taxonID	type	description	language	source
076487D8FFC0CB6052B4FACAFBFCFA56.taxon	materials_examined	Material examined in this study: 16 specimens in sample A.	en	Kaczmarek, Łukasz, Jakubowska, Natalia, Michalczyk, Łukasz (2012): Current knowledge on Turkish tardigrades with a description of Milnesium beasleyi sp. nov. (Eutardigrada: Apochela: Milnesiidae, the granulatum group). Zootaxa 3589: 49-64, DOI: 10.5281/zenodo.213616
076487D8FFC0CB6052B4FACAFBFCFA56.taxon	discussion	Remarks: Species known from many localities in Turkey but this is the first record for the Konya Province. This species is known from many localities throughout the world (McInnes 1994).	en	Kaczmarek, Łukasz, Jakubowska, Natalia, Michalczyk, Łukasz (2012): Current knowledge on Turkish tardigrades with a description of Milnesium beasleyi sp. nov. (Eutardigrada: Apochela: Milnesiidae, the granulatum group). Zootaxa 3589: 49-64, DOI: 10.5281/zenodo.213616
076487D8FFC0CB6952B4F8C6FEC4F8E8.taxon	materials_examined	Material examined. Holotype (female) and 6 paratypes (5 females and 1 male) in sample A.	en	Kaczmarek, Łukasz, Jakubowska, Natalia, Michalczyk, Łukasz (2012): Current knowledge on Turkish tardigrades with a description of Milnesium beasleyi sp. nov. (Eutardigrada: Apochela: Milnesiidae, the granulatum group). Zootaxa 3589: 49-64, DOI: 10.5281/zenodo.213616
076487D8FFC0CB6952B4F8C6FEC4F8E8.taxon	description	Description (measurements and statistics in Table 1). Body (Figs 1 – 2) yellowish (before and after preparation). Eyes were absent in 5 of the 7 examined individuals (= 71 %, including the male), after mounting in Hoyer’s medium. Cuticle without granulation or pores, but the dorsum covered with numerous tiny and shallow polygonal or rounded depressions (pseudopores), 0.1 – 0.4 μm in diameter (Figs 3 – 5). These depressions, visible under PCM as light spots with undefined edges, are a manifestation of a fine reticulum, therefore placing the species within the granulatum group (Michalczyk et al. 2012 ab); see also the Differential Diagnosis below. Weak sculpturing is also present on the outer surfaces of all legs (Fig. 5). Six peribuccal papillae (ventral papilla smallest) and six peribuccal lamellae around the mouth opening present. Two cephalic papillae positioned laterally. Bucco-pharyngeal apparatus of the Milnesium type (Figs 6 – 7). Buccal tube slightly funnel-shaped, wider anteriorly (on average the posterior diameter is 92 % of the anterior diameter). Pharyngeal bulb elongated, pearshaped and without placoids or septulum. Claws of the Milnesium type, slender (Figs 8 – 9). Primary branches on all legs with small, but distinct accessory points detaching from the branch at its greatest curvature (Fig. 9). Secondary claws of all legs without rounded basal thickenings. Secondary branches of external claws I – III and posterior claws IV with two points, and secondary branches of internal claws I – III and anterior claws IV with three points (i. e. claw configuration: [2 - 3] - [3 - 2], Figs 8 – 9). Single, long transversal, cuticular bars under claws I – III present (Fig. 8). Although eggs were not found, we should probably expect them to be oval, smooth and deposited in exuvium, i. e. following the character of all currently known Milnesium species in which eggs were described.	en	Kaczmarek, Łukasz, Jakubowska, Natalia, Michalczyk, Łukasz (2012): Current knowledge on Turkish tardigrades with a description of Milnesium beasleyi sp. nov. (Eutardigrada: Apochela: Milnesiidae, the granulatum group). Zootaxa 3589: 49-64, DOI: 10.5281/zenodo.213616
076487D8FFC0CB6952B4F8C6FEC4F8E8.taxon	discussion	Remarks. We only found one male, which was generally similar to the females. However, apart from the diagnostic secondary sexual trait in this genus (secondary branches of claws I in the shape of large and robust hooks, Fig. 10), the male also differed from the females of this species by a narrower buccal tube and shorter papillae, both in absolute and relative terms (see Table 1). N RANGE MEAN SD Holotype Male CHARACTER µm pt µm pt µm pt µm pt µm pt Although the majority of specimens had a typical claw configuration [2 - 3] - [3 - 2], two females exhibited minor claw abnormalities (Fig. 11). One female had small basal spurs on external claws I-III and posterior claws IV ([3 - 3] - [3 - 3]) and the other female had small basal spurs on posterior claws IV only ([2 - 3] - [3 - 3]). These basal spurs were always less curved (sometimes completely straight) and smaller than those on other claws (<2.0 μm vs.> 3.0 μm) and thus could be without doubt classified as developmental aberrations. Nevertheless, the presence of such claw abnormalities highlights the importance of examining a number of Milnesium specimens to ensure correct identifications. Locus typicus. 37 ° 14 ′ 52 ′ N, 31 ° 55 ′ 42 ′ E, 1480 m asl: the Tinaztepe Mağarasi cave, near the road D 695, 25 km to the south from the city Seydişehir, Konya Province, Turkey, moss from stone.	en	Kaczmarek, Łukasz, Jakubowska, Natalia, Michalczyk, Łukasz (2012): Current knowledge on Turkish tardigrades with a description of Milnesium beasleyi sp. nov. (Eutardigrada: Apochela: Milnesiidae, the granulatum group). Zootaxa 3589: 49-64, DOI: 10.5281/zenodo.213616
076487D8FFC0CB6952B4F8C6FEC4F8E8.taxon	etymology	Etymology. We dedicate this species in memory of Professor Clark Beasley (06.06.1942. - 24.06.2012.), an American tardigradologists and the first Zootaxa Associate Editor for Tardigrada. Clark was also our friend and colleague who supported us throughout our adventure with tardigrades, especially in the early stages, and for this we will always be grateful to him.	en	Kaczmarek, Łukasz, Jakubowska, Natalia, Michalczyk, Łukasz (2012): Current knowledge on Turkish tardigrades with a description of Milnesium beasleyi sp. nov. (Eutardigrada: Apochela: Milnesiidae, the granulatum group). Zootaxa 3589: 49-64, DOI: 10.5281/zenodo.213616
076487D8FFC0CB6952B4F8C6FEC4F8E8.taxon	materials_examined	Type depositories. Holotype (slide T 1 / 12) and six paratypes (slides T 1 / 3, T 1 / 11, T 1 / 13 and T 1 / 14) mounted in Hoyer’s medium are preserved at the Department of Animal Taxonomy and Ecology, A. Mickiewicz University, Umultowska 89, 61 – 614 Poznań, Poland. Differential diagnosis. Milnesium beasleyi sp. nov. by having a sculptured dorsal cuticle belongs to the granulatum group within the genus (Michalczyk et al. 2012 a, b). Including this new species, the group now consists of six species, which is equivalent to 32 % of all known Milnesium taxa. The remaining five species with sculptured cuticle are: M. alabamae Wallendorf & Miller, 2009, M. granulatum Ramazzotti, 1962, M. katarzynae Kaczmarek et al., 2004, M. krzysztofi Kaczmarek & Michalczyk, 2007 and M. reticulatum Pilato et al., 2002. Cuticular sculpture in all known species of the group appears under PCM as bright dots or polygons with blurred edges. Scanning Electron Microscope (SEM) observations revealed that these dots / polygons are in fact shallow depressions in the cuticle (pseudopores) and therefore their edges can never be fully focused under PCM (Kaczmarek & Michalczyk 2007, see also Michalczyk & Kaczmarek 2006). Interestingly, these depressions have in past been mistaken for granulation — hence the name of the nominal species for the group (see Michalczyk et al. 2012 a). In all species of the group (apart from M. beasleyi sp. nov.) the cuticular depressions are densely arranged and the cuticle between very often forms a reticulum. In the new species, however, the depressions are less dense and more irregularly scattered. Large distances between the depressions, therefore, do not create an apparent reticulum.	en	Kaczmarek, Łukasz, Jakubowska, Natalia, Michalczyk, Łukasz (2012): Current knowledge on Turkish tardigrades with a description of Milnesium beasleyi sp. nov. (Eutardigrada: Apochela: Milnesiidae, the granulatum group). Zootaxa 3589: 49-64, DOI: 10.5281/zenodo.213616
076487D8FFC8CB6852B4FD13FB33FCEB.taxon	materials_examined	Material examined in this study: 53 specimens and 25 eggs in sample A.	en	Kaczmarek, Łukasz, Jakubowska, Natalia, Michalczyk, Łukasz (2012): Current knowledge on Turkish tardigrades with a description of Milnesium beasleyi sp. nov. (Eutardigrada: Apochela: Milnesiidae, the granulatum group). Zootaxa 3589: 49-64, DOI: 10.5281/zenodo.213616
076487D8FFC8CB6852B4FD13FB33FCEB.taxon	discussion	Remarks: The first record outside the locus typicus (Cyprus) (Michalczyk & Kaczmarek 2003).	en	Kaczmarek, Łukasz, Jakubowska, Natalia, Michalczyk, Łukasz (2012): Current knowledge on Turkish tardigrades with a description of Milnesium beasleyi sp. nov. (Eutardigrada: Apochela: Milnesiidae, the granulatum group). Zootaxa 3589: 49-64, DOI: 10.5281/zenodo.213616
076487D8FFC8CB6852B4FEB2FDBBFDBB.taxon	materials_examined	Material examined in this study: 2 specimens and 1 egg in sample A.	en	Kaczmarek, Łukasz, Jakubowska, Natalia, Michalczyk, Łukasz (2012): Current knowledge on Turkish tardigrades with a description of Milnesium beasleyi sp. nov. (Eutardigrada: Apochela: Milnesiidae, the granulatum group). Zootaxa 3589: 49-64, DOI: 10.5281/zenodo.213616
076487D8FFC8CB6852B4FEB2FDBBFDBB.taxon	discussion	Remarks: Species known from many localities in Turkey but this is the first record for the Konya Province. Known from many localities throughout the world (McInnes 1994), but the majority of records need to be verified as this is believed to be a species complex.	en	Kaczmarek, Łukasz, Jakubowska, Natalia, Michalczyk, Łukasz (2012): Current knowledge on Turkish tardigrades with a description of Milnesium beasleyi sp. nov. (Eutardigrada: Apochela: Milnesiidae, the granulatum group). Zootaxa 3589: 49-64, DOI: 10.5281/zenodo.213616
076487D8FFC8CB6A52B4FBE8FA42F84A.taxon	materials_examined	Material examined in this study: 8 specimens and 6 eggs in sample B.	en	Kaczmarek, Łukasz, Jakubowska, Natalia, Michalczyk, Łukasz (2012): Current knowledge on Turkish tardigrades with a description of Milnesium beasleyi sp. nov. (Eutardigrada: Apochela: Milnesiidae, the granulatum group). Zootaxa 3589: 49-64, DOI: 10.5281/zenodo.213616
076487D8FFC8CB6A52B4FBE8FA42F84A.taxon	discussion	Remarks: Species known from many localities throughout the world (including Turkey), but as this name incorporates a species complex, the majority of records need to be verified (McInnes 1994, Guidetti et al. 2009). ...... continued on the next page ...... continued on the next page	en	Kaczmarek, Łukasz, Jakubowska, Natalia, Michalczyk, Łukasz (2012): Current knowledge on Turkish tardigrades with a description of Milnesium beasleyi sp. nov. (Eutardigrada: Apochela: Milnesiidae, the granulatum group). Zootaxa 3589: 49-64, DOI: 10.5281/zenodo.213616
