identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
D04C9A0EEA7AB96B43FE98D8E249EB19.text	D04C9A0EEA7AB96B43FE98D8E249EB19.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Swezeyana atra	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Swezeyana atra sp. n. Figures 1C, 4 </p>
            <p>Diagnosis.</p>
            <p>Medium sized, dark coloured species, with fore wing membrane unpatterned, antennae medium short, genal processes long, paramere short, and female proctiger strongly convex apically.</p>
            <p>Description.</p>
            <p> Adult. General body colour brown to black, particularly dark on the dorsum of head and thorax. Fore wing membrane generally uniformly clear or slightly fuscous, with fuscous-brown cloud usually present around vein R and basal dorsal claval margin (Fig. 1C). Fore wing apex acute; pseudopterostigma relatively long (Fig. 4S), typically no cross pseudovein in cell r1 (one individual seen with partial cross vein); surface spinules relatively sparsely distributed but denser towards wing margin, few or absent from c+sc; medium long setae on ventral margin and medium short to short setae on veins and dorsal margin. Antennae medium short (ratio AL:HW 1.30) (Fig. 4  B–C ); genal processes medium long (GP &lt;0.40 mm, ratio HW:GP&gt; 1.40) and slightly upturned at apices (Fig. 4B); medium short to short setae on vertex and thorax. Meracanthus extremely small (Fig. 4Q), genual spine reduced or absent (Fig. 4R). Male terminalia (Fig. 4  F–I ): paramere short (ratio PL:HW &lt;0.20), tapering to slightly anteriorly projecting apex with two short stout setae; distal aedeagus segment long relative to paramere (ratio PL:AEL 0.92-1.10), apex developed into a large rounded hook with blunt apex. Female terminalia (Fig. 4  J–P ): proctiger dorsal surface strongly convex apically, apex broad, blunt, bearing medial cleft and fringed with stout setae, anal ring medium long (ratio FP:RL&gt; 2.00), without or with reduced head compartment at proximal end, distal portion of ring margin smooth; subgenital plate slightly convex with short medial cleft (Fig. 4  N–O ). </p>
            <p>Egg. Unknown.</p>
            <p>Immature. Unknown.</p>
            <p>Host plant.</p>
            <p> Planchonella sandwicensis . </p>
            <p>Distribution.</p>
            <p>Oahu. Only known from the Waianae Mountains.</p>
            <p>Etymology.</p>
            <p>Named for the dark body colouration, especially the head and thorax (adjective in the nominative singular).</p>
            <p>Comments.</p>
            <p> This is the darkest of the  Swezeyana species. It has the shortest paramere in the  elongagena group, with the shape more similar to those in the  reticulata group. The distribution in the Waianae Mountains is shared with  S. oahuensis , and the molecular topology places these two species as sister taxa but with weak bootstrap support (Fig. 2). </p>
            <p>Material examined.</p>
            <p> Holotype male (slide mounted), Waianae Mnts, Oahu, USA, N21.4585, W-158.0973, ex  Planchonella sandwicensis , 5 July 2014,  “Hi65-14” D. Percy leg. (BMNH). Paratype (slide mounted) 1f, as for holotype (BMNH). Other material: 1m 1f, Puu Hapapa, Central Waianae Mnts, Oahu, USA, N21.4666, W-158.1029, ex  Planchonella sandwicensis , 17 May 2014,  “KM14-14” K. Magnacca leg. (BMNH). </p>
            <p>Gene sequences.</p>
            <p>MH001521 (COI) MH001527 (cytB) (Hi65-14); MH001522 (COI) MH001528 (cytB) (KM14-14).</p>
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	https://treatment.plazi.org/id/D04C9A0EEA7AB96B43FE98D8E249EB19	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Percy, Diana M.	Percy, Diana M. (2018): Revision of the Hawaiian psyllid genus Swezeyana, with descriptions of seven new species (Hemiptera, Psylloidea, Triozidae). ZooKeys 758: 75-113, DOI: http://dx.doi.org/10.3897/zookeys.758.23019, URL: http://dx.doi.org/10.3897/zookeys.758.23019
A93D10143B025A8463B09E04710AA987.text	A93D10143B025A8463B09E04710AA987.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Swezeyana Caldwell 1940	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Swezeyana Caldwell, 1940</p>
            <p> Swezeyana Caldwell, 1940: 389. Type species:  Swezeyana elongagena Caldwell, 1940, by original designation. </p>
            <p>Description.</p>
            <p>Adult. General colour variable ranging from pale yellow-brown, to green or yellow-green, to almost black; often with pink or reddish highlights on the fore wing as well as on the body, especially genal processes, legs, and abdomen. Fore wing membrane either with distinct darker patches or clouds of pigmentation, these range from dark brown to red, and in some cases are limited to termination of veins at wing margins and around cross veins between Rs and wing margin, if without distinct patterns of pigmentation, appearing uniformly clear, opaque yellow or fuscous; wing veins pale to red or dark brown, cross veins between Rs and ventral wing margin with or without pigmentation. Adult length including fore wing from 2-5 mm. Fore wing elongate and usually narrow (ratio WL:WW&gt; 2.80, often&gt; 3), acute to bluntly acute apically, either with trifurcation of veins R, M and Cu1, or with vein R branching anterior of bifurcation of M and Cu1; vein Rs long, reaching wing margin distad of M fork, but either with or without complete extension of Rs to wing margin, incomplete termination of Rs usually marked by pigmentation; vein R shorter than Cu1 and terminating at base of Rs, a pseudopterostigma is present between base of Rs and wing margin, and a more or less thickened wing margin (C+Sc) is present from the wing base to the pseudopterostigma, in some cases occupying part or entire area of cell c+sc; with or without one or more partial or complete cross veins traversing cell r1 between vein Rs and ventral wing margin; a single, broadly shaped marginal cluster of radular spines (Figs 3O, 6J) in cells cu1, m1, and m2; surface spinules either present in all cells, dense or sparsely distributed, or few to absent from c+sc, r2 and r1, often relatively sparsely distributed but becoming denser towards wing margin. Hind wing narrow and elongate (&gt; 0.5 length of fore wing), clear or slightly fuscous in basal half. Head moderately deflexed downwards, vertex more or less flat dorsally, with lateral ocelli lying on small tubercles, medial epicranial suture distinct; genal processes extremely long, often upturned at apices, with scattered long setae and usually a single, long subapical seta on each process. Antennae short; antennal segments 10, either entirely dark, or more usually with terminal 3(-7) segments darker, or distal part of segments 3-8 darker; a single rhinarium apically on each of segments 4, 6, 8, 9; 1-2 long setae on each of segments 3-9, terminal segment with two unequal length apical setae. Distal proboscis segment short, darker apically. Thorax somewhat flattened to only moderately arched; vertex and thorax with scattered short to moderately long setae. Legs short, hind legs robust with femur longer than tibia; hind leg with meracanthus reduced to almost absent; metafemur with several stout setae apically; metatibia with or without distinct genual spine basally and typically with 1+2 (occasionally 1+3) sclerotized apical spurs; pro- and mesotarsi subequal in length, metatarsi unequal with extremely long basal tarsus slightly expanded with concave, ridged underside (Fig. 3M). Male terminalia with somewhat elongate subgenital plate; proctiger with pronounced posterior lobes medially, 1-2 long setae usually present on posterior apices of each lobe, length shorter, subequal or longer than paramere; paramere shape variable, generally broad basally and tapering to apex, with two stout setae on the interior apex (sometimes appearing as one from lateral view); distal aedeagus segment apex hooked. Female terminalia with medium to long dorsal and ventral setae; proctiger either truncate and markedly convex apically, with apex broad, blunt, bearing small medial cleft and fringed with stout setae, or dorsal surface more or less straight, apex tapering, lacking medial cleft and distinct fringe of setae; proctiger longer than subgenital plate; anal ring hour-glass shaped (with or without a head compartment at proximal end) and composed of a, usually, uninterrupted, double row of cells, posterior/distal portion of ring margin either smooth or convoluted; subgenital plate ventral surface either convex or more or less straight, apex terminating in a variably shaped beak often bearing a short or more pronounced medial cleft spanned by a short or extended membrane; ovipositor valves small, without serrations (Fig. 9O).</p>
            <p>Egg. Known for four species. Pale or light brown, oblong-ovoid with a short, laterally positioned pedicel sub-basally on underside; distinctly hexagonal, honeycomb-like, sculpturing, to semi-hexagonal or rounded indentations dorsally; underside unsculptured, tail apparently lacking.</p>
            <p> Immature. Known for four species. 5th instar oblong-ovoid, ventro-dorsally flattened with slightly protruding wing buds and distinct humeral lobes; antennae with 3(-4) segments bearing 3(-4) rhinaria (1 on segments 2-3, and 2 on apical segment) and two long, terminal, simple setae of unequal length; tarsi with broad crescent arolia and extremely small, reduced claws; each terminal tarsus bearing a long capitate seta; anus situated ventrally, circumanal ring broad and composed of a single row of elongate cells; dorsum either with wax producing pores (see Tuthill 1966), or non-wax producing tubercles and tentacles (Fig. 12). Chaetotaxy: 5th instar with either continuous or interrupted coverage of marginal setae; overall setal types, even between closely related species (e.g.,  S. reticulata and  S. tentaculata ), highly variable (Fig. 12). Smaller instars only known for  S. reticulata and  S. tentaculata , in which tubercles are apparent from 2nd instar (Fig. 13). </p>
            <p>Biology.</p>
            <p>All species for which the biology is known have free-living immatures on the surface of leaves (either lower, or both upper and lower surfaces). Those species with immatures described here with protruding tubercles and tentacles were mostly found on the lower leaf surface among dense indumentum and often close to the mid-rib (Fig. 12N).</p>
            <p>Host plant.</p>
            <p> All  Swezeyana are host specific on a single Hawaiian endemic host plant species,  Planchonella sandwicensis (  Sapotaceae ). </p>
            <p>Comments.</p>
            <p> Two species groups are recognized,  elongagena group and  reticulata group, based primarily on the strikingly different forms of female terminalia. The  elongagena group has broad, truncate female terminalia with a strongly convex proctiger apex; the proctiger and subgenital plate bear a small to pronounced medial cleft at the apex. In contrast, the  reticulata group has tapering terminalia without a medial cleft in the proctiger apex. In both species groups the subgenital plate terminates in a more or less well developed beak with small to pronounced cleft spanned by a membrane. The underlying endoskeleton of the two different forms of female terminalia indicate distinctly different development of the apodemes in the two species groups: broad and short in  elongagena group (Fig. 4P), and long and narrow in  reticulata group (Fig. 10I). However, not all species treated here are known for both sexes, therefore current assignment to these groups relies on other characteristics and DNA barcode data (see Discussion). Due to the unknown female morphology and inconclusive placement in the molecular phylogeny,  S. magnaccai is not placed within a species group. Fore wing characteristics such as wing membrane colouration and pseudoveins/cross veins are found in both groups. The 5th instar immatures may also be diagnostic, with  elongagena group having wax producing pores dorsally (illustrated in Tuthill 1966 for  S. elongagena ) and continuous ring of marginal setae (illustrated in Caldwell 1940 for  S. elongagena ); while in contrast, the  reticulata group are characterized by non-wax producing tubercles and tentacles and lack a contiguous marginal ring of setae (Figs 12-13). However, currently immatures are known for these three species only, so it remains to be tested whether these highly distinct immature morphologies reflect species group assignments. </p>
            <p>Note on adult assignment to species group.</p>
            <p> The two species groups (  elongagena group and  reticulata group) are most easily recognized by the shape of the female terminalia, e.g., extremely convex apex of female proctiger in the  elongagena group, versus more or less dorsally straight and tapering in the  reticulata group. The  elongagena group females have a FP:HW ratio typically &lt;0.70 (range 0.51-0.69), and FP:RL ratio typically &lt;2.18 (range 1.41-2.19); whereas in the  reticulata group FP:HW ratio is typically&gt; 0.70 (range 0.69-0.95), and FP:RL ratio is typically&gt; 2.18 (range 2.17-2.59).  Swezeyana males are less easily assigned to a species group, but  elongagena group males have a distal aedeagus segment that is typically shorter than the paramere (PL:AEL ratio range 0.92-1.60), whereas  reticulata group males have a distal aedeagus segment that is longer than the paramere (PL:AEL ratio range 0.65-0.88). Notably, the fore wing characters used by Zimmerman (1948) to key out the two species described by Caldwell (  S. elongagena and  S. reticulata ), such as presence/absence of cross veins in cell r1 and presence/absence of distinct patterns of pigmentation are found in both species groups. The key below does not key to species group, rather it employs characters useful in distinguishing species, in particular those co-occurring on the same island. </p>
            <p>Note on molecular analyses.</p>
            <p> The neighbour-joining analysis of two mitochondrial DNA regions is presented in Fig. 2. Strong support is recovered for the  reticulata group, but not for the  elongagena group. The topologies recovered in comparative ML and Bayesian analyses differ only at weakly or unsupported nodes. The three nodes that group same island sister taxon pairs (on Kauai, Oahu, and Hawaii) are recovered in all analyses but with variable support (much stronger support in the Bayesian than NJ or ML analyses) (see Fig. 2 and Discussion). Maximum genetic divergence (uncorrected p-distances) among  Swezeyana species is 19.9%; maximum intraspecific divergence (3%) was found in  S. magnaccai on Oahu. </p>
            <p> Key to  Swezeyana adults </p>
            <p>Note on species descriptions.</p>
            <p> Swezeyana is a small genus with, in general, considerable morphological homogeneity. The species descriptions below provide details of species specific characteristics not supplied in the generic description above. </p>
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	https://treatment.plazi.org/id/A93D10143B025A8463B09E04710AA987	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Percy, Diana M.	Percy, Diana M. (2018): Revision of the Hawaiian psyllid genus Swezeyana, with descriptions of seven new species (Hemiptera, Psylloidea, Triozidae). ZooKeys 758: 75-113, DOI: http://dx.doi.org/10.3897/zookeys.758.23019, URL: http://dx.doi.org/10.3897/zookeys.758.23019
50EA38EF183A033381E7E0D76EA7FC74.text	50EA38EF183A033381E7E0D76EA7FC74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Swezeyana elongagena Caldwell 1940	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Swezeyana elongagena Caldwell, 1940 Figures 1A, 3 </p>
            <p> Swezeyana elongagena Caldwell, 1940: 390. </p>
            <p>Description.</p>
            <p> Adult. General body colour green to yellow-green or yellow-brown, last 3-5 antennal segments darker brown to black, apices of genae sometimes pinkish-red. Fore wing membrane uniformly pale fuscous (Fig. 1A). Fore wing apex acute to bluntly acute; pseudopterostigma relatively short to medium long (Fig. 3P), none to few (1-3) cross pseudoveins in cell r1; surface spinules sparsely distributed, apparently absent from c+sc (Fig. 3O); medium long setae on ventral margin and medium short to short setae on veins and dorsal margin. Antennae medium long (ratio AL:HW 1.54-1.78) (Fig. 3D); genal processes long (GP&gt; 0.40 mm, ratio HW:GP &lt;1.35) and often upturned at apices (Fig. 3  A–B ); short to medium short setae on vertex and thorax. Meracanthus small (Fig. 3N), genual spine reduced (Fig. 3L). Male terminalia (Fig. 3  G–I ): paramere slender, long (ratio PL:HW&gt; 0.27), tapering to anteriorly projecting apex with two short stout setae; distal aedeagus segment short relative to paramere (ratio PL:AEL 1.46-1.56), apex developed into a large rounded hook with blunt apex. Female terminalia (Fig. 3  E–F ): proctiger dorsal surface strongly convex apically, apex broad, blunt, bearing medial cleft and fringed with stout setae, anal ring long (ratio FP:RL 1.41-1.95), with reduced head compartment at proximal end, distal portion of ring margin smooth; subgenital plate convex with medial cleft pronounced (almost half length of subgenital plate). </p>
            <p>Egg. Pale with well-defined hexagonal (honeycomb-like) sculpturing dorsally (Fig. 3J).</p>
            <p> Immature. Described and illustrated by Caldwell (1940) and Tuthill (1966). Although Caldwell refers to marginal  “sectasetae” in his description, he illustrates distinctly fan-shaped and apparently unbisected setae. Tuthill provides an image of an immature with long "wax filaments" produced by dorsal pores, and notes that free-living immatures were observed mostly on the upper, but also lower, leaf surfaces. Both descriptions suggest notable morphological differences to immatures described here for  S. reticulata and  S. tentaculata sp. n. </p>
            <p>Host plant.</p>
            <p> Planchonella sandwicensis . </p>
            <p>Distribution.</p>
            <p> Maui, Molokai (a single female specimen recorded from Kauai, and apparently now missing, is queried in Zimmerman (1948), and may have been  S. magna sp. n.). Specimens from Molokai are close to the type specimens from Maui, and currently this species is considered to be restricted to these two islands. </p>
            <p>Comments.</p>
            <p> Although  Caldwell’s (1940) illustration only indicates a single cross pseudovein, examination of the type specimen revealed three unpigmented pseudoveins. A single female paratype collected from Kalalau Trail, Kauai, is likely to be  S. magna sp. n. (see also assignment in error comment in Zimmerman 1948) </p>
            <p>Material examined.</p>
            <p> Holotype female (slide mounted), Haelaau, Maui, USA, ex  Planchonella sp., 19 December 1928 (BPBM). Paratypes: 10m 5f, same data as holotype (not located). Other material: 5m 4f, Kamakou Preserve, Molokai, USA, N21.1236, W-156.9108, ex  Planchonella sandwicensis , 17 August 2003,  “Hi20-03” D. Percy leg. (BMNH). </p>
            <p>Gene sequences.</p>
            <p>MG988832 (COI) MG989153 (cytB) (Hi20-03).</p>
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	https://treatment.plazi.org/id/50EA38EF183A033381E7E0D76EA7FC74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Percy, Diana M.	Percy, Diana M. (2018): Revision of the Hawaiian psyllid genus Swezeyana, with descriptions of seven new species (Hemiptera, Psylloidea, Triozidae). ZooKeys 758: 75-113, DOI: http://dx.doi.org/10.3897/zookeys.758.23019, URL: http://dx.doi.org/10.3897/zookeys.758.23019
D9F77CF138FC99E914B8C7F21FB4798A.text	D9F77CF138FC99E914B8C7F21FB4798A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Swezeyana hawaiiensis	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Swezeyana hawaiiensis sp. n. Figures 1E, 5 </p>
            <p>Diagnosis.</p>
            <p>Medium sized, light coloured species, with fore wing membrane unpatterned, antennae and genal processes relatively short, and female proctiger strongly convex apically.</p>
            <p>Description.</p>
            <p> Adult. General body colour green to yellow-green. Fore wing membrane generally slightly fuscous, darker fuscous clouds around cross pseudoveins towards apex of cell r1, and distinct brown patches at termination of veins Cu1a, Cu1b, and M3+4 (Figs 1E, 5K). Fore wing apex acute to bluntly acute; pseudopterostigma short (Fig. 5K), 2-3 cross pseudoveins present in apical portion of cell r1; surface spinules sparsely distributed and reduced in distribution, few or absent from r1, r2, and c+sc; medium long setae on ventral margin and medium short to short setae on veins and dorsal margin. Antennae short (ratio AL:HW 0.95-1.00), terminal 3 segments darker brown (Figs 5  A–B ); genal processes relatively short (GP &lt;0.35 mm, ratio HW:GP&gt; 1.70), not or only slightly upturned at apices (Fig. 5A); short to minute setae on vertex and thorax. Meracanthus small (Fig. 5H), genual spine reduced or absent (Fig. 5I). Female terminalia (Fig. 5  D–F ): proctiger dorsal surface strongly convex apically, apex broad, blunt, bearing medial cleft and fringed with stout setae (Fig. 5E), anal ring long (ratio FP:RL 1.82), with head compartment at proximal end, distal portion of ring margin smooth; subgenital plate slightly convex with short medial cleft with beak and membrane extended (Fig. 5F). </p>
            <p>Egg. Unknown.</p>
            <p>Immature. Free-living immatures were observed mostly on the upper, but also on the lower, leaf surfaces. Specimens collected were unfortunately lost during specimen shipping.</p>
            <p>Host plant.</p>
            <p> Planchonella sandwicensis . </p>
            <p>Distribution.</p>
            <p>Hawaii. Only known from PuuWaaWaa area.</p>
            <p>Etymology.</p>
            <p>Named for its distribution on the island of Hawaii (adjective in the nominative singular).</p>
            <p>Comments.</p>
            <p> This species was collected from the same individual host tree as  S. rubra ; both species are in the  elongagena group but are easily separated in the field due to general body colour and a distinctly patterned fore wing in  S. rubra ; both species have comparatively short genal processes as well as the shortest antennae in the genus (subequal to head width). The molecular topology places these as sister taxa, but without bootstrap support. Males with the same collection data as females were unfortunately lost during specimen shipping. </p>
            <p>Material examined.</p>
            <p> Holotype female (slide mounted), PuuWaaWaa, Hawaii, USA, N19.784, W-155.833, 820m, ex  Planchonella sandwicensis , 29 July 2002,  “440A-02” D. Percy leg. (BMNH). Paratype (slide mounted) 1f, as for holotype (BMNH). </p>
            <p>Gene sequences.</p>
            <p>MG988835 (COI) MG989155 (cytB) (440A-02).</p>
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	https://treatment.plazi.org/id/D9F77CF138FC99E914B8C7F21FB4798A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Percy, Diana M.	Percy, Diana M. (2018): Revision of the Hawaiian psyllid genus Swezeyana, with descriptions of seven new species (Hemiptera, Psylloidea, Triozidae). ZooKeys 758: 75-113, DOI: http://dx.doi.org/10.3897/zookeys.758.23019, URL: http://dx.doi.org/10.3897/zookeys.758.23019
96DCAE7FE733D3A1A29BBA09DC3CCD14.text	96DCAE7FE733D3A1A29BBA09DC3CCD14.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Swezeyana magna	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Swezeyana magna sp. n. Figures 1D, 6 </p>
            <p>Diagnosis.</p>
            <p>Large, light coloured species, with fore wing membrane unpatterned, antennae and genal processes long, and paramere short.</p>
            <p>Description.</p>
            <p>Adult. General body colour yellow-green to yellow-brown, last 7-8 antennal segments darker brown. Fore wing membrane uniformly pale fuscous (Fig. 1D). Fore wing apex acute; pseudopterostigma medium long (Fig. 6I), 0-1 cross pseudoveins in basal portion of cell r1; surface spinules sparsely distributed, apparently absent from c+sc; medium long setae on ventral margin and medium short setae on veins and dorsal margin. Antennae long (AL&gt; 1 mm, ratio AL:HW 1.61) (Fig. 6A); genal processes long (GP&gt; 0.50 mm, ratio HW:GP &lt;1.40) and slightly upturned at apices (Fig. 6A); short to minute setae on vertex and thorax. Meracanthus small (Fig. 6D), genual spine reduced (Fig. 6C). Male terminalia (Fig. 6H): paramere short (ratio PL:HW 0.24), tapering to small, anteriorly projecting apex with two short stout setae; distal aedeagus segment long relative to paramere (ratio PL:AEL 1.14), apex developed into a large rounded hook with bluntly acute apex.</p>
            <p>Egg. Unknown.</p>
            <p>Immature. Unknown.</p>
            <p>Host plant.</p>
            <p> Planchonella sandwicensis . </p>
            <p>Distribution.</p>
            <p>Kauai. Only known from one location in Kokee State Park.</p>
            <p>Etymology.</p>
            <p>Named for the large body size (adjective in the nominative singular).</p>
            <p>Comments.</p>
            <p> This is the largest  Swezeyana species and is only known from a single male; fore wing type suggests it is part of the  elongagena species group, but the paramere shape is somewhat similar to other  reticulata group species on Kauai. As no molecular sequences are available for this species, the group affiliation remains uncertain, but this species and the other Kauai species may represent early divergence of the  elongagena and  reticulata species groups. </p>
            <p>Material examined.</p>
            <p> Holotype male (slide mounted), Kokee State Park, Kauai, USA, N22.1444, W-159.6477, ex  Planchonella sandwicensis , 29 October 2005,  “Hi01-05” D. Percy leg. (BMNH). </p>
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	https://treatment.plazi.org/id/96DCAE7FE733D3A1A29BBA09DC3CCD14	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Percy, Diana M.	Percy, Diana M. (2018): Revision of the Hawaiian psyllid genus Swezeyana, with descriptions of seven new species (Hemiptera, Psylloidea, Triozidae). ZooKeys 758: 75-113, DOI: http://dx.doi.org/10.3897/zookeys.758.23019, URL: http://dx.doi.org/10.3897/zookeys.758.23019
35B0E962FEC248592282F919723FB631.text	35B0E962FEC248592282F919723FB631.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Swezeyana magnaccai	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Swezeyana magnaccai sp. n. Figures 1F, 11 </p>
            <p>Diagnosis.</p>
            <p>Small, red- to yellow-brown species, with fore wing membrane patterned, antennae and genal processes medium long, and paramere short.</p>
            <p>Description.</p>
            <p> Adult. General body colour orange-red to yellow-brown, last 2-3 antennal segments darker brown. Fore wing patterned with irregular clouds of orange-brown pigmentation, darker patches indicate position of cross pseudoveins, intersections of veins and wing margin, as well as 1-2 darker patches on the dorsal claval wing margin, and a more or less distinct patch on vein R+M+Cu1 just basal to vein trifurcation, unpigmented areas surround the marginal clusters of radular spines (Figs 1F, 11M). Fore wing apex bluntly acute; pseudopterostigma relatively long (Fig. 11J, L), 2-3 cross pseudoveins in cell r1 (Fig. 11L); surface spinules densely distributed in all cells; medium long setae on ventral margin and medium short to short setae on veins and dorsal margin. Antennae medium long (ratio AL:HW 1.41) (Fig. 11B); genal processes medium long (GP &lt;0.35 mm, ratio HW:GP &lt;1.65) and not or only slightly upturned at apices (Fig. 11  A–B ); medium short to short setae on vertex and thorax. Meracanthus extremely small to almost absent, genual spine reduced or absent (Fig. 11  E–F ). Male terminalia (Fig. 11  H–I ): paramere short (ratio PL:HW &lt;0.30), tapering to apex with two short stout setae; distal aedeagus segment long relative to paramere (ratio PL:AEL &lt;1.10), apex developed into a large rounded hook with acute apex. </p>
            <p>Egg. Unknown.</p>
            <p>Immature. Unknown.</p>
            <p>Host plant.</p>
            <p> Planchonella sandwicensis . </p>
            <p>Distribution.</p>
            <p>Oahu. Only known from the Waianae Mountains.</p>
            <p>Etymology.</p>
            <p>Named for Karl Magnacca, a talented biologist who contributed several specimens for this study (noun in the genitive case).</p>
            <p>Comments.</p>
            <p> Currently only known from males, it may belong in the  reticulata species group based on fore wing characters and paramere shape, but is currently unplaced. </p>
            <p>Material examined.</p>
            <p> Holotype male (slide mounted), Mokuleia Forest Reserve, Pahole, N Waianae Mnts, Oahu, USA, N21.53208, W-158.1786, ex  Planchonella sandwicensis , 6 July 2014,  “Hi74-14” D. Percy leg. (BMNH). Paratypes (slide mounted) 2m, as for holotype (BMNH). Other material: 1m, Puu Hapapa, Central Waianae Mnts, Oahu, USA, N21.4666, W-158.1029, ex  Planchonella sandwicensis , 17 May 2014,  “KM14-14” K. Magnacca leg. (BMNH). </p>
            <p>Gene sequences.</p>
            <p> KY294144 (COI) KY294628 (cytB) (Hi74-14) [previously submitted to GenBank as  Swezeyana reticulata Caldwell, 1940 (in Percy 2017a)]; KY294145 (COI) KY294629 (cytB) (KM15-14) [previously submitted to GenBank as  Swezeyana reticulata Caldwell, 1940 (in Percy 2017a)]. </p>
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	https://treatment.plazi.org/id/35B0E962FEC248592282F919723FB631	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Percy, Diana M.	Percy, Diana M. (2018): Revision of the Hawaiian psyllid genus Swezeyana, with descriptions of seven new species (Hemiptera, Psylloidea, Triozidae). ZooKeys 758: 75-113, DOI: http://dx.doi.org/10.3897/zookeys.758.23019, URL: http://dx.doi.org/10.3897/zookeys.758.23019
696E08CBB838829C0667A703ABEA8BD3.text	696E08CBB838829C0667A703ABEA8BD3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Swezeyana oahuensis	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Swezeyana oahuensis sp. n. Figures 1B, 7 </p>
            <p>Diagnosis.</p>
            <p>Medium sized, light coloured species, with fore wing membrane unpatterned, antennae medium long, genal processes long, paramere medium long, and female proctiger strongly convex apically.</p>
            <p>Description.</p>
            <p> Adult. General body colour green to yellow-green or yellow-brown, last 5-7 antennal segments darker brown, apices of genae sometimes pinkish-red. Fore wing membrane uniformly pale fuscous (Fig. 1B). Fore wing apex acute; pseudopterostigma medium to long (Fig. 7R), no cross pseudoveins in cell r1; surface spinules sparsely distributed, reduced or absent from r1 and r2, and apparently absent from c+sc which is partly to almost entirely composed of a thickened ventral wing margin (C+Sc); medium long setae on ventral margin and medium short to short setae on veins and dorsal margin. Antennae medium long (ratio AL:HW 1.39-1.53) (Fig. 7B, D); genal processes medium long (GP ≥ 0.35 mm, ratio HW:GP ≥ 1.40) and often upturned at apices (Fig. 7  A–B , G); short to medium short setae on vertex and thorax. Meracanthus small (Fig. 7E), genual spine reduced or absent (Fig. 7E). Male terminalia (Fig. 7  M–Q ): paramere slender, medium long (ratio PL:HW ≥ 0.25), tapering to anteriorly projecting apex with two short stout setae; distal aedeagus segment short relative to paramere (ratio PL:AEL 1.19-1.60), apex developed into a large rounded hook with bluntly acute apex. Female terminalia (Figs 7  H–L ): proctiger dorsal surface strongly convex apically, apex broad, blunt, bearing medial cleft and fringed with stout setae, anal ring medium long (ratio FP:RL 1.90-2.05), with reduced head compartment at proximal end, distal portion of ring margin smooth; subgenital plate slightly convex with short medial cleft with beak and membrane extended (Fig. 7  K–L ). </p>
            <p>Egg. Unknown.</p>
            <p>Immature. Unknown.</p>
            <p>Host plant.</p>
            <p> Planchonella sandwicensis . </p>
            <p>Distribution.</p>
            <p>Oahu. Only known from the Waianae Mountains.</p>
            <p>Etymology.</p>
            <p>Named for its distribution on the island of Oahu (adjective in the nominative singular).</p>
            <p>Comments.</p>
            <p> This species and  S. atra may represent insular diversification on Oahu (see comments for  S. atra ). </p>
            <p>Material examined.</p>
            <p> Holotype male (slide mounted), Mnt Kaala road (culvert 32), Waianae Mnts, Oahu, USA, ex  Planchonella sandwicensis , 26 August 2003,  “Hi57-03” D. Percy leg. (BMNH). Paratypes (slide mounted) 2f, as for holotype (BMNH). Paratypes (slide mounted) 2m, Pahole NAR, Waianaea Mnts, Oahu, USA, N21.5364, W-158.1919, ex  Planchonella sandwicensis , 14 August 2003,  “Hi06-03” D. Percy leg. (BMNH). Other material: 1m 2f, South Mohiakea, Central Waianae Mnts, Oahu, USA, N21.4821, W-158.1247, ex  Planchonella sandwicensis , 29 January 2014,  “KM16-14” K. Magnacca leg. (BMNH). </p>
            <p> Gene sequences.KY294142 (COI) KY294626 (cytB) (KM16-14) [previously submitted to GenBank as  Swezeyana elongagena Caldwell, 1940 (in Percy 2017a)]; KY294143 (COI) KY294627 (cytB) (Hi57-03) [previously submitted to GenBank as  Swezeyana elongagena Caldwell, 1940 (in Percy 2017a)]. </p>
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	https://treatment.plazi.org/id/696E08CBB838829C0667A703ABEA8BD3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Percy, Diana M.	Percy, Diana M. (2018): Revision of the Hawaiian psyllid genus Swezeyana, with descriptions of seven new species (Hemiptera, Psylloidea, Triozidae). ZooKeys 758: 75-113, DOI: http://dx.doi.org/10.3897/zookeys.758.23019, URL: http://dx.doi.org/10.3897/zookeys.758.23019
4F08F306F809649641701BDCF3644650.text	4F08F306F809649641701BDCF3644650.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Swezeyana reticulata Caldwell 1940	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Swezeyana reticulata Caldwell, 1940 Figures 1G, 9 </p>
            <p> Swezeyana reticulata Caldwell, 1940: 390. </p>
            <p>Description.</p>
            <p> Adult. General body colour yellow-brown to darker brown, last 2-3 antennal segments darker brown. Fore wing patterned with irregular clouds of brown pigmentation (although less distinctly than in  S. rubra and  S. tentaculata ), darker brown patches indicate position of cross pseudoveins, intersections of veins and wing margin, as well as 1-2 brown patches on the dorsal claval wing margin, and a more or less distinct patch on vein R+M+Cu1 just basal to vein trifurcation, unpigmented areas surround the marginal clusters of radular spines (Fig. 1G). Fore wing apex acute to bluntly acute; pseudopterostigma short (Fig. 9S), 3-4 cross pseudoveins in cell r1 (Fig. 9T); surface spinules densely distributed in all cells; medium long setae on ventral margin and short setae on veins and dorsal margin. Antennae medium short (ratio AL:HW 1.25-1.40) (Fig. 9B, I); genal processes medium long (GP &lt;0.40 mm, ratio HW:GP &lt;1.65) and slightly upturned at apices (Fig. 9  A–B ,  E–F ); medium short to short setae on vertex and thorax. Meracanthus small, genual spine developed (Fig. 9D). Male terminalia (Fig. 9  Q–R ): paramere short (ratio PL:HW &lt;0.25), tapering to apex with two short stout setae; distal aedeagus segment long relative to paramere (ratio PL:AEL &lt;0.80), apex developed into a large rounded hook with bluntly acute apex. Female terminalia (Fig. 9  J–P ): proctiger dorsal surface more or less straight, tapering to bluntly acute apex without medial cleft, anal ring relatively short (ratio FP:RL 2.17-2.45), with well-developed head compartment at proximal end, distal portion of ring margin slightly convoluted; subgenital plate more or less straight ventrally with little or no medial cleft and with beak and membrane slightly extended (Fig. 9  L–N ). </p>
            <p>Egg. Pale with shallow hexagonal indentations dorsally.</p>
            <p> Immature. Colour: Mottled, cream and red-brown. Structure: 5th instar with circumanal ring wide, and more or less straight, with a single row of uninterrupted elongate cells (Fig. 13G). Chaetotaxy: 5th instar with marginal, narrow, blunt sectasetae, and sub-marginal longer acute, simple setae on enlarged, ridged annuli or pediments (Fig. 13  A–C ); dorsal surface rugose, with small acute, simple setae on bulbous, ridged pediments, and with distinct arrangement of 28 protruding tubercles (13 on head and thorax, 15 on abdomen, of these 10 are medial, and 18 are lateral or sub-marginal), tubercles bearing scattered small, blunt, simple setae (Fig. 13  D–F ). 1st instar with marginal narrow, blunt sectasetae (Fig. 12A); 2nd instar with marginal narrow, blunt sectasetae, sub-marginal longer acute simple setae on enlarged and ridged annuli or pediments, and 7 sub-marginal tubercles, with 3 on thorax bearing 1-2 acute simple setae, and a marginal sub-apical pair of long simple setae on abdomen (Fig. 12  B–D ); 3rd - 4th instar chaetotaxy and tubercle arrangement similar to 5th instar (Fig. 12  E–F ). In 4th and particularly 5th instar, 4 of the dorsal tubercles are more darkly pigmented (these are the distal medial tubercles on thorax and abdomen, and the proximal lateral tubercles on abdomen; Fig. 13  E–F ). </p>
            <p>Immature measurements (mm) and ratios: 5th instar (n = 4): BL 1.67-1.71; BW 1.06-1.15; WPL 0.85-0.88; CPL 0.73-0.79; CPW 0.62-0.97; RW 0.17-0.18; HW 0.52-0.57; AL 0.19-0.20; BL:BW 1.49-1.57; HW:AL 2.60-2.96; CPW:RW 3.70-5.77.</p>
            <p>Host plant.</p>
            <p> Planchonella sandwicensis . </p>
            <p>Distribution.</p>
            <p> Kauai (possibly also on Maui, see comments). Appears to be the most common of the three  Swezeyana species found on Kauai. </p>
            <p>Comments.</p>
            <p> Tuthill (1966) notes the co-occurrence of two species on Maui, and refers to these as  S. elongagena and  S. reticulata ; but  S. reticulata is here considered endemic to Kauai and the observed species on Maui may be an as yet undescribed species. The immatures as noted by Tuthill have "a remarkable armament of large proturberances [sic] on the dorsal surface" and therefore fit within the  reticulata group. </p>
            <p>Material examined.</p>
            <p> Holotype female (BPBM, not located). Other material: 1f, Nualolo Trail, Kokee State Park, Kauai, USA, on  Polyscias waimeae , 25 May 2002,  “370-02” D. Percy leg. (BMNH). 10m 4f 7i, Kokee State Park, Kauai, USA, N22.1444, W-159.6477, ex  Planchonella sandwicensis , 29 October 2005,  “Hi01-05” D. Percy leg. (BMNH). 5m 1f, Kokee State Park, Kauai, USA, N22.1503, W-159.6453, ex  Planchonella sandwicensis , 29 October 2005,  “Hi02-05” D. Percy leg. (BMNH). 12m 7f, Kokee State Park, Kauai, USA, N22.1309, W-159.6388, ex  Planchonella sandwicensis , 30 October 2005,  “Hi05-05” D. Percy leg. (BMNH). 1m 10f, Kokee State Park, Kauai, USA, N22.0948, W-159.6953, ex  Planchonella sandwicensis , 30 October 2005,  “Hi11-05” D. Percy leg. (BMNH). </p>
            <p>Gene sequences.</p>
            <p>MG988833 (COI) MG989154 (cytB) (Hi01-05); MH001524 (COI) MH001530 (cytB) (Hi05-05 adult); MH001525 (COI) MH001531 (cytB) (Hi05-05 immature); MH001526 (COI) MH001532 (cytB) (Hi11-05).</p>
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	https://treatment.plazi.org/id/4F08F306F809649641701BDCF3644650	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Percy, Diana M.	Percy, Diana M. (2018): Revision of the Hawaiian psyllid genus Swezeyana, with descriptions of seven new species (Hemiptera, Psylloidea, Triozidae). ZooKeys 758: 75-113, DOI: http://dx.doi.org/10.3897/zookeys.758.23019, URL: http://dx.doi.org/10.3897/zookeys.758.23019
0024FAE8B8BF9FFCE1A36D92A68BB39D.text	0024FAE8B8BF9FFCE1A36D92A68BB39D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Swezeyana rubra	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Swezeyana rubra sp. n. Figures 1I, 8 </p>
            <p>Diagnosis.</p>
            <p>Medium small, red-brown species, with fore wing membrane distinctly patterned, antennae and genal processes relatively short, paramere short, and female proctiger strongly convex apically.</p>
            <p>Description.</p>
            <p> Adult. General body colour red or red-brown, last 2-3 antennal segments darker brown. Fore wing distinctly patterned with irregular clouds of red pigmentation, mottled red-brown pattern in apical 2/3 of fore wing, with basal portion either clear (males) or slightly mottled (females), males have a darker almost solid red-brown area of pigmentation across middle of wing membrane, wing veins variably brown to speckled brown with darker brown patches indicating position of cross pseudoveins, intersections of veins and wing margin, as well as two brown patches on the dorsal claval wing margin, and on vein R+M+Cu1 just basal to vein trifurcation, there are distinctly unpigmented areas surrounding marginal clusters of radular spines (Fig. 1I). Fore wing apex bluntly acute, shape shorter and broader than other  Swezeyana (WL:WW &lt;3.10); pseudopterostigma long (Fig. 8V), 5-7 cross pseudoveins in cell r1 (Fig. 8W); surface spinules densely distributed in all cells, but cell c+sc often partly to almost entirely composed of a thickened ventral wing margin (C+Sc); medium long setae on ventral margin and medium short to short setae on veins and dorsal margin. Antennae short (ratio AL:HW 0.93-1.03) (Fig. 8  C–D ); genal processes relatively short (GP &lt;0.35 mm, ratio HW:GP&gt; 1.65) and not or slightly upturned at apices (Fig. 8  A–C ); medium short to short setae on vertex and thorax. Meracanthus small (Fig. 8G), genual spine reduced or absent (Fig. 8J). Male terminalia (Fig. 8  R–T ): paramere short (ratio PL:HW &lt;0.25), tapering to slightly anteriorly projecting apex with two short stout setae; distal aedeagus segment short relative to paramere (ratio PL:AEL &lt;1.10), apex developed into a large rounded hook with acute apex. Female terminalia (Fig. 8  L–Q ): proctiger dorsal surface strongly convex apically, apex broad, blunt, bearing medial cleft and fringed with stout setae, anal ring medium long (ratio FP:RL 1.71-2.19), with reduced head compartment at proximal end, distal portion of ring margin smooth; subgenital plate slightly convex with little or no medial cleft but with beak and membrane extended (Fig. 8  P–Q ). </p>
            <p>Egg. Pale with loosely structured hexagonal sculpturing dorsally (Fig. 8U).</p>
            <p>Immature. Unknown.</p>
            <p>Host plant.</p>
            <p> Planchonella sandwicensis . </p>
            <p>Distribution.</p>
            <p>Hawaii. Only known from PuuWaaWaa area.</p>
            <p>Etymology.</p>
            <p>Named for the generally red body colouration (adjective in the nominative singular).</p>
            <p>Comments.</p>
            <p> This species was collected from the same individual host tree as  S. hawaiiensis (see comments for  S. hawaiiensis ). </p>
            <p>Material examined.</p>
            <p> Holotype male (slide mounted), PuuWaaWaa, Hawaii, USA, N19.784, W-155.833, 820m, ex  Planchonella sandwicensis , 29 July 2002,  “440B-02” D. Percy leg. (BMNH). Paratypes (slide mounted) 2f, as for holotype (BMNH). Other material: 2m 8f, PuuWaaWaa Forest Reserve, Hawaii, USA, 2600 ft, ex  Planchonella sandwicensis , 23 February 2011,  “JG5B” J. Giffin leg. (BMNH). </p>
            <p>Gene sequences.</p>
            <p>MH001523 (COI) MH001529 (cytB) (JG5B); MG988834 (COI) MG989156 (cytB) (440B-02).</p>
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	https://treatment.plazi.org/id/0024FAE8B8BF9FFCE1A36D92A68BB39D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Percy, Diana M.	Percy, Diana M. (2018): Revision of the Hawaiian psyllid genus Swezeyana, with descriptions of seven new species (Hemiptera, Psylloidea, Triozidae). ZooKeys 758: 75-113, DOI: http://dx.doi.org/10.3897/zookeys.758.23019, URL: http://dx.doi.org/10.3897/zookeys.758.23019
E07FF23F7F4F80C19A418E4D6A543B64.text	E07FF23F7F4F80C19A418E4D6A543B64.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Swezeyana tentaculata	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Swezeyana tentaculata sp. n. Figures 1H, 10 </p>
            <p>Diagnosis.</p>
            <p>Medium sized, red-brown species, with fore wing membrane patterned, antennae medium long, genal processes long, paramere short, and female proctiger more or less straight dorsally.</p>
            <p>Description.</p>
            <p> Adult. General body colour red to red-brown, last 2-3 antennal segments darker brown. Fore wing distinctly patterned with irregular clouds of red-brown pigmentation, darker brown patches indicate position of cross pseudoveins, intersections of veins and wing margin, as well as two brown patches on the dorsal claval wing margin, and on vein R+M+Cu1 just basal to vein trifurcation, there are distinctly unpigmented areas surrounding marginal clusters of radular spines (Figs 1H, 10S). Fore wing apex bluntly acute; pseudopterostigma short (Fig. 10R), 4-6 cross pseudoveins in cell r1 (Fig. 10  R–S ); surface spinules densely distributed in all cells; long setae on ventral margin and medium short to short setae on veins and dorsal margin. Antennae medium long (ratio AL:HW 1.40-1.61) (Fig. 10C,  E–F ); genal processes long (GP ≥ 0.35 mm, ratio HW:GP ≤ 1.35) and upturned at apices (Figs 10  A–C ); medium short to short setae on vertex and thorax. Meracanthus reduced, almost absent, genual spine developed (Fig. 10H). Male terminalia (Fig. 10  P–Q ): paramere short (ratio PL:HW &lt;0.30), tapering to apex with two short stout setae; distal aedeagus segment long relative to paramere (ratio PL:AEL &lt;0.90), apex developed into a large rounded hook with acute apex. Female terminalia (Fig. 10  I–N ): proctiger dorsal surface more or less straight, tapering to bluntly acute apex without medial cleft, anal ring relatively short (ratio FP:RL 2.24-2.59), with well-developed head compartment at proximal end, distal portion of ring margin convoluted; subgenital plate more or less straight ventrally with little or no medial cleft and with beak and membrane slightly extended (Fig. 9  L–N ). </p>
            <p>Egg. Pale, sculpturing consisting of rounded indentations dorsally (Fig. 10O).</p>
            <p> Immature. Colour: Mostly red-brown, some cream mottling. Structure: 5th instar with circumanal ring wide, slightly constricted medially and lateral apices upturned, with a single row of uninterrupted elongate cells (Fig. 13I). Chaetotaxy: 5th instar with marginal, pointed sectasetae, and sub-marginal longer acute, simple setae on enlarged, ridged annuli or pediments (Fig. 13H,  J–K ); dorsal surface rugose, with small club setae, and with distinct arrangement of 28 protruding tubercles and tentacles (13 on head and thorax, 15 on abdomen, of these 10 are medial, and 18 are lateral or sub-marginal), all but 4 of the dorso-medial protuberances (on abdomen) are developed into elongate tentacles bearing scattered small, blunt, simple setae on the apical portion, and small club setae on the basal portion, tentacle apices bear 1-2 acute simple setae (Fig. 13L). 3rd - 4th instar chaetotaxy and tubercle/tentacle arrangement similar to 5th instar (Fig. 12  G–J ); in 3rd instars the lateral and submarginal tubercles bear 2 club setae at the apices (Fig. 12I), and the dorso-medial tubercles destined to become tentacles are larger and bear several club setae (Fig. 12H), by 4th instar elongation into tentacles is already evident. Somewhat more darkly pigmented tentacles are found in the same position as described for  S. reticulata (the distal medial tentacles on thorax and abdomen, and the proximal lateral tentacles on abdomen; Fig. 13H, K). The tentacles are also evident in photographs of immatures on the leaf surface (Fig. N). </p>
            <p>Immature measurements (mm) and ratios: 5th instar (n = 3): BL 1.48-1.52; BW 0.91-0.97; WPL 0.73-0.76; CPL 0.64; CPW 0.76; RW 0.16-0.17; HW 0.48-0.52; AL 0.18; BL:BW 1.56-1.63; HW:AL 2.61-2.95; CPW:RW 4.51-4.92.</p>
            <p>Host plant.</p>
            <p> Planchonella sandwicensis . </p>
            <p>Distribution.</p>
            <p>Kauai. Only known from Kokee State Park.</p>
            <p>Etymology.</p>
            <p>Named for the distinctly long tentacles on the dorsum of immatures (adjective in the nominative singular).</p>
            <p>Comments.</p>
            <p> Found sympatrically with  S. reticulata and  S. elongagena on the same individual plants. Immatures were observed among the ferugineous trichomes on the undersides of leaves, often along the leaf mid-rib (Fig. 13N). </p>
            <p>Material examined.</p>
            <p> Holotype male (slide mounted), Kokee State Park, Kauai, USA, N22.1444, W-159.6477, ex  Planchonella sandwicensis , 29 October 2005,  “Hi01-05” D. Percy leg. (BMNH). Paratypes (slide mounted) 3f 7i, as for holotype (BMNH). Other material: 2m 2f, Kokee State Park, Kauai, USA, N22.1309, W-159.6388, ex  Planchonella sandwicensis , 30 October 2005,  “Hi05-05” D. Percy leg. (BMNH). </p>
            <p>Gene sequences.</p>
            <p>MG989157 (cytB) (Hi01-05).</p>
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	https://treatment.plazi.org/id/E07FF23F7F4F80C19A418E4D6A543B64	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Percy, Diana M.	Percy, Diana M. (2018): Revision of the Hawaiian psyllid genus Swezeyana, with descriptions of seven new species (Hemiptera, Psylloidea, Triozidae). ZooKeys 758: 75-113, DOI: http://dx.doi.org/10.3897/zookeys.758.23019, URL: http://dx.doi.org/10.3897/zookeys.758.23019
