taxonID	type	description	language	source
022F777CFF8CAB0EB8D5FDD2CA8AFE41.taxon	materials_examined	Type: — CHINA. Hainan: epiphytic on tree trunk in evergreen broad-leaved and sunny open forest, elev. ca. 300 m, 14 May 2014 Huang 130322001 (holotype: ATCH!). Dendrobium tetrapterum is similar to D. lindleyi and D. jenkinsii, but can easily be distinguished by the petals without (vs. with) claw, slit (vs. entire) transverse septum at lip base, anther cap with (vs. without) indentation on each side and rhombic (vs. pentagonal – octagonal or oblong) pseudobulb cross-section. Epiphytic herbs with creeping rhizomes, slender, 2 – 3 mm in diam, branched. Pseudobulbs prostrate, crowded, fusiform, compressed, 1.0 – 5.0 × 0.5 – 1.5 cm, with 2 – 5 nodes, internodes 1.0 – 2.0 cm, with 4 ribs, pale yellowish brown and glossy when dry, with white sheaths. Leaf usually solitary, oblong, 3.0 – 8.0 × 0.6 – 3.0 cm, base contracted, not sheathing, apex obtuse and emarginate. Inflorescence subterminal, to 27 cm, much longer than leaves, laxly several to up to 15 - flowered; floral bracts narrowly ovate-triangular, ca. 2 mm long. Pedicel and ovary yellowish green, tinged with pale purple, 2.8 – 3.2 cm long. Flowers spreading, orange-yellow. Dorsal sepal ovate-lanceolate, 13.0 × 8.0 mm, subobtuse; lateral sepals nearly equal to dorsal sepal, 15.0 × 9.0 mm. Petals broadly elliptic, 19.0 × 12.0 mm, obtusely round. Lip transversely oblong or subreniform, usually 21.0 × 26.0 mm, entire, often emarginate; disk densely pubescent below middle, below middle contracted into a claw with the sides clasping column; near base with a transverse septum, split into two lobes. Column 3.5 mm long, 2.5 mm in diameter, column foot 4.0 mm long, at the base with an oval concavity; anther cap subglobose, glabrous, with an indentation on each side, pollinia naked, terete, 4 in two pairs. Phenology: — Flowering April, May.	en	Wei, Chun-Yi, Li, Ru-Yi, Wu, Yu-Wei, Huang, Ming-Zhong, Li, Ming-He (2024): Dendrobium tetrapterum, a cryptic species in the D. lindleyi alliance (Orchidaceae; Dendrobiinae): evidence from morphological and molecular data. Phytotaxa 655 (2): 209-218, DOI: 10.11646/phytotaxa.655.2.8, URL: http://dx.doi.org/10.11646/phytotaxa.655.2.8
022F777CFF8CAB0EB8D5FDD2CA8AFE41.taxon	distribution	Distribution: — China (Guangdong, Guangxi, Hainan, SW Guizhou, Yunnan), Vietnam (Cao Bang, Bac Can, Thai Nguyen, Ninh Binh, Quang Ninh) and Thailand (Chiang Mai). Habitat and ecology: Epiphytic, below 1800 m elevation. Other specimens studied: — THAILAND. Chiang Mai: Doi Suthep, 800 – 1200 m, Jun 1905, Hosseus 818 (MO 168775). VIETNAM. Cao Bang: Thach An district, municipality Thuy Hung, vicinities of Na Lit village (22 ° 30 ′ N, 106 ° 27 ′ E), 500 – 550 m, 4 Dec 1998, CBL 892 (LE 01077119); Thach An district, municipality Duc Xuan, (N 22 ° 21 ′ ’ E 106 ° 27 ′), 2 Dec 1998, CBL 808 (LE 01066396). Bac Can: Na Ri district, Kim Hy municipality, vicinities of Lung Hin Con village (22 ° 17 ′ N, 106 ° 03 ′ E), 27 km to 55 ° of Bac Kan city, 800 m, 19 Oct 1999, NTH 3673 (LE 01066399); Cho Don district, Ban Thi Municipality, Cho Dien village (22 ° 14 ′ 28 ″ N, 105 ° 29 ′ 50 ″ E), 250 - 350 m, 26 Dec 2002, HLF 849 (LE 01077112). Thai Nguyen: Dong Hy district, Tan Long Municipality, vicinity of Mo Ba Village (20 ° 46 ′ 26 ″ N, 105 ° 52 ′ 25 ″ E), 400 – 550 m, 14 Mar 2001, HAL 7 (MO 5686816). Ninh Binh: Cuc Phuong National Park (105 ° 43.08 ′ N, 20 ° 14.22 ′ E), 20 Dec 1933, CP 146 (L 0402526). Quang Ninh: Dam-ha district, Sai Wong Mo Shan (Sai Vong Mo Leng) Lomg Ngong Village, 18 Sep 1940, Tsang 30162 (AMES 02156601); Ching Mai district, Ku Tang Village, Pak Shik Ling and vicinity, 31 Mar 1933, Lei 496 (AMES 02156889); Yaichow, 1800 ft., May-Jul 1933, How 70525 (P 00407945). CHINA. Hainan: Apr 1932, Liang 61648 (IBSC 0626361); Apr 1932, Gao 52251 (IBK 00145549, IBSC 0626365, PE 00851167); May 1932, McClure 20080 (IBSC 0626360, PE 00293829); Apr 1932, Mar 1933, Lei 496 (IBSC 0626359, PE 00293830); 0 – 1800 m, Apr 1933, Hou 70525 (SN 002661, IBK 00145547, IBSC 0626362, PE 00293828); Apr 1936, Liu 26046 (IBK 00145546, IBSC 0626364, PE 00293826), Apr 1998, Chen 699 (IBSC 0745773); 14 May 2014, MHLi or 139 (FJFCor 139). Guangxi: no precise date, Nong-gang Comprehensive Expedition Team 10949 (IBK 00145541); no collector, 1036 (PE 00293832) & 96394 (IBK 00145542); Beijing-dui Team 893060 (PE 01997725); Jun 1935, Gao 55358 (IBK 00145534); Jul 1936, Liang 67911 (IBK 00145531) & 67932 (IBK 00145536); 0 – 1000 m, Jul 1936, Su 67911 (IBSC 0626367); Soo 67932 (PE 00293833); Jun 1937, Liang 69407 (IBK 00145535); May 1956, Wei 64 (IBK 00145532); Jul 1956, Shu A 61245 (IBK 00145544, PE 00293837); Jul 1958, Li 600971 (IBK 00145545, PE 00293835); 510 m, Jan 1959, Li 602805 (IBK 00145533, PE 00293836); May 1959, Li 403810 (IBK 00145543); Oct 1959, Zhong A 63761 (IBK 00145537); Nov 1981, Da-yao-shan Comprehensive Expedition Team 13191 (IBK 00196191); Apr 1998, Chen 699 (IBSC 0626366); Jun 2001, HK kadoorie PT 790 (PE 01997724); Jun 2007, Lu, Mo, Xue & Li 16838 (GXMG 0035395); 700 m, Jun 2010, Xu & Jiang 10746 (IBK 00281750); 154 m, Sep 2012, Lu, Nong, Huang et al 451422120907032 LY (GXMG 0102359); 412 m, Jul 2013, He & Yang YangXF 0322 (KUN 1393816). Guangdong province: no precise data, 61648 (PE 00293827); Aug 1931, Gao 51730 (IBSC 0626338); Jun 1932, Huang 32345 (IBSC 0626336); Jun 1933, Zuo 22777 (IBSC 0626337); May 1943, Chen 11886 (IBSC 0626335); 90 m, Jun 1958, Huang 110488 (PE 00293831); May 1972, Luo 667 (IBSC 0626341); Aug 1972, Luo 1212 (IBSC 0626340); Sep 1978, Yue 78 6193 (IBSC 0626334). Yunnan province: 1000 m, Nov 1980, Ji 89 (PE 00850236); Jun 1981, Ji 114 (PE 00850234); Mar 1983, Chen & Ji 1 (PE 00850231) & 22 (PE 00850228); Apr 1983, Ji 129 (PE 00850235); 1600 m, Jul 1985, Ji 258 (PE 00850227). Guizhou province: Oct 1958, Cao 1011 (PE 00293824).	en	Wei, Chun-Yi, Li, Ru-Yi, Wu, Yu-Wei, Huang, Ming-Zhong, Li, Ming-He (2024): Dendrobium tetrapterum, a cryptic species in the D. lindleyi alliance (Orchidaceae; Dendrobiinae): evidence from morphological and molecular data. Phytotaxa 655 (2): 209-218, DOI: 10.11646/phytotaxa.655.2.8, URL: http://dx.doi.org/10.11646/phytotaxa.655.2.8
022F777CFF89AB0DB8D5FB0AC989F7F3.taxon	description	Homotypic synonyms: Dendrobium aggregatum Roxb., Fl. Ind. ed. 1832, 3: 477, 1832, nom. illeg. [non Kunth 1816]. Callista aggregata (Roxb.) Kuntze, Revis. Gen. Pl. 2: 654, 1891, nom. illeg.	en	Wei, Chun-Yi, Li, Ru-Yi, Wu, Yu-Wei, Huang, Ming-Zhong, Li, Ming-He (2024): Dendrobium tetrapterum, a cryptic species in the D. lindleyi alliance (Orchidaceae; Dendrobiinae): evidence from morphological and molecular data. Phytotaxa 655 (2): 209-218, DOI: 10.11646/phytotaxa.655.2.8, URL: http://dx.doi.org/10.11646/phytotaxa.655.2.8
022F777CFF89AB0DB8D5FB0AC989F7F3.taxon	materials_examined	Type: — MYANMAR. Northern Rakhine State: Arracan, leg. Pierard s. n., icon. Roxburgh 1289 (holotype CAL; copy K). Epiphytic herbs with a creeping rhizome, slender, 2 – 3 mm in diam, branched. Pseudobulbs suberect, ovoid, crowded, 4.0 – 8.0 × 1.0 – 2.0 cm, more or less octangular, and as many prominent subacute ridges, loosely covered with a grey, shining, wrinkled cuticle, and with 2 – 4 nodes, internodes 2.0 – 2.5 cm, the outer tapering. Leaf solitary, rarely two, oblong, 5.0 – 9.0 × 2.0 – 3.0 cm, base contracted, not sheathing, margin ± undulate, apex obtuse and emarginate, twice as long as the pseudobulbs. Inflorescence (to six inches long or more) laterally from the upper internode, arching, laxly several- to 20 - flowered; floral bracts small, acute. Flowers orange-yellow, deepest towards the base of the lip; sepals ovate, united at their base where the two laterals form a mentum; petals ovate, broader and longer than the sepals, 20.0 × 12.0 mm, with a short claw; lip broader than long, 27.0 × 30.0 mm, entire, often emarginate; minutely pubescent in front, disk densely pubescent below middle, below middle contracted into a claw with the sides clasping column. Column streaked with red in front, with a truncate scale near base, slightly concave stigma, toothed at both sides of the anther, with a tooth-like filament. Anther bilocular, greenish; pollinia naked, terete, 4 in two pairs. (after Roxburgh 1832, Lindley 1835, Hooker 1839). Phenology: — Flowering March, April.	en	Wei, Chun-Yi, Li, Ru-Yi, Wu, Yu-Wei, Huang, Ming-Zhong, Li, Ming-He (2024): Dendrobium tetrapterum, a cryptic species in the D. lindleyi alliance (Orchidaceae; Dendrobiinae): evidence from morphological and molecular data. Phytotaxa 655 (2): 209-218, DOI: 10.11646/phytotaxa.655.2.8, URL: http://dx.doi.org/10.11646/phytotaxa.655.2.8
022F777CFF89AB0DB8D5FB0AC989F7F3.taxon	distribution	Distribution and habitat: — NE India, Myanmar, Thailand, Laos, Vietnam. Habitat and ecology: Epiphyte in seasonally dry forest, blow 1000 m elevation.	en	Wei, Chun-Yi, Li, Ru-Yi, Wu, Yu-Wei, Huang, Ming-Zhong, Li, Ming-He (2024): Dendrobium tetrapterum, a cryptic species in the D. lindleyi alliance (Orchidaceae; Dendrobiinae): evidence from morphological and molecular data. Phytotaxa 655 (2): 209-218, DOI: 10.11646/phytotaxa.655.2.8, URL: http://dx.doi.org/10.11646/phytotaxa.655.2.8
022F777CFF89AB0DB8D5FB0AC989F7F3.taxon	description	Specimens studied: — MYANMAR. Gulf of Martaban: Chappadong, Wallich 7411 A (K 001127306). THAILAND. Bangkok: Hui Yang (K 000596580). LAOS. Khammoune: Boualapha district, Nong Seng village, Chanh Daeng mountain (17 ° 31 ′ 11.1 ″ N, 105 ° 46 ′ 41.4 ″ E), 400 – 600 m, 11 May 2018, AL 1070 (LE 01048800). VIETNAM. Quang Tri: 2 Jul, 1924, 11069 (P 00407931).	en	Wei, Chun-Yi, Li, Ru-Yi, Wu, Yu-Wei, Huang, Ming-Zhong, Li, Ming-He (2024): Dendrobium tetrapterum, a cryptic species in the D. lindleyi alliance (Orchidaceae; Dendrobiinae): evidence from morphological and molecular data. Phytotaxa 655 (2): 209-218, DOI: 10.11646/phytotaxa.655.2.8, URL: http://dx.doi.org/10.11646/phytotaxa.655.2.8
