identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03BB87B0FFA2E938FF5AFB39B0B0FC08.text	03BB87B0FFA2E938FF5AFB39B0B0FC08.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Baikalozercon Marchenko 2022	<div><p>Genus Baikalozercon gen. nov.</p><p>Type species: Baikalozercon dracunculus sp. nov.</p><p>Other included species: Baikalozercon irbis sp. nov.</p><p>Diagnosis. The new genus is based on adult female and male material representing two newly described species. Adults of Baikalozercon are distinguished from those of other genera of the family Zerconidae in a combination of morphological attributes: very large size of idiosoma 900–1000 long; neotrichous dorsal shields; multiple gland pores gv2, dispersed over surface of opisthogastric region including ventri-anal shield; very long peritremes, extending from mid coxa IV to mid coxa I in adults as in deutonymphs; sternal region with some paired or entire fragments of sternal shields; chelicera with suboval area (“window”) with thin cuticle on paraxial side of fixed chela. Female genital shield extends under the lower edge of the posterior sternal shield, which has very specific convoluted structure of cuticle. Male with sexual dimorphism of chelicera and II legs. Male fixed digit of chelicera with extended apical edge and large apical outgrowth; male legs II with some modified enlarged setae on femur, genu, tibia or only on femur.</p><p>Etymology. The name of the genus Baikalozercon reflects the name of Baikal Lake in Eastern Siberia (Russia), because two described species co-occur in mountains of Baikal Region.</p><p>Description. Large-sized, bulky body, brownish mites, with pear-shaped idiosoma, divided into two strongly sclerotised shields.</p><p>Idiosomal dorsum. Dorsal shields neotrichous, with festoon or tuberculate ornamentation. Anterior margin of podonotal shield curved ventrally to form a vertex. Setae j1 is the longest on podonotal shield, strong, pilose, and inserted on anterior margin of idiosoma; setae z1 inserted on vertex and rx setae on peritrematal shield. Podonotal and opisthonotal shields with four pairs of pore-like structures in each. Median and lateral setae of opisthonotal shield located on elevated areas; areas without setae located in shield depressions with smooth radiate reticulation. Marginal r-R setae slightly curved and feathered, inserted on rounded tubercles. Posterodorsal 5–7 sigillae presented. Posterior part of opisthonotal shield curved on ventral side and fused with ventri-anal shield.</p><p>Idiosomal venter. Tritosternum with pair of pilose laciniae free from each other along their entire length. Female sternal region with three separate sternal platelets: first with irregular ragged edges along inner contour, second platelet divided into two small oval fragments. Third platelet large, single, with specific convoluted structure of cuticle, with medial incision in posterior part; surrounded by a hyaline membrane. Female genital shield large, with a pair of internal sclerites, surrounded by a trapeziform hyaline membrane, with mid-anterior incision opposite the same on the third sternal platelet; anterior part of genital shield extends under third sternal platelet. Female with a pair of post-genital sclerites and pair of large metapodal platelets, elongated in horizontal direction. Male sternal region with three or four sternal platelets: some divided into fragments and some entire. Male genital opening located at level between coxae II–III, covered by two sclerites, with a pair of eugenital setae on the anterior sclerite; and a pair of inner sclerites. In both sexes endopodal sclerites present around coxae III–IV; exopodal sclerites fused with peritrematal shields in region of coxae III–IV; peritrematal shields strongly sclerotised, ornamented with festoon reticulation along entire length, fused anteriorly forming a vertex, with six pairs of poroids gp1–gp3, ip1–ip3 and two pairs of short barbed setae: z1 and rx. Peritremes slightly undulating, very long, extending from mid coxa IV to mid coxa I. Gland pores gv2 multiple, dispersed over surface of opisthogastric region including ventrianal shield. Ventri-anal shield broad, fused to opisthonotal shield posteriorly, with festoon reticulation, with 9–15 smooth opisthogastric setae, and marginal rows of opisthonotal pilose setae on rounded tubercles. Anal area with para-anal and post-anal setae; with pair of glands gv3 located antero-laterally of para-anal setae; cribrum extends from ventral to dorsal sides of idiosoma between the posteriormost marginal setae. Anal opening with two pairs of lyrifissures on each valve.</p><p>Gnathosoma . Epistome with irregularly serrated lateral edges and smooth, long, bifurcated median projection. Corniculi of moderate size. Internal malae protrude distinctly beyond corniculi; with smooth elongated branches, barbed at base. Setae h1–h3 smooth, attenuate, pc serrated. Deutosternum with seven denticulate rows. Chelicerae large, with long dorsal seta, lateral (antiaxial) and dorsal lyrifissures, serrated arthrodial corona and paraxial suboval area (“window”) of thinner and lighter cuticle layer on fixed digit, pilus dentilis tubular with rounded or bifurcate top. Female fixed digit with seven medium-sized teeth, movable digit with three or four medium-sized teeth in addition to distal hook. Male fixed digit with six or seven teeth and large apical outgrowth, movable digit as in female. Palpal chaetotaxy 2–5–6–13–15, palp with five free segments; palp trochanter with seta al1 long and pilose in distal third, seta al2 short and smooth; palp genu with seta al1 smooth and al2 pilose in distal third; palp tarsal claw two-tined.</p><p>Legs. Legs of moderate length. Setae of legs heterogeneous. All legs with sclerotised claws and pulvillus with apically rounded lobes. Pretarsus of legs II–IV with ambulacral stalk, tarsi I with sessile claws. Coxae I with split on dorsal side, coxae IV with recesses on anterolateral side; coxae II–III with antero-dorsal spines: II with large sharp spine, III with small one; coxae IV with alveolar vestige of second av seta. In male legs II with sexual dimorphism in form of enlarged spine-like setae that can be located on femur— al1, al2, av1, on genu al1 and on tibia al1; or only enlarged setae on femur al1, al2. Chaetotaxy of legs I–IV: coxae 2, 2, 2, 1; trochanters 6 (1 1/3 1), 5 (1 1/3 0), 5 (1 1/3 0), 5 (1 1/3 0); femora 13 (2 5/4 2), 11 (2 5/3 1), 6 (1 4/1 0), 6 (1 4/1 0); genua 13 (2 6/3 2), 11 (2 5/2 2), 10 (2 4/2 2), 10 (2 5/2 1); tibiae 14 (2 6/4 2), 10 (2 4/2 2), 9 (2 3/2 2), 10 (2 4/2 2); tarsi I—49 (6 29/9 5), II–IV 18 (3 7/5 3). Chaetotaxy of I–IV legs in general typical for Zerconidae, except for IV tibia wich is given by Halašková (1969) and Sikora (2014) as 9 (2 3/2 2).</p><p>Differential diagnosis. Comparisons with related taxa (adults). Based on collections in ISEA there are three early derivate genera of Zerconidae in the mountains of South Siberia: Syskenozercon, Halozercon and Baikalozercon . Syskenozercon is found throughout the Palaearctic, and Halozercon and Baikalozercon inhabit the midlands and highlands of South Siberia. They are probably closely related, sharing relative a number of characters to those genera.</p><p>Shared characters of Baikalozercon with related genera Syskenozercon and Halozercon . Dorsal idiosoma: neotrichy of podonotal and opisthonotal shields; there are 4–7 posterodorsal sigillae instead of four sclerotised fossae; marginal r–R setae inserted on tubercles; podonotal shields with four pairs of pores po1–po4 and opisthonotal also with four pairs of pores Po1–Po4; anterior margin of podonotal shield curved ventrally to form a vertex with j1, z1 setae; caudal fusion of opisthonotal and ventri-anal shields.</p><p>Ventral idiosoma: sternal region of female and male (male of Syskenozercon unknown) with fragmentary sternal platelets; sternogenital region with normal five pairs of setae; genital shield of female surrounded by hyaline cuticle; tocospermous mating system; genital opening of male (for Halozercon and Baikalozercon) located at level between coxae II–III, covered by two sclerites, with a pair of eugenital setae on anterior one; expansive peritrematal shields with reticulation along entire length, fused anteriorly with podonotal shield forming vertex; with multiple gland pores gv2 dispersed over opisthogastric surface including ventri-anal shield; gland pores gv3 inserted antero-laterad anal opening; anal opening with two pairs of lyrifissures on each valve, euanal setae absent.</p><p>Gnathosoma sharing number characters of the family Zerconidae given by Halašková (1969), Moraza &amp; Lindquist (1998) and Ujvári (2011b): hypostome with four pairs of setae (h1 —the longest, h2 shortest, h3 —middle length, pc —serrated); well-developed internal malae; palps with five free segments; chaetotaxy 2–5–6–13–15, palp trochanter with seta al1 long and pilose in distal third, seta al2 short and smooth; palp tarsal claw twotined; deutosternal groove with 6–7 denticles rows with “phylliform structures” (Ujvári, 2011); chelicerae three segmented, with well-developed teeth on both digits; fixed digit with dorsal seta, dorsal and lateral lyrifissures and pilus dentilis. Internal malae in genera Baikalozercon, Halozercon and Syskenozercon similar in form—smooth and pointed anterior branches versus bifurcate in other Zerconidae .</p><p>Legs: chaetotaxy in general typical for Zerconidae, except for tibia IV in Baikalozercon, Halozercon and Syskenozercon with 10 setae (2 4/2 2) compared with 9 (2 3/2 2) in other Zerconidae (Halašková, 1969; Sikora, 2014). Other characters for these three genera are common: coxae I with split on dorsal side, coxae II with large sharp antero-dorsal spine, coxae IV with alveolar vestige of second seta av (visible in large species of Halozercon and Baikalozercon); pretarsi of legs I–IV with curved claws, pulvillus and paradactyles.</p><p>Differences between Baikalozercon and related genera. Characters based on collections of ISEA as well as by Athias-Henriot (1976), Halašková (1969), Moraza &amp; Lindquist (1998) and Cǎlugǎr (2004 /2006) are presented in Table 1.</p></div>	https://treatment.plazi.org/id/03BB87B0FFA2E938FF5AFB39B0B0FC08	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Marchenko, Irina I.	Marchenko, Irina I. (2022): Description of new genus Baikalozercon (Acari: Mesostigmata: Zerconidae) with two new species from South Siberia Mountains (Russia). Zootaxa 5120 (3): 301-333, DOI: 10.11646/zootaxa.5120.3.1
03BB87B0FFA0E92AFF5AFC65B11CFCF0.text	03BB87B0FFA0E92AFF5AFC65B11CFCF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Baikalozercon dracunculus Marchenko 2022	<div><p>Baikalozercon dracunculus sp. nov.</p><p>(Figures 1–40)</p><p>Female (Figs 1–8, 11–19, n=10)</p><p>Dorsal idiosoma (Figs 1, 5, 11). Idiosoma pear-shaped, 920–975 long, 650–700 wide, with tuberculate ornamentation, divided into two shields that cover the entire dorsal idiosoma. Anterior margin of podonotal shield curved ventrally to form a vertex. The most anterior setae j1 the longest (53–55) of all setae, pilose, inserted on anterior margin of idiosoma. Vertex with setae z1, the shortest (12–14), slightly barbed and pair of pore-like structures (glands) po1. All medial and lateral setae on dorsal shields slightly pilose (visible only with the use of oil immersion), located on rounded bases; marginal r-R setae slightly curved and feathered, inserted on rounded tubercles. Podonotal shield neotrichous, with 19–21 lateral setae of z, s -series (25–27µm), 31–36 medial setae of j -series (three pair of them longer 35–37 long, others as lateral), 18–19 setae of r -series (47–50) on each side. Podonotal shield ornamented with small tubercles forming a reticulated pattern with the exception of the midposterior region. Regions of podonotal shield between medial and lateral groups of setae with longitudinal fields of sigillae. Four pair of pore-like structures (glands) po1–po4 located in podonotum. Opisthonotal shield neotrichous, with tuberculate ornamentation except smooth reticulated regions between medial and lateral setae in anterior part and in mid-posterior region. Setae of J, Z and S -series in asymmetrical location. Medial 26–29 setae (25–27) located on raised area in the center of the shield, lateral 23–27 setae (as long as medial) located on raised ridges; and marginal 16–17 R -setae from each side (35–37); regions free of setae located in the depressions of the shield with about two pairs of sigillae. Area around five posterodorsal sigillae with smooth reticulation. Smooth reticulation radiates from the center like the petals of a flower. Four pairs of pores Po1–Po4 inserted in opisthonotum. Cribrum on the terminal end of the shield between the posteriormost marginal setae.</p><p>Ventral idiosoma. (Figs 2–4, 6, 7, 12–14). Base of tritosternum 67–73 long and 40–45 wide; paired pilose laciniae free from each other along entire length, 105–112 long (Fig. 3). First sternal platelet single, hollow inside, with sharp ragged edges along inner contour; 45–50 long (along a vertical line through setae St1) and 75–80 wide, with pair of St1 setae (45–55) and slit-form lyrifissures iv1 (Fig. 2, 12). Second sternal platelet formed by two small separate oval platelets (30–45 x 15–25) located on a larger sclerotised cuticle (35–37 x 67–75), with pair of St2 setae (35–45) and oval lyrifissures iv2 adjacent to the anterio-lateral edge. Third sternal platelet entire, the largest, 120–125 long and 117–132 wide, with convoluted structure of cuticle above level of St4 setae; with a medial incision dividing the shield into two lobes in posterior part, not reaching level of St4 setae; with two pairs of setae St3 (40–45), St4 (45–50); with slit-form lyrifissures (iv3) and pair of large rounded poroids gv1 adjacent to the anterio-lateral edge (Figs 4, 13–14). Third sternal platelet surrounded by a hyaline membrane, inside which poroids gv1 located. Genital shield axe-shaped, expanded posteriorly, 160–175 long and 160–193 at widest part; with setae St5 (35–40), lyrifissures iv5 on shield or outside. Anterior part of the genital shield has a different cuticle structure than posterior one as shown in Fig. 4. The genital shield is surrounded by a trapeziform membrane with mid-anterior incision opposite the same on the third sternal platelet. In the typical position, anterior part of the genital shield extends under the third sternal platelet; under pressure on the slide, the shields are displaced vertically as shown in Fig. 4. One pair of post-genital sclerites inserted between genital and ventri-anal shields. Endopodal sclerites presented around of coxae III–IV. The anterior part of endopodal sclerite partially extends under the third sternal platelet. Exopodal sclerites fused with peritrematal shields in region of coxae III–IV. Peritrematal shields strongly sclerotised, ornamented with festoon reticulation along entire length; fused anteriorly forming vertex. Six pairs of poroids inserted in peritrematal shields: gp1–gp3 and ip1–ip3 and two pairs of short barbed setae: z1 and of the same length rx seta (12–14), located opposite III coxae. Peritremes slightly undulating, very long, extending from mid coxa IV to mid coxae I, 320–335 long. Metapodal platelets large, elongated in horizontal direction (17–25 x 75–95). Gland pores gv2 multiple, dispersed over surface: one pair of gland openings inserted in soft cuticle postero-laterad genital shield, other 5–7 located on ventri-anal shield from each side. Ventri-anal shield broad, 315–335 long and 550–615 wide, fused to opisthonotal shield posteriorly at level of para-anal setae, with festoon reticulation, with 11–15 opisthogastric smooth setae, setae Jv1–Jv3 the longest (30–37) and others shorter (25–30), marginal rows of opisthonotal setae pilose, slightly curved, on rounded tubercles from each side. Anterior most opisthogastric setae Jv1 can be located both on the shield or on the soft cuticle above it. Setae of Jv and Zv- rows are clearly distinguished (Jv1–Jv3, Zv1–Zv3), other setae in asymmetrical location. Anal area with smooth para-anal (25–27) and post-anal (31–37) setae; anal opening (45–50 x 40–42) with two pairs of lyrifissures on each valve; cribrum extends from ventral to dorsal sides of idiosoma between the posteriormost marginal setae. Pair of glands gv3 located anterolaterally of para-anal setae.</p><p>Gnathosoma . (Figs 8, 15–19) Movable digit of chelicera 100–105 long, with three teeth in addition to apical hook; fixed digit the same length, with seven teeth in addition to apical hook and two-pointed pilus dentilis (Figs 15–16). Chelicera with long dorsal seta (40–43), lateral (antiaxial) and dorsal lyrifissures; with serrated arthrodial corona and suboval area (“window”) on paraxial side with thin cuticle. Epistome (Fig. 17) with irregularly serrated lateral edges and smooth, long, bifurcated median projection. Corniculi 55–57 long and 27–29 wide. Internal malae protrude distinctly beyond corniculi (Figs 18–19); with smooth elongated branches, barbed at base. Deutosternal groove with seven transverse denticulate rows. Subcapitulum with 2–3 paired festoon lateral transverse lines (Fig. 18). Setae h1–h3 smooth: h1 (67–69) the longest, seta h2 (37–39) shorter than h3 (55–57); pc (45–47) serrated. Dorsal side of gnathosoma with pair of paraxial lanceolate structures—“staples” between chelicera shaft and epistome in female as in male (Fig. 27) and immature stages. Palpal chaetotaxy 2–5–6–13–15, palp with five free segments; palp trochanter with seta al1 long and pilose in distal third, seta al2 short and smooth; palp genu with seta al1 smooth and al2 pilose in distal third; palp tarsal claw two-tined.</p><p>Legs. (Figs 20–23, 28–29). Lengths: I 660–710, II 560–600, III 550–600, IV 725–750 µm. Chaetotaxy of legs I–IV: coxae 2, 2, 2, 1; trochanters 6 (1 1/3 1), 5 (1 1/3 0), 5 (1 1/3 0), 5 (1 1/3 0); femora 13 (2 5/4 2), 11 (2 5/3 1), 6 (1 4/1 0), 6 (1 4/1 0); genua 13 (2 6/3 2), 11 (2 5/2 2), 10 (2 4/2 2), 10 (2 5/2 1); tibiae 14 (2 6/4 2), 10 (2 4/2 2), 9 (2 3/2 2), 10 (2 4/2 2); tarsi I—49 (6 29/9 5), II–IV 18 (3 7/5 3). All legs with sclerotised claws and pulvillus with rounded lobes (Fig. 29). Pretarsus of legs II–IV with ambulacral stalk, tarsi I with sessile claws (Fig. 28). Coxae I with split on dorsal side, coxae IV with recesses on anterolateral side; coxae II–III with antero-dorsal spines: II with large sharp spine, III with smaller spine; coxae IV with 1–3–tined postero-ventral spine and alveolar vestige of second av seta.</p><p>Male. (Figs 9–10, 24–27, 30, 31–36, n=10).</p><p>Dorsal idiosoma. Dorsal idiosoma pear-shaped, 900–950 long, 625–675 wide, ornamentation and neotrichy of dorsal shields as in female.</p><p>Ventral idiosoma. (Figs 9, 24–25, 31–34). Base of tritosterum 47–50 long and 30–32 wide, pilose laciniae 100–110 long, clamped under large subcapitular teeth (Fig. 27). Pair of St1 (42–47) setae in folded soft cuticle. Lyrifissures iv1 slit-shaped, located under setae St1. Second sternal platelet entire, in the form of a narrow strip, 12– 20 long and 55–62 wide, located exactly over the genital opening, with pair of St2 setae (35–40) and lyrifissures iv2 antero-laterally to platelet. Third sternal platelet divided in two parts on sides of genital opening, 80–87 long, 37–42 wide each part; with vertical irregular folding; with St3, St4 setae (32–37), with rounded lyrifissures iv3 and slit-shaped one iv4 from each side (Fig. 25, 34). Fourth sternal platelet (or post-genital) triangular in shape with elongated apex, 100–110 long and 65–75 wide, with relief pattern above St5 setae (25–27) (Figs 33, 34). Lyrifissures iv5 inserted on soft cuticle, postero-laterally of St5 setae. Genital opening located at level between coxae II–III, 45– 47 long and 47–49 wide, covered by two sclerites, with a pair of eugenital setae (25–27) on anterior sclerite; with pair of inner sclerites. Peritrematal shields similar to that female, with seta rx and six pore-like structures on each side. Arch of vertex with pair of slightly barbed setae z1 and pore po1. Peritremes similar to those of female, slightly undulating, very long (315–335). Pair of endopodal sclerites located at mid coxae III–IV, extends under third sternal platelets anteriorly. Exopodal sclerites fused with peritrematal shields in region of coxae III–IV. Ventri-anal shield broad, 375–380 long and 550–600 wide, fused with metapodal plates. Ornamentation, chaetotaxy and fusion with the opisthonotal shield as in female. Pair of setae Jv1 and anteriomost pair of glands gv2 inserted on anterior edge, other glands gv2 dispersed over the surface of ventri-anal shield. Pair of glands gv3 located antero-laterally to paraanal setae. Anal opening suboval (50–52 × 40–42); post-anal seta (30–32) longer than para-anal setae (22–25), with two lyrifissures on each valve.</p><p>Gnathosoma . Male chelicera with sexual dimorphism of fixed digit (Fig. 26). Fixed digit of chelicera with extended distal edge; the same length as movable digit, with six medium-sized teeth and one small tooth at large apical outgrowth; with bifurcate pilus dentilis; with large suboval “window” of thinner and lighter cuticle layer in paraxial side. Movable digit of chelicera 100–105 long, tridentate in addition to apical hook, with serrate arthrodial corona at base of digit. Chelicera with dorsal seta, antiaxial and dorsal lyrifissures. Epistome, corniculi, internal malae and palpal structures as in female. Male subcapitular structure with sexual dimorphism—one pair of massive denticles (27 x 17) protruding at an angle outward, located between h2–h3 and pc setae, laterally to deutosternal groove (Figs 10, 27, 35–36).</p><p>Legs. (Figs 9, 30, 32). Lengths: I 700–725, II 575–600, III 575–600, IV 725–750 µm. Chaetotaxy and morphology I, III–IV legs as in female. Legs II of male with sexual dimorphism. Femur II with three enlarged setae: the largest of all, elongated spine-like al2, and smaller spine-like al1, av1; genu II with modified spine-like al1 seta and tibia II with modified seta al1 as shown in figures 30, 32.</p><p>Deutonymph (Figs 37–38, n=4).</p><p>Dorsal idiosoma. Dorsal idiosoma suboval shape, 695–795 long, 500–600 wide, divided into two shields. Dorsal shields do not cover the entire body, leaving soft cuticle on the sides, with row of marginal setae. Podonotal shield does not form a vertex. Setae j1, z1 and po1 inserted on anterior margin of podonotal shield. Both shields tuberculate ornamentated over the entire surface, with neotrichy. Podonotal shield with four pairs of poroids po1– po4; opisthonotal with three pairs of poroids on shield Po1, Po3, Po4 and Po2 inserted on soft lateral cuicle. Area around five posterodorsal sigillae with smooth reticulation.</p><p>Ventral idiosoma. Sternal shield entire, slightly reticulated with setae St1–St3 and lyrifissures iv1; setae St4, St5 and lyrifissures iv5 inserted on soft cuticle. Metapodal platelets wider than long (20–25 x 40–50); pair of postgenital sclerites located posteriorly setae St5. Peritremes 265–290 long, undulated, reaching to mid coxae I. Small part of sclerotised cuticle adjoins laterally to the peritrema between coxae II and III from each side. Marginal setae rx located on soft cuticle at lateral margins of ventral idiosoma at level between coxae II–III. Opisthogastral region with four pairs of glands gv2 dispersed over surface and 12–14 simple setae on each side. Anal shield separate, wider than long (110–115 x 150–160), with pair of para-anal and post-anal setae; with cribrum and pair of glands gv3 on anterio-lateral corners of shield. Opisthonotal shield curved on ventral side postero-laterally, with nine barbed marginal setae and 2–3 setae inserted asymmetrical.</p><p>Gnathosoma . As in female.</p><p>Legs. Lengths: I 575–600, II 475–500, III 475–500, IV 585–625. Chaetotaxy and morphology of legs as in female.</p><p>Protonymph (Figs 39–40, n=7).</p><p>Dorsal idiosoma. Dorsal idiosoma 525–610 long and 355–425 wide, with punctate ornamentation of podonotal and pygidial shields. Anterior margin of podonotum not curved ventrally, with pair of pilose setae j1 at large base. Podonotal shield with five pairs of setae in j -row (j1, j3–j6), z-row with pairs of recognisable z4 and z5 setae, with 13–15 pilose setae at lateral margins from each side; two pairs of r -setae on soft cuticle and three pairs of poroids po1, po2 and po4. Pygidial region presented by one large, broad pygidial shield and two medium size mesonotal platelets. Pygidial region with three pairs of pilose setae in J -row, where J1 setae located on paired mesonotal platelets. Another lateral 19 pilose setae and 9–10 marginal curved pilose setae present on each side. Also four pairs of poroids Po1–Po4 and five or seven posterodorsal sigillae present in pygidial region.</p><p>Ventral idiosoma. Sternal region with setae St1–St3 and St5 inserted in soft folded cuticle; St5 minute. Peritrematal shields not distinct. Peritremes short (60–67), with internal cell structure; with thin ducts leading from stigmata. Two pairs of adgenital gland pores gv2 below coxae IV visible. Metapodal platelets (7–8 x 17–18) present. Opisthogastric area with five pairs of simple setae inserted in soft cuticle.Anal shield subtriangular form, ornamented with tubercles and folds, 93–95 long and 105–117 wide, with cribrum, pair of glands gv3 at antero-lateral margins; with para-anal and post-anal setae. Opisthonotal shield curved on the ventral side, not connected with anal shield; with 3–4 setae on sclerotised ornamented cuticle and eight marginal curved pilose setae on each side.</p><p>Legs. Lengths: I 465–490, II 390–415, III 390–415, IV 460–485. Chaetotaxy of legs I–IV: coxae 2, 2, 2, 1; trochanters 4, 4,4, 4; femora 10, 8, 5, 4; genua 8, 6, 6, 5; tibiae 8, 7, 7, 7; tarsi II–IV: 17, 17, 17.</p><p>Material examined. Holotype female, Russia, Baikal region, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.15&amp;materialsCitation.latitude=51.55" title="Search Plazi for locations around (long 105.15/lat 51.55)">environs of Tankhoi village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.15&amp;materialsCitation.latitude=51.55" title="Search Plazi for locations around (long 105.15/lat 51.55)">Baikal Natural Biosphere Reserve</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.15&amp;materialsCitation.latitude=51.55" title="Search Plazi for locations around (long 105.15/lat 51.55)">Khamar-Daban Ridge</a>, 51 33’ N, 105 09’ E, 700 m a.s.l., Abies sibirica - Pinus sibirica taiga, in litter, 14 August 2014, coll., L.V. Petrozhitskaya. Paratypes: 6 females, 5 males, same data as holotype; 2 males, same geographical data, 1000 m a.s.l., tall grass alpine meadow with sparse trees of Abies sibirica, in soil, 13 August 2014, coll. L. V. Petrozhitskaya; 4 females, 4 males, Baikal region, environs of Vydrino village, Khamar-Daban Ridge, 51 23’ N, 104 38’ E, 500 m a.s.l., direction to Taltsinskii Peak, Abies sibirica - Pinus sibirica taiga, in litter, 23 June 2021, coll. I.I. Marchenko; 11 females, 4 males, same geographical data, 700 m a.s.l., direction to Taltsinskii Peak, Abies sibirica - Pinus sibirica taiga, in litter, 26 June 2021, coll. I.I. Marchenko; 1 males, Chitinskaya Oblast (now—Zabaikalskii Krai), Kyra District, Sokhondinskii Nature Reserve, 49 48’ N, 111 12’ E, 1600 m a.s.l., cordon Lukovoe, Larix sibirica – Pinus sibirica taiga, in litter, 12 June 1991, coll. D. V. Logunov; 2 females, Krasnoyarskii Krai, Eastern Sayan Ridge, Natural Reserve “Stolby”, mountain taiga with Pinus sylvestris, Abies sibirica, Larix sibirica, in litter, 55 56’ N, 92 44’ E, 500 m a.s.l., 30 August, 21 September 1972, coll. T.S. Sukhova.</p><p>Other material: 4 protonymphs, Russia, Baikal Region, environs of Tankhoi village, Baikal Natural Biosphere Reserve, Khamar-Daban Ridge, 51 33’ N, 105 09’ E, 700 m a.s.l., Abies sibirica - Pinus sibirica taiga, in litter, 14 August 2014, coll., L. V. Petrozhitskaya; 2 protonymphs, same geographical data, 1000 m a.s.l., tall grass alpine meadow with sparse trees of Abies sibirica, 13 August 2014, coll. Petrozhitskaya; 4 deutonymphs, 9 protonymphs, Baikal Region, environs of Vydrino village, Khamar-Daban Ridge, 51 23’ N, 104 38’ E, 500 m a.s.l., Abies sibirica - Pinus sibirica taiga, in litter, 23 June 2021, coll. I.I. Marchenko; 2 deutonymphs, 3 protonymphs, same geographical data, 700 m a.s.l., direction to Taltsinskii Peak, Abies sibirica - Pinus sibirica taiga, in litter, 26 June 2021, coll. I.I. Marchenko.</p><p>Etymology. The specific name dracunculus is Latin, little dragon: dracon (dragon) + unculus (little). This name associated with the structural features of a pair of massive subcapitular denticles in male in form of fangs like in mythical dragons.</p><p>Remarks.Adults of Baikalozercon dracunculus differs from B. irbis as follows—first species with ornamentation of central part of opisthonotal shield in the form of a network of small tubercles; pair of posteriomost marginal setae in R -row of opisthonotal shield like other R -setae in shape and length. Female of B. dracunculus with axe-form genital shield. Movable digit of chelicera in both sexes of B. dracunculus with three teeth. Males have significant differences in sternal area, so B. dracunculus without first sternal platelet, second platelet in form of a narrow strip, fourth sternal platelet (or post-genital) triangular in shape with elongated apex. Male subcapitulum of B. dracunculus with pair of massive denticles between h2–h3 and pc setae. Male legs II of B. dracunculus with sexual dimorphism in form of some modified setae: three enlarged femur setae— al1, al2 and av1; one genu seta al1 and one tibia seta al1.</p></div>	https://treatment.plazi.org/id/03BB87B0FFA0E92AFF5AFC65B11CFCF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Marchenko, Irina I.	Marchenko, Irina I. (2022): Description of new genus Baikalozercon (Acari: Mesostigmata: Zerconidae) with two new species from South Siberia Mountains (Russia). Zootaxa 5120 (3): 301-333, DOI: 10.11646/zootaxa.5120.3.1
03BB87B0FFB2E920FF5AFC3DB271F951.text	03BB87B0FFB2E920FF5AFC3DB271F951.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Baikalozercon irbis Marchenko 2022	<div><p>Baikalozercon irbis sp. nov.</p><p>(Figures 41–63)</p><p>Female (Figs 41–53, n=10)</p><p>Dorsal idiosoma (Fig. 41). Idiosoma pear-shaped, 1025–1060 long, 675–725 wide, divided into two shields that cover the entire dorsal idiosoma. Podonotal shield with festoon ornamentation, except mid-posterior region with smooth reticulation, anterior margin curved ventrally to form a vertex. The most anterior setae j1 the longest (55–63) of all setae, pilose, inserted on large bases on anterior margin of idiosoma. Vertex with setae z1, the shortest (20–25), slightly barbed and a pair of pore-like structures (glands) po1 visible on ventral side. All medial and lateral setae on dorsal shields slightly pilose (visible only with the use of oli immersion), located on rounded bases; marginal r-R setae inserted on rounded tubercles, slightly curved and feathered. Podonotal shield neotrichous, with 25–27 lateral setae of z, s -series (25–27), 25–29 medial setae of j -series (four pairs of them longer 35–37 long, others as lateral), 15–16 marginal setae of r -series (35–37) from each side; and four pair of pore-like structures (glands) po1–po4. Regions of podonotum between medial and lateral setae with groups of sigillae of longitudinal location. Opisthonotal shield neotrichous, with smooth reticulation in median region and festoon ornamentation in lateral regions (Figs 41, 50). Setae of J, Z and S -series in asymmetrical locations. Setae of J, Z and S series in asymmetrical position. Medial setae of J- series 15–23, 22–25 long; lateral setae 31–36, 30 long; and marginal R -setae 15–18 on raised tubercles (40–43) on each side; regions free of setae located in the depressions of the shield with two pairs of sigillae from each side. Region surrounding 6–7 posterodorsal sigillae with smooth reticulation and punctate at the corners of the cells. Four pairs of pores Po1–Po4 inserted in opisthonotum. Cribrum visible on the terminal end of shield between the posteriormost marginal setae.</p><p>Ventral idiosoma. (Figs 42–47). Base of tritosternum 63–65 long and 35–37 wide, paired pilose laciniae free from each other along entire length, 107–112 long. First sternal platelet 50–52 long (along a vertical line through setae St1) and 100–107 wide, single, hollow inside; with irregular wavy edges along inner contour; with pair of St1 setae (60–63) and slit-shaped lyrifissures iv1 (Figs 42, 45). Second sternal platelet formed by two small separate rounded platelets with diagonal groove (25–27 x 25–35), with pair of St2 setae (55–57) and oval lyrifissures iv 2 in lateral position. Third sternal platelet entire, the largest, 137–142 long and 177–112 wide, with folded reticulation structure of cuticle; with a medial incision dividing the shield into two lobes in posterior part, not reaching level of St4 setae; with two pairs of setae St3, St4 of the same length (47–50); and slit-form lyrifissures iv3 (Figs 42–47). Pair of large rounded glands gv1 located at the antero-lateral edge of the hyaline membrane surrounding largest sternal shield. Genital shield sack-shaped with extension in anterior and posterior parts, with narrowing in anterior third, 125–140 long and 155–170 wide at widest part; with setae St5 (20–25), and pair of internal genital sclerites.Anterior part of the genital shield (above narrowing) has a different cuticle structure than the posterior, as shown in Fig. 43. Lyrifissures iv5 inserted on soft cuticle outside of genital shield. The genital shield is surrounded by trapeziform membrane with mid-anterior incision opposite the same on the largest sternal shield. In typical position, anterior part of genital shield located under the largest sternal shield not reaching level of St4 setae. When pressed down on a slide, the third sternal and genital shields are displaced vertically as shown in Fig. 43. One pair of post-genital sclerites inserted between genital and ventri-anal shields. Endopodal sclerites present around of coxae III–IV. The anterior part of endopodal sclerite can partially extends under third sternal platelet. Exopodal sclerites fused with peritrematal shields in region of coxae III–IV. Peritrematal shields strongly sclerotised, ornamented with festoon reticulation along entire length; fused anteriorly forming a vertex. Six pairs of poroids inserted in peritrematal shields: gp1–gp3 and ip1–ip3 and two pairs of short setae of the same length (12–14): barbed z 1 in vertex and simple rx seta located opposite coxae III. Peritremes slightly undulating, very long, extending from mid coxa IV to mid coxa I, 350–370 long, with inner partitions. Metapodal platelets large, elongated in horizontal direction (15–20 x 105–120). Gland pores gv2 multiple, dispersed over surface: one pair of glands openings inserted on soft cuticle postero-laterally of genital shield, other 7–9 located on ventri-anal shield from each side. Ventri-anal shield broad, 425–440 long and 675–725 wide, fused to opisthonotal shield posteriorly at level of post-anal setae, with festoon reticulation in region above glands gv3, and smooth reticulation surround anal opening; with 10–11 opisthogastric smooth setae (27–30) and 11 marginal opisthonotal setae, pilose, slightly curved, on rounded tubercles from each side. Setae of Jv and Zv- rows are clearly distinguished (Jv1–Jv3, Zv1–Zv3), other setae in asymmetrical location. Anal area with smooth para-anal (20–25) and post-anal (30–35) setae; anal opening (52–62 x 50–52) with two pairs of lyrifissures on each valve (Fig. 49); cribrum extends from ventral to dorsal sides of idiosoma between the longest (58–70), barbed posteriormost marginal setae (Fig.42). Pair of glands gv3 located antero-laterally of paraanal setae.</p><p>Gnathosoma . (Figs 48, 51–53) Movable digit of chelicera 112–114 long, with four teeth in addition to apical hook; fixed digit 114–116 length, with seven (six in some specimens) teeth in addition to apical hook and tubular pilus dentilis with rounded tip (Fig. 51). Chelicera with dorsal seta (45–46), lateral (antiaxial) and dorsal lyrifissures; with serrated arthrodial corona and suboval area (“window”) on paraxial side with thin cuticle. Epistome (Fig. 52) with irregularly serrated lateral edges and smooth, long, bifurcated median projection. Corniculi 55–57 long and 27–29 wide. Internal malae protrude distinctly beyond corniculi (Fig. 53) with smooth elongated branches, barbed at base. Deutosternal groove with seven transverse denticulate rows, with four paired curved lateral transverse lines. Setae h1–h3 smooth: h1 (67–69) the longest, seta h2 (43–45) shorter than h3 (63–65); pc (37–40) serrated. Dorsal side of gnathosoma with pair of paraxial lanceolate structures –“staples” between chelicera shaft and epistome. Palpal chaetotaxy 2–5–6–13–15, palps with five free segments; palp trochanter with seta al1 long and pilose in distal third, seta al2 short and smooth; palp genu with seta al1 smooth and al2 pilose in distal third; palp tarsal claw two-tined.</p><p>Legs. (Figs 44, 61). Lengths: I 750–775, II 625–675, III 625–675, IV 775–790 µm. Chaetotaxy of legs I–IV: coxae 2, 2, 2, 1; trochanters 6 (1 1/3 1), 5 (1 1/3 0), 5 (1 1/3 0), 5 (1 1/3 0); femora 13 (2 5/4 2), 11 (2 5/3 1), 6 (1 4/1 0), 6 (1 4/1 0); genua 13 (2 6/3 2), 11 (2 5/2 2), 10 (2 4/2 2), 10 (2 5/2 1); tibiae 14 (2 6/4 2), 10 (2 4/2 2), 9 (2 3/2 2), 10 (2 4/2 2); tarsi I—49 (6 29/9 5), II–IV 18 (3 7/5 3). All legs with sclerotised claws and pulvillus (Fig. 61). Pretarsus of legs II–IV with ambulacral stalk, tarsi I with sessile claws. Coxae I with split on dorsal side, coxae IV with recesses on anterolateral side; coxae II–III with antero-dorsal spines: II with large sharp spine, III with small one; IV coxae with 1–3–tined postero-ventral spine and alveolar vestige of second av seta.</p><p>Male. (Figs 54–59, n=10).</p><p>Dorsal idiosoma. (Fig. 56). Dorsal idiosoma pear-shaped, 985–1000 long, 660–700 wide, ornamentation and neotrichy of dorsal shields as in female.</p><p>Ventral idiosoma. Base of tritosterum 47–50 long and 32–37 wide, pilose laciniae 108–112 long. First sternal platelet paired, divided, irregular in form, with ragged outlines, 47–50 long and 37–50 wide, with seta St1 (47–50) and slit lyrifissures iv1 on each part (Fig. 54). Second sternal platelets entire, with irregularly rounded anterior margin and truncated posterior one, with festoon reticulation, 45–50 long and 62–65 wide, with pair of setae St2 (43–45) and lyrifissures iv2 laterad platelet, located on soft cuticle. Truncate posterior edge of second sternal platelet is adjacent to genital opening. Third sternal platelet divided in two parts, adjacent to the genital opening on the sides, with vertical reticulation, 95–105 long, 32–37 at widest part; with setae St3, St4 (32–35), with rounded glands gv1 on anterior edge and slit lyrifissures iv 3 in central region on each side. Fourth sternal platelet (or post-genital) consists of two fragments: small anterior 25–37 x 17–25 and large subtriangular posterior 55–62 x 82–100 with pair of setae St5 (25–27), and lyrifissures iv5 inserted on soft cuticle lateral to St5 setae. Genital opening located at level between coxae II–III (50–52 x 50–55), covered by two sclerites, with a pair of eugenital setae (25–27) on anterior sclerite; with pair of inner sclerites (Figs 54, 57–58). Peritrematal shields similar to those of female, with smooth rx seta and six pore-like structures on each side. Arch of vertex with pair of slightly barbed setae z1 and pore po1. Peritremes similar to those of female, slightly undulating, very long (325–350). Two pairs of endopodal sclerites almost fused with each other located around coxae III–IV; extend under third sternal platelets anteriorly. Exopodal sclerites fused with peritrematal shields in region of coxae III–IV. Ventri-anal shield broad, 415–440 long and 600–650 wide, fused with metapodal platelets. Ornamentation, chaetotaxy and fusion with the opisthonotal shield as in female. Gland pores gv2 (9–15) dispersed over the surface of ventri-anal shield. Pair of glands gv3 located antero-laterally to paraanal setae. Anal opening suboval 55–57 × 47–50. Post-anal seta (30–32) longer than para-anal setae (20–25), with two lyrifissures on each valve.</p><p>Gnathosoma . (Fig. 55). Male chelicera with sexual dimorphism of fixed digit. Fixed digit of chelicera the same length as movable digit, with six or seven (in different specimens) medium-sized teeth and apical large outgrowth; with tubular protruding pilus dentilis; with large suboval “window” of thinner and lighter cuticle layer in paraxial side. Movable digit of chelicera 97–103 long, with four teeth in addition to apical hook, with serrate arthrodial corona at base of digit. Chelicera with dorsal seta, antiaxial and dorsal lyrifissures. Epistome, corniculi, internal malae and palpal structures as in female. Male subcapitulum without sexual dimorphism.</p><p>Legs. (Figs 59–61). Lengths: I 700–735, II 580–600, III 580–600, IV 715–735 µm. Chaetotaxy and morphology of legs I, III–IV as in female. Legs II of male with sexual dimorphism due to some modified setae on femur II: the largest, elongated spine-like seta al2, and smaller and thinner spine-like al1 (Fig. 60). Other segments of legs II including genu and tibia without clearly distinct modified setae.</p><p>Deutonymph not found.</p><p>Protonymph (Figs 62–63, n=12).</p><p>Dorsal idiosoma (Fig. 62). Dorsal idiosoma 550–600 long and 385–415 wide. Podonotal shield with festoon ornamentation except area between j6–j5 setae with smooth reticulation. Anterior margin of podonotum with pair of pilose setae j1 at large base. Podonotal shield with five pairs of setae in j -row (j1, j3–j6), z -row with pairs of recognisable z4 and z5 setae, with about 15 pilose setae at lateral margins from each side; three r -setae on soft cuticle on each side and four pairs of poroids po1–po4. Pygidial shield with tuberculate and festoon ornamentation except area around 5–7 posterodorsal sigillae with smooth reticulation. Pygidial region presented by one large, broad pygidial shield and two medium sized mesonotal platelets; with three pairs of pilose setae in J -row, J1 located on paired mesonotal platelets; another 12–13 pilose lateral setae and 8–9 marginal pilose curved R setae located on each side of shield. Also poroids Po1 inserted on mesonotal platelets and Po2–Po4 on pygidial shield.</p><p>Ventral idiosoma (Fig. 63). Sternal region with setae St1–St3 and St5 inserted in soft cuticle with slight ornamentation; seta St5 minute. Peritrematal shields not distinct. Peritremes short (62–65), with internal cell structure; with thin ducts leading from stigmata. Pair of gland pores gv2 located posteriad IV coxae. Metapodal platelets (10–12 x 18–23) present. Opisthogastric area with five pairs of simple setae inserted in soft cuticle. Anal shield subtriangular, ornamented with tubercles and folds, 105–125 long and 130–150 wide with cribrum, pair of glands gv3 at antero-lateral margins; with para-anal and post-anal setae. Opisthonotal shield curved on ventral side, not connected with anal shield; with 1–3 setae on sclerotised ornamented cuticle and eight marginal curved pilose setae from each side.</p><p>Legs. Lengths: I 465–475, II 400–425, III 400–425, IV 475–500. Chaetotaxy of legs I–IV: coxae 2, 2, 2, 1; trochanters 4, 4, 4, 4; femora 10, 8, 5, 4; genua 8, 6, 6, 5; tibiae 8, 7, 7, 7; tarsi II–IV: 17, 17, 17.</p><p>Material examined. Holotype female, Russia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.15&amp;materialsCitation.latitude=51.55" title="Search Plazi for locations around (long 105.15/lat 51.55)">Baikal Region</a>, environs of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.15&amp;materialsCitation.latitude=51.55" title="Search Plazi for locations around (long 105.15/lat 51.55)">Tankhoi village</a>, Baikal Natural Biosphere Reserve, Khamar-Daban Ridge, 51 33’ N, 105 09’ E, 1100 m a.s.l., tall grass alpine meadow with sparse trees of Abies sibirica, in soil, 13 August 2014, coll. L. V. Petrozhitskaya . Paratypes: 8 females, 4 males, same data as holotype; 2 females, 2 males, same geographical data, valley of Osinovka river, 1100 m a.s.l., 13 August 2014, coll. L. V. Petrozhitskaya ; 1 male, same geographical data, alpine tundra, 1600 m a.s.l., in litter-soil, 12 August 2014, coll. I.I. Marchenko.</p><p>Other material: 11 protonymphs, same data as holotype; 1 protonymph, same geographical data as type material, alpine tundra, 1600 m a.s.l., 12 August 2014, coll. I.I. Marchenko.</p><p>Etymology. The specific name irbis refers to snow leopard in Turkic and Mongolian languages. The irbis—snow leopard inhabitant of highlands of Central Asia. New species Baikalozercon irbis also inhabits severe highlands in Siberia.</p><p>Remarks. Adults of Baikalozercon irbis differs from B. dracunculus: first species with festoon ornamentation of central part of opisthonotal shield; pair of posteriomost marginal R setae of opisthonotal shield straight and longer than others. Female of B. irbis has sack-form genital shield with constriction in anterior third. Movable digit of chelicera in both sexes of B. irbis with four teeth. Males have significant differences in sternal area, so B. irbis with first sternal platelet irregular in form, second platelet irregularly hemispherical form, fourth sternal platelet (or post-genital) fragmented, consisting of two parts: anterior smaller and posterior larger. Male gnathosoma of B. irbis without sexual dimorphism. Male II legs of B. irbis with sexual dimorphism in form of two modified enlarged setae of femur al1, al2.</p><p>Distribution. Mites of the new genus Baikalozercon are restricted in distribution to the mountains of Southern Siberia. Baikalozercon dracunculus is known from three localities: “Stolby” Nature Reserve in Eastern Sayan; Khamar-Daban Ridges in Baikal Region (Buryatia and Irkutskaya Oblast); and Sokhondinskii Nature Reserve in Zabaikalskii Krai, while B. irbis is found only in the highlands of Khamar-Daban Ridge (Fig. 64). Photos of localities along altitudinal gradient of Khamar-Daban Ridge are shown in Figures 65–71. Two species of the new genus Baikalozercon coexist on the same Khamar-Daban Ridge, but inhabit different altitudes— B. dracunculus occurs at altitudes of 500–700 m a.s.l. in mountain taiga and B. irbis at 1000–1600 m in alpine sparse forest with meadow and alpine tundra (Fig. 72).</p></div>	https://treatment.plazi.org/id/03BB87B0FFB2E920FF5AFC3DB271F951	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Marchenko, Irina I.	Marchenko, Irina I. (2022): Description of new genus Baikalozercon (Acari: Mesostigmata: Zerconidae) with two new species from South Siberia Mountains (Russia). Zootaxa 5120 (3): 301-333, DOI: 10.11646/zootaxa.5120.3.1
