identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03B98783FFDDFFA3FF26F902FB12FCA4.text	03B98783FFDDFFA3FF26F902FB12FCA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oribatodes Banks 1895	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Genus  Oribatodes Banks, 1895</p>
            <p> Type species:  Oribatodes mirabilis Banks, 1895</p>
            <p> Generic diagnosis. Adult. With character states of  Cepheusidae (e.g., Sitnikova 1975; Bernini &amp; Bernini 1990; Weigmann 2006; Norton &amp; Behan-Pelletier 2009). Body size: Medium-sized to large (length about 400–1000). Integument: Body and legs covered by layer of gel-like cerotegument, sometimes with bacillar, granulate and tuberculate inclusions. Prodorsum usually foveolate; notogaster without heavy sculpturing, but with simple foveolae which often forming reticulate or polygonate ornamentation; anogenital region with ridges, depressions and foveolae. Prodorsum: Rostrum rounded or narrowly triangular. Lamella broad, distinctly not reaching rostrum, with short cusp having lateral tooth. Translamella absent. Prolamella present or absent. Tutorium present, ridge-like. Rostral and lamellar setae medium-sized/long, setiform, with or without flagellate tip; interlamellar seta medium-sized/long, setiform/robust; bothridial seta with long stalk and slightly developed, barbed and/or shortly ciliate head; exobothridial seta absent. Interbothridial and postbothridial tubercles absent. Dorsosejugal porose area, dorsophragma and pleurophragma absent. Notogaster: Anterior notogastral margin well developed, distinctly convex medially, rarely straight. Humeral region with short tubercle-like process. Pteromorph and octotaxic system absent. Ten pairs of notogastral setae (centrodorsal part of notogaster without setae) short/medium-sized/long, mostly setiform, but sometimes some setae bacilliform or slightly phylliform.  Gnathosoma . Subcapitulum diarthric. All subcapitular setae setiform. Palp with setation: 0-2-1-3-9(+ω); solenidion bacilliform, coupled with eupathidium mediodistally. Axillary saccule absent. Chelicera chelate-dentate, with two setae. Epimeral and lateral podosomal regions: Epimeral setal formula: 3-1-3-3; epimeral setae comparatively short, setiform. Pedotectum I represented by large scale, pedotectum II represented by small scale. Humeral porose area not observed. Discidium and circumpedal carina present. Anogenital region: Six pairs of genital, one pair of aggenital, two pairs of anal, and three pairs of adanal setae comparatively short, setiform. Marginal porose area absent. Adanal lyrifissure distanced from anal plate. Legs: All legs monodactylous. Porose areas present on all femora and on trochanters III, IV. </p>
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	https://treatment.plazi.org/id/03B98783FFDDFFA3FF26F902FB12FCA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ermilov, Sergey G.;Bayartogtokh, Badamdorj	Ermilov, Sergey G., Bayartogtokh, Badamdorj (2024): Contribution to knowledge of the oribatid mite genus Oribatodes (Acari, Oribatida, Cepheusidae), with supplementary descriptions of two species and data on juvenile instars. Zootaxa 5485 (1): 73-91, DOI: 10.11646/zootaxa.5485.1.8, URL: http://dx.doi.org/10.11646/zootaxa.5485.1.8
03B98783FFDEFFA9FF26FC8EFA89F953.text	03B98783FFDEFFA9FF26FC8EFA89F953.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oribatodes mirabilis Banks 1895	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Oribatodes mirabilis Banks, 1895</p>
            <p>(Figs 1–6)</p>
            <p>
                 Material.   Nine adults and four nymphs (one deutonymph, three tritonymphs): USA, New York, Onondaga Co.,  
                <a title="Search Plazi for locations around (long -76.04788/lat 43.147816)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-76.04788&amp;materialsCitation.latitude=43.147816">Cicero Game Management Area</a>
                 , 43º08.869'N, 76º02.873'W, sphagnum moss in bog, 15.X.1994 (leg. R. A. Norton)  .   Five adults: USA, Wisconsin, Kenosha Co., Salem,  Silver Lake , sphagnum moss in bog, 23.XI.1978 (leg. W. Suter). Adults were compared indirectly with the type specimen of Nathan Banks (they were compared with specimens that had been previously compared directly to the type; pers. com. Roy A. Norton)  . 
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            <p> The material (adults and juvenile instars) is deposited in the collection of the Tyumen State  University Museum of Zoology , Tyumen, Russia. All specimens are preserved in 70% solution of ethanol with a drop of glycerol. </p>
            <p>Main morphological traits. Adult. Body length: 630–750. Rostrum narrowly triangular. Prolamella present, pair directed anteriorly, separated and close to each other. Rostral and lamellar setae setiform, with short flagellate tip, barbed; interlamellar seta robust, barbed; bothridial seta with long stalk and short, slightly developed head having barbs and short ciliae. Notogastral foveolae rounded, oval or polygonate. Notogastral setae c, la, lm, lp, h 2, h 3, p 2, p 3 setiform, barbed (h 2, h 3 ˃ lp ˃ c, la, lm ˃ p 2 ˃ p 3); h 1, p 1 bacilliform, heavily barbed; dorsal setae inserted on two longitudinal carinae. Leg trochanters III and IV with tooth dorsoanteriorly.</p>
            <p>Supplementary description of adult. Measurements. Body length: 630–750; notogastral width: 450–585. Sex not identified.</p>
            <p>Integument (Figs 1A–E; 5A–D; 6A, B). Body color brown to dark brown. Body and legs covered by thick layer of gel-like cerotegument with bacillar and microgranulate inclusions. Surface microgranulate sculpturing; notogaster with large rounded, oval or polygonal foveolae forming typical foveolate-reticulate ornamentation; prodorsum (except rostrum and basal part), lamella, pedotectum I, lateral part of epimere I with medium-sized foveolae; lateral part of body between notogaster and acetabula II, III densely tuberculate; anogenital region with ridges and depressions.</p>
            <p>Prodorsum (Figs 1A, B, D; 5A, C; 6A, B). Rostrum narrowly triangular. Lamellar cusp with small lateral tooth. Translamella absent. Prolamella simple, pair directed anteriorly, separated and close to each other. Transverse convex ridge located between rostral region and prolamellae. Tutorium ridge-like, heavily spinose dorsally. Rostral (67–79) and lamellar (90–101) setae setiform, with short flagellate tip, barbed; interlamellar seta (97–101) robust, barbed; bothridial seta (94–101) with long stalk and short, slightly developed head having barbs and short ciliae; exobothridial seta not observable.</p>
            <p>Notogaster (Figs 1A, D, E; 5A, C, D; 6A, B). Humeral process well defined. One pair of longitudinal carinae beginning from anterior notogastral margin and stretches along the insertions of dorsal setae (well developed between c and la; poorly observed between la and h 2). Notogastral setae (c, la, lm: 60–75; lp: 101–112; h 2, h 3: 131–139; p 2: 56–64; p 3: 49–56) setiform, barbed; h 1 (37–41) and p 1 (49–52) bacilliform, heavily barbed. Opisthonotal gland opening and all lyrifissures distinct (except ia not observable in dorsal view).</p>
            <p> Gnathosoma (Figs 1A, D; 5B). Similar to  O. reticulatus . Subcapitulum size: 131–135 × 97–101; subcapitular setae (a: 32–37; m: 19–22; h: 28–30) setiform, roughened; both adoral setae (9) setiform, smooth. Palp length: 82–86; postpalpal seta (9) spiniform, smooth. Chelicera length: 131–135; setae (cha: 37–41; chb: 22–24) setiform, barbed. </p>
            <p>Epimeral and podosomal regions (Figs 1C, D; 5B, C). All epimeral setae (3c, 4c: 41–45; 1b, 3b, 4a: 26–30; 4b: 22–26; 1c: 19–22; 1a, 2a, 3a: 15–19) setiform, roughened. Discidium elongate triangular, rounded distally. Circumpedal carina originating near posterior margin of acetabulum IV, then curving to approach circumgastric scissure asymptotically.</p>
            <p>Anogenital region (Figs 1C–E; 5B). Genital (26–34), aggenital (22–26), anal (19–26), and adanal (22–26) setae setiform, roughened. Adanal lyrifissure distinct.</p>
            <p> Legs (Figs 5B, C; 6A). Generally, similar to  O. reticulatus . </p>
            <p>Description of deuto- and tritonymphs. Measurements. Total length: 465 (deutonymph), 570–615 (tritonymphs); total width: 345 (deutonymph), 435–450 (tritonymphs).</p>
            <p>Integument (Figs 2, 3A, B). Body colorless with a yellowish tint. Body surface indistinctly porose covering by gel-like cerotegument with bacillar and tuberculate inclusions. Exuvial scalps foveolate.</p>
            <p>Prodorsum (Fig. 2). About 1/3 of gastronotic region (in lateral aspect). Rostrum rounded. Rostral seta (Dn: 67; Tn: 64–75) setiform, barbed, inserted on tubercle; lamellar seta (Dn: more than 140; Tn: more than 170) subflagellate, barbed, inserted on tubercle; interlamellar seta (Dn, Tn: 6) spiniform, smooth, located on tubercle; bothridial seta (Dn: more than 140; Tn: more than 170) subflagellate, barbed; exobothridial seta (Dn: 37; Tn: 41–52) setiform, slightly barbed.</p>
            <p>Gastronotic region (Figs 2, 3A, B). Exuvial scalps found in all nymphs, with nine or ten pairs of setae (c 1, c 2, da, dm, dp, la, lm, lp, h 2 in La, Pn; c 1, c 2, c 3, la, lm, lp, h 1, h 2, h 3, p 1 in Dn, Tn); all setae of larval scalp setiform, setae of protonymphal scalp setiform, with short flagellate tip (except simple c 1, c 2), setae of deutonymphal scalp subflagellate (except setiform c 2). Gastronotic setae c 1, c 3, la, lm, lp, h 1, h 2, h 3 p 1 very long (more than 200 in Dn, more than 300 in Tn), flagellate, barbed, often broken; p 2 (Dn: 150; Tn: 205), p 3 (Dn: 82; Tn: 157–165) setiform, with short flagellate tip; c 2 (Dn: 150; Tn: 169–184) setiform, barbed; all setae except p 2, p 3 inserted on large trapezoid apophyses. Opisthonotal gland opening and all cupules distinct in transmitted light.</p>
            <p>Gnathosoma. Similar to that of adult.</p>
            <p>Epimeral region (Figs 3A, B). Epimeral setal formula: Dn: 3-1-2-2; Tn: 3-1-3-3; all setae in Dn (1a, 1c, 2a, 3a: 17–19; others: 26–30) and Tn (1a, 1c, 2a, 3a: 19–22; 1b, 3b: 41–45; others: 30–34) setiform, slightly barbed.</p>
            <p>Anogenital region (Figs 3A, B). Genital (Dn: 26; Tn: 34–37), aggenital (Dn: 26; Tn: 34–37), anal (Tn: 26–30), and adanal (Dn: 26; Tn: 34–37) setae setiform, slightly barbed. Adanal cupule distinct; anal cupule not observable.</p>
            <p>Legs (Figs 4A–D). Claw of each leg strong, slightly barbed on dorsal side. Formulas of leg setation and solenidia: Dn: I (0-4-3-4-16) [1-2-2], II (0-4-3-4-13) [1-1-2], III (1-3-2-3-13) [1-1-0], IV (0-2-2-2-12) [0-1-0]; Tn: I (1-4-4-5-18) [1-2-2], II (1-4-4-5-15) [1-1-2], III (2-3-3-4-15) [1-1-0], IV (1-2-3-4-12) [0-1-0]; homology of setae and solenidia indicated in Table 1.</p>
            <p> Note: Roman letters refer to normal setae, Greek letters refer to solenidia (except ɛ—famulus), d φ and d σ— seta and solenidion coupled. One apostrophe (’) marks setae on anterior and double apostrophe (”) setae on posterior side of the given leg segment. Parentheses refer to a pair of setae. * —Leg setation in tritonymph of  O. reticulatus unknown. </p>
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	https://treatment.plazi.org/id/03B98783FFDEFFA9FF26FC8EFA89F953	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ermilov, Sergey G.;Bayartogtokh, Badamdorj	Ermilov, Sergey G., Bayartogtokh, Badamdorj (2024): Contribution to knowledge of the oribatid mite genus Oribatodes (Acari, Oribatida, Cepheusidae), with supplementary descriptions of two species and data on juvenile instars. Zootaxa 5485 (1): 73-91, DOI: 10.11646/zootaxa.5485.1.8, URL: http://dx.doi.org/10.11646/zootaxa.5485.1.8
03B98783FFD4FFAFFF26F8C1FDB8FB46.text	03B98783FFD4FFAFFF26F8C1FDB8FB46.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oribatodes reticulatus (Sitnikova 1979) Ermilov & Bayartogtokh 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Oribatodes reticulatus (Sitnikova, 1979)</p>
            <p>(Figs 7–12)</p>
            <p> Material.   Eleven adults and one deutonymph: Panama, Chiriqui Province, Finca Lenida, near  Boquete Casita Alta , ca. 690 m elev., debris in damp ravine, 17.III.1959 (leg. H.S. Dybas). As “  Oribatodes ? sp.” in the personal collection of Roy A. Norton  . </p>
            <p> The material (adults and deutonymph) is deposited in the collection of the Tyumen State  University Museum of Zoology , Tyumen, Russia. All specimens are preserved in 70% solution of ethanol with a drop of glycerol. </p>
            <p>Main morphological traits. Adult. Body length: 630–690. Rostrum rounded. Prolamella present, pair fused anteriorly. Rostral and lamellar setae setiform, with short flagellate tip, barbed; interlamellar seta setiform, barbed; bothridial seta with long stalk and short, slightly developed head having barbs and short ciliae. Notogastral foveolae rounded. Notogastral setae medium-sized, setiform, barbed. Pedotectum I with small tubercles (not foveolate). Leg trochanters III and IV with tooth dorsoanteriorly.</p>
            <p>Supplementary description of adult. Measurements. Body length: 630–690; notogastral width: 495–540. Gender not identified.</p>
            <p>Integument (Figs 7A–E; 11A–D; 12A, B). Body color brown to dark brown. Body and legs covered by thick layer of gel-like cerotegument with microgranulate inclusions. Surface microgranulate sculpturing; notogaster with large rounded foveolae forming typical foveolate-reticulate ornamentation; prodorsum (except rostrum and basal part), lamella, lateral part of epimere I with medium-sized foveolae; pedotectum I with small tubercles (not foveolate); lateral part of body between notogaster and acetabula II, III densely tuberculate; anogenital region with ridges and depressions.</p>
            <p>Prodorsum (Figs 7A, B, D; 11A, C; 12A, B). Rostrum rounded, with thin convex carina connecting insertions of rostral setae. Lamellar cusp with small lateral tooth. Translamella absent. Prolamella simple, pair fused anteriorly. Interlamellar region with longitudinal, elongate triangular ridge. Tutorium ridge-like, heavily spinose dorsally. Rostral (82–90) and lamellar (123–150) setae setiform, with short flagellate tip, barbed; interlamellar seta (60–67) setiform, barbed; bothridial seta (112–123) with long stalk and short, slightly developed head having barbs and short ciliae; exobothridial seta not observable.</p>
            <p>Notogaster (Figs 7A, D, E; 11A, C, D; 12A). Humeral process well defined. All notogastral setae (c, la, lm, lp, h 2, h 3: 60–67; h 1, p 1, p 2, p 3: 41–52) setiform, barbed. Opisthonotal gland opening and all lyrifissures distinct (except ia not observable in dorsal view).</p>
            <p>Gnathosoma (Figs 8A–C; 11B). Subcapitulum size: 142–150 × 105–120; subcapitular setae (a, h: 26–30; m: 19) setiform, roughened; both adoral setae (9–11) setiform, smooth. Palp length: 82–86; postpalpal seta (11) spiniform, smooth. Chelicera length: 142–150; setae (cha: 41–43; chb: 22–26) setiform, barbed.</p>
            <p>Epimeral and podosomal regions (Figs 7C,D; 11B,C).All epimeral setae (3c, 4c:34–41; 1b, 3b, 4a, 4b: 19–26; 1a, 1c, 2a, 3a: 11) setiform, roughened. Discidium elongate triangular, rounded distally. Circumpedal carina originating near posterior margin of acetabulum IV, then curving to approach circumgastric scissure asymptotically.</p>
            <p>Anogenital region (Figs 7C–E; 11B). Genital, aggenital, anal, and adanal setae (22–26) setae setiform, roughened. Adanal lyrifissure distinct.</p>
            <p>Legs (Figs 8D–G; 11B, C; 12A). Claw of each leg strong, slightly barbed on dorsal side. Trochanters III and IV with long tooth-like process dorsoanteriorly. Dorsoparaxial porose area on femora I–IV and on trochanters III, IV well visible. Formulas of leg setation and solenidia: I (1-5-3-4-20) [1-2-2], II (1-4-3-4-16) [1-1-2], III (2-3-2-3- 15) [1-1-0], IV (1-2-3-3-12) [0-1-0]; homology of setae and solenidia indicated in Table 1. Famulus short, slightly swollen and blunt distally, located posterior to solenidia ω 1 and ω 2; seta ft” specifically bent in the middle; seta s eupathidial; solenidion φ 1 on tibia I long, subflagellate versus other solenidia short or medium-sized, rod-like or slightly bacilliform.</p>
            <p>Description of deutonymph. Measurements. Total length: 435; total width: 315.</p>
            <p>Integument (Figs 9A–C). Body colorless with a yellowish tint. Body surface indistinctly porose covering by gel-like cerotegument with bacillar and tuberculate inclusions.</p>
            <p>Prodorsum (Figs 9A, C). About 1/2 of gastronotic region (in lateral aspect). Rostrum rounded. Rostral (64) and lamellar setae setiform, barbed, inserted on tubercles; interlamellar seta (7) spiniform, smooth, located on tubercle; bothridial seta (150) setiform, with flagellate tip, barbed; exobothridial seta (37) setiform, slightly barbed.</p>
            <p>Gastronotic region (Figs 9A, C). Exuvial scalps not found. Gastronotic setae c 1, c 3, la, lm, lp, h 1, h 2, h 3 very long (more 400), subflagellate, barbed, often broken; c 2 (37), p 1 (94), p 2 (79), p 3 (56) setiform, barbed; all setae except p 2, p 3 inserted on large trapezoid apophyses. Opisthonotal gland opening and all cupules distinct in transmitted light.</p>
            <p>Gnathosoma (Fig. 9B). Similar to that to adult.</p>
            <p>Epimeral region (Figs 9B, C). Epimeral setal formula: 3-1-2-2; all setae (17–22) setiform, slightly barbed.</p>
            <p>Anogenital region (Figs 9B, C). Genital, aggenital and adanal setae (17–22) setiform, slightly barbed. Adanal cupule distinct; anal cupule not observable.</p>
            <p>Legs (Figs 10A–D). Claw of each leg strong, slightly barbed on dorsal side. Formulas of leg setation and solenidia: I (0-4-3-4-16) [1-2-2], II (0-4-3-4-13) [1-1-2], III (1-3-2-3-13) [1-1-0], IV (0-2-2-2 -12) [0-1-0]; homology of setae and solenidia indicated in Table 1.</p>
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	https://treatment.plazi.org/id/03B98783FFD4FFAFFF26F8C1FDB8FB46	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ermilov, Sergey G.;Bayartogtokh, Badamdorj	Ermilov, Sergey G., Bayartogtokh, Badamdorj (2024): Contribution to knowledge of the oribatid mite genus Oribatodes (Acari, Oribatida, Cepheusidae), with supplementary descriptions of two species and data on juvenile instars. Zootaxa 5485 (1): 73-91, DOI: 10.11646/zootaxa.5485.1.8, URL: http://dx.doi.org/10.11646/zootaxa.5485.1.8
03B98783FFCCFFB1FF26FE33FA51FBD2.text	03B98783FFCCFFB1FF26FE33FA51FBD2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oribatodes	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Key to known species of  Oribatodes (adult) </p>
            <p> 1. Interlamellar seta very long, extending beyond tip of lamella; anterior margin of notogaster almost straight; body length: 850.............................  O. crenulatus Csiszár, 1962 (see also Walter et al. 2013). Distribution: Southeast Europe. </p>
            <p>- Interlamellar seta not extending beyond tip of lamella; anterior margin of notogaster distinctly convex medially.......... 2</p>
            <p> 2. Interlamellar seta very short (not longer than diameter of bothridium); foveolae of notogaster specifically deformed; body length: 888.................................................  O. grandis (Sitnikova, 1979) . Distribution: Ecuador. </p>
            <p>- Interlamellar seta distinctly longer than diameter of bothridium; foveolae of notogaster rounded, oval or polygonal (not deformed).......................................................................................... 3</p>
            <p>3. Notogastral setae la, lm, lp, h 2, h 3 robust, heavily barbed...................................................... 4</p>
            <p>- Notogastral setae la, lm, lp, h 2, h 3 setiform, not heavily barbed................................................. 5</p>
            <p> 4. Interlamellar area with foveolae; body length: 530–600..................................................................  O. mirabiloides (Jacot, 1938) (see also Baker &amp; Wharton 1952; Walter et al. 2013). Distribution: Eastern U.S.A. </p>
            <p> - Interlamellar area with ridges; body length: 557–635............................................  O. decoui (Vasiliu &amp; Călugăr, 1977) (see also Balogh &amp; Mahunka 1979 —as for  Cepheus (Cubacepheus) lobatus ). Distribution: Eastern U.S.A. </p>
            <p> 5. Foveolae of notogaster sparsely located, not forming foveolate-reticulate ornamentation; body length: 725........................................................................  O. areolatus (Sitnikova, 1979) . Distribution: Ecuador. </p>
            <p>- Foveolae of notogaster closely located, forming foveolate-reticulate ornamentation................................ 6</p>
            <p> 6. Interlamellar seta long, longer than distance between insertions of in–in, robust; rostrum narrowly triangular; body length: 500– 750.....  O. mirabilis Banks, 1895 (see also Berlese 1910; Banks 1915; Walter et al. 2013). Distribution: Northern Nearctic. </p>
            <p> - Interlamellar seta medium-sized, almost twice shorter than distance between insertions of in–in, setiform; rostrum rounded; body length: 630–690............................  O. reticulatus (Sitnikova, 1979) . Distribution: Northern Neotropics. </p>
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	https://treatment.plazi.org/id/03B98783FFCCFFB1FF26FE33FA51FBD2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ermilov, Sergey G.;Bayartogtokh, Badamdorj	Ermilov, Sergey G., Bayartogtokh, Badamdorj (2024): Contribution to knowledge of the oribatid mite genus Oribatodes (Acari, Oribatida, Cepheusidae), with supplementary descriptions of two species and data on juvenile instars. Zootaxa 5485 (1): 73-91, DOI: 10.11646/zootaxa.5485.1.8, URL: http://dx.doi.org/10.11646/zootaxa.5485.1.8
