identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
042C87BDFFD9FFB6FF3CBFA97608F852.text	042C87BDFFD9FFB6FF3CBFA97608F852.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tingupidae Loomis 1966	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Family  Tingupidae Loomis, 1966</p>
            <p> Tingupidae Loomis, 1966:227 . Shear, 1969:140; 1972:264; 1981:5­6. Loomis &amp; Schmitt, 1971:128. Peck &amp; Lewis, 1977:49. Hoffman, 1980:135; 1999:253. Kevan, 1983:2967. Gardner, 1986:34. Shear &amp; Hubbard, 1998:86 ­88. Shelley et al., 2000:79. </p>
            <p> Diagnosis.  Chordeumatida with two pairs of setose angiocoxites on the gonopods; 9th legs reduced to one, two, or three podomeres, coxae without prominate coxites; 10th and 11th legs with coxal glands. </p>
            <p> Components.  Tingupa Chamberlin, 1910 ;  Buotus Chamberlin, 1940 ;  Blancosoma Shear &amp; Hubbard, 1998 . </p>
            <p>Distribution. The Alaskan Panhandle; the Queen Charlotte Islands (QCI), BC, Canada; coastal Oregon; the Sierra Nevada of central California; northern Idaho and western Montana; northwestern Colorado; northern, southwestern, and southeastern Utah; southeastern Arizona; southwestern Illinois to northern Arkansas; and eastern West Virginia to northcentral North Carolina (Fig. 4) (Chamberlin 1910, 1925, 1928, 1940; Chamberlin &amp; Hoffman 1958; Loomis &amp; Schmitt 1971; Shear 1972, 1981, map 1; Shear &amp; Hubbard 1998; Hoffman 1999). Southeastern Utah is based on the obscure record of two females from Devil's Canyon, San Juan County (Chamberlin 1928) that has been missed by previous authors; its generic and specific identities are unknown.</p>
            <p> Remarks.  Tingupidae , an endemic North American milliped family, was revised by Shear (1981). He and Hoffman (1980) recognized two genera,  Tingupa and  Buotus , the latter being monotypic and transferred into the family by Shelley (1976) after being misplaced in the order  Polyzoniida , family  Polyzoniidae (Chamberlin 1940; Chamberlin &amp; Hoffman 1958). Shear and Hubbard (1998) added a second monotypic genus,  Blancosoma , and provided a key to the three component genera.  Tingupidae is closely related to the monotypic family,  Niponiosomatidae Verhoeff, 1941 , in Japan, and the two belong to the superfamily Brannerioidea Cook, 1896, suborder Craspedosomatidea Brolemann, 1935, along with eight other families (Shear 2000; Shelley 2003). Shear (1988) maintained both  Tingupidae and  Niponiosomatidae primarily because of somatic apomorphies of  Tingupa (paranota and tergal microsculpture), even though the gonopod structure of  Niponiosoma Verhoeff, 1941 , is closer to that of  Tingupa than those of either  Buotus or  Blancosoma . However,  Blancosoma does not possess these somatic features and thus resembles  Niponiosoma .  Niponiosomatidae holds taxonomic priority over the more widespread and utilized  Tingupidae , so synonymization would have the undesirable consequence of submerging this continental North American taxon under the geographically restricted and little­used Japanese name. </p>
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	https://treatment.plazi.org/id/042C87BDFFD9FFB6FF3CBFA97608F852	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shear, William A.;Shelley, Rowland M.	Shear, William A., Shelley, Rowland M. (2007): Tingupa tlingitorum, n. sp., a new milliped from Haines, Alaska, USA, with notes on the generic distribution and a revised key to species (Chordeumatida: Tingupidae). Zootaxa 1393: 53-59, DOI: 10.5281/zenodo.175290
042C87BDFFDAFFB5FF3CBDFC75FAFBE5.text	042C87BDFFDAFFB5FF3CBDFC75FAFBE5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tingupa Chamberlin 1910	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Genus  Tingupa Chamberlin, 1910</p>
            <p> Tingupa Chamberlin, 1910:238 . Chamberlin &amp; Hoffman, 1958:106. Buckett, 1964:14. Shear, 1969:141; 1972:265; 1981:6­7. Jeekel, 1971:82. Hoffman, 1980:135; 1999:254­255. Kevan, 1983:2967. Shelley et al., 2000:79. Type­species.  T. utahensis Chamberlin, 1910 , by original designation. </p>
            <p> Diagnosis.  Tingupidae with prominent, rounded paranota, metaterga with microsculpture of low tubercles interspersed among short, sharp ridges (see Shear 1981, figs. 1–2). </p>
            <p>Distribution. Same as that of the family but excluding Colorado, West Virginia, Virginia, and North Carolina (Fig. 4). Idaho and the Queen Charlotte Islands are based on the following samples of females that are unidentifiable to species; the latter site is some 450 mi (720 km) south­southwest of Haines.</p>
            <p>USA: IDAHO, Valley Co., 4 mi (6.4 km) NE McCall, Ψ, 18 October 1944, W. Ivie (NMNH). New State Record.</p>
            <p> CANADA: BRITISH  COLUMBIA : QUEEN CHARLOTTE ISLANDS, Graham I., Queen Charlotte City, 3Ψ, 9 October 1948, G. J. Spencer (RBCM). New Country and Provincial Record. </p>
            <p> Remarks.  Tingupa comprises ten species, plus the new one described herein, but only nine are keyed in Shear (1981) and cited by Hoffman (1999), who both missed  T. intergerina Loomis &amp; Schmitt, 1971 , in western Montana. Six species inhabit the northwestern Pacific Coast in the "lower 48" from east­central California to northwestern Oregon; one species occurs in western Montana; the type­species, ostensibly with two subspecies, occupies the Wasatch Mountains and Parowan Valley of Utah; another nominal species, probably a synonym of  T. utahensis , inhabits the Santa  Catalina, Pinaleno , and Chiricahua Mountains of southeastern Arizona; and a troglobitic species occupies limestone caves in southwestern Illinois, Missouri, and northern Arkansas (Chamberlin 1910, 1925, 1928; Chamberlin &amp; Hoffman 1958; Loomis &amp; Schmitt 1971; Shear 1972, 1981; Hoffman 1999). </p>
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	https://treatment.plazi.org/id/042C87BDFFDAFFB5FF3CBDFC75FAFBE5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shear, William A.;Shelley, Rowland M.	Shear, William A., Shelley, Rowland M. (2007): Tingupa tlingitorum, n. sp., a new milliped from Haines, Alaska, USA, with notes on the generic distribution and a revised key to species (Chordeumatida: Tingupidae). Zootaxa 1393: 53-59, DOI: 10.5281/zenodo.175290
042C87BDFFDBFFB3FF3CBDFC7175FE6D.text	042C87BDFFDBFFB3FF3CBDFC7175FE6D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tingupa tlingitorum	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Tingupa tlingitorum ,  n. sp.</p>
            <p>Figs. 1–3</p>
            <p>Type specimens. ɗ holotype and Ψ paratype (NCSM) collected by R. M. Shelley, 21 August 2006, in Tlingit Park, Haines, Haines Borough, Alaska; coordinates are N59°13’55.4”, W135°26’38.1”.</p>
            <p>Diagnosis: A small depigmented species with three depigmented ocelli arranged in single row; mesal angiocoxites of gonopods with three long, attenuated, sigmoidally curved, immobile macrosetae apically, lateral angiocoxites with both long and short, hooklike immobile macrosetae distad (Fig. 1).</p>
            <p>Holotype. Length 5.0 mm, width ca. 0.35 mm. Segments with small paranota; segmental setae clavate, longer anteriorly and posteriorly, becoming less clavate and more acuminate posteriorly. Antennae with 5th article inflated. Gonopods (Figs. 1,2) of typical structure; lateral angiocoxites (la) basally swollen, setose, with three or four long, attenuated, immobile macrosetae anterodistad and three strong, decurved, hooklike, immobile macrosetae on posterior surfaces of apices; mesal angiocoxites (ma) narrow, projecting anteriorly, closely appressed and basally fused, bases bulbous, shafts setose with long, attenuated, gently curved, immobile macrosetae arising from anterior surfaces at about 2/3 lengths, apices with three attenuated, sigmoidally curved, immobile macrosetae each. Anterior division of colpocoxite divided into three processes (Fig. 3, a, b, c), posterior division poorly sclerotized, bulbous and constricted in midline, lower part with fine, apparently unsocketed hairs. Ninth legs (Fig. 3) with broad, short coxae bearing anterior cuticular fimbriae and short, medially toothed processes; prefemora about twice as long as distal articles, latter pyriform.</p>
            <p>1. According to Shear (1981), these two nominal species are likely synonyms.</p>
            <p>Female paratype: Length 5.2 mm, width ca. 0.40 mm, nonsexual characters as in male.</p>
            <p> Habitat. The type specimens were discovered in Tlingit Park under logs and debris on damp, "mucky" substrate in a shallow ditch that appears to carry overflow water periodically. The ditch runs down a slope into a small wooded area but is bordered on each side by a row of cottonwood trees (  Populus balsamifera ) along the stretch where RMS discovered the specimens. </p>
            <p>Distribution. Known only from the type locality.</p>
            <p>Etymology. The species epithet is an adjective referring to the Tlingit tribe of native Alaskans.</p>
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	https://treatment.plazi.org/id/042C87BDFFDBFFB3FF3CBDFC7175FE6D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shear, William A.;Shelley, Rowland M.	Shear, William A., Shelley, Rowland M. (2007): Tingupa tlingitorum, n. sp., a new milliped from Haines, Alaska, USA, with notes on the generic distribution and a revised key to species (Chordeumatida: Tingupidae). Zootaxa 1393: 53-59, DOI: 10.5281/zenodo.175290
