identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
016D017DFFDF5F7C03A5BA80ECA78685.text	016D017DFFDF5F7C03A5BA80ECA78685.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bathynomus A. Milne-Edwards 1879	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Genus  Bathynomus A. Milne-Edwards, 1879</p>
            <p> Type species:  Bathynomus giganteus A. Milne-Edwards, 1879 , by monotypy. </p>
            <p> Remarks: The single specimen shows the ‘key’ characters for taxonomic attribution to the genus  Bathynomus , namely a semicircular pleotelson with sinuous lateral margins and distinct, approximately subequal, spines on its posterior margin, not exceeding the number of 13 (Hyžný et al. 2013). The number and arrangement of pleotelsonic spines together with the shape of uropod rami allow erection of a new species based on the studied fossil specimen. </p>
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	https://treatment.plazi.org/id/016D017DFFDF5F7C03A5BA80ECA78685	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hyžný, Matúš;Campos, Ildefonso Bajo;Carretero, Joaquín Cárdenas	Hyžný, Matúš, Campos, Ildefonso Bajo, Carretero, Joaquín Cárdenas (2020): A new species of Bathynomus A. Milne-Edwards, 1879 (Malacostraca: Peracarida Isopoda: Cirolanidae) from the upper Miocene of the Guadalquivir Basin (Spain). Zootaxa 4819 (1): 159-169, DOI: 10.11646/zootaxa.4819.1.8
016D017DFFD85F7903A5B94FE9E68757.text	016D017DFFD85F7903A5B94FE9E68757.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bathynomus civisi Hyžný & Campos & Carretero 2020	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Bathynomus civisi sp. nov.</p>
            <p>Figs 4–6</p>
            <p>urn:lsid:zoobank.org:act: 268B0BA1-02CD-4DE8-A5CD-02DBADAFD152</p>
            <p>Material examined: The species is so far represented by the only specimen (measurements: 70.0 x 51.6 mm). The holotype (MGM-6636M) represents the posterior moult consisting of pereonites 5–7, pleonites 1–5, and pleotelson; uropods were exposed with further preparation. The specimen consists of part and counterpart (Figs 4B, 5 A–B) and was collected by the authors JCC and IBC. GPS coordinates of the type locality (see below): 37°37’1.85”N, 5°47’38.42”W.</p>
            <p>Diagnosis: Pleotelson approximately 0.7 as long as wide with indistinct carina and 7 broad, short, straight pleotelsonic spines plus 2 lateral spines; central spine longest, not bifid; gap between second and third side spines wider than between other spines. Uropod with oval exopod and subtriangular endopod, distolateral corners of both elements distinctly produced into tips.</p>
            <p>Description: Pereonites 5–7 poorly preserved, pereonite 5 longer than pereonites 6 and 7 (possibly due to last pereonites being overlapped by preceding ones). Pleonites of approximately same length; pleonite 4 slightly overlapping pleonite 5 laterally; pleonites 3–5 reaching to almost same posterior level. Pleotelson subcircular in outline, approximately 0.7 as long as wide, dorsal surface finely granulated; posterior margin with 7 broad, short, straight pleotelsonic spines, and 2 smaller lateral spines; central spine longest, simple (not bifid); gap between second and third side spines wider than between other spines (Fig. 6). Uropodal exopod with elongate oval outline, lateral margin convex, distolateral corner obtuse with pointed tip, distomedial corner rounded, medial margin straight; uropodal endopod broadly subtriangular, distolateral corner obtuse, distinctly produced into sharp tip, distomedial corner broadly rounded, median margin straight.</p>
            <p>Etymology: In Memoriam of Dr. Jorge Civis, professor of Palaeontology at the University of Salamanca and great promoter of Palaeontology in the Neogene of the Guadalquivir Basin.</p>
            <p> Remarks: Although the only known specimen is incomplete, there are taxonomically important characters preserved supporting a designation of a new species. Lowry &amp; Dempsey (2006) noted that the shape of the lateral, medial, and distal margins of the uropodal endopods and exopods and the distal corners are important for species identification. The number of pleotelsonic spines are relevant in this matter, especially when combined with the shape of uropod rami (Shipley et al. 2016; Hyžný et al. 2019). These characters (pleotelsonic spines and uropod rami) are indeed present in the holotype of  Bathynomus civisi sp. nov. and they allow distinguishing the new species from other congeners. </p>
            <p> The new species clearly differs from a number of extant taxa with lanceolate uropod rami, including Bathyno- mus affinis Richardson, 1910;  B. bruscai Lowry &amp; Dempsey, 2006 ;  B. decemspinosus Bruce, 1986 ;  B. doederleini Ortmann, 1894 ;  B. immanis Bruce, 1986 ;  B. kapala Griffin, 1975 ;  B. maxeyorum Shipley, Brooks &amp; Bruce in Shipley et al., 2016 ;  B. obtusus Magalhaes &amp; Young, 2003 ; and  B. pelor Bruce, 1986 . From congeners with oval uropod rami, B. c ivisi sp. nov. is morphologically close to several of them.  Bathynomus brucei Lowry &amp; Dempsey, 2006 shares the number of pleotelsonic spines (Lowry &amp; Dempsey 2006: fig. 4F); it, however, does not have a wide gap between second and third side distal spines as the new species has. Additionally, the uropodal exopod of  B. civisi sp. nov. is more oval in outline than that of  B. brucei (Lowry &amp; Dempsey 2006: fig. 5D–E). In this respect, the new species is close to the type species,  B. giganteus A. Milne-Edwards, 1879 , but differs in the number and shape of pleotelsonic spines (Soto &amp; Mincarone 2001: fig. 5; Magalhaes &amp; Young 2003: fig. 1A). The same holds for  Bathynomus jamesi Kou, Chen, Li, He &amp; Wang, 2017 , which is close to  B. civisi sp. nov. as far as the uropod rami concerns (Lowry &amp; Dempsey 2006: fig. 22A–B; Kou et al. 2017: fig. 3d, e), but differs in the development of pleotelsonic spines (Kou et al. 2017: fig. 4). Concerning the pleotelson outline and the number and development of pleotelsonic spines, the closest extant congener is  Bathynomus miyarei Lemos de Castro, 1978 ; it, however, differs with uropodal endopod having distal margin straight (Soto &amp; Mincarone 2001: fig. 5; Magalhaes &amp; Young 2003: fig. 10B), whereas its outline is slightly concave close to distolateral corner (Figs 5C, 6). </p>
            <p> Bathynomus civisi sp. nov. differs from the fossil congeners;  Bathynomus sismondai (Ristori, 1891) from the Plio-Pleistocene of Italy has more pleotelsonic spines and uropodal exopod with less obtuse distolateral corner (Hyžný et al. 2019: figs. 6A3–A4).  Bathynomus kominatoensis Kato &amp; Tokita in Kato et al., 2016 from the Miocene of Japan has proportionally wider pleotelson and higher number of pleotelsonic spines (figs. 3.5, 4.1) than  B. civisi sp. nov. No fossil species of  Bathynomus known to date has a wide gap between second and third side distal spines as the new species has. </p>
            <p> Occurrence:  Bathynomus civisi sp. nov.  is so far known only from the  upper Tortonian marls of the Écija Formation, exposed at the type locality, an old quarry located in the close vicinity of Arroyo Trujillo (Seville, Spain)  . </p>
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	https://treatment.plazi.org/id/016D017DFFD85F7903A5B94FE9E68757	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hyžný, Matúš;Campos, Ildefonso Bajo;Carretero, Joaquín Cárdenas	Hyžný, Matúš, Campos, Ildefonso Bajo, Carretero, Joaquín Cárdenas (2020): A new species of Bathynomus A. Milne-Edwards, 1879 (Malacostraca: Peracarida Isopoda: Cirolanidae) from the upper Miocene of the Guadalquivir Basin (Spain). Zootaxa 4819 (1): 159-169, DOI: 10.11646/zootaxa.4819.1.8
