identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
5549718A75455698841B516DFA367939.text	5549718A75455698841B516DFA367939.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heptathela aha	<div><p>Heptathela aha sp. nov. Fig. 15</p> <p>Type material.</p> <p>Holotype: JAPAN · ♀; Okinawa-ken, Iheyajima Island, Mt. Aha-dake; 27.02N, 127.93E; alt. 10 m; 26 December 2012; D. Li, F.X. Liu and X. Xu leg.; XUX-2012-502.</p> <p>Paratypes: JAPAN · 6 ♀♀; same data as for holotype; XUX-2012-504 to 519.</p> <p>Diagnosis.</p> <p>Females of H. aha sp. nov. cannot be distinguished morphologically from those of H. gayozan sp. nov. but can be distinguished from those of H. kubayama sp. nov. by the receptacular clusters without genital stalks; and from those of H. mae sp. nov. by the inner receptacular clusters similar to or larger than laterals (Fig. 15 A–F). Heptathela aha sp. nov. can also be diagnosed from all other Okinawa group Heptathela species by the following unique nucleotide substitutions in the standard DNA barcode alignment: C (41), C (179), G (182), G (233), T (248), G (251), T (326), A (347), G (359), C (473), C (492), A (536), T (651).</p> <p>Description.</p> <p>Female (Holotype). Carapace brown; opisthosoma brown, brown tergites with black plaques; cheliceral groove with 12 pronounced denticles; seven spinnerets. Measurements: BL 12.60, CL 5.55, CW 5.00, OL 6.82, OW 5.50; ALE&gt; PLE&gt; PME&gt; AME; palp 9.90 (3.40 + 1.71 + 2.18 + 2.61), leg I 11.34 (3.31 + 2.05 + 2.31 + 2.35 + 1.32), leg II 11.65 (3.51 + 1.98 + 2.03 + 2.53 + 1.60), leg III 11.92 (3.30 + 1.90 + 2.05 + 2.97 + 1.70), leg IV 16.96 (4.88 + 2.22 + 2.98 + 34.45 + 2.43).</p> <p>Female genitalia. A pair of depressions on the ventro-lateral part of the genital atrium (Fig. 15 D–F). A pair of receptacular clusters along the anterior margin of bursa copulatrix, divided into two parts, both without genital stalks (Fig. 15 A–F).</p> <p>Male. Unknown.</p> <p>Etymology.</p> <p>The species epithet, a noun in apposition, refers to the type locality.</p> <p>Distribution.</p> <p>The species is endemic to the Japanese island Iheyajima (Fig. 1C).</p></div> 	https://treatment.plazi.org/id/5549718A75455698841B516DFA367939	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Xu, Xin;Ono, Hirotsugu;Kuntner, Matjaz;Liu, Fengxiang;Li, Daiqin	Xu, Xin, Ono, Hirotsugu, Kuntner, Matjaz, Liu, Fengxiang, Li, Daiqin (2019): A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands. ZooKeys 888: 1-50, DOI: http://dx.doi.org/10.3897/zookeys.888.34494, URL: http://dx.doi.org/10.3897/zookeys.888.34494
2F39360A6A615384B97F5C9A29D62EB3.text	2F39360A6A615384B97F5C9A29D62EB3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heptathela amamiensis Haupt 1983	<div><p>Heptathela amamiensis Haupt, 1983 Fig. 9</p> <p>Heptathela kimurai amamiensis Haupt, 1983: 283 (holotype: female, from Naze, Amami-oshima, Japan, collected by J. Haupt on 26 March 1980, deposited in ZMH, holotype presumably lost (Dunlop et al. 2014); Haupt 2003: 69. Heptathela amamiensis: Ono and Nishikawa 1989: 120; Ono 2009: 80; Ono and Ogata 2018: 27, 479.</p> <p>Diagnosis.</p> <p>Males of H. amamiensis differ from those of H. sumiyo sp. nov. by the wider saddle-shaped embolus in the prolateral view, and the narrower conductor base in the ventral view (Fig. 9G, J–L); from those of H. kanenoi and H. kojima sp. nov. by the spiniform conductor apex (Fig. 9F, I, J). Females of H. amamiensis resemble those of other Amami group Heptathela species but can be distinguished from those of H. kanenoi by the tuberculate outer receptacular clusters (Fig. 9 B–E). H. amamiensis can also be diagnosed from all other Amami group Heptathela species by the following unique nucleotide substitutions in the standard DNA barcode alignment: C (89), A (179), A (194), T (215), T (218), C (273), A (281), C (284), A (327), G (332), G (362), C (467), C (543), C (647).</p> <p>Description.</p> <p>Male. Carapace brown; opisthosoma light brown, with dark brown tergites; cheliceral groove with 13 denticles; 8 spinnerets. Measurements: BL 12.85, CL 6.50, CW 5.98, OL 6.65, OW 4.00; ALE&gt; PLE&gt; PME&gt; AME; leg I 18.25 (5.00 + 2.45 + 3.80 + 4.60 + 2.40), leg II 18.75 (4.85 + 2.40 + 3.75 + 5.00 + 2.75), leg III 20.10 (4.90 + 2.60 + 3.60 + 5.90 + 3.10), leg IV 25.30 (6.00 + 2.60 + 5.10 + 7.70 + 3.90).</p> <p>Palp. Prolateral side of paracymbium unpigmented and unsclerotised, numerous setae and spines at the tip of paracymbium (Fig. 9 F–H). Contrategulum with serrated margin (Fig. 9F, G, I, J). Tegulum wide, with dentate dorsal extension of terminal apophysis (Fig. 9H, L) and blunt terminal apophysis (Fig. 9H, K, L). Conductor sclerotised and rugose, with several folds and a spiniform apex (Fig. 9 F–L). Embolus sclerotised, with a wide and flat opening, the distal part slightly sclerotised, and saddle-shaped in the prolateral view (Fig. 9 F–L).</p> <p>Females (N = 5). Carapace and opisthosoma colour as in male; cheliceral groove with 12-14 pronounced denticles; tergites similar to those of male; seven or eight spinnerets. Measurements: BL 11.08-16.90, CL 5.45-6.90, CW 4.70-6.20, OL 6.61-9.10, OW 4.70-7.38; ALE&gt; PLE&gt; PME&gt; AME; palp 12.36 (4.12 + 2.11 + 2.75 + 3.38), leg I 14.12 (4.52 + 2.50 + 2.70 + 2.83 + 1.57), leg II 14.06 (4.25 + 2.37 + 2.61 + 3.15 + 1.68), leg III 15.13 (4.27 + 2.56 + 2.55 + 3.55 + 2.20), leg IV 21.86 (5.84 + 2.91 + 4.08 + 5.81 + 3.22).</p> <p>Female genitalia. A pair of indistinct depressions on the ventro-lateral part of genital atrium (Fig. 9C, D). Paired receptacular clusters along the anterior margin of bursa copulatrix, divided into two parts, the inners ovate, the outers tuberculate, without genital stalks (Fig. 9A-E).</p> <p>Material examined.</p> <p>JAPAN · 2 ♂♂, 5 ♀♀; Kagoshima-ken, Amami-Oshima, Amami-shi, Nazehirata-cho, Michinoshima Loop Bridge; 28.36N, 129.50E; alt. 60 m; 15 September 2013; D. Li and B. Wu leg.; XUX-2013-276, 278, 281 to 285.</p> <p>Distribution.</p> <p>The species is known from the Japanese island Amamioshima (Fig. 1C).</p></div> 	https://treatment.plazi.org/id/2F39360A6A615384B97F5C9A29D62EB3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Xu, Xin;Ono, Hirotsugu;Kuntner, Matjaz;Liu, Fengxiang;Li, Daiqin	Xu, Xin, Ono, Hirotsugu, Kuntner, Matjaz, Liu, Fengxiang, Li, Daiqin (2019): A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands. ZooKeys 888: 1-50, DOI: http://dx.doi.org/10.3897/zookeys.888.34494, URL: http://dx.doi.org/10.3897/zookeys.888.34494
38088D004F7D55FD8F637E6FF5900E8B.text	38088D004F7D55FD8F637E6FF5900E8B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heptathela crypta	<div><p>Heptathela crypta sp. nov. Figs 22, 23</p> <p>Type material.</p> <p>Holotype: JAPAN · ♂; Okinawa-ken, Nago-shi, Haneiji-Dam, Taira; 26.59N, 128.03E; alt. 100 m; 18 December 2012; D. Li, F.X. Liu and X. Xu leg.; XUX-2012-328.</p> <p>Paratypes: JAPAN · 1 ♂, 3 ♀♀; same data as for holotype; XUX-2012-324, 326, 327, 333A · 2 ♂♂, 4 ♀♀; Okinawa Prefecture, Nago-shi, County Road 18 south, Nago/Yofuke; 26.57N, 128.01E; alt. 150 m; 24 December 2012; D. Li, F.X. Liu and X. Xu leg.; XUX-2012-457 to 462 · 3 ♀♀; Okinawa Prefecture, Nago-shi, Mt. Nago-dake; 26.58N, 128.01E; alt. 220 m; 06 May 2014; D. Li and B. Wu leg.; XUX-2014-027 to 027B.</p> <p>Diagnosis.</p> <p>Males and females of H. crypta sp. nov. cannot be distinguished morphologically from H. unten sp. nov. (Figs 22 A–K, 23 A–L), but can be diagnosed from H. unten sp. nov. by the following unique nucleotide substitutions in the standard DNA barcode alignment: C (26), A (32), C (50), T (60), T (110), G (153), T (194), C (197), T (269), C (281), T (284), C (338), A (341), T (357), C (416), T (428), C (458), A (482), T (488), G (551), T (581), T (635), G (638), G (641), C (644), C (656), as well as from all other Okinawa group Heptathela species by the following unique nucleotide substitutions in the standard DNA barcode alignment: C (26), T (110), G (551), C (656).</p> <p>Description.</p> <p>Male (Holotype). Carapace and opisthosoma description see H. unten sp. nov.; cheliceral groove with nine denticles of variable size; seven spinnerets. Measurements: BL 7.88, CL 4.01, CW 3.51, OL 4.23, OW 3.18; ALE&gt; PLE&gt; PME&gt; AME; leg I 10.15 (3.60 + 1.50 + 2.38 + 1.00 + 1.67), leg II 13.08 (3.48 + 1.58 + 2.49 + 3.53 + 2.00), leg III 14.27 (3.38 + 1.55 + 2.65 + 4.30 + 2.39), leg IV 18.19 (4.45 + 1.63 + 3.50 + 5.50 + 3.11).</p> <p>Palp. Prolateral side of paracymbium unpigmented and unsclerotised, numerous setae and spines at the tip of paracymbium (Fig. 22 I–K). Contrategulum margin incurved nearly in the middle, and the contrategulum divided into proximally serrated and distally smooth (Fig. 22E, F, I, J). Tegulum with wide dorsal extension of terminal apophysis (Fig. 22G, H, J), blunt terminal and small marginal apophysis (Fig. 22H, I). Conductor sclerotised and ovate, prolateral conductor with one or two shallow folds, and with a serrated margin (Fig. 22F, G, K). Embolus sclerotised, with a wide opening, the distal margin slightly sclerotised, and with a saddle-shaped margin in the retrolateral view (Fig. 22F, G, K).</p> <p>Females (N = 10). Carapace and opisthosoma description see H. unten sp. nov.; chelicerae with promargin of cheliceral groove with 13-14 pronounced denticles of variable size; seven spinnerets. Measurements: BL 8.35-16.50, CL 4.07-5.10, CW 3.30-4.80, OL 4.70-6.80, OW 3.00-5.20; ALE&gt; PLE&gt; PME&gt; AME; palp 7.70 (2.87 + 1.13 + 1.68 + 2.02), leg I 9.57 (3.07 + 1.70 + 1.70 + 1.98 + 1.12), leg II 9.64 (2.95 + 1.68 + 1.61 + 2.08 + 1.32), leg III 9.60 (2.68 + 1.69 + 1.50 + 2.30 + 1.43), leg IV 14.21 (4.00 + 1.92 + 2.55 + 3.83 + 1.91).</p> <p>Female genitalia. A pair of depressions on the ventro-lateral part of genital atrium indistinct (Fig. 23F, J–L). Paired receptacular clusters along the anterior margin of bursa copulatrix, divided into two parts, the inners similar or smaller than the laterals, paired receptacular clusters tuberculate, without genital stalks (Fig. 23C, G–L).</p> <p>Etymology.</p> <p>The species epithet, a noun in apposition, refers to the cryptic nature of this species discovery.</p> <p>Distribution.</p> <p>The species is endemic to the Japanese island Okinawajima (Fig. 1C).</p></div> 	https://treatment.plazi.org/id/38088D004F7D55FD8F637E6FF5900E8B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Xu, Xin;Ono, Hirotsugu;Kuntner, Matjaz;Liu, Fengxiang;Li, Daiqin	Xu, Xin, Ono, Hirotsugu, Kuntner, Matjaz, Liu, Fengxiang, Li, Daiqin (2019): A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands. ZooKeys 888: 1-50, DOI: http://dx.doi.org/10.3897/zookeys.888.34494, URL: http://dx.doi.org/10.3897/zookeys.888.34494
2074720837D45BBEBA644BB9719F9647.text	2074720837D45BBEBA644BB9719F9647.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heptathela gayozan	<div><p>Heptathela gayozan sp. nov. Fig. 16</p> <p>Type material.</p> <p>Holotype: JAPAN · ♀; Okinawa-ken, Iheyajima Island, Mt. Gayozan; 27.02N, 127.97E; alt. 25 m; 27 December 2012; D. Li, F.X. Liu and X. Xu leg.; XUX-2012-511.</p> <p>Paratypes: JAPAN · 2 ♀♀; same data as for holotype; XUX-2012-513, 515.</p> <p>Diagnosis.</p> <p>Females H. gayozan sp. nov. cannot be distinguished morphologically from those of H. aha sp. nov. but can be distinguished from those of H. kubayama sp. nov. by the receptacular clusters without genital stalks; and from those of H. mae sp. nov. by the inner receptacular clusters that are equal in size, or slightly larger than laterals (Fig. 16 A–D). H. gayozan sp. nov. can be diagnosed from all other Okinawa group Heptathela species by the following unique nucleotide substitutions in the standard DNA barcode alignment: G (38), G (41), G (122), C (203), C (365), C (452), C (470), T (518), C (527), C (533), T (560), T (653).</p> <p>Description.</p> <p>Female (holotype). Carapace brown; opisthosoma light brown, with dark brown tergites close to each other; cheliceral groove with 12 denticles; seven spinnerets. Measurements: BL 10.00, CL 4.40, CW 3.88, OL 4.84, OW 3.48; ALE&gt; PLE&gt; PME&gt; AME; palp 7.32 (2.72 + 1.50 + 1.80 + 1.30), leg I 9.61 (2.95 + 1.62 + 1.82 + 1.97 + 1.25), leg II 9.38 (2.80 + 1.58 + 1.71 + 2.07 + 1.22), leg III 9.60 (2.65 + 1.55 + 1.62 + 2.28 + 1.50), leg IV 13.89 (3.75 + 1.82 + 2.52 + 3.75 + 2.05).</p> <p>Female genitalia. A pair of depressions on the ventro-lateral part of genital atrium (Fig. 16C, D). A pair of receptacular clusters along the anterior margin of bursa copulatrix, divided into two parts, with several granules, without genital stalks (Fig. 16).</p> <p>Male. Unknown.</p> <p>Etymology.</p> <p>The species epithet, a noun in apposition, refers to the type locality.</p> <p>Distribution.</p> <p>The species is endemic to the Japanese island Iheyajima (Fig. 1C).</p></div> 	https://treatment.plazi.org/id/2074720837D45BBEBA644BB9719F9647	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Xu, Xin;Ono, Hirotsugu;Kuntner, Matjaz;Liu, Fengxiang;Li, Daiqin	Xu, Xin, Ono, Hirotsugu, Kuntner, Matjaz, Liu, Fengxiang, Li, Daiqin (2019): A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands. ZooKeys 888: 1-50, DOI: http://dx.doi.org/10.3897/zookeys.888.34494, URL: http://dx.doi.org/10.3897/zookeys.888.34494
D5DE01B497B9536EADD01443A190F9B8.text	D5DE01B497B9536EADD01443A190F9B8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heptathela helios Tanikawa & Miyashita 2014	<div><p>Heptathela helios Tanikawa &amp; Miyashita, 2014 Fig. 24</p> <p>Heptathela helios Tanikawa &amp; Miyashita, 2014: 68 (holotype: male (NSMT-Ar 12851), from Kunigami-son, Okinawajima, Japan, collected by A. Tanikawa on 26 May 2010, matured on 9 September 2012, deposited in NMNS, examined).</p> <p>Diagnosis.</p> <p>Males of H. helios can be distinguished from those of all other Okinawa group Heptathela species by the serrated contrategulum margin and the hooked tegular marginal apophysis, the ovate, indistinctly rugose conductor with a poorly serrated margin (Fig. 24 D–G). Females of H. helios can be distinguished from those of all other Heptathela species by the receptacular clusters with the inner ones being smaller than the laterals, and the laterals with numerous small granulate tubercula (Fig. 24 H–M). H. helios can also be diagnosed from all other Okinawa group Heptathela species by the following unique nucleotide substitutions in the standard DNA barcode alignment: T (11), T (35), G (47), A (56), C (59), A (95), T (104), A (131), T (140), T (179), C (188), C (215), G (221), C (242), C (266), C (273), C (299), C (300), C (304), T (359), G (380), T (413), A (422), T (425), T (431), G (479), G (480), G (491), G (506), C (543), T (546), G (548), C (551), C (596), T (662).</p> <p>Description.</p> <p>Male. Carapace and opisthosoma brown, with dark brown tergites; cheliceral groove with 14 denticles; tergites closed to each other; seven spinnerets. Measurements: BL 9.88, CL 5.30, CW 4.81, OL 4.58, OW 2.92; ALE&gt; PLE&gt; PME&gt; AME; leg I 16.56 (4.55 + 1.90 + 3.48 + 4.33 + 2.30), leg II 17.19 (4.50 + 2.00 + 3.34 + 4.68 + 2.67), leg III 18.35 (4.35 + 1.97 + 3.40 + 5.53 + 3.10), leg IV 23.37 (5.52 + 2.10 + 4.47 + 7.48 + 3.80).</p> <p>Palp. The bulb of the two male specimens relatively distorted. Prolateral side of paracymbium unpigmented and unsclerotised, numerous setae and spines at the tip of paracymbium (Fig. 24 A–C). Contrategulum with a serrated margin (Fig. 24E, G). Tegulum with a dentate dorsal extension of terminal apophysis (Fig. 24F, G), blunt terminal and hook-like marginal apophysis (Fig. 24D, F, G). Conductor sclerotised, ovate, and wide, with indistinct rugae (Fig. 24 E–G). Embolus sclerotised, with a wide opening, the distal margin slightly sclerotised, and with a saddle-shaped margin in the retrolateral view (Fig. 24 D–G).</p> <p>Females (N = 7). Carapace and opisthosoma colour as in male; cheliceral groove with 12-14 pronounced denticles; opisthosoma with 12 well-separated tergites; seven spinnerets. Measurements: BL 11.70-14.45, CL 5.29-6.70, CW 4.29-5.81, OL 6.54-8.18, OW 4.70-6.63; ALE&gt; PLE&gt; PME&gt; AME; palp 12.29 (4.20 + 2.03 + 2.63 + 3.43), leg I 14.31 (4.51 + 2.40 + 2.65 + 3.08 + 1.67), leg II 14.04 (4.25 + 2.35 + 2.53+ 3.11 + 1.80), leg III 13.75 (3.92 + 2.40 + 2.48 + 2.67 + 2.28), leg IV 21.77 (5.90 + 2.88 + 3.75 + 6.11 + 3.13).</p> <p>Female genitalia. A pair of depressions on the ventro-lateral part of genital atrium (Fig. 24 K–M). Paired receptacular clusters along the anterior margin of bursa copulatrix, divided into two parts, inner receptacular clusters smaller than laterals, laterals with several small tubercula, with short genital stalks (Fig. 24 H–M).</p> <p>Remarks.</p> <p>We identified the specimens collected from Ginama Dam, Okinawa, as H. helios based on evidence from morphology and COI barcode genetic distance compared with the male holotype and paratype (NSMT-Ar 12851, NSMT-Ar 12855) of H. helios in Tanikawa and Miyashita (2014). K2P and p -distances between Ginama Dam specimens and the holotype (NSMT-Ar 12851) were 3.2-3.4% and 3.1-3.3%, respectively, and those between Ginama Dam specimens and the paratype (NSMT-Ar 12855) were 1.8-2.1% and 1.7-2.1%, respectively.</p> <p>Material examined.</p> <p>JAPAN · 2 ♂♂, 8 ♀♀; Okinawa-ken, Kunigami-son, Ginama Dam; 26.84N, 128.26E; alt. 150 m; 24 December 2012; D. Li, F.X. Liu and X. Xu leg.; XUX-2012-432 to 440C.</p> <p>Distribution.</p> <p>The species is endemic to the Japanese island Okinawajima (Fig. 1C).</p></div> 	https://treatment.plazi.org/id/D5DE01B497B9536EADD01443A190F9B8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Xu, Xin;Ono, Hirotsugu;Kuntner, Matjaz;Liu, Fengxiang;Li, Daiqin	Xu, Xin, Ono, Hirotsugu, Kuntner, Matjaz, Liu, Fengxiang, Li, Daiqin (2019): A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands. ZooKeys 888: 1-50, DOI: http://dx.doi.org/10.3897/zookeys.888.34494, URL: http://dx.doi.org/10.3897/zookeys.888.34494
342FD344DA735200BD2FDF1A41B67EE7.text	342FD344DA735200BD2FDF1A41B67EE7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heptathela higoensis Haupt 1983	<div><p>Heptathela higoensis Haupt, 1983 Figs 4, 5</p> <p>Heptathela kimurai higoensis Haupt, 1983: 283 (holotype: male, from Kumamoto, Kyushu, Japan, collected by M. Yoshikura on 27 September 1973, deposited in ZMH, examined); Haupt 2003: 69. Heptathela higoensis: Ono 1998: 16; Ono 2009: 80; Ono and Ogata 2018: 26, 479.</p> <p>Heptathela nishikawai Ono, 1998: 19 (holotype: female, from Hitoyoshi-shi, Kumamoto-ken, Kyushu, Japan, collected by H. Ono on 19 November 1996, deposited in NMNS, examined); Ono 2009: 83; syn. nov.</p> <p>Heptathela yaginumai Ono, 1998: 20 (holotype: female, from Honjo, Kunitomi-cho, Higashimorokata-gun, Miyazaki-ken, Kyushu, Japan, collected by T. Yaginuma on 18 June 1949, deposited in NMNS, examined); Ono 2009: 81. syn. nov.</p> <p>Diagnosis.</p> <p>Males of H. higoensis can be distinguished from those of H. kikuyai by one of the embolus peaks being longer than the other (Fig. 4H, I, K) and by the slightly blunt tegular marginal apophysis (Fig. 4J), and from those of H. yakushimaensis by the conductor with the weakly serrated prolateral margin (Fig. 4F, H, K). Females of H. higoensis can be distinguished from those of H. kimurai by the wide and flat dorso-posterior part of the genital area (Fig. 4A, B), and from those of H. kikuyai and H. yakushimaensis by the inner receptacular clusters that are larger than the outer ones (Fig. 4 A–D). Moreover, H. higoensis differs from all other Kyushu group Heptathela species by the following unique nucleotide substitutions in the standard DNA barcode alignment: G (140), C (146), A (179), C (251), C (257), C (263), G (272), A (326), C (332), T (350), G (479), G (569), C (572), A (578), G (596), G (632), G (641).</p> <p>Description.</p> <p>Males (N = 11). Carapace yellow brown; opisthosoma light brown, with dark brown tergites close to each other; cheliceral groove with 11-13 denticles; 7 or 8 spinnerets. Measurements: BL 8.80-11.00, CL 4.40-5.25, CW 4.10-4.90, OL 4.40-5.60, OW 2.90-3.40; ALE&gt; PLE&gt; PME&gt; AME; leg I 11.75 (3.45 + 1.50 + 2.60 + 2.70 + 1.50), leg II 12.40 (3.40 + 1.60 + 2.50 + 3.10 + 1.80), leg III 13.30 (3.20 + 1.60 + 2.40 + 3.80 + 2.30), leg IV 16.80 (4.40 + 1.20 + 3.30 + 5.20 + 2.70).</p> <p>Palp. Prolateral side of paracymbium unpigmented and unsclerotised, numerous setae and spines at the tip of paracymbium (Fig. 4 E–G). Contrategulum with serrated margin proximally and smooth margin distally (Fig. 4I, K). Tegulum wide, the dorsal extension of terminal apophysis and marginal apophysis with a serrated margin retrolaterally (Fig. 4J, K). Conductor wide and with an apical tooth and a deep fold (Fig. 4H, K). Embolus with two peaks, one peak longer than the other, and with a curved margin retrolaterally (Fig. 4 H–K).</p> <p>Females (N = 43). Carapace and opisthosoma colour as in male; cheliceral groove with 11-16 pronounced denticles; tergites similar to male; 7-8 spinnerets. Measurements: BL 8.00-12.80, CL 4.30-6.10, CW 3.80-5.57, OL 4.10-6.50, OW 2.70-4.90; ALE&gt; PLE&gt; PME&gt; AME; palp 8.20 (2.70 + 1.50 + 1.70 + 2.30), leg I 8.80 (2.80 + 1.75 + 1.70 + 1.55 + 1.00), leg II 8.93 (2.75 + 1.60 + 1.55 + 1.83 + 1.20), leg III 9.30 (2.70 + 1.60 + 1.50 + 2.20 + 1.30), leg IV 13.25 (3.65 + 1.80 + 2.40 + 3.40 + 2.00).</p> <p>Female genitalia. A pair of depressions on the ventro-lateral part of genital atrium (Fig. 4C, D). Paired receptacular clusters along the anterior margin of bursa copulatrix, divided into two parts, the inner main part forming a large granulate tubercle, with short genital stalks, the outers with several small granules (Fig. 4).</p> <p>Remarks.</p> <p>We examined the male holotype of H. higoensis (Fig. 5) and identified the species as H. higoensis even though the bulb of holotype male is relatively distorted compared to the specimens we collected. After we examined 11 males and 43 females collected at the type localities of H. higoensis, H. nishikawai and H. yaginumai, and compared the holotypes of H. higoensis, H. nishikawai and H. yaginumai with our specimens, we proposed synonymy of H. nishikawai and H. yaginumai with H. higoensis based on their genital morphology, molecular species delimitation (Xu et al. 2019), and intraspecific genetic distances, 0-1.19% (K2P) and 0-1.18% (p -distance) among 43 specimens, although females exhibit a considerable intraspecific variation in genitalia.</p> <p>Material examined.</p> <p>JAPAN · 1 ♂, 8 ♀♀; Kyushu, Kumamoto-ken, Hitoyoshi-shi, Fumotomachi, Hitoyoshi Ruins Park; 32.21N, 130.77E; alt. 140 m; 18 September 2013; D. Li and B. Wu leg.; XUX-2013-365 (♂ matured 19 July 2014 at CBEE), XUX-2013-360 to 364, 366 to 368 · 2 ♂♂, 5 ♀♀; Kyushu, Kumamoto-ken, Kumamoto-shi, Tatsutayama, Tatsuta National Park; 32.82N, 130.73E; alt. 60 m; 19 September 2013; D. Li and B. Wu leg.; XUX-2013-370 to 379 · 4 ♂♂, 4 ♀♀; Kyushu, Kumamoto-ken, Kumamoto-shi, Higashi-ku, Kozono 1-chome; 32.84N, 130.78E; alt. 100 m; 19 September 2013; D. Li and B. Wu leg.; XUX-2013-380 to 389 (XUX-2013-381, ♂ matured 2 August 2014 at CBEE) · 1 ♂, 3 ♀♀; Kyushu, Kumamoto-ken, Kumamoto-shi, Kasuga, Hanaokayama; 32.80N, 130.68E; alt. 120 m, 19 September 2013; D. Li and B. Wu leg.; XUX-2013-390 to 393 · 3 ♀♀; Miyazaki-ken, Nishiusuki-gun, Takachiho-cho, Mukoyama; 32.70N, 131.30E; alt. 320 m; 22 September 2013; D. Li and B. Wu leg.; XUX-2013-435 to 441 · 1 ♂, 3 ♀♀; Miyazaki-ken, Higashimorokata-gun, Kunitomi-cho, Honjo 11960-1; 32.00N, 131.34E; alt. 30 m; 23 September 2013; D. Li and B. Wu leg.; XUX-2013-456 (♂, matured 19 July 2014 at CBEE), XUX-2013-449 to 451 · 1 ♂, 12 ♀♀; Miyazaki-ken, Higashimorokata-gun, Kunitomi-cho, Honjo 4191; 31.98N, 131.33E; alt. 30 m; 23 September 2013; D. Li and B. Wu leg.; XUX-2013-457 to 467C · 1 ♀; Miyazaki-ken, Nishimorokata-gun, Takaharu-cho, Kamamuto, Lake Miike; 31.89N, 130.96E; alt. 360 m; 23 September 2013; D. Li and B. Wu leg.; XUX-2013-468 · 1 ♂, 4 ♀♀; 2 Kagoshima-ken, Kirishima-shi, Kirishima, Takachihokawara; 31.89N, 130.89E; alt. 960 m; 23 September 2013; D. Li and B. Wu leg.; XUX-2013-471 (♂, matured 8 June 2014 at CBEE), XUX-2013-472 to 474, 476.</p> <p>Distribution.</p> <p>The species is known from the following prefectures on the Japanese island Kyushu: Kumamoto-ken (Hitoyoshi-shi and Kumamoto-shi), Miyazaki-ken (Nishiusuki-gun, Higashimorokata-gun and Nishimorokata-gun), Kagoshima-ken (Kirishima-shi) (Fig. 1C).</p></div> 	https://treatment.plazi.org/id/342FD344DA735200BD2FDF1A41B67EE7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Xu, Xin;Ono, Hirotsugu;Kuntner, Matjaz;Liu, Fengxiang;Li, Daiqin	Xu, Xin, Ono, Hirotsugu, Kuntner, Matjaz, Liu, Fengxiang, Li, Daiqin (2019): A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands. ZooKeys 888: 1-50, DOI: http://dx.doi.org/10.3897/zookeys.888.34494, URL: http://dx.doi.org/10.3897/zookeys.888.34494
E5CED3E805D85B318E13363C780C18CC.text	E5CED3E805D85B318E13363C780C18CC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heptathela kanenoi Ono 1996	<div><p>Heptathela kanenoi Ono, 1996 Fig. 10</p> <p>Heptathela kanenoi Ono, 1996: 158 (holotype: male, from Mikyo, Amagi-cho, Tokunoshima, Kagoshima-ken, Japan, collected by M. Owada and S. Kaneno on 2 November 1992, deposited in NMNS, examined); Ono 2009: 80.</p> <p>Diagnosis.</p> <p>Males of H. kanenoi can be distinguished from those of all other Amami group Heptathela species by lacking a spiniform conductor apex (Fig. 10G, I). Females of H. kanenoi can be distinguished from those of all other Amami group Heptathela species by the inner receptacular clusters larger than the outers (Fig. 10B, C). H. kanenoi can also be diagnosed from all other Amami group Heptathela species by the following unique nucleotide substitutions in the standard DNA barcode alignment: A (35), G (38), C (71), C (125), C (224), G (278), T (281), C (288), T (332), G (359), C (396), C (410), G (443), T (449), A (512), C (533), G (557), C (560), C (623), T (641).</p> <p>Description.</p> <p>Males (N = 6). Carapace yellow brown; opisthosoma brown, with dark-spotted tergites close to each other; cheliceral groove with 10-13 vestigial denticles; 7 spinnerets. Measurements: BL 9.80-11.60, CL 5.10-6.00, CW 4.60-5.40, OL 5.15-5.60, OW 4.00-4.85; ALE&gt; PLE&gt; PME&gt; AME; leg I 17.85 (4.80 + 2.30 + 3.80 + 4.60 + 2.35), leg II 18.90 (4.90 + 2.30 + 3.80 + 5.20 + 2.70), leg III 20.80 (4.70 + 2.30 + 4.00 + 6.50 + 3.30), leg IV 26.00 (6.20 + 2.50 + 5.10 + 8.20 + 4.00).</p> <p>Palp. Prolateral side of paracymbium unpigmented and unsclerotised, numerous setae and spines at the tip of paracymbium (Fig. 10 D–F). Contrategulum with serrated margin (Fig. 10 G–I). Tegulum with dentate dorsal extension of terminal apophysis (Fig. 10J) and blunt tegulum terminal apophysis (Fig. 10I, J). Conductor base wide and rugose, with several folds and each fold with an apical tooth (Fig. 10 G–J). Embolus with a wide and flat opening, the distal part slightly sclerotised, and saddle-shaped in the prolateral view (Fig. 10 H–I).</p> <p>Females (N = 11). Carapace and opisthosoma colour as in male; cheliceral groove with 12-14 pronounced denticles; tergites similar to male; seven or eight spinnerets. Measurements: BL 8.30-12.90, CL 4.30-6.28, CW 3.60-5.40, OL 4.38-6.50, OW 3.70-6.20; ALE&gt; PLE&gt; PME&gt; AME; palp 7.00 (2.50 + 1.35 + 1.55 + 1.60), leg I 7.75 (2.30 + 1.25 + 1.60 + 1.50 + 1.10), leg II 7.60 (2.60 + 1.10 + 1.50 + 1.50 + 0.90), leg III 8.35 (2.45 + 1.50 + 1.10 + 2.00 + 1.30), leg IV 12.15 (3.40 + 1.50 + 2.25 + 3.20 + 1.80).</p> <p>Female genitalia. A pair of indistinct depressions on the ventro-lateral part of genital atrium. Paired receptacular clusters along the anterior margin of bursa copulatrix, divided into two parts, the inners larger than the outers, without genital stalks (Fig. 10 A–C).</p> <p>Material examined.</p> <p>JAPAN · 1 ♂, 1 ♀; Kagoshima-ken, Tokunoshima, Amagi-cho, Mikyo; 27.77N, 128.95E; alt. 180 m; 16 September 2013; D. Li and B. Wu leg.; XUX-2013-315 to 316 · 3 ♂♂, 7 ♀♀; Kagoshima-ken, Oshima-gun, Tokunoshima, Tokunoshima-cho, Tokuwase; 27.79N, 129.01E; alt. 150 m; 17 September 2013; D. Li and B. Wu leg.; XUX-2013-323 to 332 · 2 ♂♂, 3 ♀♀; Kagoshima-ken, Tokunoshima, Amagi-cho, Mikyo; 27.77N, 128.95E; alt. 130 m; 17 September 2013; D. Li and B. Wu leg.; XUX-2013-333 to 338.</p> <p>Distribution.</p> <p>The species is endemic to the Japanese island Tokunoshima (Fig. 1C).</p></div> 	https://treatment.plazi.org/id/E5CED3E805D85B318E13363C780C18CC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Xu, Xin;Ono, Hirotsugu;Kuntner, Matjaz;Liu, Fengxiang;Li, Daiqin	Xu, Xin, Ono, Hirotsugu, Kuntner, Matjaz, Liu, Fengxiang, Li, Daiqin (2019): A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands. ZooKeys 888: 1-50, DOI: http://dx.doi.org/10.3897/zookeys.888.34494, URL: http://dx.doi.org/10.3897/zookeys.888.34494
90060E34E05C5574AA14B6C27D7BF304.text	90060E34E05C5574AA14B6C27D7BF304.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heptathela kikuyai Ono 1998	<div><p>Heptathela kikuyai Ono, 1998 Fig. 6</p> <p>Heptathela kikuyai Ono, 1998: 16 (holotype: male, from Mt. Gozaga-dake, 20 km south of Oita-shi, Oita-ken, Kyushu, Japan, collected by N. Kikuya on 13 September 1979, deposited in NMNS, examined); Ono 2009: 81; Schwendinger and Ono 2011: 613; Ono and Ogata 2018: 26, 479.</p> <p>Diagnosis.</p> <p>Males of H. kikuyai differ from those of H. higoensis by the two embolus peaks of a similar height, and the hooked tegular marginal apophysis (Fig. 6J, K), and from those of H. yakushimaensis by only a weakly serrated prolateral margin of the conductor (Fig. 6H, I). Females of H. kikuyai differ from those of all other Kyushu group Heptathela species by the inner receptacular clusters with longer and slender stalks (Fig. 6 A–D). Moreover, H. kikuyai differs from all other Kyushu group Heptathela species by the following unique nucleotide substitutions in the standard DNA barcode alignment: G (11), A (33), C (41), C (59), C (68), A (83), A (89), C (116), T (194), A (212), T (222), A (239), A (251), C (260), T (276), T (287), C (305), C (329), A (345), T (365), C (470), G (491), T (546), T (578), T (581), G (635), C (656).</p> <p>Description.</p> <p>Males (N = 5). Carapace and opisthosoma brown, with dark-spotted tergites; cheliceral groove with 10-12 denticles; 6 or 7 spinnerets. Measurements: BL 7.90-9.40, CL 4.00-5.05, CW 3.55-4.30, OL 3.90-4.60, OW 2.60-3.20; ALE&gt; PLE&gt; PME&gt; AME; leg I 13.65 (3.90 + 1.90 + 2.75 + 3.30 + 1.80), leg II 13.35 (3.45 + 1.60 + 2.65 + 3.55 + 2.10), leg III 14.40 (3.30 + 1.50 + 2.75 + 4.45 + 2.40), leg IV 19.25 (4.85 + 1.90 + 3.35 + 6.00 + 3.15).</p> <p>Palp. Prolateral side of paracymbium unpigmented and unsclerotised, numerous setae and spines at the tip of paracymbium (Fig. 6 E–G). Contrategulum with serrated margin proximally and smooth margin distally (Fig. 6H, I, K). Tegulum wide, the dorsal extension of terminal apophysis and hooked marginal apophysis of tegulum with a serrated margin retrolaterally (Fig. 6J, K). Conductor wide and with an apical tooth and a deep fold (Fig. 6I, K). Embolus with two peaks at the slightly same level and a curved margin retrolaterally (Fig. 6 H–K).</p> <p>Females (N = 30). Carapace and opisthosoma as in male; cheliceral groove with 12-17 pronounced denticles; tergites similar to male; 6-8 spinnerets. Measurements: BL 8.55-15.10, CL 4.30-7.90, CW 3.80-6.70, OL 4.40-8.75, OW 2.80-6.10; ALE&gt; PLE&gt; PME&gt; AME; palp 9.15 (3.25 + 1.60 + 1.90 + 2.40), leg I 10.30 (3.30 + 1.90 + 1.90 + 2.00 + 1.20), leg II 10.15 (3.10 + 1.80 + 1.85 + 2.10 + 1.30), leg III 10.80 (3.00 + 1.80 + 1.80 + 2.60 + 1.60), leg IV 15.80 (4.50 + 2.20 + 2.80 + 4.10 + 2.20).</p> <p>Female genitalia. A pair of depressions on the ventro-lateral part of genital atrium. Paired receptacular clusters along the anterior margin of bursa copulatrix, indistinctly divided into two parts, with many small granules, inner ones with longer and slender genital stalks (Fig. 6 A–D).</p> <p>Material examined.</p> <p>JAPAN · 6 ♀♀; Oita-ken, Bungoono-shi, Totoki, Onomachi; 33.06N, 131.52E; alt. 240 m; 20 September 2013; D. Li and B. Wu leg.; XUX-2013-394 to 396, 399 to 401 · 2 ♂♂, 4 ♀♀; Oita-ken, Bungoono-shi, Onomachi-Ando, Mt. Gozaga-dake; 33.11N, 131.55E; alt. 630 m; 20 September 2013; D. Li and B. Wu leg.; XUX-2013-404, 405, 408, 409, 411, 412 (405, ♂ matured 2 May 2014 at CBEE) · 2 ♂♂, 6 ♀♀; Oita-ken, Usuki-shi, Takeyama; 33.10N, 131.72E; alt. 80 m; 20 September 2013; D. Li and B. Wu leg.; XUX-2013-414, 416 to 422 · 1 ♂, 8 ♀♀; Oita-ken, Usuki-shi, Inukai-machi Sasamuta; 33.09N, 131.67E; alt. 120 m; 21 September 2013; D. Li and B. Wu leg.; XUX-2013-424 to 430, 432, 433 · 6 ♀♀; Miyazaki-ken, Nishiusuki-gun, Takachiho-cho, Iwato; 32.73N, 131.35E; alt. 400 m; 22 September 2013; D. Li and B. Wu leg.; XUX-2013-442 to 444, 446 to 448.</p> <p>Distribution.</p> <p>The species is known from the following prefectures on the Japanese island Kyushu: Oita-ken (Bungoono-shi and Usuki-shi) and Miyazaki-ken (Nishiusuki-gun) (Fig. 1C).</p></div> 	https://treatment.plazi.org/id/90060E34E05C5574AA14B6C27D7BF304	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Xu, Xin;Ono, Hirotsugu;Kuntner, Matjaz;Liu, Fengxiang;Li, Daiqin	Xu, Xin, Ono, Hirotsugu, Kuntner, Matjaz, Liu, Fengxiang, Li, Daiqin (2019): A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands. ZooKeys 888: 1-50, DOI: http://dx.doi.org/10.3897/zookeys.888.34494, URL: http://dx.doi.org/10.3897/zookeys.888.34494
51999F5AA3295810A798EBC5826C670B.text	51999F5AA3295810A798EBC5826C670B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heptathela kimurai (Kishida 1920) Kishida 1920	<div><p>Heptathela kimurai (Kishida, 1920) Fig. 7</p> <p>Liphistius kimurai Kishida, 1920: 362 (holotype: female, from Shiroyama, Kagoshima, Kyushu, Japan, collected by A. Kimura in October 1920, lost in the Science College Museum of the Tokyo Imperial University (Haupt, 1983); neotype: male, from the same locality as for the original type specimen, collected by J. Haupt on 21 March 1982, matured in August 1982, deposited in ZMH, but the neotype may be lost according to Dunlop et al. 2014).</p> <p>Heptathela kimurai Kishida, 1923: 236; Bristowe 1933: 1030; Sawaguti and Ozi 1937: 116 (partly); Yaginuma 1954: 15; Yaginuma 1955: 35; Yaginuma 1960: 19; Yaginuma 1971: 19; Gertsch and Platnick 1979: 5; Yaginuma 1979: 1; Yaginuma 1980: 44; Haupt 1983: 283; Yoshikura 1983: 63; Haupt 1984: 163; Yaginuma 1986: 1; Yoshikura 1987: 148; Chikuni 1989 b: 18; Ono 1998: 14; Yoo and Kim 2002: 27; Haupt 2003: 69; Ono 2009: 81; Schwendinger and Ono 2011: 614; Ono and Ogata 2018: 25, 479.</p> <p>Diagnosis.</p> <p>Females of H. kimurai resemble H. higoensis females but differ by a slightly curved dorso-posterior margin of the genital area (Fig. 7A, B). Moreover, H. kimurai differs from all other Kyushu group Heptathela species by the following unique nucleotide substitutions in the standard DNA barcode alignment: C (26), T (98), G (191), G (194), G (206), C (215), T (251), T (278), G (293), C (362), T (366), G (443), T (449), C (452), G (506), G (521), C (527), G (548), T (572), C (599), A (615), C (638).</p> <p>Description.</p> <p>Females (N = 9). Carapace yellow brown; opisthosoma brown, with brown tergites close to each other; cheliceral groove with 12-14 vestigial denticles; seven spinnerets. Measurements: BL 9.30-13.80, CL 4.88-6.30, CW 4.10-5.50, OL 4.80-7.30, OW 3.45-5.40; ALE&gt; PLE&gt; PME&gt; AME; palp 7.77 (2.57 + 1.50+ 1.60 + 2.10), leg I 9.15 (2.85 + 1.70 + 1.75 + 1.65 + 1.20), leg II 8.88 (2.65 + 1.65 + 1.55 + 1.88 + 1.15), leg III 9.63 (2.70 + 1.70 + 1.60 + 2.30 + 1.33), leg IV 14.08 (3.88 + 1.90 + 2.50 + 3.60 + 2.20).</p> <p>Female genitalia. A pair of depressions on the ventro-lateral part of genital atrium. Paired of receptacular clusters along the anterior margin of bursa copulatrix, divided into two parts, the inner main part forming a large granulate tubercle, the outer part with several small granules (Fig. 7 A–D).</p> <p>Male: unknown.</p> <p>Remarks.</p> <p>We could not examine the presumably lost neotype male (Dunlop et al. 2014). The male is therefore unknown. The non-topotypical male described as H. kimurai by Ono (1998, 2009) is excluded here as we are not sure that the specimen was collected from the type locality although it was obtained in Kagoshima-shi.</p> <p>Material examined.</p> <p>JAPAN · 9 ♀♀; Kagoshima-ken, Kagoshima-shi, Shiroyama-cho, Shiroyama Park; 31.60N, 130.55E; alt. 100 m; 18 September 2013; D. Li and B. Wu leg.; XUX-2013-349, 351, 352, 354 to 359.</p> <p>Distribution.</p> <p>The species is known from the Kagoshima prefecture on the Japanese island Kyushu (Fig. 1C).</p></div> 	https://treatment.plazi.org/id/51999F5AA3295810A798EBC5826C670B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Xu, Xin;Ono, Hirotsugu;Kuntner, Matjaz;Liu, Fengxiang;Li, Daiqin	Xu, Xin, Ono, Hirotsugu, Kuntner, Matjaz, Liu, Fengxiang, Li, Daiqin (2019): A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands. ZooKeys 888: 1-50, DOI: http://dx.doi.org/10.3897/zookeys.888.34494, URL: http://dx.doi.org/10.3897/zookeys.888.34494
5D1DC7E647F650DAA232982BF0B8919B.text	5D1DC7E647F650DAA232982BF0B8919B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heptathela kojima	<div><p>Heptathela kojima sp. nov. Fig. 11</p> <p>Type material.</p> <p>Holotype: JAPAN · ♂; Kagoshima-ken, Oshima-gun, Tokunoshima, Isen-cho, Kojima; 27.74N, 128.91E; alt. 160 m; 17 September 2013; D. Li and B. Wu leg.; XUX-2013-346 (matured 10 October 2013 at CBEE).</p> <p>Paratypes: JAPAN · 2 ♂♂, 6 ♀♀; same data as for holotype; XUX-2013-339, 340, 342 to 345, 347, 348.</p> <p>Diagnosis.</p> <p>Males of H. kojima sp. nov. differ from those of H. amamiensis and H. kanenoi by a wide leaf-shaped conductor (Fig. 11I, J), and a less dentate dorsal extension of the tegular terminal apophysis (Fig. 11G, J, K), from those of H. sumiyo sp. nov. by a shallow saddle-shaped in the prolateral view, and from those of H. uken sp. nov. by embolus with two longer peaks (Fig. 11I, J). Females of H. kojima sp. nov. resemble those of other Amami group Heptathela species but differ from those of other Amami group Heptathela species by paired receptacular clusters close to each other (Fig. 11B, D). H. kojima sp. nov. can also be diagnosed from all other Amami group Heptathela species by the following unique nucleotide substitutions in the standard DNA barcode alignment: C (44), C (56), C (128), A (131), C (134), C (137), C (155), G (158), G (176), T (230), T (245), C (269), T (320), C (357), C (377), A (378), A (443), C (446), G (464), A (479), C (518), G (521), T (554), A (560), C (608), C (611).</p> <p>Description.</p> <p>Male (Holotype). Carapace brown; opisthosoma light brown, with dark brown tergites close to each other; cheliceral groove with eleven denticles; seven spinnerets. Measurements: BL 7.30, CL 3.60, CW 3.30, OL 3.60, OW 2.70; ALE&gt; PLE&gt; PME&gt; AME; leg I 14.90 (4.20 + 165 + 3.15 + 4.00 + 1.90), leg II 16.10 (4.20 + 1.85 + 3.25 + 4.40 + 2.40), leg III 16.85 (4.25 + 1.90 + 3.20 + 5.00 + 2.50), leg IV 21.70 (5.50 + 2.00 + 4.20 + 6.90 + 3.10).</p> <p>Palp. Prolateral side of paracymbium unpigmented and unsclerotised, numerous setae and spines at the tip of paracymbium (Fig. 11 E–G). Contrategulum with serrated margin (Fig. 11E, H, I). Conductor base wide, leaf-shaped and rugose, with several folds and gradually narrowing to a short spiniform apex (Fig. 11I, J). Embolus sclerotised, with a wide and flat opening (Fig. 11 H–K).</p> <p>Females (N = 6). Carapace and opisthosoma colour as in male; cheliceral groove with 13 pronounced denticles; tergites similar to male; 6-8 spinnerets. Measurements: BL 9.00-11.50, CL 4.90-6.15, CW 4.25-5.20, OL 4.30-6.10, OW 3.00-4.50; ALE&gt; PLE&gt; PME&gt; AME; palp 9.05 (3.25 + 1.65 + 1.85 + 2.30), leg I 10.50 (3.35 + 1.80 + 2.00 + 2.10 + 1.25), leg II 10.05 (3.10 + 1.80 + 1.75 + 2.10 + 1.30), leg III 10.45 (2.90 + 1.95 + 1.65 + 2.40 + 1.55), leg IV 15.20 (4.40 + 2.20 + 2.60 + 4.00 + 2.00).</p> <p>Female genitalia. A pair of indistinct depressions on the ventro-lateral part of genital atrium (Fig. 11C, D). Paired receptacular clusters separated from each other along the anterior margin of bursa copulatrix, or fused together, divided into two parts, without genital stalks (Fig. 11 A–D).</p> <p>Etymology.</p> <p>The species epithet, a noun in apposition, refers to the type locality.</p> <p>Distribution.</p> <p>The species is endemic to the Japanese island Tokunoshima (Fig. 1C).</p></div> 	https://treatment.plazi.org/id/5D1DC7E647F650DAA232982BF0B8919B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Xu, Xin;Ono, Hirotsugu;Kuntner, Matjaz;Liu, Fengxiang;Li, Daiqin	Xu, Xin, Ono, Hirotsugu, Kuntner, Matjaz, Liu, Fengxiang, Li, Daiqin (2019): A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands. ZooKeys 888: 1-50, DOI: http://dx.doi.org/10.3897/zookeys.888.34494, URL: http://dx.doi.org/10.3897/zookeys.888.34494
2C61BBF737655063A5A8113AC6EA8C44.text	2C61BBF737655063A5A8113AC6EA8C44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heptathela kubayama	<div><p>Heptathela kubayama sp. nov. Fig. 17</p> <p>Type material.</p> <p>Holotype: JAPAN · ♀; Okinawa-ken, Iheyajima Island, Mt. Kubayama Nature Conservation Area; 27.09N, 128.02E; alt. 85 m; 26 December 2012; D. Li, F.X. Liu and X. Xu leg.; XUX-2012-486.</p> <p>Paratypes: JAPAN · 8 ♀♀; same data as for holotype; XUX-2012-479, 481 to 485, 487 to 488.</p> <p>Diagnosis.</p> <p>Females of H. kubayama sp. nov. can be distinguished from those of H. gayozan sp. nov. and H. mae sp. nov. by paired receptacular clusters with short genital stalks (Fig. 17 A–B). Heptathela kubayama sp. nov. can also be diagnosed from all other Okinawa group Heptathela species by the following unique nucleotide substitutions in the standard DNA barcode alignment: G (239), G (329), G (353), C (359), G (443), G (602), G (647).</p> <p>Description.</p> <p>Female (Holotype). Carapace brown; opisthosoma light brown, with dark brown tergites close to each other; cheliceral groove with 12 pronounced denticles; six spinnerets. Measurements: BL 10.20, CL 4.89, CW 4.18, OL 4.90, OW 3.60; ALE&gt; PLE&gt; PME&gt; AME; palp 8.25 (2.82 + 1.45 + 1.80 + 2.18), leg I 9.66 (3.10 + 1.70 + 1.81 + 1.90 + 1.15), leg II 8.90 (2.90 + 1.55 + 1.75 + 1.70 + 1.00), leg III 9.59 (2.81 + 1.65 + 1.60 + 2.13 + 1.40), leg IV 13.67 (4.25 + 2.00 + 2.00 + 3.54 + 1.88).</p> <p>Female genitalia. A pair of depressions on the ventro-lateral part of the genital atrium (Fig. 17C, D). A pair of receptacular clusters along the anterior margin of bursa copulatrix, divided into two parts, with several granules; with short genital stalks (Fig. 17 A–D).</p> <p>Male. Unknown.</p> <p>Etymology.</p> <p>The species epithet, a noun in apposition, refers to the type locality.</p> <p>Distribution.</p> <p>The species is endemic to the Japanese island Iheyajima (Fig. 1C).</p></div> 	https://treatment.plazi.org/id/2C61BBF737655063A5A8113AC6EA8C44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Xu, Xin;Ono, Hirotsugu;Kuntner, Matjaz;Liu, Fengxiang;Li, Daiqin	Xu, Xin, Ono, Hirotsugu, Kuntner, Matjaz, Liu, Fengxiang, Li, Daiqin (2019): A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands. ZooKeys 888: 1-50, DOI: http://dx.doi.org/10.3897/zookeys.888.34494, URL: http://dx.doi.org/10.3897/zookeys.888.34494
99F3A3542E4655A0A8CC19EACA81F7DA.text	99F3A3542E4655A0A8CC19EACA81F7DA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heptathela mae	<div><p>Heptathela mae sp. nov. Fig. 18</p> <p>Type material.</p> <p>Holotype: JAPAN · ♀; Okinawa-ken, Iheyajima Island, Mt. Mae-dake; 27.06N, 127.99E; alt. 10 m; 26 December 2012; D. Li, F.X. Liu and X. Xu leg.; XUX-2012-497.</p> <p>Paratypes: JAPAN · 5 ♀♀; same data as for holotype; XUX-2012-494 to 496, 498, 500 · 3♀♀; Okinawa Prefecture, Iheyajima Island, Mt. Mae-dake; 27.06N, 127.99E; alt. 20 m; 10 May 2014; D. Li and B. Wu leg.; XUX-2014-079A to 079C.</p> <p>Diagnosis.</p> <p>Females of H. mae sp. nov. can be distinguished from those of H. aha sp. nov. and H. gayozan sp. nov. by the lateral receptacular clusters being larger than the inner ones (Fig. 18B, D), and from those of H. kubayama sp. nov. by the receptacular clusters without short genital stalks (Fig. 18 A–D). Heptathela mae sp. nov. can also be diagnosed from all other Okinawa group Heptathela species by the following unique nucleotide substitutions in the standard DNA barcode alignment: C (74), T (227), G (389), C (407), A (461), G (503), G (635).</p> <p>Description.</p> <p>Female (Holotype). Carapace brown; opisthosoma brown, brown tergites with black plaques; cheliceral groove with 12 denticles; seven spinnerets. Measurements: BL 9.28, CL 4.48, CW 4.00, OL 5.12, OW 3.78; ALE&gt; PLE&gt; PME&gt; AME; palp 7.80 (2.70 + 1.32 + 1.71 + 2.07), leg I 9.25 (3.00 + 1.52 + 1.80 + 1.78 + 1.15), leg II 8.55 (2.68 + 1.20 + 1.72 + 1.75 + 1.20), leg III 9.61 (2.75 + 1.60 + 1.55 + 2.31 + 1.40), leg IV 13.41 (3.90 + 1.70 + 2.31 + 3.60 + 1.90).</p> <p>Female genitalia. A pair of depressions on the ventro-lateral part of the genital atrium (Fig. 18C, D). A pair of receptacular clusters along the anterior margin of bursa copulatrix, divided into two parts, with several granules, without genital stalks, the posterior part of genital area incurved (Fig. 18 A–D).</p> <p>Male. Unknown.</p> <p>Etymology.</p> <p>The species epithet, a noun in apposition, refers to the type locality.</p> <p>Distribution.</p> <p>The species is endemic to the Japanese island Iheyajima (Fig. 1C).</p></div> 	https://treatment.plazi.org/id/99F3A3542E4655A0A8CC19EACA81F7DA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Xu, Xin;Ono, Hirotsugu;Kuntner, Matjaz;Liu, Fengxiang;Li, Daiqin	Xu, Xin, Ono, Hirotsugu, Kuntner, Matjaz, Liu, Fengxiang, Li, Daiqin (2019): A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands. ZooKeys 888: 1-50, DOI: http://dx.doi.org/10.3897/zookeys.888.34494, URL: http://dx.doi.org/10.3897/zookeys.888.34494
71A1B990F84356DBAE11F68F2953B1E2.text	71A1B990F84356DBAE11F68F2953B1E2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heptathela otoha	<div><p>Heptathela otoha sp. nov. Fig. 19</p> <p>Type material.</p> <p>Holotype: JAPAN · ♀; Okinawa-ken, Nakijin-son, Mt. Otoha-dake; 26.67N, 127.97E; alt. 80 m; 27 December 2012; D. Li, F.X. Liu and X. Xu leg.; XUX-2012-535.</p> <p>Paratypes: JAPAN · 3 ♀♀; Okinawa Prefecture, Motobu-cho, Yamazato; 26.67N, 127.91E; alt. 160 m; 11 May 2014; D. Li and B. Wu leg.; XUX-2014-094, 097, 099 · 3 ♀♀; Okinawa Prefecture, Nakijin-son, Mt. Otoha-dake; 26.67N, 127.97E; alt. 100 m; 11 May 2014; D. Li and B. Wu leg.; XUX-2014-101, 102, 103.</p> <p>Diagnosis.</p> <p>Females of H. otoha sp. nov. can be distinguished from those of H. yanbaruensis, H. unten sp. nov., and H. crypta sp. nov. by the inner receptacular clusters with several granules, albeit these inner clusters are difficult to be separated from the laterals (Fig. 19 A–D). H. otoha sp. nov. can also be diagnosed from all other Okinawa group Heptathela species by the following unique nucleotide substitutions in the standard DNA barcode alignment: G (83), G (206), G (488).</p> <p>Description.</p> <p>Female (Holotype). Carapace yellow brown; opisthosoma light brown, with brown and black-spotted tergites; cheliceral groove with 13 pronounced denticles; seven spinnerets. Measurements: BL 10.92, CL 4.75, CW 4.35, OL 6.60, OW 4.90; ALE&gt; PLE&gt; PME&gt; AME; palp 8.30 (2.78 + 1.39 + 1.90 + 2.23), leg I 9.80 (3.05 + 1.65 + 1.95 + 1.95 + 1.20), leg II 9.21 (2.90 + 1.50 + 1.81 + 1.90 + 1.10), leg III 9.84 (2.70 + 1.70 + 1.62 + 2.35 + 1.47), leg IV 14.74 (4.00 + 1.90 + 2.70 + 4.03 + 2.11).</p> <p>Female genitalia. A pair of depressions on the ventro-lateral part of genital atrium (Fig. 19C, D). Paired receptacular clusters along the anterior margin of bursa copulatrix, divided into indistinct two parts, but difficulty to separate, both with several granules, with very short genital stalks (Fig. 19 A–D).</p> <p>Male. Unknow.</p> <p>Etymology.</p> <p>The species epithet, a noun in apposition, refers to the type locality.</p> <p>Distribution.</p> <p>The species is endemic to the Japanese island Okinawajima (Fig. 1C).</p></div> 	https://treatment.plazi.org/id/71A1B990F84356DBAE11F68F2953B1E2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Xu, Xin;Ono, Hirotsugu;Kuntner, Matjaz;Liu, Fengxiang;Li, Daiqin	Xu, Xin, Ono, Hirotsugu, Kuntner, Matjaz, Liu, Fengxiang, Li, Daiqin (2019): A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands. ZooKeys 888: 1-50, DOI: http://dx.doi.org/10.3897/zookeys.888.34494, URL: http://dx.doi.org/10.3897/zookeys.888.34494
53D77AAC44A5548DB625D0A92B2BB232.text	53D77AAC44A5548DB625D0A92B2BB232.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heptathela shuri	<div><p>Heptathela shuri sp. nov. Fig. 20</p> <p>Type material.</p> <p>Holotype: JAPAN · ♀; Okinawa-ken, Naha, Shuri, Sueyoshi Park; 26.23N, 127.72E; alt. 45 m; 17 December 2012; D. Li, F.X. Liu and X. Xu leg.; XUX-2012-309.</p> <p>Paratype: JAPAN · 1 ♀; same data as for holotype; XUX-2012-308.</p> <p>Diagnosis.</p> <p>Females of H. shuri sp. nov. can be distinguished from those of H. tokashiki sp. nov. by paired receptacular clusters with larger granules (Fig. 20 A–D). Heptathela shuri sp. nov. can also be diagnosed from all other Okinawa group Heptathela species by the following unique nucleotide substitutions in the standard DNA barcode alignment: G (89), G (131), C (134), C (396), C (510), C (512), A (524), T (641).</p> <p>Description.</p> <p>Female (Holotype). Carapace yellow brown; opisthosoma brown, tergites with brown plaques; cheliceral groove with 12 pronounced denticles; seven spinnerets. Measurements: BL 10.78, CL 5.30, CW 4.12, OL 6.25, OW 4.40; ALE&gt; PLE&gt; PME&gt; AME; palp 8.93 (3.15 + 1.43 + 2.03 + 2.32), leg I 10.91 (3.48 + 1.87 + 2.05 + 2.21 + 1.30), leg II 10.64 (3.18 + 1.85 + 1.86 + 2.28 + 1.47), leg III 11.26 (3.03 + 1.81 + 1.77 + 2.90 + 1.75), leg IV 17.02 (4.62 + 2.28 + 3.07 + 4.50 + 2.55).</p> <p>Female genitalia. A pair of depressions on the ventro-lateral part of genital atrium (Fig. 20B, D). Paired receptacular clusters along the anterior margin of bursa copulatrix, divided into two parts, with several granules, the inner granules larger than laterals, both without genital stalks (Fig. 20 A–D).</p> <p>Male. Unknown.</p> <p>Etymology.</p> <p>The species epithet, a noun in apposition, refers to the type locality.</p> <p>Distribution.</p> <p>The species is endemic to the Japanese island Okinawajima (Fig. 1C).</p></div> 	https://treatment.plazi.org/id/53D77AAC44A5548DB625D0A92B2BB232	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Xu, Xin;Ono, Hirotsugu;Kuntner, Matjaz;Liu, Fengxiang;Li, Daiqin	Xu, Xin, Ono, Hirotsugu, Kuntner, Matjaz, Liu, Fengxiang, Li, Daiqin (2019): A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands. ZooKeys 888: 1-50, DOI: http://dx.doi.org/10.3897/zookeys.888.34494, URL: http://dx.doi.org/10.3897/zookeys.888.34494
4844BEF0D3BC5B0DB6ED461817EBBFBA.text	4844BEF0D3BC5B0DB6ED461817EBBFBA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heptathela sumiyo	<div><p>Heptathela sumiyo sp. nov. Fig. 12</p> <p>Type material.</p> <p>Holotype: JAPAN · ♂; Kagoshima-ken, Amami-Oshima, Amami-shi, Sumiyo-cho, Santaro-toge Pass; 28.28N, 129.42E; alt. 360 m; 15 September 2013; D. Li and B. Wu leg.; XUX-2013-293.</p> <p>Paratypes: JAPAN · 4 ♂♂, 6 ♀♀; same data as for holotype; XUX-2013-287 to 292, 294 to 296B.</p> <p>Diagnosis.</p> <p>Males of H. sumiyo sp. nov. can be distinguished from those of H. kanenoi by the spiniform conductor apex (Fig. 12A, E, F), from those of H. amamiensis by the narrow and deeper saddle-shaped embolus in the prolateral view (Fig. 12A, E, F). Females of H. sumiyo sp. nov. resemble those of the other Amami group Heptathela species but differ from those of H. kanenoi by the tuberculate lateral receptacular clusters that are equal in size, or slightly larger than the inner clusters (Fig. 12J, K). H. sumiyo sp. nov. also differs from all other Heptathela species of the Amami group by the following unique nucleotide substitutions in the standard DNA barcode alignment: G (92), C (218), A (227), G (281), C (308), A (363), T (647).</p> <p>Description.</p> <p>Male (Holotype). Carapace brown; opisthosoma light brown, with dark brown tergites; cheliceral groove with eleven denticles; seven spinnerets. Measurements: BL 15.40, CL 7.70, CW 6.80, OL 8.00, OW 5.55; ALE&gt; PLE&gt; PME&gt; AME; leg I 23.05 (6.05 + 3.10 + 4.90 + 6.10 + 2.90), leg II 24.46 (6.48 + 3.08 + 5.05 + 6.80 + 3.05), leg III 26.36 (6.10 + 3.20 + 5.05 + 8.20 + 3.81), leg IV 32.70 (8.00 + 3.50 + 6.75 + 9.70 + 4.75).</p> <p>Palp. Prolateral side of paracymbium unpigmented and unsclerotised, numerous setae and spines at the tip of paracymbium (Fig. 12 A–C). Contrategulum with serrated margin (Fig. 12A, D, E). Tegulum wide with dentate dorsal extension of terminal apophysis (Fig. 12F, G) and blunt terminal apophysis (Fig. 12B, C, F, G). Conductor sclerotised and rugose, with several folds and a spiniform apex (Fig. 12 A–B, E–F). Embolus largely sclerotised, with a wide and flat opening, the distal part slightly sclerotised, and narrow and deep saddle-shaped in the prolateral view (Fig. 12A, D–F).</p> <p>Females (N = 6). Carapace and opisthosoma colour as in male; cheliceral groove with 13 or 14 pronounced denticles; tergites similar to those of male; seven or eight spinnerets. Measurements: BL 11.70-15.60, CL 6.25-7.55, CW 5.40-6.43, OL 5.40-8.80, OW 4.00-7.35; ALE&gt; PLE&gt; PME&gt; AME; palp 12.90 (4.50 + 2.25 + 2.70 + 3.45), leg I 14.44 (4.55 + 2.68 + 2.55 + 3.08 + 1.58), leg II 13.93 (4.09 + 2.49 + 2.35 + 3.20 + 1.80), leg III 15.61 (4.40 + 2.70 + 2.68 + 3.80 + 2.03), leg IV 22.45 (6.20 + 3.20 + 4.15 + 6.10 + 2.80).</p> <p>Female genitalia. A pair of indistinct depressions on the ventro-lateral part of genital atrium (Fig. 12J, K). Two pairs of receptacular cluster along the anterior margin of bursa copulatrix, the medians ovate, the laterals tuberculate, similar or slightly larger than inners, without genital stalks (Fig. 12 H–K).</p> <p>Etymology.</p> <p>The species epithet, a noun in apposition, refers to the type locality.</p> <p>Distribution.</p> <p>The species is known from the Japanese island Amamioshima (Fig. 1C).</p></div> 	https://treatment.plazi.org/id/4844BEF0D3BC5B0DB6ED461817EBBFBA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Xu, Xin;Ono, Hirotsugu;Kuntner, Matjaz;Liu, Fengxiang;Li, Daiqin	Xu, Xin, Ono, Hirotsugu, Kuntner, Matjaz, Liu, Fengxiang, Li, Daiqin (2019): A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands. ZooKeys 888: 1-50, DOI: http://dx.doi.org/10.3897/zookeys.888.34494, URL: http://dx.doi.org/10.3897/zookeys.888.34494
E54E646D3CE3518C9723B42967CDF3A7.text	E54E646D3CE3518C9723B42967CDF3A7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heptathela tokashiki	<div><p>Heptathela tokashiki sp. nov. Fig. 21</p> <p>Type material.</p> <p>Holotype: JAPAN · ♀; Okinawa-ken, Tokashikijima Island, Aharen; 26.19N, 127.37E; alt. 100 m; 8 May 2014; D. Li and B. Wu leg.; XUX-2014-062.</p> <p>Paratypes: JAPAN · 25 ♀♀; same data as for holotype; XUX-2014-046 to 063D · 3 ♀♀; same data as for holotype; 27 December 2012; D. Li, F.X. Liu and X. Xu leg.; XUX-2012-417, 421, 425.</p> <p>Diagnosis.</p> <p>Females of H. tokashiki sp. nov. can be distinguished from H. shuri sp. nov. by the long inner receptacular clusters with several granules (Fig. 21 A–D). Heptathela tokashiki sp. nov. can also be diagnosed from all other Okinawa group Heptathela species by the following unique nucleotide substitutions in the standard DNA barcode alignment: T (68), A (200), C (290), A (362).</p> <p>Description.</p> <p>Female (Holotype). Carapace brown; opisthosoma dark brown, with dark brown tergites close to each other; cheliceral groove with eleven pronounced denticles; seven spinnerets. Measurements: BL 11.80, CL 5.15, CW 4.10, OL 6.80, OW 5.10; ALE&gt; PLE&gt; PME&gt; AME; palp 8.60 (3.05 + 1.50 + 1.95 + 2.10), leg I 10.50 (3.35 + 1.85 + 2.15 + 2.10 + 1.05), leg II 10.45 (3.40 + 1.75 + 2.00 + 2.20 + 1.10), leg III 10.95 (3.20 + 1.85 + 1.80 + 2.60 + 1.50), leg IV 15.70 (4.60 + 2.00 + 2.80 + 3.90 + 2.40).</p> <p>Female genitalia. A pair of depressions on the ventro-lateral part of the genital atrium (Fig. 21B, D). Paired receptacular clusters along the anterior margin of bursa copulatrix, divided into two parts, the inners longer than the laterals, both with several granules, without genital stalks (Fig. 21A, B).</p> <p>Male. Unknown.</p> <p>Etymology.</p> <p>The species epithet, a noun in apposition, refers to the type locality.</p> <p>Distribution.</p> <p>The species is endemic to the Japanese island Tokashikijima (Fig. 1C).</p></div> 	https://treatment.plazi.org/id/E54E646D3CE3518C9723B42967CDF3A7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Xu, Xin;Ono, Hirotsugu;Kuntner, Matjaz;Liu, Fengxiang;Li, Daiqin	Xu, Xin, Ono, Hirotsugu, Kuntner, Matjaz, Liu, Fengxiang, Li, Daiqin (2019): A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands. ZooKeys 888: 1-50, DOI: http://dx.doi.org/10.3897/zookeys.888.34494, URL: http://dx.doi.org/10.3897/zookeys.888.34494
7F077D2BA4FC5DCC95C6D09E666966EF.text	7F077D2BA4FC5DCC95C6D09E666966EF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heptathela uken	<div><p>Heptathela uken sp. nov. Fig. 13</p> <p>Type material.</p> <p>Holotype: JAPAN · ♂; Kagoshima-ken, Amami, Uken-son, Oshima-gun, Road No. 85, Redsoil Park; 28.24N, 129.34E; alt. 260 m; 15 September 2013; D. Li and B. Wu leg.; XUX-2013-297.</p> <p>Paratypes: JAPAN · 2 ♂♂, 2 ♀♀; same data as for holotype; XUX-2013-298, 301, 302, 304 · 3 ♂♂, 5 ♀♀; Kagoshima-ken, Amami, Yamato-son, Amami Forest Park; 28.31N, 129.33E; alt. 300 m; 17 September 2013; D. Li and B. Wu leg.; XUX-2013-305 to 314.</p> <p>Diagnosis.</p> <p>Males of H. uken sp. nov. can be distinguished from those of H. kanenoi by the spiniform conductor apex (Fig. 13A, B, D, E), from those of H. amamiensis by the dorsal extension of tegular terminal apophysis without dentation (Fig. 13F, G). Females of H. uken sp. nov. cannot be diagnosed from those of the other Amami group Heptathela species morphologically (Fig. 13 H–M), only by the following unique nucleotide substitutions in the standard DNA barcode alignment: T (161), T (191), G (227), C (236), T (287), T (297), A (299), C (389), T (395), G (413), C (416), G (503), C (509), C (510), T (527), T (558), G (560), G (569), C (578), T (584), C (596), T (614), G (629), G (635), C (650), C (665).</p> <p>Description.</p> <p>Male (Holotype). Carapace brown; opisthosoma light brown, with dark brown tergites; cheliceral groove with eight denticles; seven spinnerets. Measurements: BL 11.60, CL 6.10, CW 5.60, OL 6.00, OW 4.30; ALE&gt; PLE&gt; PME&gt; AME; leg I 17.47 (4.80 + 2.28 + 3.67+ 4.42 + 2.30), leg II 17.95 (4.65 + 2.30 + 3.60 + 4.90 + 2.50), leg III 18.80 (4.45 + 2.40 + 3.55 + 5.20 + 3.20), leg IV 23.60 (5.60 + 2.45 + 4.45 + 7.20 + 3.90).</p> <p>Palp. Prolateral side of paracymbium unpigmented and unsclerotised, numerous setae and spines at the tip of paracymbium (Fig. 13 A–C). Contrategulum with serrated margin (Fig. 13A, B, D, F). Tegulum with smooth dorsal extension of terminal apophysis (Fig. 13F, G). Conductor wide, sclerotised and rugose, with several folds and a spiniform apex (Fig. 13A, B, D, F). Embolus sclerotised, wide with a flat opening, and wide saddle-shaped in the prolateral view (Fig. 13A, B, D, E).</p> <p>Females (N = 8). Carapace and opisthosoma colour as in male; cheliceral groove with 13-15 pronounced denticles; tergites similar to male; 7 or 8 spinnerets. Measurements: BL 11.80-16.00, CL 5.90-8.26, CW 5.20-7.20, OL 6.00-8.10, OW 4.60-6.00; ALE&gt; PLE&gt; PME&gt; AME; palp 10.04 (3.05 + 1.89 + 2.30 + 2.80), leg I 12.05 (3.90 + 2.10 + 2.25 + 2.40 + 1.40), leg II 11.30 (3.55 + 2.05 + 1.80+ 2.45 + 1.45), leg III 12.42 (3.60 + 2.05 + 2.10 + 3.00 + 1.67), leg IV 18.33 (5.15 + 2.40 + 3.30 + 4.78 + 2.70).</p> <p>Female genitalia. A pair of indistinct depressions on the ventro-lateral part of genital atrium (Fig. 13K). Two pairs of receptacular clusters along the anterior margin of bursa copulatrix, the inners almost globose, with short genital stalks in distal view, the laterals tuberculate, without genital stalks (Fig. 13 H–M).</p> <p>Etymology.</p> <p>The species epithet, a noun in apposition, refers to the type locality.</p> <p>Distribution.</p> <p>The species is known from the Japanese island Amamioshima (Fig. 1C).</p></div> 	https://treatment.plazi.org/id/7F077D2BA4FC5DCC95C6D09E666966EF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Xu, Xin;Ono, Hirotsugu;Kuntner, Matjaz;Liu, Fengxiang;Li, Daiqin	Xu, Xin, Ono, Hirotsugu, Kuntner, Matjaz, Liu, Fengxiang, Li, Daiqin (2019): A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands. ZooKeys 888: 1-50, DOI: http://dx.doi.org/10.3897/zookeys.888.34494, URL: http://dx.doi.org/10.3897/zookeys.888.34494
5B74DB6FF1ED56B4B4B202B6EFFA3D5D.text	5B74DB6FF1ED56B4B4B202B6EFFA3D5D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heptathela unten	<div><p>Heptathela unten sp. nov. Figs 22, 23</p> <p>Type material.</p> <p>Holotype: JAPAN · ♂; Okinawa-ken, Nakijin-son, Unten Port; 26.68N, 128.00E; alt. 25 m; 27 December 2012; D. Li, F.X. Liu and X. Xu leg.; XUX-2012-522.</p> <p>Paratypes: JAPAN · 1 ♂, 2 ♀♀; same data as for holotype; XUX-2012-523, 527, 528A · 3 ♀♀; same data as for holotype; 10 May 2014; D. Li and B. Wu leg.; XUX-2014-083, 083A, 083B.</p> <p>Diagnosis.</p> <p>Males of H. unten sp. nov. can be distinguished from those of H. yanbaruensis by the blunt tegular marginal apophysis (Fig. 22A, C, D); from those of H. helios by the conductor with serrated margin and small tegular marginal apophysis (Fig. 22A, C). Females of H. unten sp. nov. cannot be distinguished morphologically form those of H. crypta sp. nov. (Fig. 23 A–L). However, H. unten sp. nov. can be diagnosed from all other Okinawa group Heptathela species by the following unique nucleotide substitutions in the standard DNA barcode alignment: G (482), C (635).</p> <p>Description.</p> <p>Male (Holotype). Carapace yellow brown; opisthosoma brown, with dark brown tergites close to each other; cheliceral groove with ten denticles; seven spinnerets. Measurements: BL 10.00, CL 5.05, CW 4.60, OL 4.60, OW 2.90; ALE&gt; PLE&gt; PME&gt; AME; leg I 15.71 (4.48 + 1.93 + 3.30 + 4.00 + 2.00), leg II 16.59 (4.45 + 1.90 + 3.37 + 4.62 + 2.25), leg III 18.29 (4.50 + 1.93 + 3.58 + 5.53 + 2.75), leg IV 23.40 (5.82 + 2.28 + 4.52 + 7.30 + 3.48).</p> <p>Palp. Prolateral side of paracymbium unpigmented and unsclerotised, numerous setae and spines at the tip of paracymbium (Fig. 22A, B). Contrategulum margin incurved nearly in the middle and the contrategulum divided into proximally serrated and distally smooth margins (Fig. 22A, D). Tegulum with wide dorsal extension of terminal apophysis (Fig. 22C, D), blunt terminal and small marginal apophysis (Fig. 22A, C, D). Conductor sclerotised and ovate, prolateral conductor with one or two shallow folds, and with a serrated margin (Fig. 22B, C). Embolus sclerotised, with a wide opening, the distal margin slightly sclerotised, and with a saddle-shaped margin in the retrolateral view (Fig. 22B, C).</p> <p>Females (N = 5). Carapace and opisthosoma colour as in male, dark brown tergites separated from each other; cheliceral groove with 12 or 13 pronounced denticles; seven or eight spinnerets. Measurements: BL 7.81-12.00, CL 3.60-4.55, CW 3.30-4.40, OL 4.10-7.60, OW 3.00-6.20; ALE&gt; PLE&gt; PME&gt; AME; palp 6.64 (2.19 + 1.22 + 1.45 + 1.78), leg I 7.48 (2.47 + 1.35 + 1.22 + 1.52 + 0.92), leg II 7.40 (2.21 + 1.30 + 1.21 + 1.65 + 1.03), leg III 7.79 (2.11 + 1.38 + 1.20 + 1.90 + 1.20), leg IV 11.55 (3.20 + 1.55 + 2.10 + 3.05 + 1.65).</p> <p>Female genitalia. A pair of depressions on the ventro-lateral part of genital atrium indistinct (Fig. 23D, E). Paired receptacular clusters along the anterior margin of bursa copulatrix, divided into two parts, the inners similar or smaller than the laterals, paired receptacular clusters tuberculate, inners with or without genital stalks (Fig. 23A, B, D, E).</p> <p>Etymology.</p> <p>The species epithet, a noun in apposition, refers to the type locality, Unten Port.</p> <p>Distribution.</p> <p>The species is endemic to the Japanese island Okinawajima (Fig. 1C).</p></div> 	https://treatment.plazi.org/id/5B74DB6FF1ED56B4B4B202B6EFFA3D5D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Xu, Xin;Ono, Hirotsugu;Kuntner, Matjaz;Liu, Fengxiang;Li, Daiqin	Xu, Xin, Ono, Hirotsugu, Kuntner, Matjaz, Liu, Fengxiang, Li, Daiqin (2019): A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands. ZooKeys 888: 1-50, DOI: http://dx.doi.org/10.3897/zookeys.888.34494, URL: http://dx.doi.org/10.3897/zookeys.888.34494
93A7B0B1F5C751FDAF1490890EE18AF2.text	93A7B0B1F5C751FDAF1490890EE18AF2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heptathela yakushimaensis Ono 1998	<div><p>Heptathela yakushimaensis Ono, 1998 Fig. 8</p> <p>Heptathela yakushimaensis Ono, 1998: 23 (holotype: female, from Mt. Kunibaidake, Yakushima Island, Kagoshima-ken, Japan, collected by A. Tanikawa on 15 July 1990, deposited in NMNS, examined); Ono 2009: 83.</p> <p>Heptathela kimurai yakushimaensis: Haupt 2003: 69.</p> <p>Diagnosis.</p> <p>Males of H. yakushimaensis differ from those of all other Kyushu group Heptathela species by a strongly serrated prolateral conductor margin (Fig. 8G, H, J, K), the tapered tegular marginal apophysis (Fig. 8L, M), and a larger tegular terminal apophysis (Fig. 8L, M). Females of H. yakushimaensis differ from those of H. kimurai and H. higoensis by the finely granulated inner receptacular clusters that are smaller than the outer ones (Fig. 8 A–F), and from those of H. kikuyai by the inner receptacular clusters lacking well defined stalks. H. yakushimaensis also differs from all other Kyushu group Heptathela species by the following unique nucleotide substitutions in the standard DNA barcode alignment: T (56), A (68), T (74), G (77), C (84), C (89), C (95), G (98), C (107), A (110), C (122), T (131), T (143), T (164), T (167), C (188), C (200), C (212), T (215), C (216), G (218), T (236), G (242), T (248), G (251), C (278), A (284), T (293), C (294), C (308), T (323), C (347), C (356), T (392), T (395), C (396), T (401), T (407), C (411), C (413), C (417), G (422), G (425), C (437), C (438), A (443), T (455), T (458), T (461), T (482), T (488), C (510), T (528), A (530), A (536), A (539), T (548), G (557), C (567), C (584), T (591), T (632), T (635), G (638), T (650), A (665).</p> <p>Description.</p> <p>Males (N = 2). Carapace brown; opisthosoma light brown, with dark brown tergites close to each other; cheliceral groove with 10-13 denticles; 7 spinnerets. Measurements: BL 8.70-10.50, CL 4.27-5.50, CW 4.00-4.90, OL 3.80-6.10, OW 2.40-4.40; ALE&gt; PLE&gt; PME&gt; AME; leg I 13.27 (3.70 + 1.60 + 2.72 + 3.50 + 1.75), leg II 13.71 (3.55 + 1.65 + 2.80 + 3.75 + 1.96), leg III 14.65 (3.52 + 1.65 + 2.78 + 4.50 + 2.20), leg IV 18.64 (4.35 + 1.70 + 3.72 + 6.20 + 2.67).</p> <p>Palp. Prolateral side of paracymbium unpigmented and unsclerotised, numerous setae and spines at the tip of paracymbium (Fig. 8 G–I). Contrategulum with serrated margin proximally and smooth margin distally, and slightly curved at the proximal 2/3 of contrategulum (Fig. 8G, H, J, K). Tegulum with serrated margin, widest in the middle (Fig. 8 K–M). Conductor prolateral margin strongly serrated (Fig. 8G, H, J, K). Embolus wide with two peaks (Fig. 8 G–K).</p> <p>Females (N = 8). Carapace and opisthosoma colour as in male; cheliceral groove with 12-15 pronounced denticles; tergites similar to male; 7 spinnerets. Measurements: BL 9.10-14.80, CL 4.50-6.80, CW 4.05-5.90, OL 4.70-8.60, OW 3.00-6.40; ALE&gt; PLE&gt; PME&gt; AME; palp 9.72 (3.22 + 1.70 + 2.10 + 2.70), leg I 11.15 (3.45 + 1.90 + 2.05 + 2.35 + 1.40), leg II 9.61 (3.07 + 1.90 + 1.00 + 2.36 + 1.28), leg III 11.75 (3.25 + 1.95 + 2.05 + 2.80 + 1.70), leg IV 16.30 (4.25 + 2.25 + 2.90 + 4.40 + 2.50).</p> <p>Female genitalia. A pair of depressions on the ventro-lateral part of genital atrium. A pair of receptacular clusters along the anterior margin of bursa copulatrix, divided into two parts, the inner part is similar or smaller than the outer part, on which there are several small granules (Fig. 8 A–F).</p> <p>Material examined.</p> <p>JAPAN · 2 ♂♂, 8 ♀♀; Kagoshima-ken, Kumage-gun, Yakushima-cho, Mt. Kankake-dake; 30.37N, 130.39E; alt. 170 m; 24 September 2013; D. Li and B. Wu leg.; XUX-2013-490, 491, 493 to 500 (500, ♂ matured 2 August 2014 at CBEE).</p> <p>Distribution.</p> <p>The species is known from the Kagoshima prefecture on the Japanese island Yakushima (Fig. 1C).</p></div> 	https://treatment.plazi.org/id/93A7B0B1F5C751FDAF1490890EE18AF2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Xu, Xin;Ono, Hirotsugu;Kuntner, Matjaz;Liu, Fengxiang;Li, Daiqin	Xu, Xin, Ono, Hirotsugu, Kuntner, Matjaz, Liu, Fengxiang, Li, Daiqin (2019): A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands. ZooKeys 888: 1-50, DOI: http://dx.doi.org/10.3897/zookeys.888.34494, URL: http://dx.doi.org/10.3897/zookeys.888.34494
F2DC85B7FA1B59F284FE30521690E460.text	F2DC85B7FA1B59F284FE30521690E460.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heptathela yanbaruensis Haupt 1983	<div><p>Heptathela yanbaruensis Haupt, 1983 Fig. 14</p> <p>Heptathela kimurai yanbaruensis Haupt, 1983: 284 (holotype: male, from Yona, Okinawa, Japan, collected by J. Haupt on 15 April 1977, deposited in ZMH, where the type may be lost (Dunlop et al. 2014); Haupt 1984: 166; Haupt 2003: 69. Heptathela yanbaruensis: Ono 2009: 80; Ono and Ogata 2018: 28, 480.</p> <p>Diagnosis.</p> <p>Males of H. yanbaruensis can be distinguished from those of H. helios by the contrategulum that is distinctly curved in the middle (Fig. 14I, J), and from those of H. unten sp. nov. and H. crypta sp. nov. by the conductor that is longer than its width (Fig. 14I, J). Females of H. yanbaruensis can be distinguished from those of H. shuri sp. nov. by the wide and straight posterior margin of the genital atrium (Fig. 14A, C). Heptathela yanbaruensis can also be diagnosed from all other Heptathela species of the Okinawa group by the following unique nucleotide substitutions in the standard DNA barcode alignment: G (53), T (327), A (356), A (443).</p> <p>Description.</p> <p>Males (N = 3). Carapace yellow brown; opisthosoma light brown, with dark-spotted tergites close to each other; cheliceral groove with 10-12 denticles; seven spinnerets. Measurements: BL 8.53-9.22, CL 4.21-4.57, CW 3.71-3.75, OL 4.20-4.48, OW 2.30-2.80; ALE&gt; PLE&gt; PME&gt; AME; leg I 14.28 (4.15 + 1.70 + 3.05 + 3.56 + 1.82), leg II 15.15 (3.97 + 1.70 + 3.07 + 4.13 + 2.28), leg III 15.24 (4.00 + 1.68 + 3.08 + 4.10 + 2.38), leg IV 19.78 (4.75 + 1.37 + 3.91 + 6.45 + 3.30).</p> <p>Palp. Prolateral side of paracymbium unpigmented and unsclerotised, numerous setae and spines at the tip of paracymbium (Fig. 14 E–G). Contrategulum margin obviously curved in the middle, the contrategulum margin proximally serrated and distally smooth (Fig. 14 H–J). Tegulum wide with a dentate dorsal extension of terminal apophysis (Fig. 14J, K), blunt terminal and marginal apophysis (Fig. 14J, K). Conductor oval, with weakly serrated margin, and a fold in prolateral view (Fig. 14 H–J). Embolus sclerotised, with a wide opening, the distal margin slightly sclerotised, and with a wide saddle-shaped margin in the retrolateral view (Fig. 14 I–K).</p> <p>Females (N = 15). Carapace and opisthosoma colour as in male; cheliceral groove with 11-14 pronounced denticles; tergites similar to male; seven spinnerets. Measurements: BL 7.78-10.33, CL 3.90-5.10, CW 3.40-4.23, OL 3.90-6.00, OW 2.80-4.70; ALE&gt; PLE&gt; PME&gt; AME; palp 8.70 (2.96 + 1.51 + 2.11 + 2.12), leg I 10.20 (3.12 + 1.75 + 1.93 + 2.12 + 1.28), leg II 9.97 (3.05 + 1.72 + 1.77 + 2.13 + 1.30), leg III 10.05 (2.91 + 1.78 + 1.48 + 2.47 + 1.41), leg IV 14.77 (4.08 + 2.10 + 2.51 + 3.97 + 2.11).</p> <p>Female genitalia. A pair of depressions on the ventro-lateral part of the genital atrium (Fig. 14C, D). The posterior margin of genital atrium wide and straight (Fig. 14 A–D). Paired receptacular clusters along the anterior margin of bursa copulatrix, divided into two parts, the size of inner ones similar to that of laterals, with several granules, with short genital stalks (Fig. 14 A–D).</p> <p>Material examined.</p> <p>JAPAN · 3 ♂♂, 9 ♀♀; Okinawa-ken, Kunigami-son, Yona, Tropical Biosphere Research Centre field station, University of the Ryukyu; 26.76N, 128.22E; alt. 20 m; 18 December 2012; D. Li, F.X. Liu and X. Xu leg.; XUX-2012-310 to 322B · 6 ♀♀; Okinawa-ken, Ogimi-son, Taiho Dam; 26.65N, 128.16E; alt. 80 m; 24 December 2012; D. Li, F.X. Liu and X. Xu leg.; XUX-2012-441 to 446.</p> <p>Distribution.</p> <p>The species is endemic to the Japanese island Okinawajima (Fig. 1C).</p></div> 	https://treatment.plazi.org/id/F2DC85B7FA1B59F284FE30521690E460	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Xu, Xin;Ono, Hirotsugu;Kuntner, Matjaz;Liu, Fengxiang;Li, Daiqin	Xu, Xin, Ono, Hirotsugu, Kuntner, Matjaz, Liu, Fengxiang, Li, Daiqin (2019): A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands. ZooKeys 888: 1-50, DOI: http://dx.doi.org/10.3897/zookeys.888.34494, URL: http://dx.doi.org/10.3897/zookeys.888.34494
