taxonID	type	description	language	source
0B13879BFFFBFFB7A9D8FBC1C96CFB4B.taxon	type_taxon	Type species. Homalenchodes jarzembowskii sp. nov.	en	LI, YAN-DA, HSIAO, YUN, YOSHITOMI, HIROYUKI, HUANG, DI-YING, CAI, CHEN-YANG (2022): Homalenchodes, a new genus of Serropalpini from mid-Cretaceous amber of northern Myanmar (Coleoptera: Tenebrionoidea: Melandryidae). Palaeoentomology 5 (3): 246-253, DOI: 10.11646/palaeoentomology.5.3.7, URL: http://dx.doi.org/10.11646/palaeoentomology.5.3.7
0B13879BFFFBFFB7A9D8FBC1C96CFB4B.taxon	etymology	Etymology. The generic name is a combination of the Greek “ homalós ”, smooth, referring to the absence of longitudinal striae on its elytra, and “ Enchodes ”, a presumably closely related genus. The name is feminine in gender.	en	LI, YAN-DA, HSIAO, YUN, YOSHITOMI, HIROYUKI, HUANG, DI-YING, CAI, CHEN-YANG (2022): Homalenchodes, a new genus of Serropalpini from mid-Cretaceous amber of northern Myanmar (Coleoptera: Tenebrionoidea: Melandryidae). Palaeoentomology 5 (3): 246-253, DOI: 10.11646/palaeoentomology.5.3.7, URL: http://dx.doi.org/10.11646/palaeoentomology.5.3.7
0B13879BFFFBFFB7A9D8FBC1C96CFB4B.taxon	discussion	The generic name Enchodes LeConte has been sometimes mistakenly treated as masculine (e. g., Jung, 2019). According to Article 30.1.4.4 of ICZN (1999), “ a compound genus-group name ending in the suffix - ites, - oides, - ides, - odes, or - istes is to be treated as masculine unless its author, when establishing the name, … treated it as such by combining it with an adjectival species-group name in another gender form ”. When establishing genus Enchodes, LeConte (1866) added a single previously described species, Dircaea sericea Haldeman, citing it as Enchodes sericea (Haldeman). Thus, the name should be feminine in gender.	en	LI, YAN-DA, HSIAO, YUN, YOSHITOMI, HIROYUKI, HUANG, DI-YING, CAI, CHEN-YANG (2022): Homalenchodes, a new genus of Serropalpini from mid-Cretaceous amber of northern Myanmar (Coleoptera: Tenebrionoidea: Melandryidae). Palaeoentomology 5 (3): 246-253, DOI: 10.11646/palaeoentomology.5.3.7, URL: http://dx.doi.org/10.11646/palaeoentomology.5.3.7
0B13879BFFFBFFB7A9D8FBC1C96CFB4B.taxon	diagnosis	Diagnosis. Body relatively small, strongly elongate. Compound eyes coarsely facetted (Fig. 2 A, B). Antennae long, filiform. Maxillary palps simple (Fig. 2 A). Pronotum widest posteriorly, with somewhat produced posterior pronotal angles (Fig. 2 B). Prosternal process complete, gradually narrowed posteriorly, slightly extending beyond posterior procoxal margin (Fig. 2 A). Procoxae narrowly separated (Fig. 2 A). Protrochantins well exposed (Fig. 2 A). Elytra without longitudinal striae (Fig. 2 E). Meso- and metatibiae without transverse rows of bristles or spines (Fig. 2 D). Metatibial spurs normal-sized (distinctly shorter than half length of metatarsomere 1) (Fig. 2 D). Penultimate tarsomeres ventrally lobed in all three pairs of legs (Figs 2 C, 3).	en	LI, YAN-DA, HSIAO, YUN, YOSHITOMI, HIROYUKI, HUANG, DI-YING, CAI, CHEN-YANG (2022): Homalenchodes, a new genus of Serropalpini from mid-Cretaceous amber of northern Myanmar (Coleoptera: Tenebrionoidea: Melandryidae). Palaeoentomology 5 (3): 246-253, DOI: 10.11646/palaeoentomology.5.3.7, URL: http://dx.doi.org/10.11646/palaeoentomology.5.3.7
0B13879BFFFBFFB7A9D8FBC1C96CFB4B.taxon	discussion	Remarks. Homalenchodes shares with Synchroidae a similar elongate habitus, lateral and oval eyes, and transverse head and pronotum. In fact, the adults of Synchroidae have an appearance similar to Melandryidae, but these two families are otherwise readily separable based on larval characters (Crowson, 1966). Nevertheless, synchroids are generally characterised by the long interfacetal setae (absent in Mallodrya Horn, although this genus might have been inappropriately placed in Synchroidae and needs to be further evaluated; Hsiao et al., 2018), unmargined posterior edge of pronotum (distinctly margined in Mallodrya), prosternal process markedly extending beyond procoxae with long oval apex (prosternal process laminate, ending just before the posterior edges of procoxae in Mallodrya), and unlobed penultimate tarsomeres (Ślipiński & Lawrence, 2010; Hsiao et al., 2018), while Homalenchodes has no interfacetal setae, a margined posterior edge of pronotum, a posteriorly gradually narrowed prosternal process with acute apex, slightly extending beyond posterior edges of procoxae, and lobed penultimate tarsomeres. Therefore, here we believe that it is appropriate to place Homalenchodes in the (currently non-monophyletic) family Melandryidae.	en	LI, YAN-DA, HSIAO, YUN, YOSHITOMI, HIROYUKI, HUANG, DI-YING, CAI, CHEN-YANG (2022): Homalenchodes, a new genus of Serropalpini from mid-Cretaceous amber of northern Myanmar (Coleoptera: Tenebrionoidea: Melandryidae). Palaeoentomology 5 (3): 246-253, DOI: 10.11646/palaeoentomology.5.3.7, URL: http://dx.doi.org/10.11646/palaeoentomology.5.3.7
0B13879BFFFBFFB4AA7AFB2DC99AF7A2.taxon	description	(Figs 1 – 3)	en	LI, YAN-DA, HSIAO, YUN, YOSHITOMI, HIROYUKI, HUANG, DI-YING, CAI, CHEN-YANG (2022): Homalenchodes, a new genus of Serropalpini from mid-Cretaceous amber of northern Myanmar (Coleoptera: Tenebrionoidea: Melandryidae). Palaeoentomology 5 (3): 246-253, DOI: 10.11646/palaeoentomology.5.3.7, URL: http://dx.doi.org/10.11646/palaeoentomology.5.3.7
0B13879BFFFBFFB4AA7AFB2DC99AF7A2.taxon	materials_examined	Material. Holotype, NIGP 180475.	en	LI, YAN-DA, HSIAO, YUN, YOSHITOMI, HIROYUKI, HUANG, DI-YING, CAI, CHEN-YANG (2022): Homalenchodes, a new genus of Serropalpini from mid-Cretaceous amber of northern Myanmar (Coleoptera: Tenebrionoidea: Melandryidae). Palaeoentomology 5 (3): 246-253, DOI: 10.11646/palaeoentomology.5.3.7, URL: http://dx.doi.org/10.11646/palaeoentomology.5.3.7
0B13879BFFFBFFB4AA7AFB2DC99AF7A2.taxon	etymology	Etymology. The species is named in honor of the palaeoentomologist Dr Edmund A. Jarzembowski.	en	LI, YAN-DA, HSIAO, YUN, YOSHITOMI, HIROYUKI, HUANG, DI-YING, CAI, CHEN-YANG (2022): Homalenchodes, a new genus of Serropalpini from mid-Cretaceous amber of northern Myanmar (Coleoptera: Tenebrionoidea: Melandryidae). Palaeoentomology 5 (3): 246-253, DOI: 10.11646/palaeoentomology.5.3.7, URL: http://dx.doi.org/10.11646/palaeoentomology.5.3.7
0B13879BFFFBFFB4AA7AFB2DC99AF7A2.taxon	diagnosis	Diagnosis. As for the genus. Locality and horizon. Amber mine located near Noije Bum Village, Tanai Township, Myitkyina District, Kachin State, Myanmar; unnamed horizon, midCretaceous, Upper Albian to Lower Cenomanian.	en	LI, YAN-DA, HSIAO, YUN, YOSHITOMI, HIROYUKI, HUANG, DI-YING, CAI, CHEN-YANG (2022): Homalenchodes, a new genus of Serropalpini from mid-Cretaceous amber of northern Myanmar (Coleoptera: Tenebrionoidea: Melandryidae). Palaeoentomology 5 (3): 246-253, DOI: 10.11646/palaeoentomology.5.3.7, URL: http://dx.doi.org/10.11646/palaeoentomology.5.3.7
0B13879BFFFBFFB4AA7AFB2DC99AF7A2.taxon	description	Description. Body strongly elongate, about 2.7 mm long, 0.6 mm wide. Surface finely punctate and with fine hairs. Head well exposed from above. Compound eyes lateral, large, emarginate in front of antennal insertions, coarsely facetted on the surface, without interfacetal setae. Antennal insertions fully concealed from above, widely separated. Subantennal grooves absent. Antennae 11 - segmented, long, filiform; antennomere 1 elongate, almost parallel-sided; antennomere 2 short, subquadrate; antennomeres 3 – 11 elongate, subequal. Clypeus with anterior margin convex. Mandibles bilobed apically. Maxillary palps 4 - segmented, simple, not clearly serrate; palpomere 1 small; palpomere 4 possibly securiform. Mentum subtrapezoidal. Submentum relatively smooth, with hairs somewhat denser than other regions. Pronotum widest posteriorly, with slightly produced posterior pronotal angles; disc without depression medially; posterior edge margined. Notosternal sutures distinct. Prosternum about as long as longitudinal procoxal diameter; prosternal process complete, gradually narrowed posteriorly, slightly extending beyond posterior procoxal margin, apically acute. Procoxal cavities narrowly separated, externally broadly open. Protrochantins well exposed. Scutellar shield transverse, subtrapezoidal, widest posteriorly; posterior edge slightly rounded, without notch. Elytra long, about 3.1 times as long as width combined, at base almost as wide as pronotum, apically separately rounded; surface without longitudinal striae. Mesocoxal cavities narrowly separated. Metacoxae almost contiguous; coxal plates absent. Legs slender. Protrochanter small, obliquely attached to profemur, with profemur abutting procoxa. All tibiae with 2 spurs, without transverse rows of bristles or spines. Metatibial spurs normal-sized. Tarsi 5 - 5 - 4; penultimate tarsomeres ventrally lobed in all three pairs of legs; metatarsomere 1 long, distinctly longer than metatarsomeres 2 – 4 combined. Pretarsal claws simple. Abdomen with five ventrites. Ratio of ventrite lengths along middle: 3.2: 2.4: 2.1: 1.7: 2.4. Ventrite 5 apically rounded, without emargination.	en	LI, YAN-DA, HSIAO, YUN, YOSHITOMI, HIROYUKI, HUANG, DI-YING, CAI, CHEN-YANG (2022): Homalenchodes, a new genus of Serropalpini from mid-Cretaceous amber of northern Myanmar (Coleoptera: Tenebrionoidea: Melandryidae). Palaeoentomology 5 (3): 246-253, DOI: 10.11646/palaeoentomology.5.3.7, URL: http://dx.doi.org/10.11646/palaeoentomology.5.3.7
