, which is parallel to the ventral margin in the new species (Figs 5, 18C), while it is inclined backwards in the two other species.</p><p>The new species differs from the other European species. It is very close to S. simililampadis and S. schellenbergi, but differs in the three following characteristics: (1) A2 with a z2 seta transformed in a claw in males (see Fig. 19A for A2 of S. simililampadis); (2) stocky shape of the hemipenis, with the dorso-distal location of the outer a lobe, the small size of the h lobe (not visible) and the curved distal sclerotized strip on the bursa copulatrix. In addition, S chellencandona capderreyae sp. nov. differs from the above-mentioned two species by (1) the A1 with a reduced number of setae, especially the lack of seta on podomere III in both males and females (seta present in S. schellenbergi), (2) the single anterior seta on podomere IV of A1 (two setae in the two other species), and (3) the lack of d and e setae on L6 (both present in S. simililampadis).</p><p>The relationships between S chellencandona capderreyae sp. nov. and the four other species described in this work are detailed in the Discussion.</p></div>	https://treatment.plazi.org/id/096BB133FF9C7E680060FA327BF59449	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Issartel, Colin;Marmonier, Pierre	Issartel, Colin, Marmonier, Pierre (2025): Description of five new species of Schellencandona Meisch, 1996 (Ostracoda: Candoninae) from the southern French Alps, a highly diversified area for groundwater ostracods. European Journal of Taxonomy 1022: 85-133, DOI: 10.5852/ejt.2025.1022.3083, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3083/13745
096BB133FF867E6F004CFDDE7B489197.text	096BB133FF867E6F004CFDDE7B489197.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Schellencandona mercantourensis Issartel & Marmonier 2025	<div><p>Schellencandona mercantourensis sp. nov.</p><p>urn:lsid:zoobank.org:act: EE6DF96A-2A5F-4345-9FA7-1D76CFB3FBD4</p><p>Figs 8–10, 18, 20; Tables 1, 3</p><p>Synonymy</p><p>Candoninae sp. 3 – Dole-Olivier et al. 2015: 537, table 2.</p><p>Diagnosis</p><p>Small trapezoid candonine of the genus Schellencandona (L = 545 µm). Carapace thin without ornamentation. Anterior margin widely rounded, while posterior margin more pointed, dorsal margin straight in LV and slightly concave in RV, dorso-posterior margin slightly convex. Greatest H of LV located in the anterior third (H/L=0.50) in male. Strong asymmetry between the two valves: LV overlaps the RV, RV 4% shorter in length than LV, resulting in a posterior gap between the two valves. Anterior and posterior calcified inner lamellas amounting to c. 14% and 9% of L, respectively. A1 without seta on the 3 rd podomere and 2 setae on the 6 th podomere. Male A2: EII and EIII separated with t2 and t3 transformed in male bristles, 3 z setae (z2 longer than z1 and z3), the longest claw (G2) represents 175% of EI length. Female A2: EII+III with 3 t and 2 z setae (z1 longer than z2). 2 nd podomere of the Mdp bears 3+2 setae. Endopodites of the maxilla (L5) developed in males into prehensile palps strongly asymmetrical, the right one hook-shaped with a ventro-distal angle marked by a sclerotized hump. Walking leg (L6) with d, e, f setae and one g seta. Cleaning leg (L7) 4-segmented, EII and EIII fused, with 2 setae (d1 and dp) on the protopodite. Zenker’s organ with 6 internal rings of spines. The outer lobe (a) of the hemipenis large and rectangular shaped, dorso-distally oriented. The inner lobe (b) is dorsally widely rounded. Lobe h short and ventrally rounded. Bursa copulatrix (e) rounded with a conical internal structure and a well-sclerotized dorsal strip. Female genital lobe anteriorly slightly rounded with a small posterior triangular expansion. Ocular structures not visible.</p><p>Etymology</p><p>The new species is named after the Mercantour National Park where the species was collected during a groundwater biodiversity survey.</p><p>Type material</p><p>Holotype</p><p>FRANCE • ♂, dissected appendages mounted in glycerine, valves stored in ethanol; Alpes de Haute Provence district, Colmars les Alpes municipality; 44.1789° N, 6.6206° E; 1225 m a.s.l.; Sep. 2010; M.- J. Dole-Olivier leg.; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=6.6206&amp;materialsCitation.latitude=44.1789" title="Search Plazi for locations around (long 6.6206/lat 44.1789)">interstitial habitat of the Verdon River</a>; MNHN-IU-2023-708.</p><p>Allotype</p><p>FRANCE • ♀; same data as for holotype; MNHN-IU-2023-709.</p><p>Paratype</p><p>FRANCE • 1 ♀, dissected appendages mounted in glycerine, valves stored in ethanol; Alpes Maritimes district, Saint Etienne de Tinée; 44.2344° N, 6.9492° E; 1080 m a.s.l.; Apr. 2009; M.-J. Dole-Olivier leg.; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=6.9492&amp;materialsCitation.latitude=44.2344" title="Search Plazi for locations around (long 6.9492/lat 44.2344)">interstitial habitat of the Tinée River</a>; MNHN-IU-2023-710 .</p><p>Other material examined</p><p>FRANCE • 3 juvs, undissected; collected in the Verdon River; UCBLZ.2012-3-217.</p><p>Description</p><p>MEASUREMENTS. Holotype, ♂ (MNHN-2023-708): LV: L = 545 µm, H = 275 µm (H/L = 0.50). RV: L = 520 µm, H = 245 µm (H/L = 0.47). W = 190 µm (W/L = 0.34). Allotype, ♀ (MNHN-2023-709): LV: L = 510 µm, H = 275 µm (H/L = 0.54). RV: L = 485 µm, H = 250 µm (H/L = 0.52). W = 185 µm (W/L = 0.36). Paratype, ♀ (MNHN-2023-710): LV: L = 520 µm, H = 260 µm (H/L = 0.50). RV: L = 495 µm, H = 235 µm (H/L = 0.47). W = 170 µm (W/L = 0.33).</p><p>CARAPACE. Whitish and thin, without ornamentation. Carapace trapezoid with marked cardinal angles (Figs 8, 18E–F). Highest H found at anterior third of animal (i.e., 42% of L). H/L = 0.50 for male and 0.50–0.54 for females. Carapace (Fig. 8C, G) moderately compressed in dorsal view, with greatest W at 50% of L, representing 0.33–0.36 of L. Anterior end weakly pointed. Posterior end moderately rounded.</p><p>VALVES. For both valves (Fig. 8), anterior margin widely rounded, posterior margin more pointed, dorso-posterior margin slightly convex. Both valves are strongly asymmetrical: LV overlaps RV, RV 4% shorter in length than LV, causing posterior gap between two valves (Fig. 8A, E). Dorsal margin straight in LV and slightly concave in RV, representing 28% of L. Ventral margin straight in LV and slightly concave in RV for both male and female (Fig. 8D, H). Anterior calcified inner lamella larger (14% and 11% of L for male and female respectively) than posterior one (9% of L for both sex). Fused marginal valve zone narrow, representing 2.5% of L in both male and female, with straight and dense radial pore canals, more numerous anteriorly.</p><p>ANTENNULE, A1 (Figs 9A, 10A). I+II: A-1l(pu), P-2l(pu) / III: 0 /IV: A-1s/ V: A-1l, P-1s /VI: A-2l/ VII: A-2l-1s(α), P-1l/VIII: D-2l-ya-1l(cs). Using IV podomere as reference, lengths of podomeres are in the ratios of 1.2-1-1.1-1-1-1.1 from III to VIII in male. ya aesthetasc very long, equal to 6× as long as IV podomere.</p><p>ANTENNA, A2 (Figs 9B–D, 10B–D). Protopodite: coxa with 3 setae, 2 long and smooth, 1 short and plumose; basis with 1 long posterior seta; exopodite with 1 long and 2 short setae; EI with 1 posterior aesthetasc Y (equalling 69% of EI length) and distally 2 setae (1s and 1m).</p><p>MALE A2 (Fig. 9B–D). EII and EIII segmented forming 2 distinct podomeres. EII with 1 short aesthetasc (y1) and 4 t setae, t1 medium, t4 short, t2 and t3 transformed in male bristles with length equal to 94% of EI length. EIII with 1 short aesthetasc (y2), 3 external z setae, z1 and z3 slightly longer than EIV length, z2 of medium size (40% of EI length). G1 reduced (72% of EI length), G2 well-developed (175% of EI length), G3 reduced to long bristle (45% of EI length). EIV with 2 claws, posteriorly 1 long (Gm, 150% of EI length) and anteriorly 1 reduced (GM, 55% of EI length), 1 aesthetasc (y3, 58% of EI length) associated with subequal seta, g seta present.</p><p>FEMALE A2 (Fig. 10B–D). EII and EIII fused, with anteriorly 2 short aesthetacs (y1 and y2), 3 t setae, distally 2 z setae (z1 150% of EIV length and z2 short). G2 claw reduced (45% of EI length). G1 and G3 claws well-developed and sub-equal (185 and 160% of EI length, respectively). EIV with anteriorly 1 long (GM, 147% of EI length) and posteriorly 1 reduced claw (Gm, 50% of EI length), 1 long aesthetasc (y3, 70% of EI length) with sub equal seta, g seta present.</p><p>MANDIBLE. Consisting of coxal plate and 4-segmented palp (Mdp). Coxa typically shaped, heavily chitinized with masticatory part. 1 st podomere of Mdp (Figs 9F, 10F) with externally exopodite plate and 2 long setae, internally with 2 long setae (1 smooth and 1 plumose S1) and 2 short setae (1 smooth, α, and 1 plumose, S2). 2 nd podomere with externally 2 setae and internally group of 3 smooth setae and second group of 2 setae (1 long and 1 short, β). 3 rd podomere with externally 3 setae, distally 1 long smooth seta (γ) and internally 3 small setae. 4 th podomere with 2 serrated and long claws (2.2 × as long as 3 rd podomere) and 3 small setae.</p><p>MAXILLULAR PALP (Mx1palp, Figs 9E, 10E). Two-segmented: 1 st segment with 4 apical plumose setae on outer corner. 2 nd segment with 2 claw-like setae (4.7 × as long as 2 nd segment) and 4 thinner setae.</p><p>MAXILLA (L5, Figs 9I–J, 10I). With protopodite bearing 1 anterior seta (a) and 2 exterior setae (b and d), masticatory process (endite) apically with group of 10 setae. Exopodite plate with 2 filaments. Male endopodites transformed in clasping organs strongly asymmetrical. Right one strongly sclerotized, distal end hook-shaped, ventro-distal angle marked by sclerotized hump. Left one stocky and slightly curved. Two endopodites bear 2 short but thick setae on ventral side and thin apical seta. In female, similar set of setae observed on protopodite and 2 filaments on exopodite. Endopodite with 3 short apical setae.</p><p>WALKING LEG (L6, Figs 9G, 10J). Five-segmented. Protopodite with one d seta, EI with one e seta (not observed in male holotype), EII with f seta and EIII with one g seta. EIV with 2 short setae (h1 and h3) and long claw (h2) serrated and equalling 145% the EI length.</p><p>CLEANING LEG (L7, Figs 9K, 10G). Four-segmented (with EII and EIII fused). Protopodite with 1 short (d1) and 1 long setae (dp, 125% of EI length). EI without seta, EII+EIII with short seta (g). EIV with 3 long setae: h1, h2 and h3 (80%, 110% and 175% of EI length, respectively).</p><p>CAUDAL RAMUS (CR, Figs 9M, 10H). Robust with medium-sized sp seta (22%–29% of anterior margin of CR, not reaching basis of Gp), short sa seta and 2 long and curved claws (Ga and Gp representing around 69–71% and 54–65% of the anterior margin of CR, respectively), both claws serrated.</p><p>FEMALE GENITAL LOBE (Fig. 10H). Anteriorly slightly rounded, with small posterior triangular expansion. Oocyte medium to large (9% of valve length).</p><p>MALE GENITAL ORGANS. Zenker’s organ (Fig. 9L) with 6 internal rings of spines representing 15% of total length of carapace. Hemipenis (Fig. 9H) with medium-sized distal outer lobe (a) and rectangular shaped, dorso-distally oriented. Inner lobe (b) dorsally widely rounded and small plication on ventral side. Lobe h short and ventrally rounded. Labyrinth well-sclerotized and divided in 4 sections, section d4 weakly reticulated. Copulatory tube thin located inside rounded bursa copulatrix (e) with marked dorsal sclerotized strip and conical internal structure. M-process flat with narrow rounded dorsal part, linked to C strip that joins base of e, and thin basal part reaching d4 section of labyrinth.</p><p>OCULAR STRUCTURES. Not visible.</p><p>Ecology and distribution</p><p>Schellencandona mercantourensis sp. nov. was sampled in the Mercantour National Park. It was collected in the interstitial habitat of two alpine rivers (i.e., the Verdon and Tinée rivers, Fig 1) at a depth of 50 cm into the riverbed sediment of two high-elevation sites (i.e., 1225 and 1080 m a.s.l., respectively). The species was not collected from the springs sampled in the same valleys and seems restricted to the interstitial habitat of the two rivers.</p><p>Specialisation to groundwater: the very long aesthetascs (ya on A1, Y and y3 on A2), the medium to large oocyte size (close to 9% of the body length) and the lack of a visible eye suggest a specialisation of the new species to groundwater. Schellencandona mercantourensis sp. nov. may be, for the moment, considered as a species specialized in riverbed sediment, with a narrow distribution area at high elevation in the Mercantour Mountains of the southern French Alps.</p><p>Taxonomic remarks</p><p>The general shape of Schellencandona mercantourensis sp. nov. is rather similar to the trapezoid S. schellenbergi and S. simililampadis, but the new species differs by its dorsal margin parallel to the ventral one (inclined backwards in the two other species), its convex dorso-posterior margin (straight in S. schellenbergi), and the asymmetry of the two valves (see below).</p><p>The new species differs from the other European and Asiatic species by the following four characteristics: (1) the strong asymmetry between the two valves (RV 4% shorter in length than LV inducing a posterior gap between the two valves), (2) by the shape of the L5 clasping organs (especially the strongly sclerotized hook-shaped right one) and (3) the shape of the hemipenis, rather thin, with a distally oriented a lobe and a very small but visible distal rounded h lobe. These three characteristics have never been observed in the genus Schellencandona before.</p><p>Despite these differences, Schellencandona mercantourensis sp. nov. may be related to S. simililampadis because of two light similarities: (1) a rather similar A1 chaetotaxy, without seta on podomere III and the two setae on podomere VI, and (2) the A2 with a z2 seta very long compared to z1, but not transformed in a claw like in S. simililampadis and S. schellenbergi .</p><p>The comparison between Schellencandona mercantourensis sp. nov. and the other species described here is detailed in the Discussion.</p></div>	https://treatment.plazi.org/id/096BB133FF867E6F004CFDDE7B489197	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Issartel, Colin;Marmonier, Pierre	Issartel, Colin, Marmonier, Pierre (2025): Description of five new species of Schellencandona Meisch, 1996 (Ostracoda: Candoninae) from the southern French Alps, a highly diversified area for groundwater ostracods. European Journal of Taxonomy 1022: 85-133, DOI: 10.5852/ejt.2025.1022.3083, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3083/13745
096BB133FF807E670019FE8478AF9071.text	096BB133FF807E670019FE8478AF9071.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Schellencandona claretae Issartel & Marmonier 2025	<div><p>Schellencandona claretae sp. nov.</p><p>urn:lsid:zoobank.org:act: 8EE9955F-D3A9-4AAF-A5E9-943602D30B46</p><p>Figs 11–14, 18, 20; Tables 1, 3</p><p>Diagnosis</p><p>Small trapezoid candonine of the genus Schellencandona (L = ~545 µm for the female and ~480 µm for the male). Carapace thin with poor ornamentation consisting of pits and a few fossae in the centre of the valves. Anterior margin widely rounded, posterior margin more pointed, dorsal margin slightly concave in both valves, dorso-posterior margin straight. Greatest H of LV located in the anterior third (H/L = 0.48). Strong asymmetry between the two valves: LV overlaps the RV, RV 5% shorter in length than LV inducing a posterior gap between the two valves. Calcified inner lamella amounting to ca 13% and 12% of L for the anterior and the posterior ones, respectively. A1 without seta on the 3 rd podomere. Male A2: EII and EIII separated with t2 and t3 transformed in male bristles, 2 z setae (z2 medium to long), the longest claw (G2) represents 180% of EI length. Female A2: EII+III with 3 t and 2 z setae. 2 nd podomere of the Mdp bears 3+2 setae. Endopodites of the maxilla (L5) developed in males into prehensile palps strongly asymmetrical, the right one hook-shaped with a ventro-distal angle marked by a sclerotized hump. Walking leg (L6) with e, f and 2 g setae. Cleaning leg (L7) 4-segmented, EII and EIII fused, with 2 setae (d1 and dp) on the protopodite. Zenker’s organ with 6 internal rings of spines. The outer lobe (a) of the hemipenis large and rectangular shaped, dorso-distally oriented. The inner lobe (b) widely rounded posteriorly and the lobe h with a sub-triangular distal expansion. Bursa copulatrix (e) conical with a ventral well-sclerotized strip. Female genital lobe rounded without posterior expansion. Ocular structures not visible.</p><p>Etymology</p><p>The new species is named after Cécile Claret who collected the species during an ecological study of the Asse River.</p><p>Type material</p><p>The only available male has an unusual shape and length of its carapace. A female is therefore chosen as the holotype.</p><p>Holotype</p><p>FRANCE • ♀, dissected appendages mounted in glycerine, valves stored in ethanol; Hautes-Alpes district, Oraison municipality; 43.8793° N, 5.9051° E; 335 m a.s.l.; Jun. 2007; C. Claret leg.; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=5.9051&amp;materialsCitation.latitude=43.8793" title="Search Plazi for locations around (long 5.9051/lat 43.8793)">interstitial habitat of the Asse River</a>; MNHN-IU-2023-711.</p><p>Allotype</p><p>FRANCE • ♂; same data as for holotype; MNHN-IU-2023-712.</p><p>Paratype</p><p>FRANCE • 1 ♀, dissected appendages mounted in glycerine, valves stored in ethanol; Alpes de Hautes Provence district, Colmars les Alpes municipality; 44.1789° N, 6.6206° E; 1225 m a.s.l.; Sep. 2010; M.-J. Dole-Olivier leg.; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=6.6206&amp;materialsCitation.latitude=44.1789" title="Search Plazi for locations around (long 6.6206/lat 44.1789)">interstitial habitat of the Verdon River</a>; MNHN-IU-2023-713 .</p><p>Other material examined</p><p>FRANCE • 5 juvs, undissected; collected in the Asse River; UCBLZ. 2012-3-168; UCBLZ • 1 juv., undissected; collected in the Verdon River; UCBLZ. 2012-3-217 .</p><p>Description</p><p>MEASUREMENTS. Holotype, ♀ (MNHN-2023-711): LV: L = 545 µm. H = 265 µm (H/L = 0.48). RV L = 515 µm, H = 250 µm (H/L = 0.48). W = 210 µm (W/L = 0.38). Paratype, ♀ (MNHN-2022-713): LV: L = 545 µm, H = 270 µm (H/L = 0.49). RV: L = 525 µm, H = 235 µm (H/L = 0.45). W = 180 µm (W/L = 0.33). Allotype, ♂ (MNHN-2023-712): LV: L = 480 µm, H = 230 µm (H/L = 0.48). RV L = 450 µm, H = 215 µm (H/L = 0.47). W = 185 µm (W/L = 0.38).</p><p>CARAPACE. Whitish and thin, with poor ornamentation of rare pits and small fossae in centre of valves. General shape of carapace trapezoid with marked cardinal angles (Figs 11A, E, G, 18G–H). Highest H located at third anterior part of animal (i.e., at 39% of L) with H/L = 0.48 for both male and female. Carapace viewed dorsally (Fig. 11D, F, H) moderately compressed, with greatest W at 50% of L in female, representing 38% of L. Carapace of male with greatest W at last third, representing 38% of L. Anterior end weakly beak-shaped in both sexes. Posterior end moderately rounded in female (Fig 11D) and more pointed in male (Fig. 11H). In female collected in Verdon River, similar general shape of carapace in lateral view (Fig. 11E; H /L = 0.49), but slenderer than holotype in dorsal view (Fig. 11F), with W representing 33% of L.</p><p>VALVES. Two valves strongly asymmetrical: LV overlaps RV, RV 5% shorter in length than LV inducing posterior gap between two valves (Fig. 11A, E, G). For both valves, anterior margin widely rounded, while posterior margin more pointed, dorso-posterior margin straight. Dorsal margin slightly concave, representing 34% of L. Ventral margin slightly convex in LV (Fig. 11C, J) and slightly concave in RV for both male and female (Fig. 11B, I). Anterior calcified inner lamella larger (13% and 14% of L for female and male respectively) than posterior one (12% of L for female and 10% for male). Fused marginal valve zone narrow, representing 2% of L for both male and female, with straight and dense radial pore canals, more numerous anteriorly. Strong difference between male and female, in size (12% difference, see below) and in shape of the carapace, especially in dorsal view. Greatest width at 50% of length in female and at last third in male. Posterior end gently rounded in female, pointed and triangular-shaped in male.</p><p>ANTENNULE, A1 (Figs 12A, 13A, 14A). I+II: A-1l(pu), P-2l(pu) / III: 0/ IV: A-1s/V: A-1l, P-1s /VI: A-2l / VII: A-2l-1s(α), P-1l/ VIII: D-2l-ya-1l(cs). Using IV podomere as reference, ratios of podomeres are 1.4-1-1.3-1.4-1.4-1.2 from III to VIII. ya aesthetasc very long, equalling 7.1× as long as IV podomere. Female of Verdon River with similar set of setae on A1 and similarly long ya (Fig. 14A). Using IV podomere as reference, ratios of podomeres are 1.6-1-1.3-1.2-1.5-1.6 from III to VIII.</p><p>ANTENNA, A2 (Figs 12B–D, 13B–D, 14B–D). Protopodite: coxa with 2 setae, only 1 long and smooth seta observed, and 1 short and plumose; basis with 1 long posterior seta; exopodite with 1 long and 2 short setae; EI with 1 posterior aesthetasc Y (equalling 68% of EI length) and distally 2 setae (1s and 1m).</p><p>MALE A2 (Fig. 12B–D). EII and EIII forming 2 distinct podomeres. EII with 1 short aesthetasc (y1) and 4 t setae, t1 and t4 short, t2 and t3 transformed in male bristles with length equal to 70% of EI length. EIII with 1 short aesthetasc (y2), 2 external z setae, z1 slightly shorter than EIV length, z2 of medium size (45% of EI length). G1 reduced (60% of EI length), G2 well-developed (180% of EI length), G3 reduced to short bristle (20% of EI length). EIV with 2 claws, 1 long posteriorly (Gm, 150% of EI length) and 1 reduced anteriorly (GM, 57% of EI length), 1 medium-sized aesthetasc (y3, 42% of EI length) associated with subequal seta, g seta not observed.</p><p>FEMALE A2 (Fig. 13B–D). EII and EIII fused, with anteriorly 2 short aesthetascs (y1 and y2), 3 t setae, distally 2 z setae (both short). G2 claw reduced (63% of EI length). G1 and G3 claws well-developed (205% and 180% of EI length, respectively). EIV with anteriorly 1 long (GM, 170% of EI length) and posteriorly 1 reduced claw (Gm, 80% of EI length), 1 long aesthetasc (y3, 83% of EI length) with subequal seta, g seta present. Female paratype of Verdon River (Figs 14B–D) with similar number of t and z setae on A2. Claws generally shorter than in the holotype: G2 reduced to 45% of EI length, G1 and G3 representing 190 and 175% of EI length. EIV with anteriorly 1 long claw (GM, 160% of EI length), posteriorly 1 reduced one (Gm, 50% of EI length) and y3 representing 57% of EI length, g seta present. MANDIBLE. Consisting of coxal plate and 4-segmented palp (Mdp). Coxa typically shaped, heavily chitinized with masticatory part. 1 st podomere of Mdp (Figs 13E, 14F) with externally exopodite plate, internally with 2 long setae (1 smooth and 1 plumose S1) and 2 short setae (1 smooth α, 1 plumose, S2). 2 nd podomere with externally 2 setae and internally group of 3 smooth setae and second group of 2 setae (1 long and 1 short, β). 3 rd podomere with externally 3 setae, distally 1 long smooth seta (γ) and internally 3 small setae. 4 th podomere with 2 serrated and long claws (166% of the 3 rd podomere length in holotype and 160% of 3 rd podomere in paratype from Verdon River) and 3 small setae.</p><p>MAXILLULAR PALP (Mx1palp, Figs 12E, 13F, 14E). Two-segmented: 1 st segment with 4 apical plumose setae on outer corner. 2 nd segment with 2 claw-like setae (370% of 2 nd segment length) and 4 thinner setae.</p><p>MAXILLA (L5, Figs 12F, 13G). With protopodite bearing 1 anterior seta (a) and 2 exterior setae (b and d), masticatory process (endite) apically with group of 11 setae. Exopodite plate with 2 filaments. Male endopodites (Fig. 12F–G) transformed in clasping organs markedly asymmetrical. Right one strongly sclerotized, distal end hook-shaped, ventro-distal angle marked by a sclerotized hump. Left one more rounded and slightly curved. Two endopodites bear 2 short but thick setae on ventral side and thin apical seta. In female (Fig 13G), similar set of setae observed on protopodite (a, b, d). Exopodite with 2 filaments and endopodite with 3 terminal setae.</p><p>WALKING LEG (L6, Figs 12I, 13I, 14G). Five-segmented. Protopodite with d seta (not observed in the male), EI with e seta, EII with f seta and EIII with 2 g setae (only one observed in male). EIV with 2 short setae (h1 and h3) and long claw (h2) equalling 270% EII length. In paratype from Verdon River, similar setal arrangement observed (d, e, f and 2 g) with claw h2 representing 300% EII length.</p><p>CLEANING LEG (L7, Figs 12J, 13K, 14H). Four-segmented (with EII and EIII fused). Protopodite with 1 short (d1) and 1 long seta (dp, 130–154% of EI length). EI without seta, EII+EIII with a short seta (g). EIV with 1 medium seta h1 (60–65% of EI length) and two long setae, h2 and h3 (130%–190% of EI length, respectively). In paratype from Verdon River, similar set of setae observed, with similar length for dp and h1, but slightly shorter h2 (120%) and h3 setae (160% of EI length).</p><p>CAUDAL RAMUS (CR, Figs 12L, 13H, 14I). Robust, with long sp seta (33% of anterior margin of CR) exceeding basis of posterior claw, short sa seta, 2 long and curved claws (Ga and Gp representing around 90% and 79% of anterior margin of CR, respectively), both claws serrated. In paratype from Verdon River, sp shorter (representing 26% of CR, not reaching basis of Gp), and Ga and Gp also shorter compared to holotype from Asse River (representing 79% and 73% of CR, respectively).</p><p>FEMALE GENITAL LOBE. Simple (Figs 13J, 14J) simply rounded for both the holotype (Fig. 13J) and paratype (Fig. 14J), without posterior expansion. Oocytes large (14% of valve length).</p><p>MALE GENITAL ORGANS. Zenker’s organ (Fig. 12K) with 6 internal rings of spines representing 15% of total length of carapace. Hemipenis (Fig. 12H) with distal outer lobe (a) large and subrectangular, dorso-distally oriented. Inner lobe (b) widely rounded dorsally and with small plication on ventral side. Lobe h short with sub-triangular distal expansion. Labyrinth well-sclerotized and divided in 4 sections, section d4 weakly reticulated. Thin copulatory tube located inside conical bursa copulatrix (e) with well-sclerotized ventral strip C. The M-process flat with narrow rounded dorsal part, linked to C strip joining base of e and thin basal part reaching d4 section of labyrinth.</p><p>OCULAR STRUCTURES. Not visible.</p><p>Ecology and distribution</p><p>Schellencandona claretae sp. nov. was sampled in the interstitial habitat of two rivers located in the Southern Alps, both tributaries of the Durance River (the Verdon River and the Asse River, Fig 1). In the Verdon River, the species occurs at high elevation (i.e., 1225 m a.s.l.), at a depth of 50 cm into the riverbed sediment, but it has never been sampled in the springs of the same valley (Dole-Olivier et al. 2015). In the Asse River, the species occurs close to the confluence with the Durance River (Fig. 1), at low elevation (335 m a.s.l.). In the Asse River, the species always occurred deeper than 20 cm in the riverbed sediment: at a depth of - 80 cm in the main channel and at depths of -50 and - 80 cm in the ‘backwaters’ and the ‘phreatic ponds’ (partially or totally isolated secondary channels fed by groundwater, Table 2), respectively.</p><p>Specialisation to groundwater: the very long aesthetascs (ya on A1, Y, and y3 on A2), the large oocyte size (close to 10% of the body length), and the lack of eyes suggest specialisation of the new species to groundwater. Schellencandona claretae sp. nov. may be considered as a species living deep in river sediment, more frequently in marginal pools than in the main active channel, with a wide elevation range. The species is known only from two tributaries of the Durance River.</p><p>Morphological remarks</p><p>Variability between male and female</p><p>We observed strong differences between the male and the two females. The only male available for this species (allotype (MNHN-IU-2023-712)) is a mature adult with complete development of the A2 (with male bristles), of the Zenker’s organ, the testes and the hemipenis, but with a surprisingly reduced size compared to the females of both the Asse and the Verdon rivers (12% shorter than the female). In addition, this male has a rather different shape in dorsal view, with a posterior end that is more pointed and triangular. It is not possible, for the moment, to be certain whether these differences between males and females are attributable to a sexual dimorphism characteristic of the species or are linked to a teratological development of the only available male (e.g., Rossetti &amp; Martens 1996).</p><p>Variability between populations We observed some differences between the female collected in the Asse River (type locality; holotype (MNHN-IU-2023-711)) and the female collected in the Verdon River (paratype (MNHN-IU-2023-713)). This latter appears slenderer in dorsal view (W/L = 0.33 vs 0.38) than the Asse River individual. It also has slightly smaller claws on A2 (i.e., G2 representing 45% vs 63%, Gm representing 50% vs 80% and ya representing 57% vs 83% of EI length, respectively) and shorter terminal setae on L7 (i.e., h2 representing 120% vs 130% and h3 representing 160% vs 190% of EI length, respectively) than the Asse River individual. Similarly, the caudal ramus of the female collected in the Verdon River is shorter and stockier than in the Asse River: Ga representing around 79% vs 90% of the anterior margin length, Gp 73% vs 79%, and Sp representing 26% vs 33%, respectively.</p><p>Taxonomic remarks</p><p>The general shape of Schellencandona claretae sp. nov. is rather similar to the trapezoid S. schellenbergi and S. simililampadis, but the new species differs by the strong asymmetry between the two valves (see below), its dorsal margin parallel to the ventral one (inclined backwards in the two other species), by its rounded anterior end in dorsal view (beak-shaped in S. schellenbergi), and its pointed posterior margin (rounded in S. simililampadis).</p><p>The new species differs from the other European and Asiatic species by the following characteristics: (1) strong asymmetry between the two valves (length of RV 5% shorter than LV with a posterior gap between the two valves), (2) the shape of the L5 clasping organs (especially the strongly sclerotized hook-shaped right one with a distal hump) and (3) the shape of the hemipenis, rather thin, with a distally oriented a lobe and a small triangular distal h lobe.</p><p>Despite these three differences, Schellencandona claretae sp. nov. may be related to S. simililampadis because of the following two characteristics: (1) a similar chaetotaxy of the A1, without seta on the podomere III in both male and female and two setae on the podomere VI and (2) the z2 seta of the A2 being very long compared to z1 (but not transformed in a claw like in S. simililampadis and S. schellenbergi).</p><p>The similarity between Schellencandona claretae sp. nov. and the other species described here is detailed in the Discussion section.</p></div>	https://treatment.plazi.org/id/096BB133FF807E670019FE8478AF9071	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Issartel, Colin;Marmonier, Pierre	Issartel, Colin, Marmonier, Pierre (2025): Description of five new species of Schellencandona Meisch, 1996 (Ostracoda: Candoninae) from the southern French Alps, a highly diversified area for groundwater ostracods. European Journal of Taxonomy 1022: 85-133, DOI: 10.5852/ejt.2025.1022.3083, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3083/13745
096BB133FF897E5C001AF9E578349215.text	096BB133FF897E5C001AF9E578349215.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Schellencandona malardi Issartel & Marmonier 2025	<div><p>Schellencandona malardi sp. nov.</p><p>urn:lsid:zoobank.org:act: C6E7925B-8EEA-4C2A-A919-23576D70C9BB</p><p>Figs 15–18, 20; Tables 1, 3</p><p>Diagnosis</p><p>Small trapezoid candonine of the genus Schellencandona (L = 485 µm). Carapace thick with strong ornamentation consisting of fossae on the entire valve surface. In lateral view, anterior margin widely rounded, while posterior margin poorly curved, dorsal margin straight or slightly concave in both valves, inclined backwards (decrease of 5.5% of H between the anterior and the posterior cardinal angle), postero-dorsal and ventral margins straight. Greatest H of LV located at the third anterior part (H/L = 0.49). In dorsal view, carapace swallen (W represents 50%–55% of L), pyriform and triangular shaped both anteriorly and posteriorly, with slightly beak-shaped ends. Calcified inner lamella amounting to c. 10% and 7% of L for the anterior and the posterior parts, respectively. A1 with a seta on the anterior side of the 3 rd podomere. Male A2: EII and EIII separated with t2 and t3 transformed in male bristles, 2 short z setae (z1 longer than z2), the longest claw (G2) represents 170% of EI length. Female A2: EII+III with 3 t and 2 z setae. 2 nd podomere of the Mdp bears 3+2 setae. Endopodites of the maxilla (L5) developed in males into prehensile palps, poorly asymmetrical, both elongated and slightly curved. Walking leg (L6) with f and 2 g setae. Cleaning leg (L7) 4-segmented, EII and EIII fused, with 2 setae (d1 and dp) on the protopodite. Zenker’s organ with 6 internal rings of spines. Hemipenis with an outer lobe (a) large and sub-rectangular, dorso-distally oriented. The inner lobe (b) dorsally pointed (close to a 90° angle). Bursa copulatrix (e) rounded with a curved distal strip and an internal sub-conical structure. Female genital lobe rounded anteriorly without posterior expansion. Ocular structures not visible.</p><p>Etymology</p><p>The new species is named after Florian Malard who collected the species during an ecological study of the braided rivers of the southern French Alps.</p><p>Type material</p><p>Holotype</p><p>FRANCE • ♂, dissected appendages mounted in glycerine, valves stored in ethanol; Drôme district, Monjoux municipality; 44.5023° N, 5.0889° E; 446 m a.s.l.; Jun.–Jul. 2010; C. Capderrey leg.; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=5.0889&amp;materialsCitation.latitude=44.5023" title="Search Plazi for locations around (long 5.0889/lat 44.5023)">interstitial habitat of the Lez River</a>; MNHN-IU-2023-714.</p><p>Allotype</p><p>FRANCE • ♀; same data as for holotype; MNHN-IU-2023-715.</p><p>Paratypes</p><p>FRANCE • 1 ♂; same data as for holotype; MNHN-IU-2023-716; MNHN • 1 ♂; same data as for holotype; UCBLZ. 2012-3-153-5; UCBLZ • 1 ♀; same data as for holotype; MNHN-IU-2023-717; MNHN • 1 juv.; same data as for holotype; UCBLZ. 2012-3-153-6 .</p><p>Other material examined</p><p>FRANCE • several juvs, undissected; same data as for holotype; UCBLZ. 2012-3-210 .</p><p>Description</p><p>MEASUREMENTS. Holotype, ♂ (MNHN-2023-714): LV: L = 485 µm, H = 240 µm (H/L = 0.49). RV=470 µm, H=230 µm (H/L=0.49). W=245 µm (W/L=0.50). Range for males (n=2): L=485 to 500 µm, H=240 to 245 µm, W=245 to 250 µm. Allotype, ♀ (MNHN-2023-715): LV: L=465 µm, H=240 µm (H/ L=0.51). RV: L= 455 µm, H=230 µm (H/L=0.50). W=255 µm (W/L=0.55). Range for females (n=2): L=465 to 490 µm, H=230 to 240 µm, W=255 to 260 µm.</p><p>CARAPACE. Whitish, with strong ornamentation (fossae) on entire carapace surface. General shape of carapace trapezoid with marked cardinal angles (Figs 15, 18I–J). Highest H located at anterior third with H/L = 0.49 and 0.51 for male and female, respectively. Carapace viewed laterally with anterior margin widely rounded, posterior margin and ventral margin straight and dorsal margin inclined backwards (5.5% decrease of H between anterior and posterior cardinal angle). Carapace viewed dorsally (Fig. 15C, F) very enlarged, pyriform and triangular-shaped both anteriorly and posteriorly, with greatest W located at last third of animal (at 60% of L in both male and female), representing 50% and 55% of L for male and female, respectively. Anterior and posterior ends slightly beak-shaped.</p><p>VALVES. LV overlaps the RV on all sides (Fig. 15A, G). Dorsal margin of LV and RV slightly concave in the male. Dorsal margin representing 37% of LV length and 33% of RV length. Dorso-posterior margin straight on both valves. Ventral margin straight in LV and RV, slightly convex near oral area (Fig. 15B, E). Anterior calcified inner lamella larger (around 10% of L) than posterior one (7% of L for both sexes). Fused marginal valve zone narrow, representing 2.5% of L for both male and female, with straight and dense radial pore canals, more numerous anteriorly (Fig. 15D).</p><p>ANTENNULE, A1 (Figs 16A, 17A). I+II: A-1l(pu), P-2l(pu) / III: A-1s / IV: A-1s / V: A-1l, P-1s / VI: A-2l / VII: A-2l-1s(α), P-1l / VIII: D-2l-ya-1l(cs). Using IV podomere as reference, podomeres are in ratios of 1.8-1-1.5-1-1.5-1.6 from III to VIII. ya aesthetasc very long, 10.7 × as long as IV podomere.</p><p>ANTENNA, A2 (Figs 16B–D, 17B–D). Protopodite: coxa with 3 setae, 2 long and smooth, 1 short and plumose; basis with 1 long posterior seta; exopodite with 1 long and 2 short setae; EI with 1 posterior aesthetasc Y (equalling 55% to 73% of EI length in male and female, respectively) and distally 2 setae (1s and 1m).</p><p>MALE A2 (Fig. 16B–D). EII and EIII segmented, forming 2 distinct podomeres. EII with 1 short aesthetasc (y1) and 4 t setae, t1 and t4 short, t2 and t3 transformed in male bristles having length equal to 90% of that of EI. EIII with 1 short aesthetasc (y2), 2 external z setae, z1 slightly longer than EIV length, z2 short (15% of EI length). G1 reduced (96% of EI length), G2 well-developed (170% of EI length), G3 reduced to short bristle (45% of EI length). EIV with 2 claws, posteriorly 1 long (Gm, 165% of EI length) and anteriorly 1 reduced (GM, 69% of EI length), 1 small aesthetasc (y3, 39% of EI length) associated with subequal seta, g seta present.</p><p>FEMALE A2 (Fig. 17B–D). EII and EIII fused, with anteriorly 2 short aesthetacs (y1 and y2), 3 t setae, distally 2 z setae (both short). G2 claw reduced (85% of EI length). G1 and G3 claws well-developed and sub-equal (196% EI length). EIV with anteriorly 1 long (GM, 163% of EI length) and posteriorly 1 medium-sized claw (Gm, 129% of EI length), 1 medium-sized aesthetasc (y3, 79% of EI length) with subequal seta, g seta present.</p><p>MANDIBLE. Consisting of coxal plate and 4-segmented palp (Mdp). Coxa typically shaped, heavily chitinized with masticatory part. 1 st podomere of Mdp (Figs 16E, 17F) with externally exopodite plate, internally with 2 long setae (1 smooth and 1 plumose S1) and 2 short setae (1 smooth (α), 1 plumose, S2). 2 nd podomere with externally 2 setae and internally group of 3 smooth setae and second group of 2 setae (1 long and 1 short, β, see detail in Fig. 16E). 3 rd podomere with externally 3 setae, distally 1 long seta (γ) and internally 3 small setae. 4 th podomere with 2 serrated and long claws (160% of the 3 rd podomere length) and 3 small setae.</p><p>MAXILLULAR PALP (Mx1palp, Figs 16F, 17E). Two-segmented: 1 st segment with 4 apical plumose setae on outer corner. 2 nd segment with 2 claw-like setae (4.7× as long as 2 nd segment) and 4 thinner setae.</p><p>MAXILLA (L5, Figs 16I–J, 17G). With protopodite bearing 1 anterior seta (a) and 2 exterior setae (b and d), masticatory process (endite) apically with group of 10 setae. Exopodite plate with 2 filaments. Male endopodites transformed in clasping organs weakly asymmetrical, both elongated and slightly curved; right one thicker than left one. Two endopodites bear 2 short but thick setae on ventral side and thin apical seta. In female, similar set of setae observed on protopodite (a, b, d). Exopodite plate with two filaments. Endopodite ending with three short setae.</p><p>WALKING LEG (L6, Figs 16G, 17J). Five-segmented. Protopodite without d seta, EI without e seta, EII with f seta and EIII with 2 g setae. EIV with 2 short setae (h1 and h3) and long claw (h2) serrated and equal to 250% of length of EII.</p><p>CLEANING LEG (L7, Figs 16K, 17I). Four-segmented (with EII and EIII fused). Protopodite with 1 short (d1) and 1 long setae (dp, 140% of EI length). EI without seta, EII+EIII with g seta (short in male and longer in female, 45% of EI length). EIV with 1 medium seta h1 (62 to 69% of EI length) and two long setae, h2 and h3 (118% and 195% of EI length, respectively).</p><p>CAUDAL RAMUS (CR, Figs 16M, 17H). Slender, with relatively short sp seta (22%–25% of anterior margin of CR, not reaching basis of Gp), short sa seta and 2 relatively short and curved claws (Ga and Gp representing around 64–66% and 47–60% of anterior margin of CR, respectively), both claws serrated.</p><p>FEMALE GENITAL LOBE. Simple (Fig. 17H) widely rounded anteriorly without posterior expansion. Oocytes large (10% of valve length).</p><p>MALE GENITAL ORGANS. Zenker’s organ (male, Fig. 16L) ith 6 internal rings of spines representing 23% of total length of carapace. Hemipenis (Fig. 16H) with distal outer lobe (a) dorso-distally oriented, large and sub-rectangular (with dorsal angle well marked in most individuals, but not all). Inner lobe (b) dorsally pointed, close to 90° angle, and small plication on ventral side. Lobe h not visible. Labyrinth well-sclerotized and divided in 4 sections, section d4 weakly reticulated. Copulatory tube thin located inside rounded bursa copulatrix (e) with well-sclerotized and curved distal strip and internal sub-conical structure. M-process flat with narrow and widely rounded dorsal part, linked to C strip joining base of e and thin basal part reaching d4 section of labyrinth.</p><p>OCULAR STRUCTURES. Not visible.</p><p>Ecology and distribution</p><p>Schellencandona malardi sp. nov. was sampled in the interstitial habitat of a single river, the Lez River, an east-side tributary of the Rhône River (Fig. 1, Table 1), at low elevation (i.e., 446 m a.s.l.). The species was exclusively sampled in groundwater upwelling zones (at the downstream end of gravel bars), never at a depth of 30 cm, rarely at a depth of 60 cm (10%) and mostly at a depth of 90 cm into the riverbed sediment (90% of the individuals). The water collected at a depth of 90 cm had a temperature ranging from 15.5 to 17.2°C, an electrical conductivity ranging from 407 to 452 µS. cm-1, a near-neutrality pH (7.4–7.7) and a wide range of dissolved oxygen concentration (2.15–7.3 mg ·L- 1).</p><p>Specialisation to groundwater: the well-developed aesthetascs (especially ya on A1 and Y on A2), the large size of the oocytes (10% of carapace length) and the lack of developed eyes suggest that Schellencandona malardi sp. nov. is specialized to groundwater life. Schellencandona malardi may thus be considered as a stygobiotic species endemic to the Lez River, living deep in the river bottom sediment and in a wide range of dissolved oxygen concentrations.</p><p>Taxonomic remarks</p><p>The new species fits with all the characteristics listed by Meisch (1996) for Schellencandona (see above) except for the surface of the carapace that is strongly ornamented with fossae on its entire surface. In the genus Schellencandona, the new species differs from all the species described in Europe and in Asia by the pyriform shape of the carapace in dorsal view. This shape is, for the moment, unique in the genus Schellencandona .</p><p>Schellencandona malardi sp. nov. has a hemipenis very similar to those of S. simililampadis and S. schellenbergi (stocky shape, with a large a lobe dorso-distally oriented, the h lobe not visible and a curved distal sclerotized strip on the bursa copulatrix), but differs from those of the above-mentioned species in (1) the presence of one seta in the 3 rd podomere of A1 (absent in the two other species) and (2) the size of the z2 seta on A2 not transformed in a claw and smaller than z1 (claw-like in the two other species).</p><p>To our knowledge, the most closely related species are two unpublished species of Schellencandona collected in the Jura Mountains (Northern French Alps, noted Schellencandona sp. J 1 and sp. J 3 in Dole-Olivier et al. 2009). These two undescribed species exhibit markedly different carapace shapes compared to those of Schellencandona malardi sp. nov., yet share a striking similarity in hemipenis shape as well as A1 and A2 chaetotaxy (for example, see A2 of Schellencandona sp. J 1 in Fig 19B). At present, Schellencandona malardi can be regarded as related to this group of Jura Mountain species; however, this classification requires a thorough evaluation following the complete description of the group.</p><p>The similarity of Schellencandona malardi sp. nov. with the other species described here is detailed in the Discussion.</p></div>	https://treatment.plazi.org/id/096BB133FF897E5C001AF9E578349215	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Issartel, Colin;Marmonier, Pierre	Issartel, Colin, Marmonier, Pierre (2025): Description of five new species of Schellencandona Meisch, 1996 (Ostracoda: Candoninae) from the southern French Alps, a highly diversified area for groundwater ostracods. European Journal of Taxonomy 1022: 85-133, DOI: 10.5852/ejt.2025.1022.3083, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3083/13745
