taxonID	type	description	language	source
0A68182DFF82F866FF39F842EA16F844.taxon	materials_examined	Type genus. Tenuipalpus Donnadieu, 1875	en	Beard, Jennifer J., Ochoa, Ronald (2011): New flat mite genera (Acari: Trombidiformes: Tenuipalpidae) associated with Australian sedges (Cyperaceae). Zootaxa 2941: 1-37, DOI: 10.5281/zenodo.204686
0A68182DFF81F864FF39FF46E965FADA.taxon	materials_examined	Type species. Gahniacarus tuberculatus Beard & Ochoa	en	Beard, Jennifer J., Ochoa, Ronald (2011): New flat mite genera (Acari: Trombidiformes: Tenuipalpidae) associated with Australian sedges (Cyperaceae). Zootaxa 2941: 1-37, DOI: 10.5281/zenodo.204686
0A68182DFF81F864FF39FF46E965FADA.taxon	diagnosis	Diagnosis. Adult female. Body elongate (approximately 4 times longer than wide); anterior margin of propodosoma without projections. All dorsal setae inserted on tubercles except h 1 – 2; setal pair c 1 inserted on single central tubercle. Dorsal setae v 2, sc 1, sc 2, c 1, c 3, d 1, d 3, e 3, f 2, f 3, h 1, h 2 present. Most dorsal setae thick, strongly barbed, except sc 1 sometimes elongate; setae h 1 minute, barbed; h 2 fine, elongate with minute club. Venter finely plicate; setae 1 a, 1 b, 2 b, 2 c, 3 a, 3 b, 4 a 1, 4 a 2, 4 b, ag 1, g 1, g 2, ps 1, ps 2 present. Setae 1 a, 1 b, 4 a 1 – 2 elongate, extremely fine distally. Palps 3 - segmented, with setal formula 0, 2, 0 (2); palp tarsus with two eupathidia. Leg chaetotaxy (Table 1): no setae are added to femora, genua or tibiae during development; ge I – IV are nude in all stages; tr I – IV with one seta, v’. Seta v’ on tr IV is the only addition to the female (i. e. female leg chaetome matches deutonymph except v’ tr IV). Male has the same leg chaetotaxy as female except for the addition of a solenidion (ω ’) to ta I – IV (3 known only for G. tuberculatus). Immatures. Larva (known only for G. tuberculatus) with posterior opisthosomal setae e 3, f 3, h 1 minute. Protonymph with posterior opisthosomal setae f 3, h 1 minute; setae e 3 minute (G. gersonus) or similar to other dorsal setae (G. tuberculatus). Deutonymph with posterior opisthosomal setae h 1 minute; setae f 3 minute (G. gersonus) or similar to other dorsal setae (G. tuberculatus). Seta v’ is added to tr I – III in deutonymph and to tr IV in adult. The leg chaetotaxies of the larva and protonymph are identical to those found on Cyperacarus gen. nov. (Table 1). continued next page Remarks. Gahniacarus gen. nov. can be separated from Cyperacarus gen. nov. by the presence of dorsal opisthosomal setae f 3 (absent in Cyperacarus), and by the size of setae d 1, which is similar in size to the other dorsal setae in Gahniacarus, but minute in Cyperacarus. The two new genera, Gahniacarus and Cyperacarus, have dorsal opisthosomal setae c 1 present, dorsal opisthosomal setae e 1 absent and nude ge I – IV. In comparison, the two previously known tenuipalpid genera associated with Cyperaceae in Australia possess the following characters: dorsal opisthosomal setae c 1 absent, dorsal opisthosomal setae e 1 present, ge I – IV Acaricis 2 - 2 - 1 - 0, Prolixus 2 - 1 - 0 - 0. See Table 2 for further morphological differences.	en	Beard, Jennifer J., Ochoa, Ronald (2011): New flat mite genera (Acari: Trombidiformes: Tenuipalpidae) associated with Australian sedges (Cyperaceae). Zootaxa 2941: 1-37, DOI: 10.5281/zenodo.204686
0A68182DFF81F864FF39FF46E965FADA.taxon	etymology	Etymology. This genus is named for the genus of plant on which it was collected, Gahnia J. R. Forst. & G. Forst. (Cyperaceae).	en	Beard, Jennifer J., Ochoa, Ronald (2011): New flat mite genera (Acari: Trombidiformes: Tenuipalpidae) associated with Australian sedges (Cyperaceae). Zootaxa 2941: 1-37, DOI: 10.5281/zenodo.204686
0A68182DFF80F86AFF39FA13E8ACFCCB.taxon	diagnosis	Diagnosis. Adult female. (Figs 1 a, 2 – 4) As per genus, in addition to: setae sc 1 similar in length to other dorsal setae, inserted on large rounded tubercles; setal pairs c 1 – c 1 and d 1 – d 1 each inserted on single tubercle; f 2 and f 3 tubercles contiguous; large tubercles (not bearing setae) present between sc 1 – c 1, laterad c 1, posterad c 1, posterad d 1. Dorsal opisthosomal setae e 3, f 2, f 3 present. Immatures. (Figs 6 – 8). Larva with posterior opisthosomal setae e 3, f 3, h 1 minute, setae f 2 similar to other dorsal setae. Protonymph with posterior opisthosomal setae f 3, h 1 minute and setae f 2, e 3 similar to other dorsal setae. Deutonymph with posterior opisthosomal setae h 1 minute and setae f 2, f 3, e 3 similar to other dorsal setae (f 3 obviously shorter than f 2, e 3). Opisthosomal setae f 3 migrate anteriorly with each nymphal moult, and are inserted anterior to f 2 in the adult (Figs 2, 6, 7). Differences in how an individual mite is mounted and the direction in which setae are flattened can alter the apparent insertion point of setae f 3 (Fig. 6). The tubercles on which setae f 2, f 3 are inserted are contiguous.	en	Beard, Jennifer J., Ochoa, Ronald (2011): New flat mite genera (Acari: Trombidiformes: Tenuipalpidae) associated with Australian sedges (Cyperaceae). Zootaxa 2941: 1-37, DOI: 10.5281/zenodo.204686
0A68182DFF80F86AFF39FA13E8ACFCCB.taxon	discussion	Remarks. Setae f 2 and f 3 are inserted on contiguous tubercles on adult Gahniacarus tuberculatus, whereas setae f 2 and f 3 are inserted on well separated tubercles on G. gersonus; prodorsal setae sc 1 are similar in size to other dorsal setae in G. tuberculatus, whereas sc 1 are elongate and tapered in G. gersonus. Adult female. (15 measured). Dorsum. (Figs 1 a, 2) Body measurements: v 2 – h 1 210 – 237 [235] (v 2 – tip of protuberance 217 – 246 [245]), sc 2 – sc 2 74 – 80 [77], c 3 – c 3 87 – 92 [89], f 3 – f 3 45 – 59 [57]. All dorsal setae, except h 1 – h 2, inserted on large tubercles: setae sc 1 inserted on a pair of large rounded tubercles; setal pairs c 1 – c 1 and d 1 – d 1 each inserted on single tubercle; tubercles f 2 – f 3 contiguous; additional non-seta bearing tubercles present posterad sc 1, laterad c 1, mesad c 3 and posterad d 1. Setae h 1 – h 2 inserted beneath a tubercle-like posterior projection. Dorsal cuticle mostly smooth, with some minor transverse grooves and plicae posterior to setae c 1 and some oblique plicae posterior to setae d 1. Most dorsal setae thick, barbed, of subequal length; setae h 1 minute; h 2 elongate, barbed with minute distal club; all other dorsal setae thick, heavily barbed; barbs on prodorsal setae longer than those on opisthosoma: v 2 37 – 41 [40 – 41], sc 1 57 – 91 [67 – 80] (two specimens with elongate sc 1, 115 – 126 (89119, 89128 )), sc 2 33 – 43 [39 – 42], c 1 52 – 66 [59 – 60], c 3 34 – 40 [35 – 40], d 1 57 – 66 [61], d 3 40 – 46 [45], e 3 46 – 53 [47 – 48], f 2 44 – 55 [49 – 54], f 3 41 – 50 [49], h 1 2 – 4 [3 – 4], h 2 126 – 148 [141]. Gnathosoma. (Fig. 2) Gnathosoma extends to genu I. Dorsal cuticle with longitudinal plicae laterally; ventral cuticle anterior to setae 1 b finely papillate, longitudinal plicae between 1 b – 1 b and 1 b – 1 a. Ventral setae m absent. Palps three segmented; setal formula: 0, 2, 0 (2); tibia with two setae (d 11 – 12; v 10 – 11); tarsus with two eupathidia (6 – 7; 5 – 6) [7; 5]. Cheliceral stylets 115 – 127. Venter. (Fig. 3) Cuticle of ventral gnathosoma finely papillate; remaining cuticle completely plicate, covered with fine, mostly transverse plicae; transverse plicae between setae 1 a – g 1; plicae longitudinal in coxal fields III – IV and laterad genital flap; plicae oblique on genital flap. Setae g 1 inserted anterior to g 2 on genital flap. All setae fine, smooth; setae 1 a, 1 b, 4 a 1, 4 a 2 elongate, fine (difficult to consistently determine full length). Setal measurements: 1 a 69 – 119 [69], 1 b 54 – 82 [67 – 82], 2 b 22 – 32 [30 – 32], 2 c 20 – 28 [24 – 25], 3 a 30 – 37 [32 – 36], 3 b 19 – 32 [20], 4 a 1 53 – 105 [76], 4 a 2 64 – 99 [99], 4 b 19 – 26 [21], ag 19 – 25 [19], g 1 23 – 32 [32], g 2 21 – 31 [27 – 30], ps 1 15 – 19 [16 – 18], ps 2 15 – 18 [16]. Spermathecal apparatus. (Fig. 3) Cylindrical vesicle, curled distally (27 – 32) [28]. Legs. (Fig. 4) Setal formulae for legs I – IV: 1 - 1 - 3 - 0 - 5 - 7 (1), 2 - 1 - 3 - 0 - 5 - 7 (1), 1 - 1 - 2 - 0 - 3 - 3, 1 - 1 - 2 - 0 - 3 - 3 respectively. Tarsi I and II each with one antiaxial solenidion ω ” (5 – 6, 4 – 5 respectively) and two distal eupathidia p ζ ’ - p ζ ” (7 – 8, 7 – 8; 6 – 7, 7 respectively); ta I – IV with u’ - u ” asymmetrically barbed. Femur I with long, barbed dorsal seta (31 – 40) [32 – 37]. Colour. This species is yellow with small black spots visible internally (presumed to be food in the gut). Adult male. (8 measured). Dorsum. (Fig. 5) Body measurements: v 2 – h 1 181 – 200, sc 2 – sc 2 68 – 72, c 3 – c 3 70 – 75, f 3 – f 3 45 – 46. Dorsal cuticle mostly smooth; band of weak transverse plicae anterior to setae c 1. Setae sc 1, c 3, d 3, e 3, f 2 inserted on large rounded tubercles; setae v 2, sc 2, c 1, d 1, f 3 inserted on smaller tubercles (tubercles f 2, f 3 contiguous); setae h 1 – 2 inserted under posterior tubercle-like protuberance. Setae h 1 short; h 2 elongate with minute distal club; all other dorsal setae thick, heavily barbed. Setal measurements: v 2 29 – 32, sc 1 27 – 30, sc 2 30 – 39, c 1 24 – 28, c 3 29 – 32, d 1 20 – 26, d 3 30 – 34, e 3 36 – 40, f 2 35 – 39, f 3 33 – 36, h 1 5 – 7, h 2 114 – 148. Gnathosoma. (Fig. 5) Gnathosoma extends just past genu I. Dorsal cuticle with longitudinal plicae laterally; ventral cuticle with fine longitudinal plicae anterior to setae 1 b. Ventral setae m absent. Palps three segmented; setal formula: 0, 2, 0 (2); tibia with two setae (d 10 – 11, v 9); tarsus with two eupathidia (6 – 7; 4 – 6). Cheliceral stylets 89 – 92. Venter. Cuticle completely plicate, covered with fine, mostly transverse plicae; plicae longitudinal between gnathosoma – 1 a, transverse between 1 a – 4 a 2; band of strong plicae and folds between coxae IV; plicae longitudinal in coxal fields III – IV; plicae oblique posterior 4 a 1 – 2, oblique to transverse ag – g 1 (plicae longitudinal / oblique laterad anus, arching anteriorly to become transverse anterior g 1 – 2). Setae ps 1 modified to short thick spurs. All setae fine; setae 1 a, 1 b, 4 a 1, 4 a 2 elongate (difficult to consistently determine full length). Setal measurements: 1 a 77 – 117, 1 b 44 – 76, 2 b 22 – 26, 2 c 19 – 24, 3 a 28 – 31, 3 b 17 – 20, 4 a 1 63 – 78, 4 a 2 42 – 72, 4 b 15 – 20, ag 14 – 17, g 1 13 – 19, g 2 14 – 15, ps 1 6 – 7, ps 2 11 – 15. Aedeagus. (Fig. 5) Sclerotised elongate, narrow arrow-head shaped aedeagus (28 – 33), with long, narrow membranous tube (up to 85), indistinguishable distally. Legs. (Fig. 5) Setal formulae same as adult female except ω ’ present on ta I – IV: 1 - 1 - 3 - 0 - 5 - 8 (2), 2 - 1 - 3 - 0 - 5 - 8 (2), 1 - 1 - 2 - 0 - 3 - 4 (1), 1 - 1 - 2 - 0 - 3 - 4 (1) respectively. Tarsi I and II each with two solenidion (ta I paraxial ω ’ 8 – 9, antiaxial ω ” 7 – 8; ta II paraxial ω ’ 8, antiaxial ω ” 6 – 7) and two distal eupathidia p ζ ’ - p ζ ” (7 – 8, 8 – 9; 8, 6 – 7 respectively); tarsi III and IV each with one paraxial solenidion ω ’ (8, 8 – 9 respectively); ta I – IV u’ - u ” asymmetrically barbed. Femur I with long, barbed dorsal seta (27 – 32). Deutonymph. (5 measured). Dorsum. (Fig. 6) Body measurements: v 2 – h 1 170 – 228, sc 2 – sc 2 64 – 73, c 3 – c 3 70 – 82, f 3 – f 3 31 – 43. Cuticle mostly smooth with some transverse plicae mesally sc 2 – d 1. Setae sc 1, c 1, c 3, d 1, d 3, e 3, f 2 inserted on rounded tubercles (tubercles f 2, f 3 contiguous); additional tubercles between setae sc 1, anterior c 3, on posterior margin of opisthosoma. Setae h 1 – 2 inserted under posterior tubercle-like protuberance. Setae f 3 short; h 1 minute; setae h 2 elongate, barbed, with minute club; all other setae thick, heavily barbed. Setal measurements: v 2 30 – 35, sc 1 31 – 47, sc 2 24 – 31, c 1 24 – 34, c 3 28 – 30, d 1 23 – 35, d 3 27 – 32, e 3 29 – 36, f 2 27 – 32, f 3 13 – 18 (one specimen 6), h 1 1 – 2, h 2 112 – 126. Gnathosoma. Gnathosoma extends to genu I. Ventral cuticle anterior to setae 1 b finely papillate; cuticle with longitudinal plicae between setae 1 b – 1 b and 1 b – 1 a. Ventral setae m absent. Palp with three segments; setal formula: 0, 2, 0 (2); tibia with two setae (d 10, v 8); tarsus with two eupathidia (6, 5). Cheliceral stylets 97 – 103. Venter. Cuticle completely plicate, covered mostly with transverse plicae; plicae longitudinal from gnathosoma to setae 1 a; plicae transverse between 1 a – g 1; plicae oblique laterad ps 1 – 2. All ventral setae fine; setae 1 a, 1 b, 4 a 1, 4 a 2 elongate (difficult to determine full length). Setal measurements: 1 a 63 – 104, 1 b 37 – 56, 2 b 15 – 20, 2 c 16 – 20, 3 a 13 – 23, 3 b 13 – 20, 4 a 1 38 – 53, 4 b 10 – 12, ag 7 – 11, g 1 5 – 12, ps 1 6 – 7, ps 2 6. Legs. Setal formulae same as adult female except tr IV nude: 1 - 1 - 3 - 0 - 5 - 7 (1), 2 - 1 - 3 - 0 - 5 - 7 (1), 1 - 1 - 2 - 0 - 3 - 3, 1 - 0 - 2 - 0 - 3 - 3 respectively. Tarsi I and II each with one antiaxial solenidion ω ” (4 – 5; 4 respectively) and two distal eupathidia p ζ ’ - p ζ ” (6, 6; 6, 6 respectively); ta I – IV u’ - u ” asymmetrically barbed. Femur I with long, barbed dorsal seta (25 – 34). Protonymph. (3 measured). Dorsum. (Fig. 7) Body measurements: v 2 – h 1 161, sc 2 – sc 2 65, c 3 – c 3 66, f 3 – f 3 29. Cuticle with smooth area anteriorly on prodorsum and posteriorly between setae d 1 – h 1; transverse plicae between setae sc 2 – d 1. Setae sc 1, c 1, d 3, e 3, f 2 inserted on rounded tubercles (tubercles e 3, f 2 contiguous); additional tubercles between setae sc 1, laterad c 1, posterior margin of opisthosoma. Setae h 1, h 2 inserted under posterior tubercle-like projection. Setae f 3, h 1 minute; h 2 elongate, barbed, with minute club; all other setae thick, strongly barbed: v 2 28 – 31, sc 1 34 – 35, sc 2 31 – 32, c 1 25 – 29, c 3 26 – 27, d 1 22 – 23, d 3 27 – 28, e 3 24 – 28, f 2 20 – 21, f 3 1 – 2, h 1 1, h 2 95. Gnathosoma. Gnathosoma extends beyond genu I. Ventral cuticle anterior setae 1 b finely papillate; cuticle with longitudinal plicae between setae 1 b – 1 b and 1 b – 1 a. Ventral setae m absent. Palps with three segments; setal formula: 0, 2, 0 (2); tibia with two setae (d 8 – 9, v 7); tarsus with two eupathidia (6, 4). Cheliceral stylets 88. Venter. Cuticle completely plicate, mostly covered with transverse striae; plicae longitudinal from gnathosoma to 1 a; plicae transverse 1 a – ag; plicae oblique laterad ps 1 – 2. Setae fine; setae 1 a, 1 b elongate (difficult to determine full length). Setal measurements: 1 a 57 – 61, 1 b 43, 2 c 7 – 10, 3 a 30, 3 b 8, ag 4, ps 1 4, ps 2 4. Legs. Setal formulae for legs I – IV: 1 - 0 - 3 - 0 - 5 - 7 (1), 1 - 0 - 3 - 0 - 5 - 7 (1), 1 - 0 - 2 - 0 - 3 - 3, 0 - 0 - 2 - 0 - 3 - 3 respectively. Tarsi I and II each with one antiaxial solenidion ω ” (4; 3 respectively) and two distal eupathidia p ζ ’ - p ζ ” (6, 6; 5, 5 respectively); ta I – IV u’ - u ” asymmetrically barbed. Femur I with long, barbed dorsal seta (24 – 25). Larva. (3 measured) Dorsum. (Fig. 8) Body measurements: v 2 – h 1 132, sc 2 – sc 2 63, c 3 – c 3 70, f 3 – f 3 26. Cuticle almost entirely plicate with mostly transverse plicae between setae sc 2 – d 1; central prodorsum and posterior opisthosoma with smooth cuticle. Dorsum with large rounded tubercle anterior to setae c 1. Most opisthosomal setae set on small tubercles, except e 3, f 3, h 1. Most dorsal setae of subequal length, thick, strongly barbed, except setae e 3, f 3, h 1 minute; h 2 elongate, lightly barbed basally: v 2 23, sc 1 29 – 31, sc 2 28 – 29, c 1 22 – 23, c 3 25 – 26, d 1 26 – 29, d 3 26 – 27, e 3 2, f 2 26 – 27, f 3 1 – 2, h 1 1 – 2, h 2 98 – 100. Gnathosoma. (Fig. 8) Gnathosoma extends beyond tibia I. Ventral cuticle longitudinal; ventral setae m absent. Palps with three segmented; 0, 2, 0 (2); tarsus with two eupathidia (6; 2); tarsus with two setae (d 7 – 9, v 6 – 7). Cheliceral stylets 80. Venter. Plicae on cuticle fine, difficult to see, mostly transverse with longitudinal plicae between setae 1 b – 1 a; transverse plicae between setae 1 a – ps 2, oblique laterad ps 1 – 2. Setal measurements: 1 a 79 – 94, 1 b 32, 3 a 25, ps 1 2, ps 2 3. Legs. (Fig. 8) Setal formulae for legs I – IV: 1 - 0 - 3 - 0 - 5 - 6 (1), 0 - 0 - 3 - 0 - 5 - 6 (1), 0 - 0 - 2 - 0 - 3 - 3 respectively. Tarsi I and II each with one antiaxial solenidion ω ” (3, 3 respectively) and two distal eupathidia p ζ ’ - p ζ ” (5, 6; 5, 5 respectively); ta I – IV u’ - u ” asymmetrically barbed. Femur I with long, barbed dorsal seta (17 – 18). Host. Rough saw sedge, Gahnia aspera sensu lato (Cyperaceae) growing along margin of notophyll vineforest.	en	Beard, Jennifer J., Ochoa, Ronald (2011): New flat mite genera (Acari: Trombidiformes: Tenuipalpidae) associated with Australian sedges (Cyperaceae). Zootaxa 2941: 1-37, DOI: 10.5281/zenodo.204686
0A68182DFF80F86AFF39FA13E8ACFCCB.taxon	distribution	Distribution. Mt Hedley, Far North Queensland.	en	Beard, Jennifer J., Ochoa, Ronald (2011): New flat mite genera (Acari: Trombidiformes: Tenuipalpidae) associated with Australian sedges (Cyperaceae). Zootaxa 2941: 1-37, DOI: 10.5281/zenodo.204686
0A68182DFF80F86AFF39FA13E8ACFCCB.taxon	discussion	Remarks. Live individuals of this species are yellow in colour and live within grooves on the ventral surface of the leaf blade of its sedge host plant.	en	Beard, Jennifer J., Ochoa, Ronald (2011): New flat mite genera (Acari: Trombidiformes: Tenuipalpidae) associated with Australian sedges (Cyperaceae). Zootaxa 2941: 1-37, DOI: 10.5281/zenodo.204686
0A68182DFF80F86AFF39FA13E8ACFCCB.taxon	etymology	Etymology. This species is named for the tubercles present over the dorsum. The Latin word “ tuber ” means swelling.	en	Beard, Jennifer J., Ochoa, Ronald (2011): New flat mite genera (Acari: Trombidiformes: Tenuipalpidae) associated with Australian sedges (Cyperaceae). Zootaxa 2941: 1-37, DOI: 10.5281/zenodo.204686
0A68182DFF8EF869FF39FC05EC9CF868.taxon	diagnosis	Diagnosis. Adult female (Figs 1 b, 9). As per genus, in addition to: setae sc 1 inserted on weakly developed tubercles, elongate; setal pair c 1 - c 1 inserted on tubercle; f 2 and f 3 tubercles separated; large tubercles (not bearing setae) present between sc 1 - c 1, posterad c 1. Dorsal opisthosomal setae e 3, f 2, f 3 present. Immatures (Figs 10 – 11). Larva not known. Protonymph with posterior opisthosomal setae e 3, f 3, h 1 minute and setae f 2 similar to other dorsal setae. Deutonymph with posterior opisthosomal setae f 3, h 1 minute and setae d 1, e 3, f 2 similar to other dorsal setae.	en	Beard, Jennifer J., Ochoa, Ronald (2011): New flat mite genera (Acari: Trombidiformes: Tenuipalpidae) associated with Australian sedges (Cyperaceae). Zootaxa 2941: 1-37, DOI: 10.5281/zenodo.204686
0A68182DFF8EF869FF39FC05EC9CF868.taxon	discussion	Remarks. Setae f 2 and f 3 are inserted on well separated tubercles in G. gersonus, whereas setae f 2 and f 3 are inserted on contiguous tubercles in Gahniacarus tuberculatus; prodorsal setae sc 1 are elongate and tapered in G. gersonus, whereas setae sc 1 are similar in size to other dorsal setae in G. tuberculatus. Adult female. (15 measured). Dorsum. (Figs 1 b, 9) Body measurements: v 2 – h 1 287 – 315, sc 2 – sc 2 75 – 86, c 3 – c 3 81 – 92, f 3 – f 3 37 – 42. Dorsal cuticle variable, with some indication of a prodorsal shield: oblique plicae mesad setae sc 2 and eyes; transverse plicae posterad prodorsal shield to level of c 3; cuticle between c 1 – e 2 mostly smooth with some oblique grooves and plicae associated with tubercles; longitudinal plicae present between f 3 – f 3. Setae c 1 inserted together on large single central tubercle; setae sc 1, c 3, d 3, e 3, f 2 on large tubercles; single smooth tubercle present posterad c 1. Setae sc 1 obviously elongate, strongly barbed; h 1 minute, barbed; h 2 elongate, smooth with minute distal club; all other dorsal setae thick with dorsal surface strongly barbed, almost pubescent (especially basally) and ventral surface smooth: v 2 37 – 43, sc 1 171 – 215, sc 2 32 – 38, c 1 43 – 58, c 3 31 – 35, d 1 45 – 53, d 3 35 – 41, e 3 33 – 40, f 2 48 – 55, f 3 28 – 38, h 1 3, h 2 150 – 166. Gnathosoma. (Fig. 9) Gnathosoma elongate, as long as leg I. Dorsal cuticle with longitudinal plicae; ventral cuticle finely papillate anterad setae 1 b. Ventral setae m absent. Palps with three segments; 0, 2, 0 (2); tibia with two setae (d 13 – 16, v 10 – 13); tarsus with two eupathidia (5 – 6, 7 – 9). Cheliceral stylets 147 – 158. Venter. Cuticle with fine, mostly transverse, plicae; plicae longitudinal on ventral gnathosoma to setae 1 a; plicae transverse between 1 a - ag, becoming thicker and more widely spaced laterally between leg II – III (similar to dorsal plicae); plicae on genital flap longitudinal laterally, arching anteriorly around g 1; plicae longitudinal to oblique laterad genito-anal region. Setae g 1 and g 2 on genital flap. All ventral setae fine, smooth. Setal measurements: 1 a 63 * – 146, 1 b 64 – 84, 2 b 18 – 30, 2 c 15 – 22, 3 a 20 – 25, 3 b 13 – 21, 4 a 1 49 – 77, 4 a 2 48 – 80, 4 b 10 – 19, ag 18 – 20, g 1 18 – 25, g 2 16 – 24, ps 1 13 – 19, ps 2 10 – 15 (* = broken). Setae 1 a, 1 b, 4 a 1, 4 a 2 elongate, fine (difficult to consistently determine full length); setae 2 b and 2 c widely separated. Spermatheca. (Fig. 9) A long narrow tube from the genital opening (anterad seta ps 2) terminates in a small blunt-tipped, narrow, cylindrical vesicle (10 – 11). Legs. (Fig. 9) Setal formula for legs I – IV: 1 - 1 - 3 - 0 - 5 - 7 (1), 2 - 1 - 3 - 0 - 5 - 7 (1), 1 - 1 - 2 - 0 - 3 - 3, 1 - 1 - 2 - 0 - 3 - 3 respectively. Tarsi I and II each with one antiaxial solenidion ω ” (5 – 7, 4 – 5 respectively) and two distal eupathidia p ζ ’ - p ζ ” (7 – 8, 6 – 7; 6 – 8, 6 respectively); ta I – IV u’ - u ” asymmetrically barbed. Femur I with long, barbed dorsal seta (50 – 62). Tenent hairs increasing in length, shortest basally to longest distally. Colour. This species is lemon green with small black spots internally (presumably food in the gut). Adult male. Unknown. Deutonymph. (1 measured). Dorsum. (Fig. 10) Body measurements: v 2 – h 1 258, sc 2 – sc 2 78, c 3 – c 3 82, f 3 – f 3 43. Cuticle completely plicate, mostly transverse plicae, with some longitudinal plicae on prodorsum between v 2 – sc 1 and posteriorly between setae d 1 – h 1. Setae f 3, h 1 minute; setae h 2 elongate, smooth; all other setae thick, strongly barbed: v 2 23 – 24, sc 1 52, sc 2 27 – 28, c 1 23, c 3 25 – 27, d 1 24, d 3 26 – 29, e 3 20 – 22, f 2 29 – 30, f 3 3, h 1 2, h 2 91. Gnathosoma. (Fig. 10) Gnathosoma elongate, as long as leg I. Ventral setae m absent. Palps with three segments; formula 0, 2, 0 (2); tibia with two setae (d 11, v 8); tarsus with two eupathidia (5, 8). Cheliceral stylets 119. Venter. Cuticle completely plicate; longitudinal plicae on ventral gnathosoma to setae 1 a, transverse plicae 1 a – g 1; oblique plicae laterad ps 1 – 2 (anal region); lateral plicae between legs II – III stronger and more widely separated than medial plicae. Setal measurements: 1 a 47 * – 92, 1 b 32, 2 b 17 – 18, 2 c 14 – 15, 3 a 16, 3 b 11, 4 a 1 41, 4 b 8, ag 9, g 1 7 – 8, ps 1 6 – 7, ps 2 4 (* = broken). Setae 4 a 2, g 2 absent. Legs. (Fig. 10) Setal formulae for legs I – IV: 1 - 1 - 3 - 0 - 5 - 7 (1), 2 - 1 - 3 - 0 - 5 - 7 (1), 1 - 1 - 2 - 0 - 3 - 3, 1 - 0 - 2 - 0 - 3 - 3 respectively. Tarsis I and II each with one antiaxial solenidion ω ” (4, 3 respectively) and two distal eupathidia p ζ ’ - p ζ ” (7, 6; 6, 6 respectively); ta I – IV u’ - u ” asymmetrically barbed. Femur I with long, barbed dorsal seta (31). Protonymph. (1 measured). Dorsum. (Fig. 11) Body measurements: v 2 – h 1 202, sc 2 – sc 2 72, c 3 – c 3 77, f 3 – f 3 31. Cuticle completely plicate, mostly transverse plicae, with some longitudinal plicae on prodorsum between setae v 2 – sc 1 and posteriorly between setae d 1 – h 1. Setae e 3, f 3, h 1 minute; h 2 elongate with minute club; all other setae thick, strongly barbed: v 2 8 – 10, sc 1 29, sc 2 21 – 22, c 1 16, c 3 23 – 24, d 1 12 – 15, d 3 24 – 25, e 3 3 – 5, f 2 18 – 19, f 3 1 – 2, h 1 1 – 2, h 2 74. Gnathosoma. (Fig. 11) Gnathosoma almost as long as leg I. Band of oblique plicae ventro-lateral margins. Ventral setae m absent. Palps with three segments; formula 0, 2, 0 (2); tibia with two setae (d 9 – 10, v 7 – 8); tarsus with two eupathidia (5, 7). Cheliceral stylets 93. Venter. Cuticle completely plicate; plicae longitudinal gnathosoma to setae 1 a, transverse 1 a – ag, oblique laterad ps 1 – 2. Setal measurements: 1 a 42 – 48, 1 b 36 – 42, 2 c 9, 3 a 11, 3 b 5, ag 3, ps 1 3, ps 2 3. Setae 2 b, 4 a 1, 4 a 2, 4 b, g 1, g 2 absent. Legs. (Fig. 11) Setal formulae for legs I – IV: 1 - 0 - 3 - 0 - 5 - 7 (1), 1 - 0 - 3 - 0 - 5 - 7 (1), 1 - 0 - 2 - 0 - 2 - 3, 0 - 0 - 2 - 0 - 3 - 3 respectively. Tarsus I and II each with one antiaxial solenidion (3, 2 – 3 respectively) and two distal eupathidia p ζ ’ - p ζ ” (5, 5; 5, 4 respectively). Femur I with long, barbed dorsal seta (19). Larva. Unknown. Host. Red-fruited saw sedge, Gahnia sieberiana (Cyperaceae), growing in open eucalypt forest along a watercourse, in clay soil.	en	Beard, Jennifer J., Ochoa, Ronald (2011): New flat mite genera (Acari: Trombidiformes: Tenuipalpidae) associated with Australian sedges (Cyperaceae). Zootaxa 2941: 1-37, DOI: 10.5281/zenodo.204686
0A68182DFF8EF869FF39FC05EC9CF868.taxon	distribution	Distribution. Mt Alexander Reserve, Mittagong, New South Wales, Australia.	en	Beard, Jennifer J., Ochoa, Ronald (2011): New flat mite genera (Acari: Trombidiformes: Tenuipalpidae) associated with Australian sedges (Cyperaceae). Zootaxa 2941: 1-37, DOI: 10.5281/zenodo.204686
0A68182DFF8EF869FF39FC05EC9CF868.taxon	discussion	Remarks. Individuals of this species wedge their bodies into the longitudinal grooves on the ventral surface of the leaf blades of the host plant. Once in the grooves, the adult mites extend their elongate anterior (sc 1) and posterior setae (h 2) out over the surface of the leaf blade. Numbers of mites were higher towards the tip of the leaf blades than near the base.	en	Beard, Jennifer J., Ochoa, Ronald (2011): New flat mite genera (Acari: Trombidiformes: Tenuipalpidae) associated with Australian sedges (Cyperaceae). Zootaxa 2941: 1-37, DOI: 10.5281/zenodo.204686
0A68182DFF8EF869FF39FC05EC9CF868.taxon	etymology	Etymology. This species is named for our colleague and fellow tenuipalpiphile, Uri Gerson.	en	Beard, Jennifer J., Ochoa, Ronald (2011): New flat mite genera (Acari: Trombidiformes: Tenuipalpidae) associated with Australian sedges (Cyperaceae). Zootaxa 2941: 1-37, DOI: 10.5281/zenodo.204686
0A68182DFF91F875FF39FF46ECACFB55.taxon	materials_examined	Type species. Cyperacarus naomae Beard & Ochoa	en	Beard, Jennifer J., Ochoa, Ronald (2011): New flat mite genera (Acari: Trombidiformes: Tenuipalpidae) associated with Australian sedges (Cyperaceae). Zootaxa 2941: 1-37, DOI: 10.5281/zenodo.204686
0A68182DFF91F875FF39FF46ECACFB55.taxon	diagnosis	Diagnosis. Adult female. Body elongate (approximately 3 times longer than wide); anterior margin of propodosoma with three prominent projections — a single median projection without setae, and a pair of prominent lateral projections each bearing setae v 2. Dorsal setae sc 1, h 2 elongate, tapered. Dorsal lateral setae sc 1, c 3, d 3, e 3, f 2 inserted on prominent tubercles; setal pairs c 1 - c 1 and d 1 - d 1 not inserted on single tubercles. Dorsal setae v 2, sc 1, sc 2, c 1, c 3, d 1, d 3, e 3, f 2, h 1, h 2 present; setae f 3 absent. Most dorsal setae broad, with strong lateral barbs, finely pubescent on dorsal surface and smooth ventrally, except setae sc 1 elongate barbed; setae c 1 short; setae d 1, h 1 minute, barbed; h 2 with minute club. Venter finely plicate; setae 1 a, 1 b, 2 b, 2 c, 3 a, 3 b, 4 a 1, 4 a 2, 4 b, ag, g 1, g 2, ps 1, ps 2 present. Setae 1 a, 1 b, 4 a 1 - 2 elongate, extremely fine distally. Palps 3 - segmented, with setal formula 0, 2, 0 (2); palp tarsus with two eupathidia. Leg chaetotaxy: nude trochanters and genua (male with v’ on tr I); no additions from deutonymph to female (v’ tr I, l’ fe I, ω’ ta I – IV are added from deutonymph to male); female fe I with 3 setae (male fe I with 4 setae). Leg chaetotaxy in all stages almost identical to that of Gahniacarus, except Cyperacarus with nude trochanters in all stages and male adds one seta to fe I (l’) (Table 1). Immatures. Opisthosomal setal pair c 1, inserted on single central tubercle, is much longer in immature stages than in the adult. Larva with posterior opisthosomal setae d 1, e 3, h 1 minute, and setae f 2 short. Protonymph with posterior opisthosomal setae d 1, e 3, h 1 minute, and setae f 2 similar to other dorsal setae. Deutonymph with posterior opisthosomal setae d 1, h 1 minute, and setae e 3, f 2 similar to other dorsal setae. Leg chaetotaxy: no setae added to the larval complement on the femora, genua or tibiae during development; tr I – IV and ge I – IV are nude in all stages. The leg chaetotaxies of the larva and protonymph are identical to those found on Gahniacarus gen. nov. (Table 1).	en	Beard, Jennifer J., Ochoa, Ronald (2011): New flat mite genera (Acari: Trombidiformes: Tenuipalpidae) associated with Australian sedges (Cyperaceae). Zootaxa 2941: 1-37, DOI: 10.5281/zenodo.204686
0A68182DFF91F875FF39FF46ECACFB55.taxon	discussion	Remarks. Cyperacarus can be separated from Gahniacarus gen. nov. by the absence of dorsal opisthosomal setae f 3 (present in Gahniacarus), and by the size of setae d 1 which is minute in Cyperacarus, but similar in size to other dorsal setae in Gahniacarus. The two new genera Gahniacarus and Cyperacarus both have dorsal opisthosomal setae c 1 present, dorsal opisthosomal setae e 1 absent and nude ge I-IV. In comparison, the two previously known tenuipalpid genera associated with Cyperaceae in Australia possess the following characters: dorsal opisthosomal setae c 1 absent, dorsal opisthosomal setae e 1 present, ge I-IV Acaricis 2 - 2 - 1 - 0, Prolixus 2 - 1 - 0 - 0. See Table 2 for further morphological differences.	en	Beard, Jennifer J., Ochoa, Ronald (2011): New flat mite genera (Acari: Trombidiformes: Tenuipalpidae) associated with Australian sedges (Cyperaceae). Zootaxa 2941: 1-37, DOI: 10.5281/zenodo.204686
0A68182DFF91F875FF39FF46ECACFB55.taxon	etymology	Etymology. This genus is named for the family of plants on which it was collected, Cyperaceae.	en	Beard, Jennifer J., Ochoa, Ronald (2011): New flat mite genera (Acari: Trombidiformes: Tenuipalpidae) associated with Australian sedges (Cyperaceae). Zootaxa 2941: 1-37, DOI: 10.5281/zenodo.204686
0A68182DFF91F87BFF39FAE4ECD3FA5D.taxon	diagnosis	Diagnosis. Adult female (Figs 1 c, 12 – 13). As per genus, in addition to: setae c 1 short, thick, barbed; setae d 1 short to minute, barbed; male with dorsal setae sc 1 not elongate, subequal in length with setae v 2. Immatures (Figs 16 – 19). Larva with posterior opisthosomal setae d 1, e 3 and h 1 minute, setae f 2 short. Protonymph with posterior opisthosomal setae d 1, e 3 and h 1 minute, setae f 2 similar to other dorsal setae. Deutonymph with posterior opisthosomal setae d 1 and h 1 minute, setae e 3, f 2 similar to other dorsal setae. Adult female. (9 measured). Dorsum. (Figs 1 c, 12) Body measurements: v 2 – h 1 265 – 301, sc 2 – sc 2 91 – 102, c 3 – c 3 106 – 122, f 2 – f 2 58 – 63. Dorsal cuticle mostly smooth, with some minor transverse grooves and plicae between setae sc 2 and c 1. A dorsal shield or dorsal thickening is evident, marked laterally by a change in cuticle. Anterior margin of prodorsum with three projections, one large rounded central and two smaller lateral projections, on which setae v 2 are inserted (on antiaxial margin of projections). Setae sc 1, c 3, d 3, e 3, f 2 inserted on large rounded tubercles. Setae sc 1 thick, elongate, barbed; c 1 short, bushy; d 1, h 1 minute, barbed; h 2 elongate, barbed with minute distal club; all other dorsal setae broad, thick, dorsoventrally flattened, with strong lateral barbs; dorsal surface of setae finely spiculate (Fig. 12, detail), ventral surface smooth, with a median keel: v 2 22 – 30 [26], sc 1 192 – 216 [198 – 200], sc 2 33 – 42 [33 – 35], c 1 9 – 14 [10 – 11], c 3 34 – 41 [35 – 38], d 1 3 – 6 [6], d 3 34 – 42 [36 – 42], e 3 36 – 41 [38 – 40], f 2 34 – 41 [34 – 40], h 1 3 – 7 [5], h 2 156 – 190 [156 – 167]. Gnathosoma. (Fig. 12) Gnathosoma extends to distal margin of genu I. Dorsal cuticle with longitudinal plicae; ventral cuticle finely papillate with some transverse striae anterior to setae 1 b. Ventral setae m absent. Palps with three segments; formula 0, 2, 0 (2); tibia with two setae (d 13 – 14, v 9 – 11); tarsus with two eupathidia (8 – 9; 4 – 5). Cheliceral stylets 109 – 116. Venter. (Fig. 13) Cuticle completely plicate, covered with fine, mostly transverse plicae, between setae 1 a – g 2; some anterior longitudinal plicae between setae 1 b – 1 a; plicae longitudinal and oblique laterad genital flap, becoming transverse anterior to genital flap. Setae g 1 inserted in an anterior position to g 2 on genital flap. All setae fine, smooth; setae 1 a, 1 b, 4 a 1, 4 a 2 elongate, fine (difficult to consistently determine full length). Setal measurements: 1 a 63 * – 119 [119], 1 b 62 – 84 [64], 2 b 18 – 34 [32], 2 c 15 – 26 [22], 3 a 18 – 29 [25], 3 b 22 – 30 [24], 4 a 1 37 * – 103 [62 – 81], 4 a 2 40 * – 88 [60 – 83], 4 b 18 – 28 [23], ag 18 – 21 [21], g 1 17 – 25 [20], g 2 15 – 25 [25], ps 1 11 – 16 [11], ps 2 13 – 19 [13] (* = broken). Spermatheca. (Fig. 13) An elongate, cylindrical vesicle (15 – 18) [16]. Legs. (Figs 12, 13) Setal formulae for legs I – IV: 1 - 0 - 3 - 0 - 5 - 7 (1), 2 - 0 - 3 - 0 - 5 - 7 (1), 1 - 0 - 2 - 0 - 3 - 3, 1 - 0 - 2 - 0 - 3 - 3 respectively. Tarsi I and II each with one antiaxial solenidion ω ” (5 – 7, 4 – 5 respectively) and two distal eupathidia p ζ ’ - p ζ ” (7 – 8, 6 – 7; 6 – 8, 6 respectively); ta I – IV u’ - u ” asymmetrically barbed. Femur I with long, barbed dorsal seta (24 – 34). Colour. This species is yellow to orange with small black spots internally (presumably food in gut). Adult male. (6 measured). Dorsum. (Fig. 14) Body measurements: v 2 – h 1 217 – 221, sc 2 – sc 2 90 – 93, c 3 – c 3 107 – 108, f 2 – f 2 55 – 56. Dorsal cuticle mostly smooth. Three dorsal shields evident. Setae c 3, d 3, e 3, f 2 inserted on tubercles. Setae sc 1 not elongate; d 1, h 1 short, barbed; h 2 elongate, barbed with minute distal club; all other dorsal setae thick, dorsoventrally flattened, with strong lateral barbs; dorsal surface finely spiculate, ventral surface smooth. Setal measurements: v 2 40 – 49, sc 1 32 – 47, sc 2 35 – 41, c 1 11 – 20, c 3 28 – 38, d 1 6 – 12, d 3 35 – 41, e 3 36 – 46, f 2 37 – 43, h 1 3 – 8, h 2 163 – 194. Gnathosoma. (Fig. 14) Gnathosoma extends to distal end of femur leg I. Ventral cuticle finely papillate with some transverse plicae anterior to setae 1 b. Ventral setae m absent. Palps with three segments; formula 0, 2, 0 (2); tarsus with two eupathidia (4 – 7; 6 – 8); tibia with two setae (d 12 – 14, v 9 – 10). Cheliceral stylets 85 – 87. Venter. (Fig. 15) Cuticle completely plicate, covered with fine, mostly transverse, plicae; plicae longitudinal between setae 1 b – 1 a, transverse between 1 a – 4 a 2; band of strong plicae and striae between 4 a 1 – ag posterad level with coxae IV; transverse plicae posterior to g 1 – g 2 becoming longitudinal lateral to ps 1 – 2 around anal opening. Setae ps 1 modified to short thick spurs. All setae fine; setae 1 a, 1 b, 4 a 1, 4 a 2 elongate (difficult to consistently determine full length). Setal measurements: 1 a 63 – 109, 1 b 46 – 84, 2 b 19 – 33, 2 c 18 – 24, 3 a 20 – 29, 3 b 19 – 25, 4 a 1 64 – 86, 4 a 2 49 – 69, 4 b 15 – 24, ag 14 – 23, g 1 12 – 24, g 2 11 – 20, ps 1 6 – 9, ps 2 9 – 14. Aedeagus. (Fig. 15) Sclerotised, elongate arrow-head shaped organ (31 – 35) with a narrow membranous tube, widened into funnel-shape distally (11 long, 10 wide), opening into a membranous vesicle. Legs. (Fig. 14) Setal formulae same as female except additions v’ on trI, l’ fe I and ω’ on taI – IV: 1 - 1 - 4 - 0 - 5 - 8 (2), 2 - 0 - 3 - 0 - 5 - 8 (2), 1 - 0 - 2 - 0 - 3 - 4 (1), 1 - 0 - 2 - 0 - 3 - 4 (1) respectively. Tarsi I and II each with two solenidion (ta I paraxial ω ’ 10 – 12, antiaxial ω ” 7 – 9; ta II paraxial ω ’ 8 – 10, antiaxial ω ” 6 – 9) and two distal eupathidia p ζ ’ - p ζ ” (6 – 7, 6 – 7; 5 – 6, 6 – 7 respectively); tarsi III and IV each with one paraxial solenidion ω ’ (9 – 10; 10 – 11 respectively); ta I – IV u’ - u ” asymmetrically barbed. Femur I with long, barbed dorsal seta (59 – 70). Deutonymph. (7 measured) Dorsum. (Figs 16, 17) Body measurements: v 2 – h 1 199 – 231, sc 2 – sc 2 73 – 86, c 3 – c 3 90 – 103, f 2 – f 2 42 – 51. Cuticle mostly smooth with some transverse plicae between setae sc 2 – c 3. Projections on anterior margin of propodosoma absent (Fig. 16) or present (Fig. 17). Central tubercles with setal pairs c 1 and d 1 vestigial (Fig. 16) or developed (Fig. 17). Setae sc 1 elongate, barbed; d 1, h 1 minute; setae h 2 elongate, barbed, with minute club; all other setae thick, dorsoventrally flattened, strongly barbed; dorsal surface finely spiculate, ventral surface smooth. Setal measurements: v 2 15 – 19, sc 1 140 – 172, sc 2 27 – 33, c 1 15 – 23, c 3 24 – 29, d 1 4 – 5, d 3 24 – 29, e 3 21 – 29, f 2 25 – 29, h 1 2 – 4, h 2 118 – 151. Gnathosoma. (Fig. 16) Gnathosoma extends to distal margin genu leg I. Ventral cuticle finely papillate. Ventral setae m absent. Palps with three segments; formula 0, 2, 0 (2); tibia with two setae (d 11 – 12, v 9 – 10); tarsus with two eupathidia (6 – 7, 2 – 4). Cheliceral stylets 84 – 95. Venter. Cuticle completely plicate, covered mostly with transverse striae; plicae longitudinal between setae 1 b – 1 a, transverse 1 a – g 1 with some mixed plicae anterior to 3 a; plicae oblique g 1 – ps 2. All ventral setae fine; setae 1 a, 1 b, 4 a 1, 4 a 2 elongate (difficult to determine full length). Setal measurements: 1 a 68 – 91, 1 b 36 – 46, 2 b 10 – 16, 2 c 12 – 18, 3 a 12 – 18, 3 b 11 – 15, 4 a 1 36 – 60, 4 b 10 – 15, ag 7 – 10, g 1 6 – 10, ps 1 6 – 8, ps 2 8 – 11. Legs. (Fig. 16) Setal formulae same as female, and same as protonymph except additions cx II 2 c, cx IV 4 b: 1 - 0 - 3 - 0 - 5 - 7 (1), 2 - 0 - 3 - 0 - 5 - 7 (1), 1 - 0 - 2 - 0 - 3 - 3, 1 - 0 - 2 - 0 - 3 - 3 respectively. Tarsi I and II each with one antiaxial solenidion ω ” (3 – 4; 3 respectively) and two distal eupathidia p ζ ’ - p ζ ” (4 – 6, 4 – 6; 4 – 6, 4 – 6 respectively); ta I – IV u’ - u ” asymmetrically barbed. Femur I with long, barbed dorsal seta (24 – 32). Protonymph. (4 measured). Dorsum. (Fig. 18) Body measurements: v 2 – h 1 180, sc 2 – sc 2 69, c 3 – c 3 66, f 2 – f 2 29. Cuticle with smooth area anteriorly on prodorsum and posteriorly between setae d 1 – h 1; strong transverse plicae between setae sc 2 – d 1. Dorsum with 2 central seta-bearing tubercles — anterior tubercle with setal pair c 1, posterior tubercle with setal pair d 1. Setae sc 1 elongate, barbed; v 2, e 3 short; setae d 1, h 1 minute; h 2 elongate, barbed, with minute club; all other setae thick, strongly barbed: v 2 9 – 12, sc 1 120 – 127, sc 2 23 – 33, c 1 21 – 28, c 3 23 – 27, d 1 3 – 4, d 3 23 – 28, e 3 5 – 16, f 2 19 – 26, h 1 2 – 3, h 2 76 – 116. Gnathosoma. (Fig. 18) Gnathosoma extends to tibia leg I. Ventral setae m absent. Palps with three segmented; formula 0, 2, 0 (2); tibia with two setae (d 8, v 7); tarsus with two eupathidia (6, 3). Cheliceral stylets 78. Venter. Cuticle completely plicate, mostly covered with transverse striae; plicae longitudinal between setae 1 b – 1 a, transverse 1 a – ag with some mixed plicae anterior to 3 a; plicae oblique laterad ps 1 – 2, mixed between ag – ps 2. Setae fine; setae 1 a, 1 b elongate (difficult to determine full length). Setal measurements: 1 a 31 – 74, 1 b 25 – 47, 2 c 10 – 14, 3 a 11 – 18, 3 b 10 – 12, ag 3 – 9, ps 1 3 – 5, ps 2 3 – 4. Legs. (Fig. 18) Setal formulae for legs I – IV: 1 - 0 - 3 - 0 - 5 - 7 (1), 1 - 0 - 3 - 0 - 5 - 7 (1), 1 - 0 - 2 - 0 - 3 - 3, 0 - 0 - 2 - 0 - 3 - 3 respectively. Tarsi I and II each with one antiaxial solenidion ω ” (2 – 3, 2 – 3 respectively) and two distal eupathidia p ζ ’ - p ζ ” (5, 5; 4 – 5, 4 – 5 respectively); ta I – IV u’ - u ” asymmetrically barbed. Femur I with long, barbed dorsal seta (12). Larva. Dorsum. (Fig. 19) Body measurements: v 2 – h 1 135 – 136, sc 2 – sc 2 57 – 58, c 3 – c 3 66 – 69, e 3 – e 3 28 – 29. Cuticle entirely plicate with mostly transverse plicae, some longitudinal plicae on prodorsum. Dorsum with three rounded tubercles aligned medially - anterior tubercle without setae; setae c 1 inserted together on central tubercle; setae d 1 inserted together on posterior tubercle. Setae sc 1 elongate, barbed; v 2, f 2 short; setae d 1, e 3, h 1 minute; h 2 elongate, barbed; all other setae thick, strongly barbed: v 2 4 – 7, sc 1 53 – 68, sc 2 24 – 34, c 1 38 – 43, c 3 31 – 38, d 1 3 – 4, d 3 32 – 38, e 3 3 – 4, f 2 7 – 9, h 1 1 – 3, h 2 92 – 138. Gnathosoma. (Fig. 19) Gnathosoma extends beyond proximal margin of genu I. Ventral setae m absent. Palps with three segments; formula 0, 2, 0 (1); tibia with two setae (d 9, v 7); tarsus with one eupathidion (6). Cheliceral stylets 68 – 74. Venter. Plicae on cuticle fine, difficult to see, mostly transverse with longitudinal plicae between setae 1 b – 1 a. Setal measurements: 1 a 38 – 77, 1 b 16 – 31, 3 a 8 – 16, ps 1 3 – 4, ps 2 3 – 4. Legs. (Fig. 19) Setal formulae for legs I – IV: 1 - 0 - 3 - 0 - 5 - 6 (1), 0 - 0 - 3 - 0 - 5 - 6 (1), 0 - 0 - 2 - 0 - 3 - 3 respectively. Tarsi I and II each with one short antiaxial solenidion ω ” (2, 2 respectively) and two distal eupathidia p ζ ’ - p ζ ” (4 – 5, 5; 4, 4 respectively); ta I – IV u’ - u ” asymmetrically barbed. Femur I with long, barbed dorsal seta (11). Host. Red-fruited saw sedge, Gahnia sieberiana (Cyperaceae).	en	Beard, Jennifer J., Ochoa, Ronald (2011): New flat mite genera (Acari: Trombidiformes: Tenuipalpidae) associated with Australian sedges (Cyperaceae). Zootaxa 2941: 1-37, DOI: 10.5281/zenodo.204686
0A68182DFF91F87BFF39FAE4ECD3FA5D.taxon	distribution	Distribution. Windsor Tableland and Lamb Range, Far North Queensland.	en	Beard, Jennifer J., Ochoa, Ronald (2011): New flat mite genera (Acari: Trombidiformes: Tenuipalpidae) associated with Australian sedges (Cyperaceae). Zootaxa 2941: 1-37, DOI: 10.5281/zenodo.204686
0A68182DFF91F87BFF39FAE4ECD3FA5D.taxon	discussion	Remarks. Two adult female specimens have abnormally elongate dorsal setae e 3, 41 – 76 compared with 36 – 41. The larval and protonymphal leg chaetotaxy of this species match that of G. gersonus and G. tuberculatus. However the deutonymph differs in that no setae are added to the trochanters (whereas v’ is added to tr I – III in G. gersonus and G. tuberculatus). The leg chaetotaxy of the adult female matches that of the deutonymph, i. e. there are no additions (whereas v’ is added to tr IV in G. gersonus, G. tuberculatus). Like male G. tuberculatus (male G. gersonus not known), v’ is added to tr I and solenidion ω ’ is added to all tarsi of male C. naomae. species name in italics; however this species differs in also adding l’ to fe I. The ventral grooves on the leaf blades of the host plant are shallow. Mites were more commonly found at the tip of the blade than elsewhere on the blade. At the tip of the blade the leaf curls over on itself forming a tube and the longitudinal grooves are slightly wider and appear to be slightly deeper. Mites were more common on outer leaf blades than on the inner unfurled leaves.	en	Beard, Jennifer J., Ochoa, Ronald (2011): New flat mite genera (Acari: Trombidiformes: Tenuipalpidae) associated with Australian sedges (Cyperaceae). Zootaxa 2941: 1-37, DOI: 10.5281/zenodo.204686
0A68182DFF91F87BFF39FAE4ECD3FA5D.taxon	etymology	Etymology. This species is named in loving memory of Naomi Armstrong-Beard. † genua IV nude in original description (Meyer 1979), genua IV with one seta in Mesa et al. (2009).	en	Beard, Jennifer J., Ochoa, Ronald (2011): New flat mite genera (Acari: Trombidiformes: Tenuipalpidae) associated with Australian sedges (Cyperaceae). Zootaxa 2941: 1-37, DOI: 10.5281/zenodo.204686
0A68182DFF9FF878FF39F9E8E92EFC83.taxon	diagnosis	Diagnosis. Adult female (Figs 20 – 21). As per genus, in addition to: setae c 1 and d 1 minute, smooth; dorsal lateral opisthosomal setae broad. Male unknown. Immatures. Unknown. Adult female. (1 measured) Dorsum. (Fig. 20) Body measurements: v 2 – h 1 286 (tip of anterior projection to h 1 324), sc 2 – sc 2 91, c 3 – c 3 108, f 2 – f 2 56. Dorsal cuticle mostly smooth, with few transverse grooves and folds between setae sc 2 and c 1. Dorsal shield or dorsal thickening evident, marked laterally by a change in cuticle. Anterior margin of prodorsum with three projections — a broad, rounded central projection, flanked by two smaller lateral projections each bearing setae v 2. Lateral setae sc 1, c 3, d 3, e 3, f 2 inserted on rounded tubercles. Setae sc 1, h 2 elongate, barbed with minute distal club; c 1, d 1, h 1 minute; all other dorsal setae broad, dorsoventrally flattened, with strong lateral barbs; dorsal surface of setae finely spiculate, ventral surface smooth: v 2 25 – 27, sc 1 242 – 258, sc 2 34 – 35, c 1 4, c 3 32 – 33, d 1 4, d 3 35 – 37, e 3 37, f 2 36, h 1 2 – 3, h 2 173 – 176. Gnathosoma. (Fig. 20) Gnathosoma extends to middle of genu I. Dorsal cuticle with longitudinal plicae; ventral cuticle finely papillate medially; longitudinal plicae between setae 1 b – 1 a. Ventral setae m absent. Palps 3 - segmented; 0, 2, 0 (2); tibia with two setae (d 14, v 10 – 11); tarsus with two eupathidia (8 – 9; 4). Cheliceral stylet length from curve of hook to anterior tip 107. Venter. (Fig. 21) Cuticle completely plicate, covered with fine, transverse plicae between setae 1 a – g 2; some anterior longitudinal plicae between setae 1 b – 1 a; plicae on genital flap transverse, arching anteriorly around setae g 1 – g 2. Setae g 1 inserted in an anterior position to g 2 on genital flap. All setae fine, smooth; setae 1 a, 1 b, 4 a 1, 4 a 2 elongate, fine (difficult to consistently determine full length). Setal measurements: 1 a 56 – 72, 1 b 50, 2 b 14 – 18, 2 c 17 – 20, 3 a 20 – 24, 3 b 13 – 16, 4 a 1 78, 4 a 2 71, 4 b 16 – 18, ag 12 – 17, g 1 14 – 17, g 2 19 – 21, ps 1 9 – 10, ps 2 7 – 8. Spermatheca. (Fig. 21) Cylindrical vesicle (14). Legs. (Fig. 20) Setal formulae for legs I – IV: 1 - 0 - 3 - 0 - 5 - 7 (1), 2 - 0 - 3 - 0 - 5 - 7 (1), 1 - 0 - 2 - 0 - 3 - 3, 1 - 0 - 2 - 0 - 3 - 3 respectively. Tarsi I and II each with one short antiaxial solenidion ω ’ (4, 3 respectively) and two distal eupathidia p ζ ’ - p ζ ’’ (7 – 8, 6 – 7; 7 – 8, 6 – 7 respectively); ta I – IV with u’ - u’ ’ asymmetrically barbed. Colour. This species is orange with small black spots internally (presumed to be food in gut).	en	Beard, Jennifer J., Ochoa, Ronald (2011): New flat mite genera (Acari: Trombidiformes: Tenuipalpidae) associated with Australian sedges (Cyperaceae). Zootaxa 2941: 1-37, DOI: 10.5281/zenodo.204686
0A68182DFF9FF878FF39F9E8E92EFC83.taxon	description	Male. Unknown.	en	Beard, Jennifer J., Ochoa, Ronald (2011): New flat mite genera (Acari: Trombidiformes: Tenuipalpidae) associated with Australian sedges (Cyperaceae). Zootaxa 2941: 1-37, DOI: 10.5281/zenodo.204686
0A68182DFF9FF878FF39F9E8E92EFC83.taxon	discussion	Remarks. As only one specimen is known for this species, the true range and variability of measurements remains unknown. The tip of the spermathecal apparatus is truncate and presumed to be damaged.	en	Beard, Jennifer J., Ochoa, Ronald (2011): New flat mite genera (Acari: Trombidiformes: Tenuipalpidae) associated with Australian sedges (Cyperaceae). Zootaxa 2941: 1-37, DOI: 10.5281/zenodo.204686
0A68182DFF9FF878FF39F9E8E92EFC83.taxon	etymology	Etymology. The masculine Latin word “ foliatus ” means “ leafy ” and refers to the leaf-like setae on the opisthosoma of this species.	en	Beard, Jennifer J., Ochoa, Ronald (2011): New flat mite genera (Acari: Trombidiformes: Tenuipalpidae) associated with Australian sedges (Cyperaceae). Zootaxa 2941: 1-37, DOI: 10.5281/zenodo.204686
