taxonID	type	description	language	source
016D017DFFDF5F7C03A5BA80ECA78685.taxon	type_taxon	Type species: Bathynomus giganteus A. Milne-Edwards, 1879, by monotypy.	en	Hyžný, Matúš, Campos, Ildefonso Bajo, Carretero, Joaquín Cárdenas (2020): A new species of Bathynomus A. Milne-Edwards, 1879 (Malacostraca: Peracarida Isopoda: Cirolanidae) from the upper Miocene of the Guadalquivir Basin (Spain). Zootaxa 4819 (1): 159-169, DOI: 10.11646/zootaxa.4819.1.8
016D017DFFDF5F7C03A5BA80ECA78685.taxon	discussion	Remarks: The single specimen shows the ‘ key’ characters for taxonomic attribution to the genus Bathynomus, namely a semicircular pleotelson with sinuous lateral margins and distinct, approximately subequal, spines on its posterior margin, not exceeding the number of 13 (Hyžný et al. 2013). The number and arrangement of pleotelsonic spines together with the shape of uropod rami allow erection of a new species based on the studied fossil specimen.	en	Hyžný, Matúš, Campos, Ildefonso Bajo, Carretero, Joaquín Cárdenas (2020): A new species of Bathynomus A. Milne-Edwards, 1879 (Malacostraca: Peracarida Isopoda: Cirolanidae) from the upper Miocene of the Guadalquivir Basin (Spain). Zootaxa 4819 (1): 159-169, DOI: 10.11646/zootaxa.4819.1.8
016D017DFFD85F7903A5B94FE9E68757.taxon	description	Figs 4 – 6 urn: lsid: zoobank. org: act: 268 B 0 BA 1 - 02 CD- 4 DE 8 - A 5 CD- 02 DBADAFD 152	en	Hyžný, Matúš, Campos, Ildefonso Bajo, Carretero, Joaquín Cárdenas (2020): A new species of Bathynomus A. Milne-Edwards, 1879 (Malacostraca: Peracarida Isopoda: Cirolanidae) from the upper Miocene of the Guadalquivir Basin (Spain). Zootaxa 4819 (1): 159-169, DOI: 10.11646/zootaxa.4819.1.8
016D017DFFD85F7903A5B94FE9E68757.taxon	materials_examined	Material examined: The species is so far represented by the only specimen (measurements: 70.0 x 51.6 mm). The holotype (MGM- 6636 M) represents the posterior moult consisting of pereonites 5 – 7, pleonites 1 – 5, and pleotelson; uropods were exposed with further preparation. The specimen consists of part and counterpart (Figs 4 B, 5 A – B) and was collected by the authors JCC and IBC. GPS coordinates of the type locality (see below): 37 ° 37 ’ 1.85 ” N, 5 ° 47 ’ 38.42 ” W.	en	Hyžný, Matúš, Campos, Ildefonso Bajo, Carretero, Joaquín Cárdenas (2020): A new species of Bathynomus A. Milne-Edwards, 1879 (Malacostraca: Peracarida Isopoda: Cirolanidae) from the upper Miocene of the Guadalquivir Basin (Spain). Zootaxa 4819 (1): 159-169, DOI: 10.11646/zootaxa.4819.1.8
016D017DFFD85F7903A5B94FE9E68757.taxon	diagnosis	Diagnosis: Pleotelson approximately 0.7 as long as wide with indistinct carina and 7 broad, short, straight pleotelsonic spines plus 2 lateral spines; central spine longest, not bifid; gap between second and third side spines wider than between other spines. Uropod with oval exopod and subtriangular endopod, distolateral corners of both elements distinctly produced into tips.	en	Hyžný, Matúš, Campos, Ildefonso Bajo, Carretero, Joaquín Cárdenas (2020): A new species of Bathynomus A. Milne-Edwards, 1879 (Malacostraca: Peracarida Isopoda: Cirolanidae) from the upper Miocene of the Guadalquivir Basin (Spain). Zootaxa 4819 (1): 159-169, DOI: 10.11646/zootaxa.4819.1.8
016D017DFFD85F7903A5B94FE9E68757.taxon	description	Description: Pereonites 5 – 7 poorly preserved, pereonite 5 longer than pereonites 6 and 7 (possibly due to last pereonites being overlapped by preceding ones). Pleonites of approximately same length; pleonite 4 slightly overlapping pleonite 5 laterally; pleonites 3 – 5 reaching to almost same posterior level. Pleotelson subcircular in outline, approximately 0.7 as long as wide, dorsal surface finely granulated; posterior margin with 7 broad, short, straight pleotelsonic spines, and 2 smaller lateral spines; central spine longest, simple (not bifid); gap between second and third side spines wider than between other spines (Fig. 6). Uropodal exopod with elongate oval outline, lateral margin convex, distolateral corner obtuse with pointed tip, distomedial corner rounded, medial margin straight; uropodal endopod broadly subtriangular, distolateral corner obtuse, distinctly produced into sharp tip, distomedial corner broadly rounded, median margin straight.	en	Hyžný, Matúš, Campos, Ildefonso Bajo, Carretero, Joaquín Cárdenas (2020): A new species of Bathynomus A. Milne-Edwards, 1879 (Malacostraca: Peracarida Isopoda: Cirolanidae) from the upper Miocene of the Guadalquivir Basin (Spain). Zootaxa 4819 (1): 159-169, DOI: 10.11646/zootaxa.4819.1.8
016D017DFFD85F7903A5B94FE9E68757.taxon	etymology	Etymology: In Memoriam of Dr. Jorge Civis, professor of Palaeontology at the University of Salamanca and great promoter of Palaeontology in the Neogene of the Guadalquivir Basin.	en	Hyžný, Matúš, Campos, Ildefonso Bajo, Carretero, Joaquín Cárdenas (2020): A new species of Bathynomus A. Milne-Edwards, 1879 (Malacostraca: Peracarida Isopoda: Cirolanidae) from the upper Miocene of the Guadalquivir Basin (Spain). Zootaxa 4819 (1): 159-169, DOI: 10.11646/zootaxa.4819.1.8
016D017DFFD85F7903A5B94FE9E68757.taxon	discussion	Remarks: Although the only known specimen is incomplete, there are taxonomically important characters preserved supporting a designation of a new species. Lowry & Dempsey (2006) noted that the shape of the lateral, medial, and distal margins of the uropodal endopods and exopods and the distal corners are important for species identification. The number of pleotelsonic spines are relevant in this matter, especially when combined with the shape of uropod rami (Shipley et al. 2016; Hyžný et al. 2019). These characters (pleotelsonic spines and uropod rami) are indeed present in the holotype of Bathynomus civisi sp. nov. and they allow distinguishing the new species from other congeners. The new species clearly differs from a number of extant taxa with lanceolate uropod rami, including Bathyno- mus affinis Richardson, 1910; B. bruscai Lowry & Dempsey, 2006; B. decemspinosus Bruce, 1986; B. doederleini Ortmann, 1894; B. immanis Bruce, 1986; B. kapala Griffin, 1975; B. maxeyorum Shipley, Brooks & Bruce in Shipley et al., 2016; B. obtusus Magalhaes & Young, 2003; and B. pelor Bruce, 1986. From congeners with oval uropod rami, B. c ivisi sp. nov. is morphologically close to several of them. Bathynomus brucei Lowry & Dempsey, 2006 shares the number of pleotelsonic spines (Lowry & Dempsey 2006: fig. 4 F); it, however, does not have a wide gap between second and third side distal spines as the new species has. Additionally, the uropodal exopod of B. civisi sp. nov. is more oval in outline than that of B. brucei (Lowry & Dempsey 2006: fig. 5 D – E). In this respect, the new species is close to the type species, B. giganteus A. Milne-Edwards, 1879, but differs in the number and shape of pleotelsonic spines (Soto & Mincarone 2001: fig. 5; Magalhaes & Young 2003: fig. 1 A). The same holds for Bathynomus jamesi Kou, Chen, Li, He & Wang, 2017, which is close to B. civisi sp. nov. as far as the uropod rami concerns (Lowry & Dempsey 2006: fig. 22 A – B; Kou et al. 2017: fig. 3 d, e), but differs in the development of pleotelsonic spines (Kou et al. 2017: fig. 4). Concerning the pleotelson outline and the number and development of pleotelsonic spines, the closest extant congener is Bathynomus miyarei Lemos de Castro, 1978; it, however, differs with uropodal endopod having distal margin straight (Soto & Mincarone 2001: fig. 5; Magalhaes & Young 2003: fig. 10 B), whereas its outline is slightly concave close to distolateral corner (Figs 5 C, 6). Bathynomus civisi sp. nov. differs from the fossil congeners; Bathynomus sismondai (Ristori, 1891) from the Plio-Pleistocene of Italy has more pleotelsonic spines and uropodal exopod with less obtuse distolateral corner (Hyžný et al. 2019: figs. 6 A 3 – A 4). Bathynomus kominatoensis Kato & Tokita in Kato et al., 2016 from the Miocene of Japan has proportionally wider pleotelson and higher number of pleotelsonic spines (figs. 3.5, 4.1) than B. civisi sp. nov. No fossil species of Bathynomus known to date has a wide gap between second and third side distal spines as the new species has. Occurrence: Bathynomus civisi sp. nov. is so far known only from the upper Tortonian marls of the Écija Formation, exposed at the type locality, an old quarry located in the close vicinity of Arroyo Trujillo (Seville, Spain).	en	Hyžný, Matúš, Campos, Ildefonso Bajo, Carretero, Joaquín Cárdenas (2020): A new species of Bathynomus A. Milne-Edwards, 1879 (Malacostraca: Peracarida Isopoda: Cirolanidae) from the upper Miocene of the Guadalquivir Basin (Spain). Zootaxa 4819 (1): 159-169, DOI: 10.11646/zootaxa.4819.1.8
