taxonID	type	description	language	source
042C87BDFFD9FFB6FF3CBFA97608F852.taxon	diagnosis	Diagnosis. Chordeumatida with two pairs of setose angiocoxites on the gonopods; 9 th legs reduced to one, two, or three podomeres, coxae without prominate coxites; 10 th and 11 th legs with coxal glands. Components. Tingupa Chamberlin, 1910; Buotus Chamberlin, 1940; Blancosoma Shear & Hubbard, 1998.	en	Shear, William A., Shelley, Rowland M. (2007): Tingupa tlingitorum, n. sp., a new milliped from Haines, Alaska, USA, with notes on the generic distribution and a revised key to species (Chordeumatida: Tingupidae). Zootaxa 1393: 53-59, DOI: 10.5281/zenodo.175290
042C87BDFFD9FFB6FF3CBFA97608F852.taxon	distribution	Distribution. The Alaskan Panhandle; the Queen Charlotte Islands (QCI), BC, Canada; coastal Oregon; the Sierra Nevada of central California; northern Idaho and western Montana; northwestern Colorado; northern, southwestern, and southeastern Utah; southeastern Arizona; southwestern Illinois to northern Arkansas; and eastern West Virginia to northcentral North Carolina (Fig. 4) (Chamberlin 1910, 1925, 1928, 1940; Chamberlin & Hoffman 1958; Loomis & Schmitt 1971; Shear 1972, 1981, map 1; Shear & Hubbard 1998; Hoffman 1999). Southeastern Utah is based on the obscure record of two females from Devil's Canyon, San Juan County (Chamberlin 1928) that has been missed by previous authors; its generic and specific identities are unknown.	en	Shear, William A., Shelley, Rowland M. (2007): Tingupa tlingitorum, n. sp., a new milliped from Haines, Alaska, USA, with notes on the generic distribution and a revised key to species (Chordeumatida: Tingupidae). Zootaxa 1393: 53-59, DOI: 10.5281/zenodo.175290
042C87BDFFD9FFB6FF3CBFA97608F852.taxon	discussion	Remarks. Tingupidae, an endemic North American milliped family, was revised by Shear (1981). He and Hoffman (1980) recognized two genera, Tingupa and Buotus, the latter being monotypic and transferred into the family by Shelley (1976) after being misplaced in the order Polyzoniida, family Polyzoniidae (Chamberlin 1940; Chamberlin & Hoffman 1958). Shear and Hubbard (1998) added a second monotypic genus, Blancosoma, and provided a key to the three component genera. Tingupidae is closely related to the monotypic family, Niponiosomatidae Verhoeff, 1941, in Japan, and the two belong to the superfamily Brannerioidea Cook, 1896, suborder Craspedosomatidea Brolemann, 1935, along with eight other families (Shear 2000; Shelley 2003). Shear (1988) maintained both Tingupidae and Niponiosomatidae primarily because of somatic apomorphies of Tingupa (paranota and tergal microsculpture), even though the gonopod structure of Niponiosoma Verhoeff, 1941, is closer to that of Tingupa than those of either Buotus or Blancosoma. However, Blancosoma does not possess these somatic features and thus resembles Niponiosoma. Niponiosomatidae holds taxonomic priority over the more widespread and utilized Tingupidae, so synonymization would have the undesirable consequence of submerging this continental North American taxon under the geographically restricted and little­used Japanese name.	en	Shear, William A., Shelley, Rowland M. (2007): Tingupa tlingitorum, n. sp., a new milliped from Haines, Alaska, USA, with notes on the generic distribution and a revised key to species (Chordeumatida: Tingupidae). Zootaxa 1393: 53-59, DOI: 10.5281/zenodo.175290
042C87BDFFDAFFB5FF3CBDFC75FAFBE5.taxon	diagnosis	Diagnosis. Tingupidae with prominent, rounded paranota, metaterga with microsculpture of low tubercles interspersed among short, sharp ridges (see Shear 1981, figs. 1 – 2).	en	Shear, William A., Shelley, Rowland M. (2007): Tingupa tlingitorum, n. sp., a new milliped from Haines, Alaska, USA, with notes on the generic distribution and a revised key to species (Chordeumatida: Tingupidae). Zootaxa 1393: 53-59, DOI: 10.5281/zenodo.175290
042C87BDFFDAFFB5FF3CBDFC75FAFBE5.taxon	distribution	Distribution. Same as that of the family but excluding Colorado, West Virginia, Virginia, and North Carolina (Fig. 4). Idaho and the Queen Charlotte Islands are based on the following samples of females that are unidentifiable to species; the latter site is some 450 mi (720 km) south­southwest of Haines. USA: IDAHO, Valley Co., 4 mi (6.4 km) NE McCall, Ψ, 18 October 1944, W. Ivie (NMNH). New State Record.	en	Shear, William A., Shelley, Rowland M. (2007): Tingupa tlingitorum, n. sp., a new milliped from Haines, Alaska, USA, with notes on the generic distribution and a revised key to species (Chordeumatida: Tingupidae). Zootaxa 1393: 53-59, DOI: 10.5281/zenodo.175290
042C87BDFFDAFFB5FF3CBDFC75FAFBE5.taxon	description	CANADA: BRITISH COLUMBIA: QUEEN CHARLOTTE ISLANDS, Graham I., Queen Charlotte City, 3 Ψ, 9 October 1948, G. J. Spencer (RBCM). New Country and Provincial Record.	en	Shear, William A., Shelley, Rowland M. (2007): Tingupa tlingitorum, n. sp., a new milliped from Haines, Alaska, USA, with notes on the generic distribution and a revised key to species (Chordeumatida: Tingupidae). Zootaxa 1393: 53-59, DOI: 10.5281/zenodo.175290
042C87BDFFDAFFB5FF3CBDFC75FAFBE5.taxon	discussion	Remarks. Tingupa comprises ten species, plus the new one described herein, but only nine are keyed in Shear (1981) and cited by Hoffman (1999), who both missed T. intergerina Loomis & Schmitt, 1971, in western Montana. Six species inhabit the northwestern Pacific Coast in the " lower 48 " from east­central California to northwestern Oregon; one species occurs in western Montana; the type­species, ostensibly with two subspecies, occupies the Wasatch Mountains and Parowan Valley of Utah; another nominal species, probably a synonym of T. utahensis, inhabits the Santa Catalina, Pinaleno, and Chiricahua Mountains of southeastern Arizona; and a troglobitic species occupies limestone caves in southwestern Illinois, Missouri, and northern Arkansas (Chamberlin 1910, 1925, 1928; Chamberlin & Hoffman 1958; Loomis & Schmitt 1971; Shear 1972, 1981; Hoffman 1999).	en	Shear, William A., Shelley, Rowland M. (2007): Tingupa tlingitorum, n. sp., a new milliped from Haines, Alaska, USA, with notes on the generic distribution and a revised key to species (Chordeumatida: Tingupidae). Zootaxa 1393: 53-59, DOI: 10.5281/zenodo.175290
042C87BDFFDBFFB3FF3CBDFC7175FE6D.taxon	materials_examined	Type specimens. ɗ holotype and Ψ paratype (NCSM) collected by R. M. Shelley, 21 August 2006, in Tlingit Park, Haines, Haines Borough, Alaska; coordinates are N 59 ° 13 ’ 55.4 ”, W 135 ° 26 ’ 38.1 ”.	en	Shear, William A., Shelley, Rowland M. (2007): Tingupa tlingitorum, n. sp., a new milliped from Haines, Alaska, USA, with notes on the generic distribution and a revised key to species (Chordeumatida: Tingupidae). Zootaxa 1393: 53-59, DOI: 10.5281/zenodo.175290
042C87BDFFDBFFB3FF3CBDFC7175FE6D.taxon	diagnosis	Diagnosis: A small depigmented species with three depigmented ocelli arranged in single row; mesal angiocoxites of gonopods with three long, attenuated, sigmoidally curved, immobile macrosetae apically, lateral angiocoxites with both long and short, hooklike immobile macrosetae distad (Fig. 1).	en	Shear, William A., Shelley, Rowland M. (2007): Tingupa tlingitorum, n. sp., a new milliped from Haines, Alaska, USA, with notes on the generic distribution and a revised key to species (Chordeumatida: Tingupidae). Zootaxa 1393: 53-59, DOI: 10.5281/zenodo.175290
042C87BDFFDBFFB3FF3CBDFC7175FE6D.taxon	materials_examined	Holotype. Length 5.0 mm, width ca. 0.35 mm. Segments with small paranota; segmental setae clavate, longer anteriorly and posteriorly, becoming less clavate and more acuminate posteriorly. Antennae with 5 th article inflated. Gonopods (Figs. 1,2) of typical structure; lateral angiocoxites (la) basally swollen, setose, with three or four long, attenuated, immobile macrosetae anterodistad and three strong, decurved, hooklike, immobile macrosetae on posterior surfaces of apices; mesal angiocoxites (ma) narrow, projecting anteriorly, closely appressed and basally fused, bases bulbous, shafts setose with long, attenuated, gently curved, immobile macrosetae arising from anterior surfaces at about 2 / 3 lengths, apices with three attenuated, sigmoidally curved, immobile macrosetae each. Anterior division of colpocoxite divided into three processes (Fig. 3, a, b, c), posterior division poorly sclerotized, bulbous and constricted in midline, lower part with fine, apparently unsocketed hairs. Ninth legs (Fig. 3) with broad, short coxae bearing anterior cuticular fimbriae and short, medially toothed processes; prefemora about twice as long as distal articles, latter pyriform.	en	Shear, William A., Shelley, Rowland M. (2007): Tingupa tlingitorum, n. sp., a new milliped from Haines, Alaska, USA, with notes on the generic distribution and a revised key to species (Chordeumatida: Tingupidae). Zootaxa 1393: 53-59, DOI: 10.5281/zenodo.175290
042C87BDFFDBFFB3FF3CBDFC7175FE6D.taxon	description	Female paratype: Length 5.2 mm, width ca. 0.40 mm, nonsexual characters as in male. Habitat. The type specimens were discovered in Tlingit Park under logs and debris on damp, " mucky " substrate in a shallow ditch that appears to carry overflow water periodically. The ditch runs down a slope into a small wooded area but is bordered on each side by a row of cottonwood trees (Populus balsamifera) along the stretch where RMS discovered the specimens.	en	Shear, William A., Shelley, Rowland M. (2007): Tingupa tlingitorum, n. sp., a new milliped from Haines, Alaska, USA, with notes on the generic distribution and a revised key to species (Chordeumatida: Tingupidae). Zootaxa 1393: 53-59, DOI: 10.5281/zenodo.175290
042C87BDFFDBFFB3FF3CBDFC7175FE6D.taxon	distribution	Distribution. Known only from the type locality.	en	Shear, William A., Shelley, Rowland M. (2007): Tingupa tlingitorum, n. sp., a new milliped from Haines, Alaska, USA, with notes on the generic distribution and a revised key to species (Chordeumatida: Tingupidae). Zootaxa 1393: 53-59, DOI: 10.5281/zenodo.175290
042C87BDFFDBFFB3FF3CBDFC7175FE6D.taxon	etymology	Etymology. The species epithet is an adjective referring to the Tlingit tribe of native Alaskans.	en	Shear, William A., Shelley, Rowland M. (2007): Tingupa tlingitorum, n. sp., a new milliped from Haines, Alaska, USA, with notes on the generic distribution and a revised key to species (Chordeumatida: Tingupidae). Zootaxa 1393: 53-59, DOI: 10.5281/zenodo.175290
