taxonID	type	description	language	source
791099101D915C118A0546BAFB55238D.taxon	diagnosis	Diagnosis. Leptopilina australis is a relatively small species (1.3 – 1.4 mm ♀ body length in a small number of measured females) with somewhat slender appearance and a relatively short antenna (Fig. 2 A). The species is unique in the combination of the mesoscutum having rows of setae mediolaterally (Fig. 11 A, susceptible to damage), a state shared with L. clavipes (mesoscutum glabrous in other species), and the short metapleural ridge 1 (Fig. 2 D, see Fig. 9 A for an overview on the metapleural ridges 1 – 3). The short ridge is shared with L. fimbriata, but in other species it is either absent (L. boulardi) or at least half the length of the metapleuron. The metapleural ridge 2 is shorter than half the length of the metapleuron, albeit longer than ridge 1 (Fig. 2 D). The ridge 2 is equally long in L. boulardi, L. clavipes and L. fimbriata but at least half the length of the metapleuron in the other species. A setal patch on the base of the hind coxa is present (Fig. 2 D), in contrast to L. japonica and L. boulardi, where there are at most a few singular setae that are not arranged in a patch.	en	Vogel, Jonathan, Martin, Jakob, Forshage, Mattias, Salden, Tobias, Staverløkk, Arnstein, Verheyde, Fons, Nordlander, Göran, Herz, Annette, Peters, Ralph S. (2025): The Leptopilina Förster, 1869 of the Western Palearctic, and an updated list of the world species (Hymenoptera, Figitidae, Eucoilinae). Journal of Hymenoptera Research 98: 915-969, DOI: 10.3897/jhr.98.165583
791099101D915C118A0546BAFB55238D.taxon	distribution	Distribution. Western Palearctic but sparsely distributed, possibly spreading towards northwest but slowly and still uncommon. Armenia (locus typicus of Rhoptromeris australis), Belgium (new record), Denmark, the Netherlands and Slovenia.	en	Vogel, Jonathan, Martin, Jakob, Forshage, Mattias, Salden, Tobias, Staverløkk, Arnstein, Verheyde, Fons, Nordlander, Göran, Herz, Annette, Peters, Ralph S. (2025): The Leptopilina Förster, 1869 of the Western Palearctic, and an updated list of the world species (Hymenoptera, Figitidae, Eucoilinae). Journal of Hymenoptera Research 98: 915-969, DOI: 10.3897/jhr.98.165583
13E9BDC71FD05AC49EE3162B4F1ADE6D.taxon	diagnosis	Diagnosis. Leptopilina boulardi is a small to medium-sized species (up to 1.5 mm ♀ body length) with a stout appearance and relatively short appendages (Fig. 3 A). The species is unique by its smooth metapleuron, having at most a short ridge 2 and the other ridges absent (Fig. 3 D). In all other species, the three ridges are at least somewhat developed. Also, the propodeal carinae are diverging to a varying degree in the posterior half of the propodeum (Fig. 3 E). Other species have their propodeal carinae running uniformly straight or outwardly curved from the anterior to the posterior margin of the propodeum. The sculpture on the dorsal surface of the mesoscutellum is striate. The striae are radiating from the base of the mesoscutellar plate (Fig. 3 C) and the posterior surface is areolate. The sculpture of L. heterotoma is similar, though the striae are more dispersed elong the entire length of the mesoscutellar plate and the posterior surface is foveate-reticulate. In other species, the mesoscutellum is sculptured entirely foveate-reticulate. Just as in L. japonica, the setal patch on the base of the metacoxa of L. boulardi is absent. There are at most a few singular setae (Fig. 3 D). The forewing is usually relatively small with a narrow, elongate marginal cell, but can be different and is thereby not sufficient as a diagnostic character.	en	Vogel, Jonathan, Martin, Jakob, Forshage, Mattias, Salden, Tobias, Staverløkk, Arnstein, Verheyde, Fons, Nordlander, Göran, Herz, Annette, Peters, Ralph S. (2025): The Leptopilina Förster, 1869 of the Western Palearctic, and an updated list of the world species (Hymenoptera, Figitidae, Eucoilinae). Journal of Hymenoptera Research 98: 915-969, DOI: 10.3897/jhr.98.165583
13E9BDC71FD05AC49EE3162B4F1ADE6D.taxon	distribution	Distribution. Cosmopolitan species; in Western Palearctic with a center in the Mediterranean region and not extending far north: found in the Canary Islands, France, Greece, Iran, Italy, Madeira, Portugal, Serbia, Spain, Switzerland, Tunisia and Turkey. Also present in the Afrotropics: Democratic Republic of the Congo, Republic of the Congo, Gambia, Ivory Coast, Kenya, Madagascar, Seychelles (locus typicus of Charips mahensis), South Africa, Zambia and Zimbabwe; in North America: Canada and USA (east and west); South America: Argentina, Brazil, Guadeloupe (locus typicus of Cothonaspis boulardi), and Mexico; and in Australia and Vanuatu. We have not seen records from tropical Asia, but the species is probably also present there.	en	Vogel, Jonathan, Martin, Jakob, Forshage, Mattias, Salden, Tobias, Staverløkk, Arnstein, Verheyde, Fons, Nordlander, Göran, Herz, Annette, Peters, Ralph S. (2025): The Leptopilina Förster, 1869 of the Western Palearctic, and an updated list of the world species (Hymenoptera, Figitidae, Eucoilinae). Journal of Hymenoptera Research 98: 915-969, DOI: 10.3897/jhr.98.165583
2012FC72C15F5DE684CF89AB98F0A56E.taxon	diagnosis	Diagnosis. Leptopilina clavipes is a size-variable species with relatively short appendages (Fig. 4 A). The species has a unique mesoscutellum which is not subdivided into a dorsal and posterior surface by a circumscutellar carina or varying sculpture dorsally and posteriorly (Fig. 4 E). In other species, there is a more or less clear division by the circumscutellar carina and different sculpture dorsally and posteriorly. The mesoscutum has rows of setae mediolaterally, just as in the smaller L. australis (Fig. 11 A, susceptible to damage). None of the other species exhibit these rows of setae. The metapleural ridge 1 reaches to about half the length of the metapleuron (Fig. 4 D), as in L. longipes. The other species either possess a longer ridge 1 (L. japonica and L. heterotoma), a shorter (L. australis and L. fimbriata), or an absent one (L. boulardi). The metapleural ridge 2 is shorter than half the length of the metapleuron (Fig. 4 D) as in L. australis, L. boulardi and L. fimbriata. In all other species, ridge 2 is at least half the length of the metapleuron.	en	Vogel, Jonathan, Martin, Jakob, Forshage, Mattias, Salden, Tobias, Staverløkk, Arnstein, Verheyde, Fons, Nordlander, Göran, Herz, Annette, Peters, Ralph S. (2025): The Leptopilina Förster, 1869 of the Western Palearctic, and an updated list of the world species (Hymenoptera, Figitidae, Eucoilinae). Journal of Hymenoptera Research 98: 915-969, DOI: 10.3897/jhr.98.165583
2012FC72C15F5DE684CF89AB98F0A56E.taxon	description	Molecular characterisation. Maximum intraspecific barcode-distance: - % (1). Minimum interspecific barcode-distance: 14.3 % (L. longipes). CO 1 barcode sequence: 658 bp. 5 ’ - TTTAATATATTTTATATTTGGAATTTGGTCAGGGATAGTAGGAGCAAGATTAAGAATAATTATTCGATTAGAGTTAGGAACTCCTGGGCAGTTAATTAATAATGACCAGATTTATAATTCTATAGTGACTGTTCATGCTTTTGTTATAATTTTTTTTATAGTTATGCCTATTATAGTAGGAGGATTTGGTAATTATTTAGTTCCTTTAATAGTTACAGTTCCTGATATGGCTTTTCCTCGTTTAAATAATATGAGATTATGACTTTTATTTCCTTCTTTAATTTTAATGTTAGCTAGTATATTTATTGATCAAGGAGCAGGAACTGGGTGAACTGTGTATCCTCCTCTTTCTTTAAGTGTAAGGCATCCTGGAGTAGCTGTAGATTTAATTATTTTTTCTTTACATTTAAGAGGGGTTTCATCAATTTTAGGGTCTATTAATTTTATTTCTACAATTTTTAATATTCGTCCATTGTTAATAGGGATAGATAAAATTACTTTATTTTTATGATCTATTTTTTTAACAACTATTTTATTATTACTTTCTTTACCTGTATTAGCAGGAGGGATTACAATATTATTATTTGACCGTAATTTAAATACTTCTTTTTATGATCCAGTTGGGGGTGGGGATCCAATTTTGTATCAACATTTATTT- 3 ’.	en	Vogel, Jonathan, Martin, Jakob, Forshage, Mattias, Salden, Tobias, Staverløkk, Arnstein, Verheyde, Fons, Nordlander, Göran, Herz, Annette, Peters, Ralph S. (2025): The Leptopilina Förster, 1869 of the Western Palearctic, and an updated list of the world species (Hymenoptera, Figitidae, Eucoilinae). Journal of Hymenoptera Research 98: 915-969, DOI: 10.3897/jhr.98.165583
2012FC72C15F5DE684CF89AB98F0A56E.taxon	distribution	Distribution. Possibly Holarctic: in Europe mainly in Northern and central parts, records from Austria, Belgium (new record), Czech Republic, Denmark, Estonia, Finland, Germany (locus typicus of Cothonaspis clavipes), the Netherlands, Norway, Spain, Sweden, and United Kingdom. Also present in eastern USA (and likely Canada, but none of the records confirmed), Japan, and Argentina.	en	Vogel, Jonathan, Martin, Jakob, Forshage, Mattias, Salden, Tobias, Staverløkk, Arnstein, Verheyde, Fons, Nordlander, Göran, Herz, Annette, Peters, Ralph S. (2025): The Leptopilina Förster, 1869 of the Western Palearctic, and an updated list of the world species (Hymenoptera, Figitidae, Eucoilinae). Journal of Hymenoptera Research 98: 915-969, DOI: 10.3897/jhr.98.165583
74160433797A517DAED54382B5904E0B.taxon	diagnosis	Diagnosis. Leptopilina fimbriata is a small to medium-sized species (up to 1.6 mm ♀ body length) with slender appearance and remarkably long and filiform antennae in both sexes reaching more than 1.0 times the body length in females and more than 1.9 times the body length in males (Fig. 5 A). The metasoma is significantly paler than head and mesosoma, a pattern similar as in L. longipes, but more distinct. Also, the legs are strikingly stramineous (Fig. 5 A), while they are generally darker (brownish to reddish) in all other species. Additionally, the mesoscutellar plate is usually notably circular and short in dorsal view (Fig. 5 C), and elevated and strongly sloping posteriorly in lateral view (Fig. 5 A). Other species possess either a rhombic (L. heterotoma) or drop-shaped (all other species), not notably short or elevated mesoscutellar plate, only slightly sloping posteriorly in L. longipes. The metapleural ridge 1 is shorter than half the length of the metapleuron (Fig. 5 D) as in L. australis and L. clavipes. In other species, ridge 1 is at least half as long as the metapleuron, if present at all. The metapleural ridge 2 is shorter than half the length of the metapleuron (Fig. 5 D), as in L. australis, L. boulardi, and L. clavipes. In other species, ridge 2 is at least half as long as the metapleuron. The forewing is long with a narrow marginal cell. The forewing vein Rs is clearly longer than 2 r, and the accessory veins (M, Rs + M CU 1 and CU 1 a) are usually very distinct (Fig. 12 B), while they are faint or absent in other species. Superficially, L. fimbriata is quite similar to Ganaspis seticornis Hellén, 1960. This is another eucoiline species from a genus potentially associated with Drosophila and the most slenderly built European species in that genus. However, these genera are not closely related. Ganaspis is having far more of a hairy ring, less of a petiolar rim, a long row of setae on the metacoxae, and modified F 1 instead of F 2 in the male antenna.	en	Vogel, Jonathan, Martin, Jakob, Forshage, Mattias, Salden, Tobias, Staverløkk, Arnstein, Verheyde, Fons, Nordlander, Göran, Herz, Annette, Peters, Ralph S. (2025): The Leptopilina Förster, 1869 of the Western Palearctic, and an updated list of the world species (Hymenoptera, Figitidae, Eucoilinae). Journal of Hymenoptera Research 98: 915-969, DOI: 10.3897/jhr.98.165583
74160433797A517DAED54382B5904E0B.taxon	description	Molecular characterisation. Maximum intraspecific barcode-distance: 1.4 % (37). Minimum interspecific barcode-distance: 13.0 % (L. japonica). Consensus barcode sequence: 658 bp. 5 ’ - AGTTATATATTTTATTTTTGGGATTTGATCTGGGATAGTGGGGGCGAGATTGAGGATAATTATTCGTATAGAATTGGGGATACCGGGGCAGTTAATTAATAATGATCAAGTTTATAATACTATTGTTACGGCTCATGCATTTATTATAATTTTTTTTATAGTGATACCTATTATAGTTGGTGGGTTTGGGAACTATTTAATTCCTTTAATAATTACAGTTCCTGATATGGCGTTTCCTCGATTAAATAATATAAGATTATGACTTTTATTTCCTTCTTTATTTTTAATGTTAGCTAGAATATTTATTGATCAGGGGGCCGGGACAGGATGAACTGTTTATCCCCCTTTATCTTTAAGAATTGGGCATCCGGGGGTTTCTGTTGATTTAGTGATTTTTTCGTTACATTTAAGGGGGGTTTCTTCTATTTTGGGGTCAATTAATTTTATTTCTACTATTTTAAATGTTCGTCCAAATTTAATAATAATGGATAAAGTTACTTTATTTATTTGGTCTATTTTTTTAACAACTATTTTATTACTGTTATCTTTACCGGTATTAGCTGGGGGGATTACAATATTATTATTTGATCGTAATTTAAATACTTCTTTTTATGATCCTGTGGGAGGGGGGGATCCAATTTTGTATCAACATTTATTT- 3 ’.	en	Vogel, Jonathan, Martin, Jakob, Forshage, Mattias, Salden, Tobias, Staverløkk, Arnstein, Verheyde, Fons, Nordlander, Göran, Herz, Annette, Peters, Ralph S. (2025): The Leptopilina Förster, 1869 of the Western Palearctic, and an updated list of the world species (Hymenoptera, Figitidae, Eucoilinae). Journal of Hymenoptera Research 98: 915-969, DOI: 10.3897/jhr.98.165583
74160433797A517DAED54382B5904E0B.taxon	distribution	Distribution. Palearctic species. Present in Austria, Belgium, the Canary Islands, Czech Republic, Denmark, Estonia, Finland (locus typicus of Episoda dolichocera), France (locus typicus of Eucoela fimbriata, Psilosema xanthopum, Psilosema filicorne and Psilosema longicornis), Georgia, Germany, Greece, Italy, Lithuania, Madeira, the Netherlands, Norway, Poland, Portugal, Russia, Slovenia, Spain (locus typicus of Erisphagia longipes), Sweden, Switzerland, Ukraine, and the United Kingdom. Further East, the species was also recorded from Kyrgysztan and China (BOLD). The published record from the Afrotropical region (van Noort et al. 2015) may be a mistake and requires substantiation.	en	Vogel, Jonathan, Martin, Jakob, Forshage, Mattias, Salden, Tobias, Staverløkk, Arnstein, Verheyde, Fons, Nordlander, Göran, Herz, Annette, Peters, Ralph S. (2025): The Leptopilina Förster, 1869 of the Western Palearctic, and an updated list of the world species (Hymenoptera, Figitidae, Eucoilinae). Journal of Hymenoptera Research 98: 915-969, DOI: 10.3897/jhr.98.165583
13E31EB69EB5510989F484D558B80670.taxon	type_taxon	Type species. Leptopilina longipes (Hartig, 1841).	en	Vogel, Jonathan, Martin, Jakob, Forshage, Mattias, Salden, Tobias, Staverløkk, Arnstein, Verheyde, Fons, Nordlander, Göran, Herz, Annette, Peters, Ralph S. (2025): The Leptopilina Förster, 1869 of the Western Palearctic, and an updated list of the world species (Hymenoptera, Figitidae, Eucoilinae). Journal of Hymenoptera Research 98: 915-969, DOI: 10.3897/jhr.98.165583
13E31EB69EB5510989F484D558B80670.taxon	diagnosis	Diagnosis. Leptopilina species are recognized from other cynipoid wasps by having a mesoscutellar plate (as do all members of the Eucoilinae subfamily), the dorsally broadly interrupted to almost absent hairy ring of the metasoma (dorsally shortly interrupted, absent, or complete in other eucoiline genera) in combination with the postpetiolar rim (absent in most genera, e. g. Ganaspis Förster, 1869 and Hexacola Förster, 1869, present in Cothonaspis Hartig, 1840, Diglyphosematini, and Zaeucoilini; these usually lack a hairy ring). Female specimens have 13 antennomeres, while male specimens have 15. In males, the second flagellomere (F 2) is curved and somewhat irregular (unlike in several other groups of Eucoilinae, where the first flagellomere (F 1) is irregular). Leptopilina species can have their hairy ring reduced and species of Diglyphosematini, Zaeucoilini or Cothonaspis can developed an unusually distinct hairy ring. In these cases, Leptopilina can be differentiated from the Diglyphosematini and Zaeucoilini by having a regularly wide pronotum without distinct shoulders (very broad pronotum with distinct shoulders in Diglyphosematini and Zaeucoilini). Additionally, the male F 1 is modified in Diglyphosematini and Zaeucoilini and the metasoma is roundish, not longer than high in Diglyphosematini and Zaeucoilini (more elongate in Leptopilina). In comparison to Cothonaspis, Leptopilina species have a much less elongate appearance, especially the mesosoma is less elongate in Leptopilina, with a mesoscutum about as long as wide (clearly longer than wide in Cothonaspis), and the propodeal area is much more setose in Leptopilina (hardly any seta in Cothonaspis).	en	Vogel, Jonathan, Martin, Jakob, Forshage, Mattias, Salden, Tobias, Staverløkk, Arnstein, Verheyde, Fons, Nordlander, Göran, Herz, Annette, Peters, Ralph S. (2025): The Leptopilina Förster, 1869 of the Western Palearctic, and an updated list of the world species (Hymenoptera, Figitidae, Eucoilinae). Journal of Hymenoptera Research 98: 915-969, DOI: 10.3897/jhr.98.165583
8BE5EAD9D28E5D0880650B05D5BA0AC9.taxon	diagnosis	Diagnosis. Leptopilina heterotoma is a size-variable species but on average large (frequently around 2 mm ♀ body length) with a robust appearance and medium-long antennae (Fig. 6 A). The species possesses a uniquely large mesoscutellar plate which is widest in its anterior half, making it rhombic in shape (Fig. 6 C). However, the shape is varying slightly and can overlap with that of L. japonica and vice versa. The dorsal profile of the mesoscutellar plate usually appears s-shaped in L. heterotoma and is more evenly convex in L. japonica. The mesoscutellar plate is either more or less circular in L. fimbriata or less elongate drop-shaped in all other species. The metapleural ridges 1 and 2 are reaching (or almost so) the anterior margin of the metapleuron (Fig. 6 E), as in L. japonica. The ridges 1 and 2 of the other species, if present, reach at most to half the length of the metapleuron. The sculpture of the dorsal surface of the mesoscutellum is mediolaterally areolate. This is somewhat similar to the striae of L. boulardi, though the striae in L. heterotoma are more equally distributed along the mesoscutellar plate (Fig. 6 C), while they are radiating from the base of the mesoscutellar plate in L. boulardi. The sculpture of all other species is usually foveate-reticulate. The base of the metacoxa has a setal patch (Fig. 6 E), like most other species. Only L. japonica and L. boulardi have either no patch or at most a few singular setae. L. heterotoma can be confused with small species of Trybliographa, which are similar and closely related but they have a full hairy ring and lack the petiolar rim.	en	Vogel, Jonathan, Martin, Jakob, Forshage, Mattias, Salden, Tobias, Staverløkk, Arnstein, Verheyde, Fons, Nordlander, Göran, Herz, Annette, Peters, Ralph S. (2025): The Leptopilina Förster, 1869 of the Western Palearctic, and an updated list of the world species (Hymenoptera, Figitidae, Eucoilinae). Journal of Hymenoptera Research 98: 915-969, DOI: 10.3897/jhr.98.165583
8BE5EAD9D28E5D0880650B05D5BA0AC9.taxon	description	Molecular characterisation. Maximum intraspecific barcode-distance: 1.1 % (45). Minimum interspecific barcode-distance: 11.7 % (L. japonica). Consensus barcode sequence: 658 bp. 5 ’ - TATTATATATTTTATATTTGGAATTTGATCAGGGATAGTAGGGGCAGGGTTAAGGTTGATTGTTCGGATAGAGTTAGGTATACCAGGTCAATTAATTAATAATGATCAAATTTATAATTCTATTGTTACTGCTCATGCATTTATTATAATTTTTTTTATAGTTATACCAATTATAGTTGGAGGATTTGGGAATTATTTAATTCCATTAATACTTACAGTTCCTGATATAGCATTTCCACGTTTAAATAATATAAGTTTATGACTTTTATTTCCTTCTATGATTTTAATATTAGCAAGAATAATAATTGACCAAGGGGCAGGAACAGGATGAACTGTTTACCCTCCTTTATCTCTTAGAGATAGACATCCTGGGGTTTCAACTGATTTAGTAATTTTTTCATTACATTTAAGGGGGGTATCTTCAATTTTAGGGTCTATTAATTTTATTTCAACAATTATTAATATACGACCTTATTTAATATCAATAGATAAAATTACATTATTTGTTTGAGCAATTTTTTTAACAACTATTCTTTTATTGTTATCATTACCTGTTTTAGCAGGAGGAATTACAATATTATTATTTGATCGAAATTTAAATACTTCTTTTTATGATCCTGTTGGAGGAGGAGATCCAATTTTGTATCAACATTTATTT- 3 ’.	en	Vogel, Jonathan, Martin, Jakob, Forshage, Mattias, Salden, Tobias, Staverløkk, Arnstein, Verheyde, Fons, Nordlander, Göran, Herz, Annette, Peters, Ralph S. (2025): The Leptopilina Förster, 1869 of the Western Palearctic, and an updated list of the world species (Hymenoptera, Figitidae, Eucoilinae). Journal of Hymenoptera Research 98: 915-969, DOI: 10.3897/jhr.98.165583
8BE5EAD9D28E5D0880650B05D5BA0AC9.taxon	distribution	Distribution. Cosmopolitan species, widespread in the Western Palearctic: Austria, the Azores, Belgium, Bulgaria, the Canary Islands, Czech Republic, Finland, France (locus typicus of Ganaspis monilicornis), Germany, Greece, Ireland, Israel, Italy, Madeira, the Netherlands, Norway, Slovenia, Spain, Sweden (locus typicus of Eucoila heterotoma), Switzerland, Tunisia, Turkey, and the United Kingdom. In Eastern Palearctic: Japan; in North America: eastern USA (locus typicus of Pseudeucoila bochei), and Canada; in tropical Asia: the Philippines (locus typicus of Erisphagia philippinensis), in the Afrotropics: Democratic Republic of the Congo, Madagascar, and St Helena; and in Australia and Vanuatu.	en	Vogel, Jonathan, Martin, Jakob, Forshage, Mattias, Salden, Tobias, Staverløkk, Arnstein, Verheyde, Fons, Nordlander, Göran, Herz, Annette, Peters, Ralph S. (2025): The Leptopilina Förster, 1869 of the Western Palearctic, and an updated list of the world species (Hymenoptera, Figitidae, Eucoilinae). Journal of Hymenoptera Research 98: 915-969, DOI: 10.3897/jhr.98.165583
89E8D9F073495BD684F794BA56F9F296.taxon	diagnosis	Diagnosis. Leptopilina japonica is a large species (usually around 2 mm ♀ body length) with a robust appearance and medium-long antennae (Fig. 7 A). The species is, together with L. boulardi, characteristic in having at most a few singular setae on the base of the metacoxa instead of a distinct setal patch (Fig. 7 D), which is typical for all other species. Leptopilina japonica differs from L. boulardi by the distinct metapleural ridges 1 and 2 that almost reach the anterior metapleural margin (Fig. 7 D). The extension of the ridges 1 and 2 is shared with L. heterotoma, while all other species have significantly shorter or no ridges. The mesoscutellar plate is similar in shape to that of L. heterotoma, being more elongate than those of the other species, but L. heterotoma has a typically rhombic mesoscutellar plate, while it is more drop-shaped, being widest in the posterior half, in L. japonica (Fig. 7 C). The shape is however rather variable and may overlap between the two species. The lateral view of the mesoscutellar plate usually appears s-shaped in L. heterotoma and is more evenly convex in L. japonica. The sculpture of the dorsal surface of the mesoscutellum is mediolaterally foveate-reticulate, as in most other species (Fig. 7 C). Only L. boulardi and L. heterotoma are areolate instead. The mesoscutellar surface sometimes has additional concentric striae (as in Fig. 7 C).	en	Vogel, Jonathan, Martin, Jakob, Forshage, Mattias, Salden, Tobias, Staverløkk, Arnstein, Verheyde, Fons, Nordlander, Göran, Herz, Annette, Peters, Ralph S. (2025): The Leptopilina Förster, 1869 of the Western Palearctic, and an updated list of the world species (Hymenoptera, Figitidae, Eucoilinae). Journal of Hymenoptera Research 98: 915-969, DOI: 10.3897/jhr.98.165583
89E8D9F073495BD684F794BA56F9F296.taxon	description	Molecular characterisation. Maximum intraspecific barcode-distance: 0.6 % (6). Minimum interspecific barcode-distance: 11 % (L. heterotoma). Consensus barcode sequence: 658 bp. 5 ’ - TGTAATGTATTTTGTTTTTGGTATTTGGTCTGGGATAGTGGGGGCTGGGTTAAGATTCCTTGTTCGTACAGAATTAGGGATACCTGGGCAGTTGATTAATAATGATCAAATTTATAATTCAATTGTAACTGCTCATGCTTTTGTTATAATTTTTTTTATAGTGATACCAATTATGGTTGGGGGGTTTGGCAATTATTTAGTACCATTAATATTAACTGTTCCTGACATAGCTTTCCCTCGATTGAATAATATAAGATTATGATTATTATTTCCTTCAATGATTTTAATGGTGGCAAGGATGATAATTGATCAAGGGGCAGGGACAGGGTGAACGGTTTATCCTCCTTTATCTTTAATAGATAGTCATCCTGGGGTTTCTACTGATTTAGTAATTTTTTCATTACATTTAAGAGGGGTATCTTCGATTTTAGGGTCAATTAATTTTATTTCTACTATTATTAATATACGTCCTTATTTAATAACAATAGATAAAATTACTTTATTTATTTGAGCTATTTTTTTAACAACAATTCTTTTATTATTATCTTTACCTGTTTTAGCAGGGGGGATTACTATATTATTATTTGATCGTAATTTAAATACTTCTTTTTATGATCCTGTTGGAGGGGGGGACCCAATTTTGTATCAACATTTATTT- 3 ’.	en	Vogel, Jonathan, Martin, Jakob, Forshage, Mattias, Salden, Tobias, Staverløkk, Arnstein, Verheyde, Fons, Nordlander, Göran, Herz, Annette, Peters, Ralph S. (2025): The Leptopilina Förster, 1869 of the Western Palearctic, and an updated list of the world species (Hymenoptera, Figitidae, Eucoilinae). Journal of Hymenoptera Research 98: 915-969, DOI: 10.3897/jhr.98.165583
89E8D9F073495BD684F794BA56F9F296.taxon	distribution	Distribution. Non-native species in the Western Palearctic, originally from East Asia. In Europe since 2019: present in Belgium (since 2022, new record), France (since 2022, Rousse et al. 2023), Germany (since 2021, Martin et al. 2023), Italy (since 2019, Puppato et al. 2020), Switzerland (since 2021, Rossi-Stacconi et al. 2025), and the United Kingdom (since 2024, Powell et al. 2025). In Asia present in China (Daane et al. 2016), Japan (Novković et al. 2011), and South Korea (Giorgini et al. 2019). It has spread to Canada since 2016 (Abram et al. 2020) and to the USA since 2020 (Beers et al. 2022). Records in 2023 and 2024 from Northern Germany in the state of Brandenburg represent the globally northernmost detections of this species so far (Rossi-Stacconi et al. 2025).	en	Vogel, Jonathan, Martin, Jakob, Forshage, Mattias, Salden, Tobias, Staverløkk, Arnstein, Verheyde, Fons, Nordlander, Göran, Herz, Annette, Peters, Ralph S. (2025): The Leptopilina Förster, 1869 of the Western Palearctic, and an updated list of the world species (Hymenoptera, Figitidae, Eucoilinae). Journal of Hymenoptera Research 98: 915-969, DOI: 10.3897/jhr.98.165583
9E9BDCA514485231A547C168B41F61A1.taxon	diagnosis	Diagnosis. Leptopilina longipes is a size-variable species with relatively slender appearance, medium-long antennae, and the metasoma is notably paler than the head and mesosoma (Fig. 8 A). The species is unique in having the propodeal carina well separated from the mesoscutellum by a part of the metanotum (Fig. 8 E). Other species show at most an insignificant gap. The metapleural ridge 1 and 2 are of similar length, reaching about half the length of the metapleuron (Fig. 8 D). In other species, the lengths of ridge 1 and 2 are dissimilar, or they reach the anterior margin of the metapleuron (L. japonica and L. heterotoma). Whereas other species have a more or less unicoloured body, the metasoma of L. longipes, especially of the females, is distinctly paler amber-coloured than the head and mesosoma (Fig. 8 A). This colouration pattern is similar to that of L. fimbriata, where it is usually even more distinct. The surface anterior to the glandular pit of the mesoscutellar plate is concave and areolate (Fig. 8 C), as it is in L. australis and L. clavipes. That area is mostly smooth in all other species. The female antenna is usually more uniformly dark, with all flagellomeres brown to dark brown (Fig. 8 B), while all other species have at least a few proximal flagellomeres pale brown or even yellow.	en	Vogel, Jonathan, Martin, Jakob, Forshage, Mattias, Salden, Tobias, Staverløkk, Arnstein, Verheyde, Fons, Nordlander, Göran, Herz, Annette, Peters, Ralph S. (2025): The Leptopilina Förster, 1869 of the Western Palearctic, and an updated list of the world species (Hymenoptera, Figitidae, Eucoilinae). Journal of Hymenoptera Research 98: 915-969, DOI: 10.3897/jhr.98.165583
9E9BDCA514485231A547C168B41F61A1.taxon	description	Molecular characterisation. Maximum intraspecific barcode-distance: 0.5 % (2). Minimum interspecific barcode-distance: 14.3 % (L. clavipes). Consensus barcode sequence: 658 bp. 5 ’ - TATAATATATTTTATATTTGGTATTTGATCAAGTATAGTAGGGGCAAGGCTAAGAATAATTATTCGAATAGAGTTAGGGACTGTAACTCAGTTAATTAATAATGATCAGATTTATAATTCTATTGTTACGGCTCATGCATTTGTAATAATTTTTTTTATAGTTATACCTATTATAGTGGGAGGGTTTGGTAATTATTTGGTTCCTTTAATAGTTAGAGTTCCTGACATAGCTTTTCCTCGTCTTAATAATATAAGGTTATGATTATTATTTCCTTCTTTAATTTTAATAATTACAAGAATATTTATTGATCAGGGGGCAGGGACTGGGTGAACGGTGTATCCTCCTTTATCTTTATCTATAAGGCATCCTGGTGTGGCACCTGATTTAGTAATTTTTTCTTTACATTTGAGGGGGGTATCTTCAATTTTAGGGGCAATTAATTTTATTACTACAATTATAAATATACGACCAAAAATAATATCTATAGATAAAATTTCTTTATTTGTTTGATCTATTTTTTTAACTACAATTTTACTTTTATTATCTTTACCTGTGTTAGCTGGAGGAATTACAATATTATTATTTGATCGTAATTTAAATACTTCTTTTTATGATCCWATTGGAGGGGGGGAWCCTATTTTGTATCARCATTTATTT- 3 ’.	en	Vogel, Jonathan, Martin, Jakob, Forshage, Mattias, Salden, Tobias, Staverløkk, Arnstein, Verheyde, Fons, Nordlander, Göran, Herz, Annette, Peters, Ralph S. (2025): The Leptopilina Förster, 1869 of the Western Palearctic, and an updated list of the world species (Hymenoptera, Figitidae, Eucoilinae). Journal of Hymenoptera Research 98: 915-969, DOI: 10.3897/jhr.98.165583
9E9BDCA514485231A547C168B41F61A1.taxon	distribution	Distribution. Restricted to the Palearctic; mainly northern and central Europe: found in Austria (locus typicus of Eucoila pusilla), Bulgaria, Croatia, Czech Republic, Denmark, Finland, Georgia, Germany (locus typicus of Cothonaspis longipes), Moldova (locus typicus of Rhoptromeris rutilus), the Netherlands, Norway, Sweden, Switzerland, and the United Kingdom. Outside of Europe only known from Japan.	en	Vogel, Jonathan, Martin, Jakob, Forshage, Mattias, Salden, Tobias, Staverløkk, Arnstein, Verheyde, Fons, Nordlander, Göran, Herz, Annette, Peters, Ralph S. (2025): The Leptopilina Förster, 1869 of the Western Palearctic, and an updated list of the world species (Hymenoptera, Figitidae, Eucoilinae). Journal of Hymenoptera Research 98: 915-969, DOI: 10.3897/jhr.98.165583
