identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
ED14ECDE65F852FBABC12D9997C01BFF.text	ED14ECDE65F852FBABC12D9997C01BFF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyleoglomeris alba Nguyen, Kuroda & Eguchi 2022	<div><p>Hyleoglomeris alba Nguyen, Kuroda &amp; Eguchi sp. nov.</p><p>Figs 6, 7, 8, 9, 10, 11</p><p>Material examined.</p><p>Holotype. Vietnam: male; Cao Bang Province, Tra Linh District, Quoc Toan commune, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.2911&amp;materialsCitation.latitude=22.765" title="Search Plazi for locations around (long 106.2911/lat 22.765)">Thang Hen lake</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.2911&amp;materialsCitation.latitude=22.765" title="Search Plazi for locations around (long 106.2911/lat 22.765)">Ky Rang cave</a>, 22.7650N, 106.2911E, 2 Nov. 2021, leg. AD Nguyen (IEBR-Myr 919) . Paratypes. Vietnam: 2 females, Cao Bang Province, Tra Linh District, Quoc Toan commune, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.2911&amp;materialsCitation.latitude=22.765" title="Search Plazi for locations around (long 106.2911/lat 22.765)">Thang Hen lake</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.2911&amp;materialsCitation.latitude=22.765" title="Search Plazi for locations around (long 106.2911/lat 22.765)">Ky Rang cave</a>, 22.7650N, 106.2911E, 12 Oct. 2020, leg. AD Nguyen, VT Mai &amp; VD Dang (IEBR-Myr 917) ; 1 male, Cao Bang Province, Tra Linh District, Quoc Toan commune, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.2911&amp;materialsCitation.latitude=22.765" title="Search Plazi for locations around (long 106.2911/lat 22.765)">Thang Hen lake</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.2911&amp;materialsCitation.latitude=22.765" title="Search Plazi for locations around (long 106.2911/lat 22.765)">Ky Rang cave</a>, 22.7650N, 106.2911E, 17 March 2022, leg. AD Nguyen &amp; DD Nguyen (IEBR-Myr 928) .</p><p>Diagnosis.</p><p>The species can be recognized by a completely troglobiotic form with no eyes, an unpigmented body, and a roundly triangular syncoxital lobe.</p><p>According to Golovatch et al. (2006, 2013), the new species seems to belong to the troglobiont species group containing H. speophila, H. spelaea, H. cavernicola, H. differens Golovatch, Geoffroy &amp; Mauriès, 2006, H. reducta Golovatch, Geoffroy &amp; Mauriès, 2006, and H. albicorporis Zhang &amp; Zhang, 1995. These species are characterized by a totally unpigmented body. However, the new species differs from these species in lacking ocelli and the telopods bearing a roundly triangular syncoxital lobe. In contrast, the other species have convex ocelli, telopods with a roundly subtraperziform syncoxital lobe ( H. speophila, H. cavernicola, H. reducta, H. albicorporis), or a subquadrate syncoxital lobe ( H. spelaea), or a roundly triangular syncoxital lobe ( H. differens).</p><p>Etymology.</p><p>From the Latin Hyleoglomeris alba, meaning white. It was used to emphasize the unpigmented body of the new species.</p><p>Description.</p><p>Body length 4.38 mm, width of the second segment ~ 2.02 mm. Color entirely white, unpigmented (Figs 6 - 8). Ocelli totally absent. TÖmÖsvary’s organ transverse, strongly horseshoe-shaped, ~ 2 × as wide as long (Fig. 8B). Antennae long and slender, antennomere, ~ 3 × longer than wide, antennal tip with four apical sensory cones (Fig. 8A).</p><p>Collum semicircular, with a trace of a transverse oval spot in the center and two distinctly transverse striae (Fig. 7D). Second tergum with a narrow hyposchism, not reaching the caudal margin, with seven or eight striae, five or six of which cross the dorsum. Anal shield rounded, very slightly concave medio-caudally (Fig. 8D).</p><p>Leg-pair 17 (Figs 9A, 11A) strongly reduced, with four podomeres, with a high, regularly rounded, outer coxal lobe; coxa with an apical setiferous spine; leg-pair 18 (Figs 9B, 11B) also strongly reduced, but more developed in comparison with leg-pair 17, with four podomeres, and a simple V-shaped syncoxital notch.</p><p>Telopods (Figs 9C, D, 10, 11C-E) with a roundly triangular, sparsely setose, central syncoxital lobe (syl) accompanying two setiferous syncoxital horns (syh), each directed ventrad, slightly longer than the syncoxital lobe, tip crowned with an apical setoid. Prefemur (pre) and femur (fe) with long trichosteles (pret and fet), prefemoral one (pret) longer than femoral one (fet). Prefemur without additional processes. Distomesal process of femur (dpf) large, long, straight, rectangular; distal part tuberculiform and strongly curved downwards, directed laterodorsad. Tibia (ti) with a shorter triangular distolateral process (dpt). Tarsus (ta) slightly sigmoid anteriomesad, subacuminate apically, with a seta distoventrally.</p><p>DNA barcode.</p><p>We failed to amplify the COI fragments of this species.</p><p>Remarks.</p><p>The species was collected from the totally dark region in the cave. The completely unpigmented body without ocelli, and with long, slender antennae suggest that this species is a true troglobiont. These characters were also mentioned in Liu et al. (2017) who reviewed the morphological adaptations seen in troglobitic glomerids and other millipedes.</p></div>	https://treatment.plazi.org/id/ED14ECDE65F852FBABC12D9997C01BFF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Kuroda, Mai;Eguchi, Katsuyuki;Oguri, Emiko;Nguyen, Anh D.	Kuroda, Mai, Eguchi, Katsuyuki, Oguri, Emiko, Nguyen, Anh D. (2022): Two new cave Hyleoglomeris species (Glomerida, Glomeridae) from northern Vietnam. ZooKeys 1108: 161-174, DOI: http://dx.doi.org/10.3897/zookeys.1108.85423, URL: http://dx.doi.org/10.3897/zookeys.1108.85423
2B6FD3D5229159F0A2452FD410E545D2.text	2B6FD3D5229159F0A2452FD410E545D2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyleoglomeris halang Kuroda, Nguyen & Eguchi 2022	<div><p>Hyleoglomeris halang Kuroda, Nguyen &amp; Eguchi sp. nov.</p><p>Figs 1, 2, 3, 4, 5</p><p>Material examined.</p><p>Holotype. Vietnam: male, Cao Bang Province, Ha Lang District, Duc Quang commune, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.6692&amp;materialsCitation.latitude=22.7208" title="Search Plazi for locations around (long 106.6692/lat 22.7208)">Quang Hoai village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.6692&amp;materialsCitation.latitude=22.7208" title="Search Plazi for locations around (long 106.6692/lat 22.7208)">Nguom Hang cave</a>, 22.7208N, 106.6692E, 10 Oct 2020, coll. AD Nguyen, VD Dang &amp; VT Mai (IEBR-Myr 898H) . Paratypes. Vietnam: 1 male, 1 female; Cao Bang Province, Ha Lang District, Duc Quang commune, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.6692&amp;materialsCitation.latitude=22.7208" title="Search Plazi for locations around (long 106.6692/lat 22.7208)">Quang Hoai village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.6692&amp;materialsCitation.latitude=22.7208" title="Search Plazi for locations around (long 106.6692/lat 22.7208)">Nguom Hang cave</a>, 22.7208N, 106.6692E, 10 Oct 2020, coll. AD Nguyen, VD Dang &amp; VT Mai (IEBR-Myr 898P) ; 1 male, 1 female; Cao Bang Province, Ha Lang District, Duc Quang commune, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.6692&amp;materialsCitation.latitude=22.7208" title="Search Plazi for locations around (long 106.6692/lat 22.7208)">Quang Hoai village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.6692&amp;materialsCitation.latitude=22.7208" title="Search Plazi for locations around (long 106.6692/lat 22.7208)">Nguom Hang cave</a>, 22.7208N, 106.6692E, 15 March 2022, coll. AD Nguyen &amp; DD Nguyen (IEBR-Myr 926) .</p><p>Diagnosis.</p><p>The species differs from its congeners in having a distinct body color pattern of white with oval-black spots on terga 4 and 5 and tadpole-shaped black bands on the thoracic shield and terga 6-9; telopods with a large, square/rectangular, slightly concave medially, sparsely setose, central syncoxital lobe; and syncoxital horns ~ 1.5 × as long as the lobe.</p><p>The new species can be keyed out at the 26th node in Golovatch et al. (2006), characterized by a thoracic shield and pygidium that are entirely or mainly light and whitish to brown, as well as spots or markings, when present, that is darker than the background. Thus, this species can be grouped with Hyleoglomeris triangulifera Attems, 1938 and Hyleoglomeris siamensis (Silvestri, 1917). However, its smaller size distinguishes it from both species (3.5 mm wide vs. 6.5 mm and 5.0 mm wide).</p><p>Etymology.</p><p>The new species is named after the Ha Lang District where the types were found. Noun in apposition.</p><p>Description.</p><p>Body length 6 mm, width of the second segment ca. 3.5 mm. Coloration shown as in Figs 2 and 3. Generally white with a line of symmetrical, marbled, black, oval spots at terga 4 and 5, tadpole-shaped black bands on thoracic shield and terga 6-9. Anal shield white, with two laterally symmetrical triangular black spots.</p><p>Head. Ocelli 6+1; lenses convex, black contrasting against to a pale blackish background of the head. Tömösváry’s organs transverse and strongly horseshoe-shaped, ~ 2 × as wide and long. Antennae clavate apically; antennomere 6 large, ~ 2.5 × longer than wide; antennal tip with four large, apical sensory cones.</p><p>Collum semi-circular, with a very large marbled white spot in the center and two transverse striae; the other parts of the collum pale black. The thoracic shield has a narrow hyposchism, not reaching the caudal margin, with 12 superficial transverse striae, eight of which cross the dorsum.</p><p>Leg-pair 17 strongly reduced, 4-segmented with a high, regularly rounded, outer coxal lobe (Figs 3D, 5A). Leg-pair 18 also reduced, but more developed in comparison with leg-pair 17, 4-segmented with a simple V-shaped syncoxital notch.</p><p>Telopods (Figs 4, 5B-D) with a large, quadrate, slightly concave medially, sparsely setose, central syncoxital lobe (syl) accompanying two setiferous horns (syh), each directed subventrad, 1.5 × longer than the syncoxital lobe, tip crowned with an apical setoid. Prefemur (pre) and femur (fe) with long trichosteles (pret and fet), prefemoral one (pret) longer than femoral one (fet). Prefemur without additional processes. Distomesal process of femur (dpf) large, long, straight, lamelliform; distal part tuberculiform and strongly curved down, directed laterodorsad. Tibia (ti) with a shorter triangular distolateral process (dpt). Tarsus (ta) slightly sigmoid anteriomesad, subacuminate apically with a seta distoventrally.</p><p>Variability.</p><p>Syncoxital horns can be 2.0 × longer than the syncoxital lobe, each horn directed ventromesad. Two trichosteles are almost of the same in length.</p><p>DNA barcode.</p><p>The COI barcode data (679 bp fragment of the COI) for the paratype was uploaded to GenBank under the accession numbers ON704753 and ON704754. The new species shares 86.98% and 85.89% of its identity with Hyleoglomeris lobus Nguyen, Sierwald &amp; Marek, 2019 (MT749402) and Hyleoglomeris hoanglien Nguyen, Eguchi &amp; Hwang, 2019 (MH248038), respectively.</p><p>Remarks.</p><p>This species is not a true cave inhabitant. However, it was discovered in the cave entrance (Fig. 1), and its body is less pigmented with large white areas, suggesting that this species is adapting to a cave-dwelling life.</p></div>	https://treatment.plazi.org/id/2B6FD3D5229159F0A2452FD410E545D2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Kuroda, Mai;Eguchi, Katsuyuki;Oguri, Emiko;Nguyen, Anh D.	Kuroda, Mai, Eguchi, Katsuyuki, Oguri, Emiko, Nguyen, Anh D. (2022): Two new cave Hyleoglomeris species (Glomerida, Glomeridae) from northern Vietnam. ZooKeys 1108: 161-174, DOI: http://dx.doi.org/10.3897/zookeys.1108.85423, URL: http://dx.doi.org/10.3897/zookeys.1108.85423
