taxonID	type	description	language	source
037387810D0AFFA2618C5D47D35EF7CE.taxon	type_taxon	Type species: Graziaphylus joceliae Schuh and Salas	en	Schuh, Randall T., Salas, Ruth (2021): Graziaphylus, a new genus and two new species of Eremophila-feeding (Scrophulariceae) Phylinae (Insecta: Heteroptera: Miridae) from Australia. Zootaxa 4958 (1): 319-326, DOI: 10.11646/zootaxa.4958.1.18
037387810D0AFFA2618C5D47D35EF7CE.taxon	etymology	Etymology: From the surname Grazia and the generic name Phylus; masculine.	en	Schuh, Randall T., Salas, Ruth (2021): Graziaphylus, a new genus and two new species of Eremophila-feeding (Scrophulariceae) Phylinae (Insecta: Heteroptera: Miridae) from Australia. Zootaxa 4958 (1): 319-326, DOI: 10.11646/zootaxa.4958.1.18
037387810D0AFFA2618C5D47D35EF7CE.taxon	diagnosis	Diagnosis: Male: Small, total length 2.56 – 3.18, weakly elongate, broadly ovate, weakly to distinctly flattened, width across pronotum 1.05 – 1.11, membrane weakly exceeding apex of abdomen; coloration castaneous to nearly black, posterior margin of vertex pale, antennae pale (Fig. 1, 3), legs, except tibiae and tarsi ranging from pale to black / castaneous, tibiae pale, always with black spines with black bases, tarsi pale; head short, broad, eyes somewhat bulging as viewed from above, posterior margin of vertex rounded, nearly straight across (Figs. 1, 3); antennae short, slender, not sexually dimorphic, pale (Figs. 1, 3); claws only moderately elongate, nearly straight over much of length; pulvilli elongate, adnate to claw over most of length of the latter (Figs. 7, 8); parempodia setiform. Dimensions and proportions as in Table 1; interocular distance relative to width of head apparently distinctive for two known species. Vestiture composed of reclining simple setae intermixed with flattened lanceolate setae, the latter either dark or silvery in appearance (Figs. 1, 3, 9, 10). Legs short, femora flat (Figs. 1, 3). GENITALIA (Figs. 12 – 15, 18): Endosoma with a single strap, apical attenuation single, about the length of the gonopore and with one or two barbs; gonopore relatively large; phallotheca with apical portion at right angles relative to proximal portion; left paramere conventionally phyline, without distinctive structural variation; right paramere lanceolate. Female (Figs. 2, 4) similar to male in coloration and vestiture, but somewhat larger and more broadly ovoid. Among Australian Phylinae Graziaphylus is most similar in coloration and shape to Xiphoidellus gilvus Weirauch and Schuh (2011), but the latter taxon lacks scalelike setae, has small flaplike pulvilli placed near the midpoint of the claw, and the endosoma forms a complete coil and with a fingerlike process at the apex of the secondary gonopore.	en	Schuh, Randall T., Salas, Ruth (2021): Graziaphylus, a new genus and two new species of Eremophila-feeding (Scrophulariceae) Phylinae (Insecta: Heteroptera: Miridae) from Australia. Zootaxa 4958 (1): 319-326, DOI: 10.11646/zootaxa.4958.1.18
037387810D0AFFA2618C5D47D35EF7CE.taxon	description	Description: Male: Moderately small, broad-bodied, somewhat flattened, total length 2.56 – 3.18, width across pronotum 1.05 – 1.11. COLORATION (Figs. 1, 3): Body castaneous to nearly black; antennae pale; legs pale or castaneous / black; tibiae pale, spines with dark bases; membrane of wing, including veins, fumose. SURFACE AND VESTITURE (Figs. 1, 3, 5, 9, 10): Dorsum smooth, shining, covered with recumbent, dark, simple setae and some dark or silvery scalelike setae. STRUCTURE: Curvature of posterior margin of vertex and eyes more or less conforming to anterior margin of pronotum (Figs. 1, 3); posterior margin of vertex and eyes smoothly and rather broadly rounded (Figs. 1, 3); vertex and frons moderately rounded; clypeus barely visible from above; dorsal margin of antennal fossa at level of ventral margin of eye; head short in lateral view, eyes occupying much of height of head; labium reaching to middle trochanters; costal margin of hemelytra broadly convex (Figs. 1, 3), cuneal fracture pronounced, hemelytra weakly deflexed at fracture (Figs. 1, 3); pronotum flattened, nearly straight on lateral and posterior margins; mesoscutum moderately exposed, scutellum flattened, weakly elevated anteriorly; claws of moderate length, nearly straight along dorsal margin, sharply bent near apex, pulvilli long, adnate to ventral surface of claw over most of its length (Figs. 7, 8), parempodia setiform (Figs. 7, 8). GENITALIA (Figs. 12, 15, 18): Endosoma formed of a single strap, more-or-less J-shaped, with a slender, nearly straight, apical attenuation about length of secondary gonopore with one or two barbs; secondary gonopore well sclerotized, subtending apical endosomal spine, gonopore sclerite lacking; phallotheca with apical portion conical, at right angle to long, broad basal flap; left paramere of characteristic phyline type; right paramere small, lanceolate. Female: Similar to male, but more broadly ovoid; total length 2.98 – 3.24; width pronotum 1.11 – 1.26. COLORATION (Figs. 2, 4): As in male. SURFACE AND VESTITURE (Figs. 9 – 11): As in male. STRUCTURE (Fig. 2, 4): Macropterous; eyes apparently lacking sexual dimorphism. GENITALIA: Figures 16, 17. Discussion: Among Australian taxa, Graziaphylus will be most easily confused with the uniformly dark-colored species of Xiphoidellus Weirauch and Schuh; nonetheless, the male genitalia in the two groups are distinctive and the latter lacks the scalelike setae seen in the former. On a world basis, Graziaphylus species are similar in size and appearance to the multiple dark-colored species of Chlamydatus Fieber in the Northern Hemisphere, but the latter have minute, flaplike pulvilli situated at the midpoint of the claw (Schuh & Schwartz, 2005). Among other Phylinae, genera such as Arctostaphylocoris Schuh and Schwartz (2004) from the western Nearctic, the Holarctic Phoenicocoris Reuter (Schwartz & Stonedahl, 2004), members of the widely distributed Sthenaridea Reuter from the Southern Hemisphere, and several of the New World Rhinacloa Reuter (Schuh & Schwartz, 1985) species also share a resemblance, including their size and the fact that they all have some scalelike setae on the dorsum and usually on the thoracic pleuron. We tentatively place Graziaphylus in the tribe Nasocorini (see Schuh & Menard, 2013) based on the size, structure of the male genitalia, and form of the claws and pulvilli. This is the first Australian taxon attributed to the tribe, whose monophyly and accommodation of Graziaphylus will strongly benefit from further testing using DNA sequence data.	en	Schuh, Randall T., Salas, Ruth (2021): Graziaphylus, a new genus and two new species of Eremophila-feeding (Scrophulariceae) Phylinae (Insecta: Heteroptera: Miridae) from Australia. Zootaxa 4958 (1): 319-326, DOI: 10.11646/zootaxa.4958.1.18
037387810D09FFA4618C5912D0F9F641.taxon	description	(Figs 1, 2, 12 – 17, Table 1)	en	Schuh, Randall T., Salas, Ruth (2021): Graziaphylus, a new genus and two new species of Eremophila-feeding (Scrophulariceae) Phylinae (Insecta: Heteroptera: Miridae) from Australia. Zootaxa 4958 (1): 319-326, DOI: 10.11646/zootaxa.4958.1.18
037387810D09FFA4618C5912D0F9F641.taxon	materials_examined	Holotype. Male. AUSTRALIA: South Australia: 8 km S of Martins Well, 31.40001 ° S 139.0753 ° E, 187 m, 08 Nov 2001, Cassis, Schuh, and Schwartz, Eremophila sp. (Scrophulariaceae), det. NSW staff NSW 666362, 1 ♂ (AMNH _ PBI 00415222) (SAMA). Paratypes: AUSTRALIA: South Australia: 8 km S of Martins Well, 31.40001 ° S 139.0753 ° E, 187 m, 08 Nov 2001, Cassis, Schuh, and Schwartz, Eremophila sp. (Scrophulariaceae), det. NSW staff NSW 666362, 7 ♂ (00415236 – 00415242), 14 ♀ (00415276 – 00415289) (AM); 10 ♂ (00415221, 00415223, 00415231 – 00415235, 00415251, 00415252, 00415255), 20 ♀ (00415253, 00415254, 00415256, 00415257, 00415298, 00415299, 00415308 – 00415321) (AMNH); 1 ♂ (00415245), 2 ♀ (00415294, 00415295) (ANIC); 1 ♂ (00415247), 2 ♀ (00415300, 00415301) (CNC); 7 ♂ (00415224 – 00415230), 18 ♀ (00415258 – 00415275) (SAMA); 1 ♂ (00415249), 2 ♀ (00415304, 00415305) (UCR); 2 ♂ (00415243, 00415244), 4 ♀ (00415290 – 00415293) (UNSW); 1 ♂ (00415250), 2 ♀ (00415306, 00415307) (USNM); 1 ♂ (00415246), 2 ♀ (00415296, 00415297) (WAMP); 1 ♂ (00415248), 2 ♀ (00415302, 00415303) (ZISP).	en	Schuh, Randall T., Salas, Ruth (2021): Graziaphylus, a new genus and two new species of Eremophila-feeding (Scrophulariceae) Phylinae (Insecta: Heteroptera: Miridae) from Australia. Zootaxa 4958 (1): 319-326, DOI: 10.11646/zootaxa.4958.1.18
037387810D09FFA4618C5912D0F9F641.taxon	etymology	Etymology: Named in honor of Jocelia Grazia.	en	Schuh, Randall T., Salas, Ruth (2021): Graziaphylus, a new genus and two new species of Eremophila-feeding (Scrophulariceae) Phylinae (Insecta: Heteroptera: Miridae) from Australia. Zootaxa 4958 (1): 319-326, DOI: 10.11646/zootaxa.4958.1.18
037387810D09FFA4618C5912D0F9F641.taxon	diagnosis	Diagnosis: Recognized by the attributes given in the generic diagnosis, the overall length of the male (2.72), the legs entirely pale except for the numerous castaneous spots on the distal half of the hind femur in both males and females, the black bases of the tibial spines, and the dorsum and thoracic pleuron with a few scattered, dark-colored, scalelike setae (Figs. 1, 2, 9, 10).	en	Schuh, Randall T., Salas, Ruth (2021): Graziaphylus, a new genus and two new species of Eremophila-feeding (Scrophulariceae) Phylinae (Insecta: Heteroptera: Miridae) from Australia. Zootaxa 4958 (1): 319-326, DOI: 10.11646/zootaxa.4958.1.18
037387810D09FFA4618C5912D0F9F641.taxon	description	Description: Male: Body conformation as in generic description; total length 2.56 – 2.87, width across pronotum 1.05 – 1.11. COLORATION (Fig. 1): As in generic description; legs entirely pale except for the numerous castaneous spots on the distal half of the hind femur and dark bases of tibial spines. SURFACE AND VESTITURE (Figs. 1, 5, 9 – 11): As in generic description, with recumbent, dark, simple setae intermixed with relatively dark, scalelike setae on dorsum; dorsal margin of meso- and metapleuron broadly with a few silvery, shining, scalelike setae; abdominal venter with numerous recumbent shining setae (Fig. 11). STRUCTURE: As in generic description. GENITALIA (Figs. 12 – 15): Apical endosomal spine with two lateral barbs; otherwise as in generic description. Female: As in generic description; total length 2.98 – 3.06, width pronotum 1.11 – 1.17. COLORATION (Fig. 2): As in male. SURFACE AND VESTITURE (Figs. 2, 5, 9 – 11): As in male. STRUCTURE (Figs. 2, 5 – 11): Similar to male, but more broadly ovoid; metathoracic scent-efferent system extensive, as in Figure 6. GENITALIA: Figures 16, 17.	en	Schuh, Randall T., Salas, Ruth (2021): Graziaphylus, a new genus and two new species of Eremophila-feeding (Scrophulariceae) Phylinae (Insecta: Heteroptera: Miridae) from Australia. Zootaxa 4958 (1): 319-326, DOI: 10.11646/zootaxa.4958.1.18
037387810D09FFA4618C5912D0F9F641.taxon	distribution	Distribution: Known only from the type locality in the Flinders Ranges, South Australia, Australia.	en	Schuh, Randall T., Salas, Ruth (2021): Graziaphylus, a new genus and two new species of Eremophila-feeding (Scrophulariceae) Phylinae (Insecta: Heteroptera: Miridae) from Australia. Zootaxa 4958 (1): 319-326, DOI: 10.11646/zootaxa.4958.1.18
037387810D09FFA4618C5912D0F9F641.taxon	biology_ecology	Host: Eremophila sp. (Scrophulariaceae).	en	Schuh, Randall T., Salas, Ruth (2021): Graziaphylus, a new genus and two new species of Eremophila-feeding (Scrophulariceae) Phylinae (Insecta: Heteroptera: Miridae) from Australia. Zootaxa 4958 (1): 319-326, DOI: 10.11646/zootaxa.4958.1.18
037387810D0FFFA4618C5CB7D18AF271.taxon	description	(Figs 3, 4, 18, Table 1)	en	Schuh, Randall T., Salas, Ruth (2021): Graziaphylus, a new genus and two new species of Eremophila-feeding (Scrophulariceae) Phylinae (Insecta: Heteroptera: Miridae) from Australia. Zootaxa 4958 (1): 319-326, DOI: 10.11646/zootaxa.4958.1.18
037387810D0FFFA4618C5CB7D18AF271.taxon	materials_examined	Holotype. Male. AUSTRALIA: Western Australia: 46.5 km W of Yalgoo, 28.41302 ° S 116.2151 ° E, 600 m, 27 Oct 1996, Schuh and Cassis, Eremophila forrestii F. Muell. subsp. forrestii (Scrophulariaceae), det. PERTH staff PERTH 05120705, 1 ♂ (AMNH _ PBI 00087344) (WAMP). Paratypes: AUSTRALIA: Western Australia: 46.5 km W of Yalgoo, 28.41302 ° S 116.2151 ° E, 600 m, 27 Oct 1996, Schuh and Cassis, Eremophila forrestii F. Muell. subsp. forrestii (Scrophulariaceae), det. PERTH staff PERTH 05120705, 4 ♀ (00087345, 00415326 – 00415328), 1 ♂ (00415322) (AM); 4 ♀ (00415324, 00415325, 00415332, 00415333), 1 ♂ (00415323) (AMNH); 3 ♀ (00415329 – 00415331) (WAMP).	en	Schuh, Randall T., Salas, Ruth (2021): Graziaphylus, a new genus and two new species of Eremophila-feeding (Scrophulariceae) Phylinae (Insecta: Heteroptera: Miridae) from Australia. Zootaxa 4958 (1): 319-326, DOI: 10.11646/zootaxa.4958.1.18
037387810D0FFFA4618C5CB7D18AF271.taxon	etymology	Etymology: Named for the dark-colored legs. From the Latin, niger, black, and pes, foot.	en	Schuh, Randall T., Salas, Ruth (2021): Graziaphylus, a new genus and two new species of Eremophila-feeding (Scrophulariceae) Phylinae (Insecta: Heteroptera: Miridae) from Australia. Zootaxa 4958 (1): 319-326, DOI: 10.11646/zootaxa.4958.1.18
037387810D0FFFA4618C5CB7D18AF271.taxon	diagnosis	Diagnosis: Recognized by the attributes given in the generic diagnosis, the overall length of the male (2.91), the legs castaneous / black except for the tibiae and tarsi, and the dorsum and thoracic pleuron with numerous shining, silvery, scalelike setae (Figs. 3, 4).	en	Schuh, Randall T., Salas, Ruth (2021): Graziaphylus, a new genus and two new species of Eremophila-feeding (Scrophulariceae) Phylinae (Insecta: Heteroptera: Miridae) from Australia. Zootaxa 4958 (1): 319-326, DOI: 10.11646/zootaxa.4958.1.18
037387810D0FFFA4618C5CB7D18AF271.taxon	description	Description: Male: Body conformation as in generic description; total length 2.74 – 3.18, width across pronotum 1.16 – 1.21. COLORATION (Fig. 3): As in generic description; legs almost entirely black / castaneous, except tibiae and tarsi. SURFACE AND VESTITURE (Fig. 3): As in generic description, with recumbent, dark, simple setae intermixed with relatively numerous shining, silvery, scalelike setae on dorsum (Fig. 3); dorsal margin of meso- and metapleuron broadly with numerous silvery, shining, scalelike setae; abdominal venter with numerous recumbent shining setae. STRUCTURE: As in generic description. GENITALIA (Fig. 18): Apical endosomal spine with one lateral barb; otherwise as in generic description. Female: As in generic description; total length 2.94 – 3.24, width pronotum 1.16 – 1.33. COLORATION (Fig. 4): As in male. SURFACE AND VESTITURE (Fig. 4): As in male. STRUCTURE (Fig. 4): Similar to male, but more broadly ovoid.	en	Schuh, Randall T., Salas, Ruth (2021): Graziaphylus, a new genus and two new species of Eremophila-feeding (Scrophulariceae) Phylinae (Insecta: Heteroptera: Miridae) from Australia. Zootaxa 4958 (1): 319-326, DOI: 10.11646/zootaxa.4958.1.18
037387810D0FFFA4618C5CB7D18AF271.taxon	distribution	Distribution: Known from the Northern Goldfields region, Western Australia.	en	Schuh, Randall T., Salas, Ruth (2021): Graziaphylus, a new genus and two new species of Eremophila-feeding (Scrophulariceae) Phylinae (Insecta: Heteroptera: Miridae) from Australia. Zootaxa 4958 (1): 319-326, DOI: 10.11646/zootaxa.4958.1.18
037387810D0FFFA4618C5CB7D18AF271.taxon	biology_ecology	Host: Eremophila forrestii F. Muell. subsp. forrestii (Scrophulariaceae).	en	Schuh, Randall T., Salas, Ruth (2021): Graziaphylus, a new genus and two new species of Eremophila-feeding (Scrophulariceae) Phylinae (Insecta: Heteroptera: Miridae) from Australia. Zootaxa 4958 (1): 319-326, DOI: 10.11646/zootaxa.4958.1.18
