taxonID	type	description	language	source
03BB87B9095CFFE9FF32035AFC10A1C4.taxon	description	Figs. 1 A, 1 B and 3 A. External morphology. — Lot MLP DB 4806, Figs. 4 A and 4 B. Body depressed (BMH / BMW = 0.86 ± 0.04); body length little longer than one third of total length (BL / TL = 0.34 ± 0.01); body shape ovoid in dorsal view; widest at posterior third of body, behind spiracle. In lateral view, ventral contour of body flat in gular and branchial regions, convex in abdominal region. Dorsal contour of body slightly convex from eyes to origin of dorsal fin. Snout semicircular in dorsal view, rounded in lateral view. Nostrils oval, with thin pigmented marginal rim, with a small subtriangular fleshy projection in medial margin (Fig. 18 A); nostrils dorsolaterally located (EN / BWN = 0.58 ± 0.04), placed in a depression and closer to eyes than to the tip of the snout (FN / END = 1.45 ± 0.18), more visible in dorsal than in lateral view. Eyes large (E / BWE = 0.24 ± 0.02), dorsally positioned (EO / BWE = 0.73 ± 0.03), dorsolaterally directed, not visible in ventral view. Spiracle single, lateral and sinistral; its inner wall fused to body except for its distal end (Fig. 17 A); its opening oval, slightly elevated, with a diameter smaller than tube diameter, located between second and posterior thirds of body (RSD / BL = 0.64 ± 0.02), posterodorsally directed, visible in lateral and dorsal views. Lateral line system visible. Intestinal assa located at centre of abdominal region. Vent tube starts at midline, at posterior end of body, reaching free margin of lower fin, opening dextral. Tail large (TaL / TL = 0.66 ± 0.01), with both fins a little higher than body height (MTH / BMH = 1.07 ± 0.07). Dorsal fin originates at tail-body junction. Edge of dorsal fin slightly convex, free margin of ventral fin almost flat. Tail axis straight, tail tip pointed; tail musculature reaching tail tip. Oral disc (Fig. 8 A) anteroventral, small (OD / BMW = 0.35 ± 0.02, disc measured folded), with a well marked infra – angular constriction at each side (also a less marked supra-angular constriction in two specimens). Marginal papillae simple, longer than wide, with rounded or pointed tip. Dorsal gap present, medium-sized (DG / OD = 0.31 ± 0.05). Row of marginal papillae single or double; some infraangular submarginal papillae present. Upper jaw sheath widely arch-shaped, with a slightly convex medial region in one specimen. Lower jaw sheath with free margin U-shaped. Both jaw sheaths well developed, serrated and heavily pigmented in the distal half (upper one) or third (lower one). Labial tooth row formula 2 (2) / 4 (1). A 1 bent with an angle directed to the front. Length of P 4 about half to one third of P 3. In four specimens, P 4 appears fragmented. Some lateral flaps or submarginal papillae with labial teeth are present. Tooth of all rows with spatulate and convex heads bearing 6 – 8 well-marked cusps (Fig. 10 A). Coloration in preservative. — Body light brown, with dark brown spots on dorsal and dorsolateral regions of body. A dark brown, triangular interocular spot, with the base directed backwards. Perinasal region dark brown. Ventral region transparent. Caudal musculature brown in dorsal view, whitish in lateral view with small dark blotches that coalesce forming a reticulation. A black horizontal band runs between epi- and hypaxial musculature for first third of the tail’s length. Fins opalescent with few dark flecks, more abundant in dorsal fin. Variation. — Lot MLP DB 5520. In two specimens (stages 33 and 36) the A 1 and A 2 tooth row are abnormally arranged. Oral cavity morphology. — (Figs. 11 A and 13 A). Buccal roof: Prenarial arena presents a low transversal crest and some pustules surrounding it. Choanae transversely oriented, with anteromedial and rounded vacuities, posterior margins forming the narial valves, and both margins undulated by the presence of small prenarial pustules, in particular the anterior one. Postnarial arena presents 8 conical papillae forming an arch between the lateral ridges. There are some pustules arranged among the postnarial papillae. The low, rounded and wide median ridge presents short undulations on its free margin. Each square-shaped lateral ridge papilla possess about 4 – 5 projections on the free margin, none of them predominant in size. There are approximately 160 pustules and 40 long and conical papillae on the buccal roof arena (BRA). The pustules are mainly on the central region and the papillae on the posterior and most lateral areas of the arena placed in a “ V ” arrangement. Posterior area of the roof with secretory pits and medially interrupted dorsal velum with lobated margins. Buccal floor: Infralabial papillae on infrarostral cartilages absent. There are two long and digitiform infralabial papillae placed at the level of cartilago meckeli. The lingual anlage bears 2 long and conical lingual papillae placed very close to each other and practically fused at the midline in some specimens. Buccal pockets mostly transversal to the axial axis. Prepocket area with 4 long and 14 short conical papillae and about 80 pustules. There are 20 long and conical papillae (two of them bifid), 26 short and conical papillae and about 190 pustules on the buccal floor arena (BFA). The papillae arrange in one anterior and other posterior Vshaped patterns. The ventral velum has a weakly-marked median notch, three poorly-developed marginal projections at each side of the posterior margin and many secretory pits.	en	Kolenc, Francisco, Borteiro, Claudio, Alcalde, Leandro, Baldo, Diego, Cardozo, Dario, Faivovich, Julián (2008): Advertisement call and female sexual cycle in Uruguayan populations of Physalaemus henselii (Anura, Leiuperidae). Zootaxa 1927 (2): 1-66
03BB87B90952FFEAFF320050FB3EA47C.taxon	description	Figs. 1 C and 1 D External morphology. — Lot MLP DB 4258, Figs. 4 C and 4 D. Body slightly depressed (BMH / BMW = 0.90 ± 0.04); body length little less than one third of total length (BL / TL = 0.32 ± 0.01); body ovoid in dorsal view; widest at posterior third of body. In lateral view, ventral contour of body flat or slightly concave in gular and branchial regions, convex in abdominal region. Dorsal contour of body convex. Snout acutely rounded in dorsal view, rounded in lateral view. Nostrils oval, with a subtriangular or rounded elevated projection in medial margin (Fig. 18 B); nostrils dorsolaterally located (EN / BWN = 0.68 ± 0.04), placed in a depression, closer to eyes than to the tip of snout (FN / END = 1.38 ± 0.24), more visible in dorsal than in lateral view. Eyes large (E / BWE = 0.27 ± 0.01), dorsally positioned (EO / BWE = 0.91 ± 0.02), laterally oriented, external edge of corneas visible in ventral view. Spiracle single, lateral and sinistral; it projects a tube posterodorsally directed, with its internal wall free from body; its opening oval, with a diameter smaller than or equal to tube diameter, located in posterior third of body (RSD / BL = 0.76 ± 0.05), visible in dorsal and lateral views. Lateral line system visible. Infraorbital line runs in a groove which is evident from anterior edge of nostrils to behind posterior edge of eyes (Fig. 19 A). Intestinal assa located at centre of abdominal region. Vent tube starts at midline, at the posterior end of body, reaches margin of the lower fin and opens dextrally. Tail large (TaL / TL = 0.68 ± 0.01), with both fins higher than body height (MTH / BMH = 1.10 ± 0.06). Dorsal fin originates at the end of body; its origin is abrupt in some specimens. Ventral fin origin concealed by vent tube. Tail axis straight, tail tapering markedly in last half; end acute; tail musculature reaching tail tip. Oral disc (Figs. 8 B and 9 A) anteroventral, small (OD / BMW = 0.35 ± 0.05, disc measured folded) with a well marked infraangular constriction on each side. Marginal papillae simple, longer than wide, subconical, with rounded tip. Dorsal gap present, but small (DG / OD = 0.20 ± 0.03). Row of marginal papillae single on both sides of the dorsal gap, double at angular regions and simple or double at posterior margin of oral disc; some infraangular submarginal papillae present. Upper jaw sheath widely arch-shaped. Lower jaw sheath with free margin V-shaped. Both jaw sheaths well developed, serrated but sparsely pigmented distally. Labial tooth row formula 2 (1,2) / 3 (1) (six specimens), 2 (1,2) / 3 (two specimens), 2 (1,2) / 3 (1,2) (one specimen) or 2 (1,2) / 0 (one specimen, posterior rows likely lost). Length of P 3 about one third the length of other rows of labial teeth. Coloration in life. — Body yellowish brown in dorsal view, with small dark blotches that coalesce forming a reticulation which is more noticeable laterally. A dark brown, triangular interocular spot, with the base directed backwards. Nostrils surrounded by a dark brown region. Lateral sides of body, head and proximal third of caudal musculature with golden, copper and silver spots and blotches. Gular region transparent. Branchial region with white spots. Abdominal surface white. Caudal musculature yellowish brown, with a black horizontal line between hypaxial and epaxial musculature, that runs for about one third of its length. Distal third of caudal epaxial musculature with dark blotches. Tail fins translucent, with rounded dark brown spots. Iris golden, with four dark zones at the edges of the eye, at the end of imaginary horizontal and vertical rays arising from the pupil. Coloration in preservative. — Body dark brown with unpigmented regions. Small dark spots on dorsal and dorsolateral regions of body. Ventral region transparent. Caudal musculature with dark irregular bands in lateral and dorsal views. Fins opalescent with irregular dark blotches. Variation. — Lot MLP DB 5052. LTRF 2 (1,2) / 3 (1) in all specimens. Submarginal papillae absent in two specimens (stages 27 and 34), present in the other two (stages 35 and 38). Some labial teeth between upper jaw sheath and A 2 on one side of the oral disc in one individual (stage 34). Oral cavity morphology. — (Fig. 11 B and 13 B). Buccal roof: There are 8 blunt pustules irregularly distributed on the prenarial arena. These pustules do not form a clear prenarial transversal crest, however there are 3 – 4 of them arranged in the same position that the crest of the previous species. Choanae transversely oriented, with anteromedial and rounded vacuities, posterior margins forming the narial valves, and both margins undulated by the presence of small prenarial pustules, in particular the anterior one. On the postnarial arena, there are 16 pustules and 6 short and blunt papillae. The low, rounded and wide median ridge presents 17 short undulations on its free margin. Each square-shaped lateral ridge papilla possess about 5 – 6 projections on the free margin, one of them very large. The BRA is not defined, because of the absence of papillae. There are approximately 180 pustules and a few short and conical papillae on the buccal floor; the pustules are mostly on the central region, but there are some in the lateral region forming a continuum with the former. Posterior area of the roof with secretory pits, and dorsal velum bearing lobated margins. Buccal floor: Infralabial papillae on infrarostral cartilages absent. There are two long and digitiform infralabial papillae placed at the level of cartilago meckeli (not figured). The lingual anlage bears 2 long and conical lingual papillae placed very close to each other. Buccal pockets mostly transversal to the axial axis. Prepocket area scattered with 42 pustules and 24 conical papillae, four of them long. There are 5 long and 7 short conical papillae, and about 160 pustules on the BFA. The papillae are arranged in two V-shaped patterns, one anterior and other posterior. The ventral velum has a weakly-marked median notch, three poorly-developed marginal projections at each side of the posterior margin and many secretory pits.	en	Kolenc, Francisco, Borteiro, Claudio, Alcalde, Leandro, Baldo, Diego, Cardozo, Dario, Faivovich, Julián (2008): Advertisement call and female sexual cycle in Uruguayan populations of Physalaemus henselii (Anura, Leiuperidae). Zootaxa 1927 (2): 1-66
03BB87B90951FFEBFF3206B8FE28A74C.taxon	description	Figs. 1 E, 1 F, 3 B and 3 C External morphology. — Lot MLP DB 4722, Figs. 5 A and 5 B. Body slightly depressed (BMH / BMW = 0.94 ± 0.03); body length little less than one third of total length (BL / TL = 0.31 ± 0.01); body oval in dorsal view, widest at middle or posterior thirds of body. In lateral view, ventral contour of body flat in gular and branchial regions, slightly convex in abdominal region. Dorsal contour of body almost flat and sloping from nostrils to origin of dorsal fin. Snout semicircular in dorsal view and rounded in lateral view. Nostrils oval, with a thin marginal rim, with a small rounded fleshy projection in medial margin; nostrils dorsolaterally located (EN / BWN = 0.51 ± 0.03), placed in a depression and closer to eyes than to the tip of snout (FN / END = 1.10 ± 0.17), more visible in dorsal than in lateral view. Eyes large (E / BWE = 0.23 ± 0.02), dorsally positioned (EO / BWE = 0.83 ± 0.03), laterally directed, not visible in ventral view. Spiracle single, lateral and sinistral; its inner wall fused to body except for its very distal end; its opening oval, slightly elevated, with a diameter smaller than tube diameter, located in posterior third of body (RSD / BL = 0.76 ± 0.05), posterodorsally directed and visible in lateral and dorsal views. Lateral line system visible. Intestinal assa located at centre of abdominal region. Vent tube starts at midline, at posterior end of body, reaching free margin of lower fin, opening dextral. Tail large (TaL / TL = 0.69 ± 0.01), with both fins higher than body height (MTH / BMH = 1.09 ± 0.08). Dorsal fin originates on end of body. Ventral fin origin concealed by vent tube. Edges of both fins subparallel for first and second thirds, convergent in last third. Tail axis straight, tail tip acute or slightly rounded; tail musculature reaching tail tip. Oral disc (Fig. 8 C) anteroventral, small (OD / BMW = 0.39 ± 0.04, disc measured folded), with a well marked infraangular and a less evident supraangular constriction. Marginal papillae simple, longer than wide, subconical, with rounded tip. Dorsal gap present, but small (DG / OD = 0.29 ± 0.06). Row of marginal papillae single on both sides of dorsal gap, double at angular regions and double or single at posterior margin of oral disc; some infraangular submarginal papillae usually present. Upper jaw sheath widely arch-shaped. Lower jaw sheath with V-shaped free margin. Both jaw sheaths well developed, but upper one appears smooth and sparsely pigmented distally, while inferior one is finely serrated. Labial tooth row formula 2 (2) / 3 (1). A 1 bent at the middle with an angle directed to the front. Length of P 3 about half the length of P 2. Most specimens have short flaps with labial teeth at angular regions, unconnected with main rows. Tooth of all rows with spatulate, convex heads bearing 8 – 10 well marked cusps (Fig. 10 B). Coloration in life. — Body light brown with small dark dots, more abundant in central region, a dark triangle between the eyes. Perinasal region dark brown. Gular region with small dark dots. Abdominal region whitish. Caudal musculature dark brown in dorsal view, orange-brown in lateral view, with a black horizontal band between epi- and hypaxial musculature running for first third of the tail’s length. Small golden flecks on dorsal and lateral sides of body and tail musculature. Fins translucid with golden dots. Iris gold, with three radial black dots arising from the pupil (a fourth dorsal dot is not evident). Coloration in preservative. — Body dark brown in dorsal and lateral views with barely evident small dots. Abdominal region whitish. Caudal musculature in dorsal view dark brown; yellowish brown in lateral view, with rounded unpigmented regions and distal third entirely dark brown. A black horizontal band runs between epi- and hypaxial musculature for first third of the tail’s length. Fins opalescent with few dark flecks, more abundant in dorsal fin. Variation. — Lot MLP DB 3280. Supraangular constrictions on both sides of oral disc are more evident than in lot MLP DB 4722. Perioral region pigmented. Some specimens lack submarginal papillae. Jaw sheaths smooth and almost lacking pigmentation. All specimens lacking lateral flaps with labial teeth. Oral cavity morphology. — (Fig. 11 C and 13 C). Buccal roof: Prenarial arena with a low and wide transversal crest and some pustules surrounding it. Choanae transversely oriented, with anteromedial and rounded vacuities, posterior margins forming the narial valves, and both margins undulated by the presence of small prenarial pustules, in particular the anterior one (Fig. 16 A, B). There are about 16 pustules and 8 short papillae on the postnarial arena. The set of postnarial papillae are arranged transversely. The most lateral papilla of each side is bifid. Each square-shaped lateral ridge papilla possesses 4 – 5 digitiform projections on the free margin, any of them predominant in size. The low, rounded and wide median ridge presents 17 undulations on its free margin. There are about 96 pustules and 44 short and conical papillae placed in the central region of the BRA. Almost all papillae are placed in the lateral regions and on the posterior margin of the arena. Posterior area of the roof with secretory pits and dorsal velum bearing lobated margins. Buccal floor: Infralabial papillae on the infrarostral cartilages absent. There are two long and digitiform infralabial papillae placed at the level of cartilago meckeli (not figured). The lingual anlage bears 2 long and conical lingual papillae. Buccal pockets mostly transversal to the axial axis. Prepocket area bears 40 pustules and 44 conical papillae, two of them longer and bifid. There are 25 conical papilla of moderate size (6 of them longer than the rest), and about 50 pustules on the BFA. The papillae arranged in two V-shaped patterns but not as clear as in other species. There is a flap (that resembles the lateral ridges of the roof), bearing 5 projections on its free end, and placed posterior to the medial flange of each pocket. The ventral velum has a wellmarked median notch, three poorly-developed marginal projections at each side of the posterior margin and many secretory pits.	en	Kolenc, Francisco, Borteiro, Claudio, Alcalde, Leandro, Baldo, Diego, Cardozo, Dario, Faivovich, Julián (2008): Advertisement call and female sexual cycle in Uruguayan populations of Physalaemus henselii (Anura, Leiuperidae). Zootaxa 1927 (2): 1-66
03BB87B90957FFEDFF3202E0FB0BA2B4.taxon	description	Figs. 2 A and 3 D External morphology. — Lot MACN 35116, (Figs. 5 C – E). Body depressed (BMH / BMW = 0.86 ± 0.03); body length little less than one third of total length (BL / TL = 0.31 ± 0.01); body shape oval in dorsal view, widest behind the eyes. In lateral view, ventral contour of body slightly concave in gular and branchial regions, convex in abdominal region. Snout rounded in dorsal and lateral views. Dorsal contour of body almost flat from eyes to origin of dorsal fin. Nostrils oval, rimmed, with triangular elevated projection in the medial margin (Fig. 18 C); nostrils dorsolaterally located (EN / BWN = 0.67 ± 0.05), placed in a depression, closer to eyes than to the tip of snout (FN / END = 1.20 ± 0.17), more visible in dorsal than in lateral view. Eyes large (E / BWE = 0.19 ± 0.02), dorsally positioned (EO / BWE = 0.79 ± 0.03), dorsolaterally directed, not visible in ventral view. Spiracle single, lateral, sinistral and short; its inner wall fused to body except for a variable length at its distal end; its opening oval, elevated, with diameter slightly smaller than tube diameter, located in posterior third of body (RSD / BL = 0.74 ± 0.03), posterodorsally directed, visible in lateral and posterior views. Lateral line system visible with a unique pattern of ventrolateral aggregations of neuromasts at each side of the body, near the body-tail junction (Figs. 15 A – E and 20 A). Intestinal assa located approximately at centre of abdominal region. Vent tube starts at midline, at the posterior end of the body, reaches free margin of lower fin, opening dextral. Tail large (TAL / TL = 0.69 ± 0.01), with both fins higher than body height (MTH / BMH = 1.12 ± 0.05). Dorsal fin originates at tail-body junction. Ventral fin origin concealed by vent tube. Tail axis straight, edges of both fins subparallel, converging at posterior third, end acutely rounded; tail musculature reaching tail tip. Oral disc (Figs. 8 D and 9 B) anteroventral, not visible dorsally, small (OD / BMW = 0.39 ± 0.04), ventrolateral folds present, well marked infraangular constriction at each side of the oral disc. Marginal papillae simple, longer than wide, subconical, with rounded tip. Dorsal gap present, but small (DG / OD = 0.23 ± 0.03). Row of marginal papillae single or alternate on both sides of the dorsal gap, usually double at the angular regions and double or single in the posterior margin of the oral disc except at midline where they are sparse and separated in most specimens, very short ventral gap present in two specimens (stages 28 and 33); some infraangular submarginal papillae are usually present. Upper jaw sheath widely arch-shaped. Lower jaw sheath with V-shaped free margin. Jaw sheaths well developed, serrated and heavily pigmented distally. Labial tooth row formula 2 (2) / 4 (1). Length of P 4 one quarter to half the length of P 2, usually fragmented. Most of the specimens have short flaps with labial teeth at the angular regions, unconnected with main rows. Tooth of all rows spatulate and slightly concave bearing 5 – 10 poorly-marked distal cusps (Fig. 10 C). Coloration in preservative. — Tadpoles at stages 25 – 27 body brown, venter translucent, tail and fins black, specially the distal half (Fig. 5 E). Tadpoles at later stages (33 – 37) body dark brown with inconspicuous dark dots in head and snout, tail light brown, with scattered dark blotches in proximal third, with dark reticulation in distal third (tail light brown with rounded dark blotches in some specimens). Belly dark. All specimens have an unpigmented spot ventrolaterally on each side of the abdomen representing aggregations of neuromasts randomly distributed. Variation. — Lot MLP DB 5410, N = 8, stages 25 – 27; and MLP DB 3890, N = 3, stages 28 – 34. All tadpoles LTRF 2 (2) / 4 (1) — except for one tadpole stage 33 in which oral structures are destroyed. P 4 very short. Row of marginal papillae single or double, usually double in angular regions, dorsal gap present in all specimens, very short ventral gap in three specimens (with one papilla in the middle in two of them, stages 26 and 28), papillae sparse in the middle of the posterior border of the oral disc in all other specimens, usually with submarginal papillae laterally. Lateral flaps with labial teeth in 7 specimens. Oral cavity morphology. — (Figs. 11 D and 13 D). Buccal roof: Prenarial arena scattered with at least 5 blunt pustules and a clear transversal crest. Choanae transversely oriented, with anteromedial and rounded vacuities, posterior margins forming the narial valves, and both margins undulated by the presence of small prenarial pustules, in particular the anterior one. There are about 48 pustules, 9 short and 10 long conical postnarial papillae between the lateral ridge papillae. The postnarial pustules are scattered among the papillae and also near the posterior margins of the choana. Each square-shaped lateral ridge papilla possesses 5 projections on the free margin, none predominant in size. The low, rounded and wide median ridge presents 9 undulations on its free margin. BRA is not defined; there are approximately 330 pustules and only 10 lateral short and bifid papillae on the buccal roof. Posterior area of the roof with secretory pits and dorsal velum bearing lobated margins. Buccal floor: There are 3 – 5 pairs of short and digitated infralabial papillae at the level of the infrarostral cartilages (not figured). There are two long and digitiform infralabial papillae placed at the level of cartilago meckeli (not figured). The lingual anlage bears 2 long and ramified lingual papillae placed very close to each other and with a pustule near their basis. Buccal pockets mostly transversal to the axial axis. Prepocket area with 18 short and conical papillae and about 100 pustules. There are 22 long and conical papillae (4 of them bifid), 16 short and conical papillae and about 260 pustules on the BFA. The papillae are arranged in clear two V-shaped patterns, one anterior and other posterior. The ventral velum has a well-marked median notch, three weakly-marked undulations at each side of the posterior margin and many secretory pits.	en	Kolenc, Francisco, Borteiro, Claudio, Alcalde, Leandro, Baldo, Diego, Cardozo, Dario, Faivovich, Julián (2008): Advertisement call and female sexual cycle in Uruguayan populations of Physalaemus henselii (Anura, Leiuperidae). Zootaxa 1927 (2): 1-66
03BB87B90956FFEFFF320160FA38A4E4.taxon	description	Figs. 2 C, 2 D and 3 E External morphology. — Lot ZVCB 16147, Fig. 6 A and 6 B. Body slightly depressed (BMH / BMW = 0.93 ± 0.07); body length little less than one third of total length (BL / TL = 0.31 ± 0.02); body oval in dorsal view, widest at middle third of body. In lateral view, ventral contour of body flat in gular and branchial regions, slightly convex in abdominal region. Dorsal contour of body smoothly convex. Snout semicircular in dorsal view, rounded in lateral view. Nostrils oval, with thin marginal rim, with a subtriangular or rounded elevated fleshy projection in medial margin; nostrils dorsolaterally located (EN / BWN = 0.59 ± 0.04), placed in a depression, closer to eyes than to the tip of snout (FN / END = 1.11 ± 0.13), more visible in dorsal than in lateral view. Eyes large (E / BWE = 0.27 ± 0.01), laterally positioned (EO / BWE = 0.87 ± 0.01), laterally directed, corneas visible in ventral view. Spiracle single, lateral and sinistral; its inner wall fused to body except for its distal end; its opening oval, elevated, with a diameter smaller than tube diameter, located in posterior third of body (RSD / BL = 0.73 ± 0.02), posterodorsally directed, visible in lateral and dorsal views. Lateral line system visible. Intestinal assa located at centre of abdominal region. Vent tube starts at midline, reaching free margin of lower fin, opening dextral. Tail large (TaL / TL = 0.69 ± 0.02), with both fins higher than body height (MTH / BMH = 1.25 ± 0.06). Dorsal fin originates on end of body. Ventral fin origin concealed by vent tube. Tail axis straight, dorsal fin edge slightly convex and ventral fin edge almost straight, tail tapering markedly in last third, end acute; tail musculature reaching tail tip. Oral disc (Figs. 8 E and 9 C) anteroventral, small (OD / BMW = 0.43 ± 0.04, disc measured folded), with a well marked infraangular constriction on each side. Marginal papillae simple, longer than wide, with rounded tip. Dorsal gap present, small (DG / OD = 0.29 ± 0.06). Row of marginal papillae single except for angular regions on which it is double; some wider and shorter lateral submarginal papillae present. Upper jaw sheath widely arch shaped with a convex central portion. Lower jaw sheath with V-shaped free margin. Jaw sheaths well developed, serrated and pigmented on their distal third. Labial tooth row formula 2 (2) / 3 (1). A 1 bent with an angle directed to the front. Length of P 3 about half the length of P 2. Tooth of all rows spatulated and concave bearing 8 – 10 well-marked distal cusps (Fig. 10 D). Coloration in preservative. — Body yellowish-brown in dorsal view, dark brown on head and next to inner margins of nostrils, with small dark spots on head and body. Dark brown laterally. Venter translucent. Caudal musculature yellowish tan, with irregular darker regions that in some specimens made an alternating dark and light transversal pattern in dorsal view. Fins opalescent, with dark blotches. Variation. — MLP DB 7144: LTRF 2 (2) / 3 (1), except for one example 2 (2) / 3 and another one 2 (1,2) / 3 (1). Three specimens with labial teeth placed on lateral submarginal papillae, on a infraangular lateral flap, or on the anterior labium on both sides of the dorsal gap. MLP DB 5051: Body ovoid in dorsal view, widest on last third of body. Two specimens (stages 30 and 31) with scarcely evident supraangular constrictions on both sides of oral disc. LTRF 2 (2) / 3 (1). MLP DB 4257: In two specimens (stage 31) LTRF 2 (1,2) / 3 (1), being 2 (2) / 4 (1) in another one (stage 33), which has a short and fragmented P 4. Row of marginal papillae double on posterior margin of oral disc in some specimens. P 2 has a medial angulation (but is not divided) in six specimens. MLP DB 5306: Row of marginal papillae single, sparse submarginal papillae laterally. LTRF 2 (2) / 3 (2). P 3 about 1 / 3 the length of P 2. Oral cavity morphology. — (Figs. 12 A and 14 A). Buccal roof: Prenarial arena with a compact group of 3 short and blunt transversely oriented pustules, not fused forming a crest. Choanae transversely oriented, with anteromedial and rounded vacuities, posterior margins forming the narial valves, and both margins undulated by the presence of small prenarial pustules, in particular the anterior one. There are about 16 pustules, and 6 short papillae on the postnarial arena. The postnarial papillae are transversely oriented between the two lateral ridge papillae. The most lateral papilla of each side is bifid. Each square-shaped lateral ridge papilla possesses 4 – 5 digitiform projections on the free margin, none of them predominant in size. The low, rounded and wide median ridge presents 9 – 11 undulations on its free margin. There are approximately 140 pustules and only 10 – 12 short and conical papillae on the BRA. Most of them are uniformly distributed on the central region of the arena, posterior to the median ridge and anterior to the secretory pits. However, 10 pustules and 2 – 3 papillae are placed in each lateral area of the arena, separated from the main group by an area without papillation. Some of the lateral papillae are bifid. The longest papillae of the arena form a V-shaped pattern on the posterior margin of the central region. Posterior area of the roof with secretory pits and dorsal velum bearing lobated margin. Buccal floor: Infralabial papillae on infrarostral cartilages absent. There are two long and digitiform infralabial papillae placed at the level of cartilago meckeli (not figured). The lingual anlage bears 2 long and conical lingual papillae and some pustules. Buccal pockets mostly transversal to the axial axis. Prepocket area scattered with 20 pustules and 16 short and conical papillae. There are 18 long and conical papillae, 4 short and conical papillae and about 100 pustules on the BFA. There is a flap (that resembles the lateral ridges of the roof) bearing 3 projections on its free end and placed posterior to the medial ridge of the pocket. The papillae are arranged in two V-shaped patterns, one anterior and other posterior. The ventral velum has a well-marked median notch, three weakly-marked projections at each side on the posterior margin and many secretory pits.	en	Kolenc, Francisco, Borteiro, Claudio, Alcalde, Leandro, Baldo, Diego, Cardozo, Dario, Faivovich, Julián (2008): Advertisement call and female sexual cycle in Uruguayan populations of Physalaemus henselii (Anura, Leiuperidae). Zootaxa 1927 (2): 1-66
03BB87B90954FFF0FF320730FC33A764.taxon	description	Figs. 2 B and 3 F External morphology. — Lot MLP DB 4548, Figs. 6 C and 6 D. Body depressed (BMH / BMW = 0.81 ± 0.03); little longer than one third of total length (BL / TL = 0.35 ± 0.01); body shape oval in dorsal view, widest at spiracle level. In lateral view, ventral contour of body flat in gular and concave in branchial regions, slightly convex in abdominal region. Dorsal contour of body slightly convex from the nostrils to beginning of dorsal fin. Snout rounded in dorsal and lateral views. Nostrils oval, with slightly elevated marginal rim and a rounded projection in medial margin; nostrils dorsolaterally located (EN / BWN = 0.57 ± 0.03), placed in a depression, closer to eyes than to the tip of snout (FN / END = 1.37 ± 0.30), visible in dorsal and lateral views. Eyes large (E / BWE = 0.23 ± 0.03), dorsally positioned (EO / BWE = 0.70 ± 0.04) and dorsolaterally directed, not visible in ventral view. Spiracle single, lateral and sinistral, its inner wall fused to body except for its distal end; its opening oval, elevated, with diameter smaller than or equal to tube diameter, located in posterior third of body (RSD / BL = 0.71 ± 0.03), visible in dorsal and lateral views. Lateral line system visible. Intestinal assa located at the centre of abdominal region. Vent tube starts at midline, at posterior end of body, reaching free margin of lower fin, opening dextral. Tail large (TaL / TL = 0.65 ± 0.01), with both fins higher than body height (MTH / BMH = 1.12 ± 0.03). Dorsal fin originates at tail-body junction. Ventral fin origin concealed by vent tube. Dorsal fin contour slightly convex; ventral fin contour almost flat, both fins tapering to tip of tail in last third. Tail axis straight, end acute; tail musculature reaching tail tip. Oral disc (Figs. 8 F and 9 D) ventral, small (OD / BMW = 0.46 ± 0.02, disc measured folded) with a well marked infraangular constriction on each side. Marginal papillae simple; those at anterior lip short and blunt, while the ones at posterior lip longer and thinner, with pointed tip. Dorsal gap present, small (DG / OD = 0.29 ± 0.11). Row of marginal papillae single or biserially disposed in angular and infraangular regions; some infraangular submarginal papillae present, shorter and wider than marginal ones. Upper jaw sheath slightly convex in the middle and concave laterally, pigmented in distal third and finely serrated. Lower jaw sheath with V-shaped free margin, pigmented in distal ¼, finely serrated. Labial tooth row formula 2 (2) / 4 (1). A 1 bent with an angle directed to the front. P 4 located very close to papillae of posterior border of oral disc, with its labial teeth smaller than other ones. Tooth of all rows spatulate and concave, bearing 8 – 10 well-marked distal cusps (Fig. 10 E). Variation. — One tadpole (stage 28) has one submarginal papilla on each side of oral disc, bearing labial teeth; P 4 fragmented. Another one (stage 29) with some short and conical papillae making a double row in anterior lip, row of marginal papillae double at angular and infraangular regions, while row is single and discontinuous at posterior margin of oral disc. P 4 well developed and continuous, with labial teeth smaller than the other ones. At the infraangular regions there are some submarginal papillae and some flaps with labial teeth. In another specimen (stage 36) there is a double fold with papillae on both sides of the rostral gap, and P 4 has multiple interruptions. One premetamorph (stage 41) has double or multiple rows of papillae in the infraangular regions and some submarginal papillae, one bearing labial teeth. Row of papillae in the posterior lip discontinuous and most of P 4 is lost due to the metamorphosis. Coloration in preservative. — Body light brown in dorsal and lateral views, with fine melanophores. Inner margin of nostrils dark brown. Abdomen dark brown. Caudal musculature little pigmented with a marbled pattern, darker in dorsal view, unpigmented in ventral view. Dorsal fin translucent with dark flecks. Ventral fin translucent, with some melanophores in the last third. Oral cavity morphology. — (Fig. 12 B and 14 B). Buccal roof: Prenarial arena with 20 blunt pustules and a clear transversal crest. The pustules are arranged surrounding the crest. Choanae transversely oriented, with anteromedial and rounded vacuities, posterior margins forming the narial valves, and both margins undulated by the presence of small prenarial pustules, in particular the anterior one. There are about up to 20 pustules, 11 short and simple papillae, and 8 long and bifid papillae on the postnarial arena. The series of longest papillae are arranged forming an arch between the lateral ridge papillae. Each square-shaped lateral ridge papilla possesses 7 projections, some of them longer, on the free margin. The tall, distally narrow but proximally wide square-shaped median ridge presents 8 short undulations on its free margin, lateral borders free of undulations and one short papilla at each side of the base. BRA not defined; there are approximately 264 pustules, and no more than 2 very short and conical papillae on central region of the buccal roof. Posterior area of the roof with secretory pits and dorsal velum bearing lobated margin. Buccal floor: Infralabial papillae on infrarostral cartilages absent. There are two long and digitiform infralabial papillae placed at the level of cartilago meckeli (not figured). The lingual anlage bears 2 long and conical lingual papillae. Buccal pockets mostly transversal. Prepocket area with 22 long and ramified papillae, 10 short conical papillae and 48 pustules. There are 12 long and conical papillae, 32 short and conical papillae and about 120 pustules on the BFA. The papillae are not arranged in a V-shaped pattern. There is a flap (that resembles the lateral ridges of the roof) bearing 3 long projections on its free end and placed posterior to the medial flange of the pocket. The ventral velum has a well-marked median notch, three few-marked projections at each side on the posterior margin and many secretory pits.	en	Kolenc, Francisco, Borteiro, Claudio, Alcalde, Leandro, Baldo, Diego, Cardozo, Dario, Faivovich, Julián (2008): Advertisement call and female sexual cycle in Uruguayan populations of Physalaemus henselii (Anura, Leiuperidae). Zootaxa 1927 (2): 1-66
03BB87B90949FFF7FF320544FD34A7A3.taxon	description	Figs. 2 E and 3 G External morphology. — Lot MLP DB 5303, Figs. 7 A, 7 B and 18 D. The external morphology of the studied specimens is in agreement with the detailed description presented by Rossa-Feres & Nomura (2006), except that we observed a very short ventral gap in one specimen, and a short P 4 row in another one (Figs. 8 G and 9 E). We provide a SEM photograph of the oral disc (Fig. 9 E) and a description of the oral cavity morphology. Oral cavity morphology. — (Fig. 12 C and 14 C). Buccal roof: There are four blunt pustules on the central region of the prenarial arena; three of them are transversely arranged forming a crest. Choanae transversely oriented, with anteromedial and rounded vacuities, posterior margins forming the narial valves, and both margins undulated by the presence of small prenarial pustules, in particular the anterior one. There are about 12 short conical papillae on the postnarial arena forming a transversal line between the lateral ridge papillae. Each square-shaped lateral ridge papilla possesses 4 digitiform projections on the free margin. The low, rounded and wide median ridge presents 11 undulations on its free margin. BFA not defined; there are approximately 130 pustules lying throughout the central area of the buccal roof. The lateral regions possess two pustules per side. Posterior area of the roof with secretory pits and dorsal velum bearing lobated margin. Buccal floor: Infralabial papillae on infrarostral cartilages absent. There are 2 pairs of digitiform infralabial papillae, both placed on the cartilago meckeli. The anterior pair is shorter than the posterior one which is twofold higher. The lingual anlage bears 2 long and conical lingual papillae on each side of the midline, and some pustules and short papillae on the margins of the anlage. Buccal pockets mostly transversal to the axial axis. Prepocket area with about 20 short papillae. There are 24 long and conical papillae and about 34 pustules on the BFA. The papillae are arranged in two V-shaped patterns. The papillae of the anterior V are forming 2 ridges. The ventral velum lacks the median notch, shows 2 – 3 marginal projections (the medial ones less evident) on each side, and many secretory pits.	en	Kolenc, Francisco, Borteiro, Claudio, Alcalde, Leandro, Baldo, Diego, Cardozo, Dario, Faivovich, Julián (2008): Advertisement call and female sexual cycle in Uruguayan populations of Physalaemus henselii (Anura, Leiuperidae). Zootaxa 1927 (2): 1-66
03BB87B90943FFFCFF3204B1FB93A7FA.taxon	description	Figs. 2 F and 3 H External morphology. — Lot MLP DB 2440, Figs. 7 C and 7 D. Body depressed (BMH / BMW = 0.86 ± 0.02); body length little longer than one third of total length (BL / TL = 0.35 ± 0.02); body shape ovoid in dorsal view with a constriction behind eyes; widest at posterior third of body, behind spiracle. In lateral view, ventral contour of body flat in gular and branchial regions, convex in abdominal region. Dorsal contour of body almost flat from eyes to origin of dorsal fin. Snout semicircular in dorsal view and rounded in lateral view. Nostrils oval, with thin marginal rim, with a small rounded or subtriangular fleshy projection in medial margin; nostrils dorsolaterally located (EN / BWN = 0.54 ± 0.06), placed in a depression, closer to eyes than to the tip of snout (FN / END = 1.32 ± 0.15), more visible in dorsal than in lateral view. Eyes large (E / BWE = 0.21 ± 0.01), dorsally positioned (EO / BWE = 0.75 ± 0.03), dorsolaterally directed, not visible in ventral view. Spiracle single, lateral and sinistral; its inner wall fused to body except for its distal end; its opening oval, slightly elevated, with a diameter smaller than tube diameter, located between second and posterior thirds of body (RSD / BL = 0.65 ± 0.03), posterodorsally directed, visible in lateral and dorsal views. Lateral line system visible. Intestinal assa located at centre of abdominal region. Vent tube starting at midline, at posterior end of body, reaching margin of lower fin, opening dextral. Tail large (TaL / TL = 0.65 ± 0.02), with both fins slightly lower than body height (MTH / BMH = 0.98 ± 0.04). Dorsal fin originates at tail-body junction. Ventral fin origin concealed by vent tube. Edges of both fins sub-parallel for first and second thirds, convergent in last one. Tail axis straight, tail tip rounded; tail musculature not reaching tail tip. Oral disc (Figs. 8 H and 9 F) anteroventral, small (OD / BMW = 0.33 ± 0.02, disc measured folded), with a well marked infraangular constriction on each side of oral disc (also a less marked supraangular constriction in one example). Marginal papillae simple, longer than wide, with rounded or pointed tip. Dorsal gap present, medium-sized (DG / OD = 0.39 ± 0.09). Row of marginal papillae single or double; some infraangular submarginal papillae present. Upper jaw sheath widely arch-shaped. Lower jaw sheath with U-shaped free margin. Jaw sheaths well developed, but appear smooth and sparsely pigmented distally. Labial tooth row formula 2 (2) / 3 (1). P 3 almost as long as P 2. Tooth of all rows spatulated and concave bearing 8 – 10 well-marked distal cusps (Fig. 10 F). Coloration in preservative. — Body dark brown in dorsal view, even darker next to the inner margins of nostrils, with small dark spots on the head and body. Almost black laterally. Venter translucent. Caudal musculature in dorsal view dark brown, except at the insertion of the dorsal fin; in lateral view yellowish, with a dark brown marbled pattern. Fins translucent, delicately marbled in dark brown. Variation. — MLP DB 2440. Three specimens (stages 28 and 31) exhibited some labial teeth anterior to A 1, at one or both sides of the dorsal gap. In many specimens, labial teeth are partially lost from some rows. One specimen (stage 35) with some labial teeth arranged as a short P 4, and another one (stage 28) with one lateral submarginal flap with labial teeth. Two additional specimens at later developmental stages (37 and 40) with LTRF 2 (2) / 3 (1), without extra labial teeth. Lot MLP DB 3303. LTRF 2 (2) / 3 (1) in four specimens (stages 26 – 37), 2 (2) / 4 (1) in another three (stages 35 – 39). When present, P 4 appears irregularly fragmented. A 1 is bent at the middle, with an angle directed to the front. Some lateral submarginal flaps with labial teeth are present in five specimens (stages 26 – 37), absent in another two (stages 27 and 39). Oral cavity morphology. — (Figs. 12 D and 14 D). Buccal roof: There are about 11 blunt pustules on the central region of the prenarial arena. Three of them are transversely arranged, tending to but not forming a crest. Choanae transversely oriented, with anteromedial and rounded vacuities, posterior margins forming the narial valves, and both margins undulated by the presence of small prenarial pustules, in particular the anterior one. There are about 12 pustules, 2 short and 6 long conical papillae on the postnarial arena. The six longest postnarial papillae are placed anteriorly and between the two lateral ridge papillae, forming a discontinuous arch. None of these papillae seems to be bifid. Each square-shaped lateral ridge papilla possesses 6 digitiform projections on the free margin. The high, rounded and narrow median ridge presents 6 undulations distributed in all the free margin and one short papillae per side. There are approximately 200 pustules and 25 conical papillae on the BFA. The pustules are occupying all the area of the arena. The papillae are mostly arranged in a V-shaped pattern on the posterior margin of the arena, but some of them are scattered among the central pustules. The lateral regions of the arena possess few pustules. Posterior area of the roof with secretory pits and dorsal velum bearing lobated margin. Buccal floor: Infralabial papillae on infrarostral cartilages absent. There are 2 long and digitiform infralabial papillae placed at the level of cartilago meckeli (not figured). The lingual anlage bears 2 long and conical lingual papillae very near to each other, and some pustules surrounding them. Buccal pockets mostly transversal to the axial axis. Prepocket area with 14 conical papillae of moderate size and 48 pustules. There are 40 long and conical papillae, 24 short and conical papillae and about 100 pustules on the BFA. The papillae are not arranged in two V-shaped patterns. The ventral velum has a well-marked median notch, three weaklymarked serrations on each side on the posterior margin and many secretory pits.	en	Kolenc, Francisco, Borteiro, Claudio, Alcalde, Leandro, Baldo, Diego, Cardozo, Dario, Faivovich, Julián (2008): Advertisement call and female sexual cycle in Uruguayan populations of Physalaemus henselii (Anura, Leiuperidae). Zootaxa 1927 (2): 1-66
