taxonID	type	description	language	source
066E87D24969FFF7FF5388DDFD2AFE62.taxon	materials_examined	Material examined. BRASIL, Amazonas, Ipixuna, Rio Gregrório com [munity] Lago Grande, 07 ° 10 ’ 11.7 ’’ S 70 ° 49 ’ 10.3 ’’ W, 18 – 20. v. 2011, Malaise [trap], J. A. Rafael, J. T. Câmara, F. Silva, A. Somavilla, C. Gonçalves leg. (3 INPA).	en	Chagas-Vieira, Cínthia (2012): New records of Tonnoira Enderlein (Diptera, Psychodidae, Psychodinae) from the Brazilian Amazon with descriptions of two new species. Zootaxa 3318: 51-56, DOI: 10.5281/zenodo.213567
066E87D24969FFF7FF5388DDFD2AFE62.taxon	diagnosis	Diagnosis. Eyes separated by little less than 1 facet diameter; eye bridge with 4 facet rows except median margin with only 3 rows; interocular suture as inverted Y-shaped, stem longer than arm (Fig. 1). Flagellomere 1 elongate, fusiform, nearly as long as scape and pedicel combined (Fig. 2). Proportion of palp segments = 1.0: 2.3: 2.7: 2.8 (Fig. 3). Wing brown with radial fork and medial fork complete (Fig. 4). Gonocoxal apodeme with pair of slender, acute processes flanking aedeagus and extending nearly to tip of aedeagus (Fig. 5); paramere bluntly rounded, slightly shorter than gonocoxal processes (Fig. 5); epandrium rather subrectangular with two foramina near anterior margin (Fig. 6); cercus with 2 tenacula, basal separated from apical by more than width of cercus (Fig. 7).	en	Chagas-Vieira, Cínthia (2012): New records of Tonnoira Enderlein (Diptera, Psychodidae, Psychodinae) from the Brazilian Amazon with descriptions of two new species. Zootaxa 3318: 51-56, DOI: 10.5281/zenodo.213567
066E87D24969FFF7FF5388DDFD2AFE62.taxon	description	Variation. Male: Aedeagus as long as paramere.	en	Chagas-Vieira, Cínthia (2012): New records of Tonnoira Enderlein (Diptera, Psychodidae, Psychodinae) from the Brazilian Amazon with descriptions of two new species. Zootaxa 3318: 51-56, DOI: 10.5281/zenodo.213567
066E87D24969FFF7FF5388DDFD2AFE62.taxon	distribution	Distribution. Panama and Brazil (new record).	en	Chagas-Vieira, Cínthia (2012): New records of Tonnoira Enderlein (Diptera, Psychodidae, Psychodinae) from the Brazilian Amazon with descriptions of two new species. Zootaxa 3318: 51-56, DOI: 10.5281/zenodo.213567
066E87D2496BFFF7FF538C25FB6CFA6E.taxon	materials_examined	HOLOTYPE 3. BRASIL, Amazonas, Ipixuna, Rio Gregrório com Lago Grande, 07 ° 10 ’ 11.7 ’’ S 70 ° 49 ’ 10.3 ’’ W, 18 – 20. v. 2011, Malaise [trap], J. A. Rafael, J. T. Câmara, F. Silva, A. Somavilla, C. Gonçalves leg. (INPA).	en	Chagas-Vieira, Cínthia (2012): New records of Tonnoira Enderlein (Diptera, Psychodidae, Psychodinae) from the Brazilian Amazon with descriptions of two new species. Zootaxa 3318: 51-56, DOI: 10.5281/zenodo.213567
066E87D2496BFFF7FF538C25FB6CFA6E.taxon	etymology	Etymology. From Latin spina, meaning “ spine ”, in reference to the tip of the dorsal paramere.	en	Chagas-Vieira, Cínthia (2012): New records of Tonnoira Enderlein (Diptera, Psychodidae, Psychodinae) from the Brazilian Amazon with descriptions of two new species. Zootaxa 3318: 51-56, DOI: 10.5281/zenodo.213567
066E87D2496BFFF7FF538C25FB6CFA6E.taxon	diagnosis	Diagnosis. Eyes separated by 1 facet diameter; eye bridge with 4 facet rows; interocular suture inverted Y shaped; flagellomeres cylindrical; first flagellomere 1.3 X length of second; base of R 2 + 3 not attached to R 4; hypandrium narrow; gonostyli with divergent apex; cercus with 3 tenacula; aedeagus bipartite; paramere lying close to the hypandrium.	en	Chagas-Vieira, Cínthia (2012): New records of Tonnoira Enderlein (Diptera, Psychodidae, Psychodinae) from the Brazilian Amazon with descriptions of two new species. Zootaxa 3318: 51-56, DOI: 10.5281/zenodo.213567
066E87D2496BFFF7FF538C25FB6CFA6E.taxon	description	Description. Head (Fig. 8): eyes separated by 1 facet diameter; eye bridge with 4 facet rows; interocular suture as inverted Y shaped, stem as long as branches. Antenna (Fig. 9): scape cylindrical; pedicel spherical; flagellomeres elongate, cylindrical; first flagellomere 1.3 X length of second; ascoids not visible. Proportion of palp segments = 1.0: 1.8: 1.9: 2 (Fig. 10). Wing (Fig. 11) light brown; Sc short; base of R 2 + 3 not attached to R 4; radial fork basal to medial fork; medial fork complete; R 5 ending at wing apex. Male terminalia: hypandrium a distinct band connecting gonocoxites, narrow and archlike; gonocoxite 0.9 X length of the gonostylus with long setae along inner margin (Fig. 12); gonostyli with apices curved laterally, pointed (Fig. 12); aedeagus bipartite (Fig. 12); paramere lying close to the hypandrium (Fig. 12); gonocoxal apodeme fused at midline and articulated with the ejaculatory apodeme (Fig. 12). Epandrium subrectangular with one oval foramen near anterior margin; cercus 1.1 X length of epandrium, slightly curved, with 3 tenacula (Fig. 13); tergite 10 with posterior margin rounded (Fig. 13). Female. Unknown.	en	Chagas-Vieira, Cínthia (2012): New records of Tonnoira Enderlein (Diptera, Psychodidae, Psychodinae) from the Brazilian Amazon with descriptions of two new species. Zootaxa 3318: 51-56, DOI: 10.5281/zenodo.213567
066E87D2496BFFF7FF538C25FB6CFA6E.taxon	discussion	Remarks. Tonnoira spina sp. n. appears to be closely related to T. mirabilis based on eyes separated by 1 facet diameter, eye bridge with 4 facet rows, flagellomeres elongate, cylindrical, aedeagus bipartite, epandrium wider than long and cercus with 3 tenacula, but T. spina has interocular suture inverted Y-shaped; dorsal margin of cercus without projections and tergite 10 with posterior margin rounded. T. mirabilis has interocular suture without median spur; dorsal margin of cercus with 3 short projections and 3 – 4 bristles near base and tergite 10 subtriangular. Moreover, the aedeagus of T. spina is longer and thinner than the aedeagus of T. mirabilis.	en	Chagas-Vieira, Cínthia (2012): New records of Tonnoira Enderlein (Diptera, Psychodidae, Psychodinae) from the Brazilian Amazon with descriptions of two new species. Zootaxa 3318: 51-56, DOI: 10.5281/zenodo.213567
066E87D2496BFFF1FF538819FAE5F8CE.taxon	materials_examined	HOLOTYPE 3. BRASIL, Amazonas, Ipixuna, Rio Gregrório com [munity] Lago Grande, 07 ° 10 ’ 11.7 ’’ S 70 ° 49 ’ 10.3 ’’ W, 18 – 20. v. 2011, Malaise [trap], J. A. Rafael, J. T. Câmara, F. Silva, A. Somavilla, C. Gonçalves leg. (3 INPA).	en	Chagas-Vieira, Cínthia (2012): New records of Tonnoira Enderlein (Diptera, Psychodidae, Psychodinae) from the Brazilian Amazon with descriptions of two new species. Zootaxa 3318: 51-56, DOI: 10.5281/zenodo.213567
066E87D2496BFFF1FF538819FAE5F8CE.taxon	etymology	Etymology. From Latin tripenis, meaning “ three aedeagus ”, in reference to aedeagus tripartite.	en	Chagas-Vieira, Cínthia (2012): New records of Tonnoira Enderlein (Diptera, Psychodidae, Psychodinae) from the Brazilian Amazon with descriptions of two new species. Zootaxa 3318: 51-56, DOI: 10.5281/zenodo.213567
066E87D2496BFFF1FF538819FAE5F8CE.taxon	diagnosis	Diagnosis. Eyes separated by little less than 1 facet diameter; eye bridge with 4 facet rows; hypandrium a distinct band connecting gonocoxites; aedeagus trifid; no parameres; cercus with 1 apical tenaculum.	en	Chagas-Vieira, Cínthia (2012): New records of Tonnoira Enderlein (Diptera, Psychodidae, Psychodinae) from the Brazilian Amazon with descriptions of two new species. Zootaxa 3318: 51-56, DOI: 10.5281/zenodo.213567
066E87D2496BFFF1FF538819FAE5F8CE.taxon	description	Description. Head (Fig. 14): eyes separated by little less than 1 facet diameter; eye bridge with 4 facet rows; interocular suture as inverted Y shaped. Palp segment 1 without sensilla. Wing (Fig. 15) light brown; Sc short; base of R 2 + 3 not attached to R 4; radial fork basal to medial fork; medial fork complete; R 5 ending at wing apex. Male terminalia: hypandrium a distinct band connecting gonocoxites; gonocoxite 0.8 X length of the gonostylus (Fig. 16); gonostyli with pointed apex (Fig. 16); aedeagus tripartite, with L-shaped lateral shaft, central part broad over basal half and tapering to apex over distal half and the last shaft with basal 2 / 3 wide and apical 1 / 3 narrow (Fig. 16); paramere absent (Fig. 16); gonocoxal apodeme fused (Fig. 16). Epandrium subrectangular with one larger oval foramen near anterior margin; cercus 3.1 X length of epandrium, slightly curved, with 1 tenaculum (Fig. 17); tergite 10 rounded (Fig. 17). Female. Unknown.	en	Chagas-Vieira, Cínthia (2012): New records of Tonnoira Enderlein (Diptera, Psychodidae, Psychodinae) from the Brazilian Amazon with descriptions of two new species. Zootaxa 3318: 51-56, DOI: 10.5281/zenodo.213567
066E87D2496BFFF1FF538819FAE5F8CE.taxon	discussion	Remarks. Tonnoira tripenis sp. n. appears to be closely related to T. sicilis based on eyes separated by little less than 1 facet diameter, eye bridge with 4 facet rows, hypandrium a distinct band connecting gonocoxites, aedeagus tripartite and cercus with 1 tenaculum, but T. tripenis has paramere absent, hypandrium long and wide, gonocoxite 0.8 X length of the gonostylus and aedeagus tripartite, with L-shaped lateral shaft, central part broad over basal half and tapering to apex over distal half and the last shaft with basal 2 / 3 wide and apical 1 / 3 narrow. T. sicilis has aedeagus tripartite, with short basal spur, a sickle-shaped lateral shaft, and broad, paddle-shaped central part twisted in center, paramere slender, hypandrium short and narrow and gonocoxite as longer as gonostylus.	en	Chagas-Vieira, Cínthia (2012): New records of Tonnoira Enderlein (Diptera, Psychodidae, Psychodinae) from the Brazilian Amazon with descriptions of two new species. Zootaxa 3318: 51-56, DOI: 10.5281/zenodo.213567
